26 Turczaninowia 2013, 16 (4) : 26–46

УДК 582.594 (597) DOI: http:/dx.doi.org/10.14258/turczaninowia.16.4.7

L.V. Averyanov Л.В. Аверьянов

NEW AND RARE ORCHIDS () IN THE FLORA OF CAMBODIA AND LAOS

НОВЫЕ И РЕДКИЕ ВИДЫ ОРХИДНЫХ ВО ФЛОРЕ КАМБОДЖИ И ЛАОСА

Summary. Herbarium material collected in 2009–2013 in Cambodia and Laos provides 240 new localities for 156 orchid species (from 73 genera). Among them, 13 and 45 species respectively are new for the flora of each country. One species (Bulbophyllum konstantinovii) discovered in Cambodia is described as new for science. Eight genera (Acanthephippium, Didymoplexiopsis, Eclecticus, Herpysma, Hetaeria, Lecanorchis, Neuwiedia, and Trichosma) were found in Laos at first. Key words: Cambodia, flora, Laos, Orchidaceae, diversity. Аннотация. Гербарные сборы 2009–2013 годов документируют 240 новых нахождений 156 видов орхидных (из 73 родов) в Камбодже и Лаосе. Среди них 13 и 45 видов, соответственно, приводятся впервые для этих стран. Один вид (Bulbophyllum konstantinovii), найденный в Камбодже, описывается в качестве нового для науки. Восемь родов (Acanthephippium, Didymoplexiopsis, Eclecticus, Herpysma, Hetaeria, Lecanorchis, Neuwiedia и Trichosma) приводятся для флоры Лаоса впервые. Ключевые слова: Камбоджа, Лаос, орхидные, разнообразие растений, флора, Orchidaceae. INTRODUCTION tables 1 & 2) represent significant addition to our previous knowledge (Callaghan, 2004; Gagnepain, The orchid flora of Cambodia and Laos Guillaumin, 1932–1934; Schuiteman, Vogel, still remains one of the less studied orchid floras in 2000; Seidenfaden, 1966, 1973, 1975; Svengsuksa, Tropical Asia. In comparison to other countries of Lamxay, 2005; Vidal, 1959) summarized in last Indochina Peninsula, such as or accounts of Cambodian (Seidenfaden, 1992) and each including at least 1100 species (Averyanov, Laotian floras (Newman et al., 2007; Schuiteman et al., 2003; Pedersen et al., 2011; Seidenfade, 1992), only 2008). One species, B. konstantinovii Aver., discovered 163 orchid species were recorded for Campodia in Cambodia is described as new for science. (Seidenfaden, 1992) and about 485 species for Laos (Newman et al., 2007; Schuiteman et al., Table 1. Orchid species recorded 2008). Meanwhile, according to their square and as new for the flora of Cambodia* various natural conditions all countries of Indochina should certainly conceal similar orchid richness Acriopsis liliifolia (J. Koenig) Seidenf. and diversity. Present paper summarizes some Aerides crassifolia Par. ex Burb. consequent steps in studies of the orchid flora of Bulbophyllum parviflorum Par. et Rchb. f. Cambodia and Laos based on herbarium specimens B. rufilabrum Par. ex Hook. f. collected by different collectors in the field during B. tricornoides Seidenf. 2009–2013. Calanthe cardioglossa Schltr. The paper provides 240 new localities Dendrobium indivisum (Blume) Miq. for 156 species (from 73 genera) among which D. keithii Ridl. 13 species are new for the flora of Cambodia and 45 species represent new records for Laos. Eight *Acriopsis liliifolia, Dendrobium indivisum, Eria lasiopetala, and Porpax elwesii were recently mentioned genera are recorded for the Laotian flora at first: in the amateur site “Orchids of Cambodia” (http://www. Acanthephippium, Didymoplexiopsis, Eclecticus, orchidcambodia.com) without citation of locality and Herpysma, Hetaeria, Lecanorchis, Neuwiedia, voucher specimens proving species occurrence in the and Trichosma. Mentioned new records (see country.

Ботанический институт им. В.Л. Комарова РАН, ул. Проф. Попова, 2; 197376, Санкт-Петербург, Россия; e-mail: [email protected]; [email protected] Komarov Botanical Institute, Russian Academy of Sciences, Prof. Popovа str., 2; 197376, St. Petersburg, Russia Поступило в редакцию 20.12.2013 г. Submitted 20.12.2013 Принято к публикации 24.12.2013 г. Accepted 24.12.2013 Averyanov L.V. New and rare orchids (Orchidaceae) in the flora of Cambodia and Laos 27

D. scabrilingue Lindl. Zeuxine flava (Wall. ex Lindl.) Trimen D. setifolium Ridl. Z. nervosa Trimen Eria lasiopetala (Willd.) Ormerod E. muscicola (Lindl.) Lindl. The list of all discoveries is presented Porpax elwesii (Rchb. f.) Rolfe below. The orchid names in the list are arranged in alphabetic order. Maximally abbreviated and Table 2. Orchid species recorded unified text of herbarium labels is reported for as new for the flora of Lao PDR each collecting number and includes geographical locality, coordinates and elevation (when available), Acanthephippium striatum Lindl. collection date, collectors’ names, collection number Agrostophyllum callosum Rchb. f. and acronym of Herbarium where mentioned Bulbophyllum apodum Hook. f. specimen is housed. B. psychoon Rchb. f. The following standard abbreviations are B. umbellatum Lindl. used in the text: prov. (province), distr. (district), vic. Cheirostylis griffithii Lindl. (vicinity, vicinities), elev. (elevation, elevations), C. pusilla Lindl. mt. (mountain, mountains), N.P. (National Park), C. spathulata J.J. Sm. N.P.A. (National Protected Area). The names of C. thailandica Seidenf. orchid species and genera reported for Cambodia Coelogyne virescens Rolfe and Laos at first are marked with crosslet (+) and Cymbidium eburneum Lindl. asterisk (*) respectively. Name of a new species Dendrobium kontumense Gagnep. is marked by double asterisk (**). The following Didymoplexiopsis khiriwongensis Seidenf. Herbarium acronyms are used in the text: Eclecticus chungii P. O’Byrne CPC – Herbarium of the Center for Plant Epigeneium chapaense Gagnep. Conservation, Vietnam Union of Science and Eria calcarea V.N. Long et Aver. Technology Associations, Hanoi, Vietnam. E. corneri Rchb. f. LE – Herbarium of Komarov Botanical E. thao Gagnep. Institute, Russian Academy of Sciences, St. Flickingeria angustifolia (Blume) A.D. Hawkes Petersburg, Russian Federation. Gastrochilus calceolaris (Buch.-Ham. ex Sm.) D. Don NHOL – National Herbarium of Laos, G. pseudodistichus (King et Pantl.) Schltr. Vientiane, Lao PDR. Goodyera fumata Thwaites NUOL – Herbarium of National University G. hispida Lindl. of Laos, Vientiane, Lao PDR. Herpysma longicaulis Lindl. Prefixes “CPC” and “LA-VN” at some Hetaeria affinis (Griff.) Seidenf. et Ormerod collecting numbers represent abbreviations: Holcoglossum subulifolium (Rchb. f.) Christenson CPC – The Center for Plant Conservation. Lecanorchis vietnamica Aver. LA-VN – Lao-Vietnamese expedition. Liparis balansae Gagnep. Both collecting series are attributed to L. pumila Aver. herbaria collected in expedition managed by the Micropera poilanei (Guillaumin) Garay Center for Plant Conservation, Vietnam Union of Nephelaphyllum tenuiflorum Blume Science and Technology Associations with main Neuwiedia zollingeri Rchb. f. collectors – Prof. Leonid V. Averyanov and Nguyen Oberonia cavaleriei Finet Tien Hiep. Otochilus pseudoporrectus Aver. Paphiopedilum canhii Aver. et O. Gruss New and rare orchids in Cambodia and in Laos Papilionanthe biswasiana (Ghose et Mukerjee) Garay Phalaenopsis wilsonii Rolfe Acampe Lindl. Podochilus oxystophylloides Ormerod A. ochracea (Lindl.) Hochr. Sunipia rimannii (Rchb. f.) Seidenf. LAOS, Vientiane prov., Phou Khao Thelasis khasiana Hook. f. Khouay N.P., Tad Leuk waterfall, 18°23’99.1 N, Thrixspermum sutepense (Rolfe ex Downie) Tang 103°04’03.1 E, 28 Nov. 2012, E.L. Konstantinov et et F.T. Wang al., 52OR-L-GPS005 (LE). LAOS, Vientiane prov., Trichosma simondii Gagnep. Kasi distr., Tam Tai village, about 10 km to the NW T. suavis Lindl. of Kasi town, 19°12’57” N, 102°14’59” E, 500–650

28 Averyanov L.V. New and rare orchids (Orchidaceae) in the flora of Cambodia and Laos m, 24 March 2013, L. Averyanov, N.S. Khang, S. Agrostophyllum Blume Lorphengsy, LA-VN 767 (CPC, NHOL, NUOL). *A. callosum Rchb. f. A. praemorsa (Roxb.) Blatt. et McCann LAOS, Kham Mouan prov., Bounlapha LAOS, Nam Lik, 18°36’80.8 N, 102°24’60.5 distr., Thong Sam village, Hinnam No N.P.A., Pu E, 24 Nov. 2011, E.L. Konstantinov et al., 5 (LE). Pha Song mt., 17°35’13.2” N, 105°47’49.8” E, 1400 A. rigida (Buch.-Ham. ex Sm.) P.F. Hunt m, 9 March 2013, L. Averyanov et al., LA-VN 323, LAOS, Vientiane prov., Kasi distr., Tam Fig. 1, B (CPC, NHOL, NUOL). LAOS, Vientiane Tai village, about 10 km to the NW of Kasi town, prov., Kasi distr., Thong Mout village, 19°23’27.3” 19°12’57”N 102°14’59”E, 500-650 m, 24 March N, 102°08’21.3” E, 1800 m, 20 March 2013, N.T. 2013, L. Averyanov, N.S. Khang, S. Lorphengsy, LA- Hiep et al., LA-VN 656 (CPC, NHOL, NUOL). VN 766 (CPC, NHOL, NUOL). A. planicaule (Lindl.) Rchb. f. LAOS, Vientiane prov., Vang Vieng distr., *Acanthephippium Blume ex Endl. Na Khun Village, around point 18°52’28” N, *A. striatum Lindl. 102°24’21” E, 300 m, 21 Jan. 2009, O. Souliya et LAOS, Saravan prov., Sa Mouay distr., to al., LA-VN 58 (CPC). LAOS, Vientiane prov., Vang the SEE of Sa Mouay town, elev. from Ta Oy village, Vieng distr., Nathong village, about 5 km to the W 1350 m, 16°08’47.0” N, 106°52’35.4” E, 4 March of Vang Vieng town, Tham Kang mt., 18°55’51.4” 2013, L. Averyanov et al., LA-VN 161 (CPC, NHOL, N, 102°23’50.6” E, 300–500 m, 14 March 2013, NUOL). LAOS, Kham Mouan prov., Bounlapha N.T. Hiep et al., LA-VN 376 (CPC, NHOL, NUOL). distr., Thong Sam village, Hinnam No N.P.A., Pu LAOS, Vientiane prov., Kasi distr., Tam Tai village, Pha Song mt, 17°35’13.2” N, 105°47’49.8” E, about 10 km to the NW of Kasi town, 19°12’57” 1000–1300 m, 9 March 2013, L. Averyanov et al., N, 102°14’59” E, 500–650 m, 24 March 2013, L. LA-VN 298, Fig. 1, A (CPC, NHOL, NUOL). Averyanov, N.S. Khang, S. Lorphengsy, LA-VN 745 (CPC, NHOL, NUOL). Acriopsis Reinw. ex Blume A. indica C. Wright Appendicula Blume LAOS, Saravan prov., Ta Oy distr., to the A. hexandra (J. Koenig) J.J. Sm. SW of Sa Mouay town, near Thong Hang village, CAMBODIA, Bokor, 24 Nov. 2012, T. 15°37’30.9” N, 106°12’26.5” E, 650–700 m, 4 Maisak et al., 922 (LE). March 2013, L. Averyanov et al., LA-VN 217 (CPC, A. torta Blume NHOL, NUOL). LAOS, Vientiane prov., Vang LAOS, Vientiane prov., Vang Vieng distr., Vieng distr., Phoudindeng village, 1–2,5 km to the E Phol Xai village, about 8 km to the W of Vang Vieng of Vang Vieng town, 18°57’48.7” N, 102°27’34.0” town, Phar Poon mt., 18°56’11.5” N, 102°20’05.0” E, 400 m, 13 March 2013, N.T. Hiep et al., LA-VN E, 800–900 m, 15 March 2013, N.T. Hiep et al., LA- 343 (CPC, NHOL, NUOL). VN 474 (CPC, NHOL, NUOL). +A. liliifolia (J.Koenig) Seidenf. CAMBODIA, to the N of Sianukwille Arachnis Blume town, Kaoh Rong Samoloem Khong Island in Siam A. labrosa (Lindl. et Paxton) Rchb. f. Gulf, 10°36’45” N, 103°18’38” E, 20 Nov. 2011, T. LAOS, Vientiane prov., Vang Vieng distr., Maisak, L. Osinovez, M. Telepova s.n. (LE). Na Khun Village, around point 18°52’28” N, 102°24’21” E, 300 m, 21 Jan. 2009, O. Souliya Aerides Lour. et al., LA-VN 81 (CPC). LAOS, Attapeu prov., +A. crassifolia Par. ex Burb. Saysettha distr., Dong Am Pham N.P.A., to the CAMBODIA, eastern part of Cardamom N of Paosamphanmixay village, 14°51’53.4” N, Mountains, Phnom Aural mt, 23 Nov. 2012. T. 107°11’38.2” E, 178 m, 28 February 2013, L. Maisak et al., 961 (LE). Averyanov et al., LA-VN 116 (CPC, NHOL, NUOL). A. rosea Lodd. LAOS, Saravan prov., Sa Mouay distr., to the SEE LAOS, Vientiane prov., Kasi distr., Tam of Sa Mouay town, elev. from Ta Oy village, 700 Tai village, about 10 km to the NW of Kasi town, m, 16°08’47.0” N, 106°52’35.4” E, 4 March 2013, 19°12’57” N, 102°14’59” E, 500–650 m, 24 March L. Averyanov et al., LA-VN 164 (CPC, NHOL, 2013, L. Averyanov, N.S. Khang, S. Lorphengsy, LA- NUOL). LAOS, Vientiane prov., Kasi distr., Thong VN 758 (CPC, NHOL, NUOL). Mout village, 19°26’51.5” N, 102°07’45.8” E,

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1550–1650 m, 20 March 2013, N.T. Hiep et al., LA- Protected Authority (WMPI) region, 17°57’32” N, VN 608 (CPC, NHOL, NUOL). 105°01’05” E, 538 m, 18 Jan. 2009, S. Bounphanmy et al., LA-VN 19 (CPC). LAOS, Saravan prov., Ta Arundina Blume Oy distr., to the SW of Sa Mouay town, near Thong A. graminifolia (D. Don) Hochr. Hang village, 15°37’30.9” N, 106°12’26.5” E, 650– CAMBODIA, Kampot prov., Phnom Bokor 700 m, 4 March 2013, L. Averyanov et al., LA-VN N.P., 10°37’92.3 N, 104°01’14.3 E, 21 Dec. 2012, 215 (CPC, NHOL, NUOL). E.L. Konstantinov et al., 45P-C-GPS042 (LE). LAOS, **B. konstantinovii Aver., sp. nov. (sect. Attapeu prov., Saysettha distr., Dong Am Pham N.P.A., Macrocaulia (Blume) Aver.) to the E of Paosamphanmixay village, 14°48’24.8” Described from S. CAMBODIA N, 107°24’23.2” E, 600–700 m, 28 Feb. 2013, L. (“CAMBODIA, Kampot prov., Phnom Bokor N.P. Averyanov et al., LA-VN 111 (CPC, NHOL, NUOL). Epiphyte. 10°37’92.3 N, 104°01’14.3 E). Type (“21 Dec. 2012. E.L. Konstantinov et al., 051P-C- Brachypeza Garay GPS042”) – LE (holotype). B. laotica (Seidenf.) Seidenf. Dwarf creeping epiphyte. Rhizome thin, LAOS, Vientiane prov., Vang Vieng distr., wiry, young greenish, later whitish-gray, 0.5–1 mm Nathong village, about 5 km to the W of Vang Vieng wide, with numerous pseudobulbs distant on 1–5 cm town, Tham Kang mt., 18°55’51.4” N, 102°23’50.6” and 3–4 wiry, whitish flexuose roots at pseudobulb E, 300–400 m, 14 March 2013, N.T. Hiep et al., LA- base. Pseudobulbs light olive, yellowish-green to VN 425 (CPC, LE, NHOL, NUOL). green, ovate, 4–8(10) mm tall, 2–6 mm wide, often oblique, slightly bending to rhizome, longitudinally Bromheadia Lindl. irregularly grooved and wrinkled, with 1 apical B. finlaysoniana (Lindl.) Miq. leaf. Leaves coriaceous, narrowly ovate to ovate, CAMBODIA, Kampot prov., Phnom Bokor 8–22 × 3–8 mm, conduplicate, obtuse, subsessile or N.P., 10°39’63.9 N, 104°03’24.8 E, 17 Dec. 2011, with very short petiole. Inflorescence arising from E.L. Konstantinov et al., 2b (LE). the base of pseudobulb, 1.5–2.5 cm long, with 1 terminal flower, on ascending, filiform, sometime Bulbophyllum Thouars slightly flexuose, light yellowish-green, glabrous B. affine Wall. ex Lindl. stalk 1–1.8 cm long, with one small bract at the base. LAOS, Saravan prov., Ta Oy distr., to the SW of Bract hyaline, goblet-shaped, 0.5–1 mm long, 0.3 Sa Mouay town, near Thong Hang village, 15°37’30.9” mm in diam., embracing apex of inflorescence stalk. N, 106°12’26.5” E, 650–700 m, 4 March 2013, L. Pedicel and ovary light green, glabrous, filiform, Averyanov et al., LA-VN 219 (CPC, NHOL, NUOL). 5–10 mm long; ovary 0.2 mm wide. Flowers *B. apodum Hook. f. widely opening, 1 cm across spreading lateral LAOS, Vientiane prov., Vang Vieng distr., sepals. Sepals sub-similar, light dull-yellowish with Phol Xai village, about 8 km to the W of Vang Vieng broad purple-brown stripes, 5–6 × 1.5–1.8 mm, town, Phar Poon mt., 18°56’11.5” N, 102°20’05.0” acute, with 3 distinct nerves; median sepal elliptic, E, 800–900 m, 15 March 2013, N.T. Hiep et al., LA- directed forward; lateral sepals narrowly ovate, VN 482 (CPC, NHOL, NUOL). LAOS, Vientiane spreading, slightly longer and wider, oblique at prov., Kasi distr., Tam Tai village, about 10 km to the the base. Petals directed forward, brightly yellow, NW of Kasi town, 19°12’57” N, 102°14’59” E, 500– oblique ovate, acute, 1.6–2 × 1–1.2 mm, with 1 650 m, 24 March 2013, L. Averyanov, N.S. Khang, S. nerve. Lip brightly yellow, simple, without auricles, Lorphengsy, LA-VN 765 (CPC, NHOL, NUOL). elliptic, 3–4 × 1–1.5 mm, at the base with narrowing B. blepharistes Rchb. f. strongly bending neck, jointed with column foot LAOS, Attapeu prov., Saysettha distr., Dong apex by semi-movable articulation; lip blade with 3 Am Pham N.P.A., to the N of Paosamphanmixay thick nerves, fleshy along margin. Column brightly village, 14°51’53.4” N, 107°11’38.2” E, 200 m, 1 yellow, erect, 0.8 × 0.5 mm, at the apex with 2 long March 2013, L. Averyanov et al., LA-VN 133 (CPC, erect, straight, acute stelidia 0.8 mm long; column NHOL, NUOL). LAOS, Saravan area, 100–200 foot forward curved, apically broadening into cup- m, 6 March 2013, L. Averyanov et al., LA-VN 325 like cordate apex 1 mm long and wide. Operculum (CPC, NHOL, NUOL). yellow, cup-like, half ovoid, 0.4 mm long, with short B. dissitiflorum Seidenf. broad beak. Pollinia 2, globular. Fig. 1, C; 2. LAOS, Borikhamxai prov., Khamkeut distr., Etymology: Species is named after its Dan Khum Ngan locality, Watershed Managing discoverer – Dr. E.L. Konstantinov.

30 Averyanov L.V. New and rare orchids (Orchidaceae) in the flora of Cambodia and Laos

Flowering time: November – December. Thong Mout village, Pudang Ridge, 19°23’33.7” N, Ecology: Epiphyte, 100–200 m a.s.l. 102°08’05.7” E, 1950–2000 m, 22 March 2013, L. Distribution: S. CAMBODIA, Kampot Averyanov, N.S. Khang, S. Lorphengsy, LA-VN 703 prov. (Phnom Bokor N.P.). Endemic. (CPC, NHOL, NUOL). Notes: Morphologically new species may +B. rufilabrum Par. ex Hook. f. be most close to Bulbophyllum simplicilabellum CAMBODIA, Koh Rong island, 10°45’90.9 Seidenf. and B. didymotropis Seidenf. described N, 103°13’33.1 E, 15 Dec. 2011, E. Konstantinov et from Thailand, but distinctly differs in 3-nerved al., 3 (LE). LAOS, Phu Khao Khuay N.P., Tad Leuk sepals, broadly ovate, oblique, acute petals, broad waterfall, 18°23’72.3 N, 103°04’32.8 E, 12 Nov. cup-like, cordate broadening of column foot apex 2011, E. Konstantinov et al., 1a (LE). LAOS, Nam and inflated fleshy margin of the lip. Lik, 18°36’80.8 N, 102°24’60.5 E, 24 Nov. 2011, B. lasiochilum Parish et Rchb. f. E. Konstantinov et al., 4 (LE). LAOS, Vientiane LAOS, Borikhamxai prov., Khamkeut distr., prov., Phou Khao Khouay N.P., Tad Leuk waterfall, Phone Sa At village, 18°03’38” N, 105°02’28” E, 18°23’99.1 N, 103°04’03.1 E, 28 Nov. 2012, E.L. 635 m, 18 Jan. 2009, S. Bounphanmy et al., LA-VN Konstantinov et al., 53OR-L-GPS005 (LE). 26 (CPC, LE). B. rufinum Rchb. f. B. orectopetalum Garay, Hamer et Siegerist LAOS, Attapeu prov., Saysettha distr., Dong CAMBODIA, Kampot prov., Phnom Bokor Am Pham N.P.A., to the N of Paosamphanmixay N.P., 10°37’92.3 N, 104°01’14.3 E, 21 Dec. 2012, village, 14°51’53.4” N, 107°11’38.2” E, 200 m, 1 E.L. Konstantinov et al., 28P-C-GPS042 (LE). March 2013, L. Averyanov et al., LA-VN 134 (CPC, LAOS, Vientiane prov., Vang Vieng distr., Phol Xai NHOL, NUOL). village, about 8 km to the W of Vang Vieng town, +B. tricornoides Seidenf. Phar Poon mt., 18°56’11.5” N, 102°20’05.0” E, CAMBODIA, to the N of Sianukwille 800–900 m, 15 March 2013, N.T. Hiep et al., LA- town, Kaoh Rong Samoloem Khong Island in Siam VN 464 (CPC, NHOL, NUOL). LAOS, Vientiane Gulf, 10°36’45” N, 103°18’38” E, 20 Nov. 2011, T. prov., Kasi distr., Namken village, Phachao mt., Maisak, L. Osinovez, M. Telepova, s.n. (LE). 19°18’45.5” N, 102°22’31.4” E, 1200 m, 24 March *B. umbellatum Lindl. 2013, L. Averyanov, N.S. Khang, S. Lorphengsy, LA- LAOS, Vientiane prov., Kasi distr., Thong VN 790 (CPC, NHOL, NUOL). Mout village, 19°26’51.5” N, 102°07’45.8” E, +B. parviflorum Par. et Rchb. f. 1550–1650 m, 20 March 2013, N.T. Hiep et al., LA- CAMBODIA, to the N of Sianukwille town, VN 612, Fig. 1, F (CPC, LE, NHOL, NUOL). Kaoh Rong Samoloem Khong Island in Siam Gulf, 10°36’45” N, 103°18’38” E, 19 Nov. 2011, T. Maisak, Calanthe Ker Gawl. M. Telepova, L. Osinovets, 847, Fig. 1, D (LE). +C. cardioglossa Schltr. *B. psychoon Rchb. f. CAMBODIA, Kampot prov., Phnom Bokor LAOS, Kham Mouan prov., Bounlapha N.P., 10°37’92.3 N, 104°01’14.3 E, 21 Dec. 2012, E.L. distr., Thong Sam village, Hinnam No N.P.A., Pu Konstantinov et al., 44P-C-GPS042, Fig. 1, G (LE). Pha Song mt., 17°35’13.2” N, 105°47’49.8” E, 800 m, 9 March 2013, L. Averyanov et al., LA-VN Callostylis Blume 319 (CPC, LE, NHOL, NUOL). LAOS, Vientiane C. rigida Blume prov., Kasi distr., Thong Mout village, 19°26’51.5” LAOS, Vientiane prov., Vang Vieng distr., N, 102°07’45.8” E, 1550–1650 m, 20 March 2013, Phol Xai village, about 8 km to the W of Vang Vieng N.T. Hiep et al., LA-VN 613, Fig. 1, E (CPC, LE, town, Phar Poon mt., 18°56’11.5”N 102°20’05.0”E, NHOL, NUOL). LAOS, Vientiane prov., Kasi 800-900 m, 15 March 2013, N.T. Hiep et al., LA-VN distr., Namken village, Phachao mt., 19°18’45.5” 476 (CPC, NHOL, NUOL). N, 102°22’31.4” E, 1600 m, 24 March 2013, L. Averyanov, N.S. Khang, S. Lorphengsy LA-VN 795 Cephalantheropsis Guillaumin (CPC, NHOL, NUOL). C. obcordata (Lindl.) Ormerod B. reptans (Lindl.) Lindl. ex Wall. LAOS, Saravan prov., Sa Mouay distr., LAOS, Saravan prov., Sa Mouay distr., to the SEE of Sa Mouay town, elev. from Ta Oy to the SEE of Sa Mouay town, elev. from Ta Oy village, 1350 m, 16°08’47.0” N, 106°52’35.4” E, 4 village, 1350 m, 16°08’47.0” N, 106°52’35.4” E, 4 March 2013, L. Averyanov et al., LA-VN 160 (CPC, March 2013, L. Averyanov et al., LA-VN 175 (CPC, NHOL, NUOL). NHOL, NUOL). LAOS, Vientiane prov., Kasi distr.,

Turczaninowia 2013, 16 (4) : 26–46 31

Fig. 1. New orchids in the flora of Cambodia and Laos. A – Acanthephippium striatum Lindl. (L. Averyanov et al., LA-VN 298). B – Agrostophyllum callosum Rchb. f. (L. Averyanov et al., LA-VN 323). C – Bulbophyllum konstantinovii Aver. (type specimen, E.L. Konstantinov et al., 051P-C-GPS042). D – B. parviflorum Par. et Rchb. f. (T. Maisak, M. Telepova, L. Osinovets, 847). E – B. psychoon Rchb. f. (N.T. Hiep et al., LA-VN 613). F – B. umbellatum Lindl. (N.T. Hiep et al., LA-VN 612). G: Calanthe cardioglossa Schltr. (E.L. Konstantinov et al., 44P-C-GPS042). H – Cheirostylis griffithii Lindl. (L. Averyanov, N.S. Khang, S. Lorphengsy, LA-VN 782). I – C. pusilla Lindl. (E.L. Konstantinov et al., 14bOR-L-GPS018). Photos by Nguyen Sinh Khang (A, H), Pham Van The (B), E. Konstantinov (C, G, I) and L. Averyanov (D, E, F).

32 Averyanov L.V. New and rare orchids (Orchidaceae) in the flora of Cambodia and Laos

Fig. 2. Bulbophyllum konstantinovii Aver. a – ; b – flattened flower, half-side view; c – flattened flower, frontal view; d – lip, half-side view; e – column, side view. All drawings from the type (Konstantinov et al. 051P-C-GPS042) by L. Averyanov. Ceratostylis Blume *C. griffithii Lindl. C. himalaica Hook. f. LAOS, Vientiane prov., Kasi distr., Namken LAOS, Saravan prov., Sa Mouay distr., village, Phachao mt., 19°18’45.5” N, 102°22’31.4” to the SEE of Sa Mouay town, elev. from Ta Oy E, 1500–1750 m, 24 March 2013, L. Averyanov, N.S. village, 1350 m, 16°08’47.0” N, 106°52’35.4” E, 4 Khang, S. Lorphengsy, LA-VN 782, Fig. 1, H (CPC, March 2013, L. Averyanov et al., LA-VN 174 (CPC, LE, NHOL, NUOL). NHOL, NUOL). *C. pusilla Lindl. C. subulata Blume LAOS, Luang Namtha prov., Ban Nam CAMBODIA, Kampot prov., Phnom Bokor Eng, 21°43’49.7 N, 101°09’23.9 E, 3 Dec. 2012, E.L. N.P., 10°37’92.3 N, 104°01’14.3 E, 21 Dec. 2012, Konstantinov et al., 14bOR-L-GPS018, Fig. 1, I (LE). E.L. Konstantinov et al., 27P-C-GPS042 (LE). *C. spathulata J.J. Sm. LAOS, Vientiane prov., Vang Vieng distr., Cheirostylis Blume Phol Xai village, about 8 km to the W of Vang Vieng C. chinensis Rolfe town, Phar Poon mt., 18°56’11.5” N, 102°20’05.0” LAOS, Vientiane prov., Vang Vieng distr., E, 300–900 m, 15 March 2013, N.T. Hiep et al., Na Khun Village, 18°52’28” N, 102°24’21” E, 300 LA-VN 487 (CPC, LE, NHOL, NUOL). LAOS, m, 21 Jan. 2009, O. Souliya et al., LA-VN 78 (CPC). Vientiane prov., Vang Vieng distr., Na Po village,

Turczaninowia 2013, 16 (4) : 26–46 33 about 12 km to the W of Vang Vieng town, Pa Nang Phachao mt., 19°18’45.5” N, 102°22’31.4” E, Oua mt., 18°55’36.4” N, 102°20’44.6” E, 350–450 1500–1750 m, 24 March 2013, L. Averyanov, N.S. m, 17 March 2013, N.T. Hiep et al., LA-VN 509, Fig. Khang, S. Lorphengsy, LA-VN 768 (CPC, LE, 3, A (CPC, NHOL, NUOL). LAOS, Vientiane prov., NHOL, NUOL). Kasi distr., Tam Tai village, about 10 km to the NW *C. virescens Rolfe of Kasi town, 19°12’57” N, 102°14’59” E, 500–650 LAOS, Attapeu prov., Saysettha distr., Dong m, 24 March 2013, L. Averyanov, N.S. Khang, S. Am Pham N.P.A., to the N of Paosamphanmixay Lorphengsy, LA-VN 734 (CPC, NHOL, NUOL). village, 14°51’53.4” N, 107°11’38.2” E, 178 m, 28 *C. thailandica Seidenf. Feb. 2013, L. Averyanov et al., LA-VN 113, Fig. 3, C LAOS, Vientiane prov., Vang Vieng distr., (CPC, LE, NHOL, NUOL). Nathong village, about 5 km to the W of Vang Vieng C. viscosa Rchb. f. town, 18°55’34.2” N, 102°22’58.4” E, 300 m, 13 LAOS, Vientiane prov., Vang Vieng distr., March 2013, N.T. Hiep et al., LA-VN 351a, Fig. 3, Na Khun Village, 18°52’28” N, 102°24’21” E, 400– B (CPC, LE, NHOL, NUOL). LAOS, Vientiane 600 m, 21 Jan. 2009, O. Souliya et al., LA-VN 101 prov., Vang Vieng distr., Na Po village, about 12 km (CPC). to the W of Vang Vieng town, Pa Nang Oua mt., 18°55’44.9” N, 102°20’20.1” E, 500–1000 m, 17 Collabium Blume March 2013, N.T. Hiep et al., LA-VN 513 (CPC, LE, C. chinense (Rolfe) Tang et F.T. Wang NHOL, NUOL). LAOS, Kham Mouan prov., Bounlapha distr., Thong Sam village, Hinnam No N.P.A., Pu Cleisostoma Raf. Pha Song mt., 17°35’13.2” N, 105°47’49.8” E, 900 C. fuerstenbergianum Kraenzl. m, 9 March 2013, L. Averyanov et al., LA-VN 304 LAOS, Borikhamxai prov., Khamkeut distr., (CPC, NHOL, NUOL). Phone Sa At village, 18°03’38” N, 105°02’28” E, 635 m, 18 Jan. 2009, S. Bounphanmy et al., LA-VN Corymborkis Thouars 37 (CPC). C. veratrifolia (Reinw.) Blume C. rostratum (Lindl.) Garay LAOS, Vientiane prov., Vang Vieng distr., LAOS, Vientiane prov., Phou Khao Khouay Na Khun Village, 18°52’28” N, 102°24’21” E, 300 N.P., Tad Leuk waterfall, 18°23’99.1 N, 103°04’03.1 m, 21 Jan. 2009, O. Souliya et al., LA-VN 74 (CPC). E, 28 Nov. 2012, E.L. Konstantinov et al., 11OR-L- LAOS, Vientiane prov., Vang Vieng distr., Na Po GPS005 (LE). LAOS, Vientiane prov., Vang Vieng village, about 12 km to the W of Vang Vieng town, distr., Nathong village, about 5 km to the W of Pa Nang Oua mt., 18°55’36.4” N, 102°20’44.6” E, Vang Vieng town, Tham Kang mt., 18°55’51.4” N, 350–450 m, 17 March 2013, N.T. Hiep et al., LA- 102°23’50.6” E, 300–500 m, 14 March 2013, N.T. VN 494 (CPC, NHOL, NUOL). LAOS, Vientiane Hiep et al., LA-VN 384 (CPC, NHOL, NUOL). prov., Kasi distr., Namken village, Phachao mt., C. simondii (Gagnep.) Seidenf. 19°18’45.5” N, 102°22’31.4” E, 650–800 m, LAOS, Phu Khao Khuay N.P., Tad Leuk 24 March 2013, L. Averyanov, N.S. Khang, S. waterfall, 18°23’72.3 N, 103°04’32.8 E, 12 Nov. Lorphengsy LA-VN 789 (CPC, NHOL, NUOL). 2011, E. Konstantinov et al., 1b (LE). Cymbidium Sw. Coelogyne Lindl. C. aloifolium (L.) Sw. C. fimbriata Lindl. LAOS, Kham Mouan prov., Bounlapha LAOS, Saravan prov., Sa Mouay distr., distr., Nam Boua Sam village, Hinnam No N.P.A., to the SEE of Sa Mouay town, elev. from Ta Oy Pu Pha Song mt., 17°33’43.6” N, 105°45’08.3” E, village, 1350 m, 16°08’47.0” N, 106°52’35.4” E, 4 230 m, 10 March 2013, L. Averyanov et al., LA- March 2013, L. Averyanov et al., LA-VN 154 (CPC, VN 328 (CPC, NHOL, NUOL). LAOS, Vientiane NHOL, NUOL). prov., Vang Vieng distr., Nathong village, about 5 C. flaccida Lindl. km to the W of Vang Vieng town, Tham Kang mt., LAOS, Vientiane prov., Vang Vieng distr., 18°55’51.4” N, 102°23’50.6” E, 300–500 m, 14 Na Po village, about 12 km to the W of Vang March 2013, N.T. Hiep et al., LA-VN 390 (CPC, Vieng town, Pa Nang Oua mt., 18°55’44.9” N, NHOL, NUOL). 102°20’20.1” E, 500–1000 m, 17 March 2013, N.T. *C. eburneum Lindl. var. eburneum Hiep et al., LA-VN 515 (CPC, LE, NHOL, NUOL). LAOS, Vientiane prov., Kasi distr., Namken LAOS, Vientiane prov., Kasi distr., Namken village, village, Phachao mt., 19°18’45.5” N, 102°22’31.4”

34 Averyanov L.V. New and rare orchids (Orchidaceae) in the flora of Cambodia and Laos

E, 1500–1750 m, 24 March 2013, L. Averyanov, N.S. March 2013, L. Averyanov et al., LA-VN 130 (CPC, Khang, S. Lorphengsy, LA-VN 804, Fig. 3, D (CPC, NHOL, NUOL). LE, NHOL, NUOL). D. draconis Rchb. f. *C. eburneum Lindl. var. longzhouense CAMBODIA, eastern part of Cardamom Z.J. Liu et S.C. Chen Mountains, Phnom Aural mt., 23 Nov. 2012, T. LAOS, Vientiane prov., Kasi distr., Namken Maisak et al., 944 (LE). village, Phachao mt., 19°18’45.5” N, 102°22’31.4” D. findleyanum Par. et Rchb. f. E, 1500–1750 m, 24 March 2013, L. Averyanov, N.S. LAOS, Vientiane prov., Kasi distr., Namken Khang, S. Lorphengsy LA-VN 803, Fig. 3, E (CPC, village, Phachao mt., 19°18’45.5” N, 102°22’31.4” LE, NHOL, NUOL). E, 1600 m, 24 March 2013, L. Averyanov, N.S. C. lancifolium Hook. Khang, S. Lorphengsy LA-VN 792 (CPC, NHOL, LAOS, Saravan prov., Sa Mouay distr., to NUOL). the SEE of Sa Mouay town, elev. from Ta Oy village, D. gratiosissimum Rchb. f. 1350 m, 16°08’47.0” N, 106°52’35.4” E, 4 March LAOS, Borikhamxai prov., Khamkeut 2013, L. Averyanov et al., LA-VN 177 (CPC, NHOL, distr., Dan Khum Ngan locality, Watershed NUOL). LAOS, Kham Mouan prov., Bounlapha Managing Protected Authority region, 17°57’32” N, distr., Thong Sam village, Hinnam No N.P.A., Pu 105°01’05” E, 538 m, 18 Jan. 2009, S. Bounphanmy Pha Song mt., 17°35’13.2” N, 105°47’49.8” E, 1100 et al., LA-VN 21b (CPC, LE). LAOS, Vientiane m, 9 March 2013, L. Averyanov et al., LA-VN 300 prov., Kasi distr., Namken village, Phachao mt., (CPC, NHOL, NUOL). LAOS, Vientiane prov., Kasi 19°18’45.5” N, 102°22’31.4” E, 1200 m, 24 March distr., Namken village, Phachao mt., 19°18’45.5” N, 2013, L. Averyanov, N.S. Khang, S. Lorphengsy, LA- 102°22’31.4” E, 1500–1750 m, 24 March 2013, L. VN 772 (CPC, LE, NHOL, NUOL). Averyanov, N.S. Khang, S. Lorphengsy LA-VN 786 +D. indivisum (Blume) Miq. (CPC, NHOL, NUOL). CAMBODIA, eastern part of Cardamom C. lowianum (Rchb. f.) Rchb. f. Mountains, Phnom Aural mt., 23 Nov. 2012, T. LAOS, Vientiane prov., Kasi distr., Thong Maisak et al., 953, 955, Fig. 3, F (LE). Mout village, 19°26’51.5” N, 102°07’45.8” E, +D. keithii Ridl. 1550–1650 m, 20 March 2013, N.T. Hiep et al., LA- CAMBODIA, to the N of Sianukwille town, VN 604 (CPC, LE, NHOL, NUOL). Kaoh Rong Samoloem Khong Island in Siam Gulf, 10°36’45” N, 103°18’38” E, 20 Nov. 2011, T. Maisak, Dendrobium Sw. L. Osinovez, M. Telepova, s.n., Fig. 3, G (LE). D. cariniferum Rchb. f. *D. kontumense Gagnep. LAOS, Borikhamxai prov., Khamkeut distr., LAOS, Borikhamxai prov., Khamkeut Phone Sa At village, 18°03’38” N, 105°02’28” E, distr., Dan Khum Ngan locality, Watershed 635 m, 18 Jan. 2009, S. Bounphanmy et al., LA-VN Managing Protected Authority region, 17°57’32” N, 29 (CPC). LAOS, Phongsali prov., Muong May 105°01’05” E, 538 m, 18 Jan. 2009, S. Bounphanmy distr., 21°12’32” N, 102°53’44” E, 1250–1400 m, 22 et al., LA-VN 21a, Fig. 3, H (CPC, LE). April 2011, L. Averyanov, P.V. The, CPC 2429 ((CPC). D. monticola Hunt et Summerh. D. crepidatum Lindl. LAOS, Borikhamxai prov., Khamkeut distr., LAOS, Vientiane prov., Vang Vieng distr., Phone Sa At village, 18°02’09” N, 105°02’31” E, Phol Xai village, about 8 km to the W of Vang Vieng 618 m, 18 Jan. 2009, S. Bounphanmy et al., LA-VN town, Phar Poon mt., 18°56’11.5” N, 102°20’05.0” 39 (CPC). E, 800–900 m, 15 March 2013, N.T. Hiep et al., LA- D. multilineatum Kerr VN 481 (CPC, NHOL, NUOL). LAOS, Kham Mouan prov., Bounlapha D. cumulatum Lindl. distr., Thong Sam village, Hinnam No N.P.A., Pu CAMBODIA, to the N of Sianukwille Pha Song mt., 17°35’13.2” N, 105°47’49.8” E, town, Kaoh Rong Samoloem Khong Island in Siam 1300–1425 m, 9 March 2013, L. Averyanov et al., Gulf, 10°36’45” N, 103°18’38” E, 20 Nov. 2011, T. LA-VN 256 (CPC, LE, NHOL, NUOL). Maisak, L. Osinovez, M. Telepova, 835, 836 (LE). D. nathanielis Rchb. f. D. dalacourii Guillaumin LAOS, Attapeu prov., Saysettha distr., Dong LAOS, Attapeu prov., Saysettha distr., Dong Am Pham N.P.A., to the N of Paosamphanmixay Am Pham N.P.A., to the N of Paosamphanmixay village, 14°51’53.4” N, 107°11’38.2” E, 178 m, 28 village, 14°51’53.4” N, 107°11’38.2” E, 200 m, 1 Feb. 2013, L. Averyanov et al., LA-VN 114 (CPC, LE, NHOL, NUOL).

Turczaninowia 2013, 16 (4) : 26–46 35

Fig. 3. New orchids in the flora of Cambodia and Laos. A – Cheirostylis spathulata J.J. Sm. (N.T. Hiep et al., LA-VN 509). B – C. thailandica Seidenf. (N.T. Hiep et al., LA-VN 351a). C – Coelogyne virescens Rolfe (L. Averyanov et al., LA-VN 113). D – Cymbidium eburneum Lindl. var. eburneum (L. Averyanov, N.S. Khang, S. Lorphengsy, LA- VN 804). E – C. eburneum Lindl. var. longzhouense Z.J. Liu et S.C. Chen (L. Averyanov, N.S. Khang, S. Lorphengsy LA-VN 803). F – Dendrobium indivisum (Blume) Miq. (T. Maisak et al., 953, 955). G – D. keithii Ridl. (T. Maisak, L. Osinovez, M. Telepova, s.n.). H – D. kontumense Gagnep. (S. Bounphanmy et al., LA-VN 21a). I – D. scabrilingue Lindl. (E.L. Konstantinov et al., 29P-C-GPS042). Photos by L. Averyanov (A, B, C, F, G, H), Nguyen Sinh Khang (D, E) and E. Konstantinov (I).

36 Averyanov L.V. New and rare orchids (Orchidaceae) in the flora of Cambodia and Laos

D. oligophyllum Gagnep. Doritis Lindl. LAOS, Vientiane prov., Vang Vieng distr., D. pulcherrima Lindl. Nathong village, about 5 km to the W of Vang Vieng LAOS, Borikhamxai prov., Khamkeut town, Tham Kang mt., 18°55’51.4” N, 102°23’50.6” distr., Dan Khum Ngan locality, Watershed E, 300–500 m, 14 March 2013, N.T. Hiep et al., LA- Managing Protected Authority region, 17°57’32” N, VN 383 (CPC, LE, NHOL, NUOL). 105°01’05” E, 538 m, 18 Jan. 2009, S. Bounphanmy D. parcum Rchb. f. et al., LA-VN 21 (CPC). LAOS, Borikhamxai prov., Khamkeut distr., Dan Khum Ngan locality, Watershed *Eclecticus P. O’Byrne Managing Protected Authority region, 17°57’32” N, *E. chungii P. O’Byrne 105°01’05” E, 538 m, 18 Jan. 2009, S. Bounphanmy LAOS, Vientiane prov., Vang Vieng distr., et al., LA-VN 20 (CPC). LAOS, Saravan prov., Ta Nathong village, about 5 km to the W of Vang Vieng Oy distr., to the SW of Sa Mouay town, near Thong town, Tham Kang mt., 18°55’51.4” N, 102°23’50.6” Hang village, 15°37’30.9”N 106°12’26.5”E, 650- E, 300–500 m, 14 March 2013, N.T. Hiep et al., LA- 700 m, 4 March 2013, L. Averyanov et al., LA-VN VN 385c, Fig. 4, C, D (LE). 222 (CPC, NHOL, NUOL). +D. scabrilingue Lindl. Epigeneium Gagnep. CAMBODIA, Kampot prov., Phnom Bokor *E. chapaense Gagnep. N.P., 10°37’92.3 N, 104°01’14.3 E, 21 Dec. 2012, E.L. LAOS, Saravan prov., Sa Mouay distr., Konstantinov et al., 29P-C-GPS042, Fig. 3, I (LE). to the SEE of Sa Mouay town, elev. from Ta Oy D. secundum (Blume) Lindl. village, 1350 m, 16°08’47.0” N, 106°52’35.4” E, 4 LAOS, Attapeu prov., Saysettha distr., Dong March 2013, L. Averyanov et al., LA-VN 193 (CPC, Am Pham N.P.A., to the N of Paosamphanmixay NHOL, NUOL). village, 14°51’53.4” N, 107°11’38.2” E, 200 m, 1 March 2013, L. Averyanov et al., LA-VN 132 (CPC, Epipogium J.G. Gmel. ex Borkh. NHOL, NUOL). E. roseum (D. Don) Lindl. +D. setifolium Ridl. LAOS, Vientiane prov., Vang Vieng distr., CAMBODIA, Kampot prov., Phnom Bokor Phol Xai village, about 8 km to the W of Vang Vieng N.P., 10°37’92.3 N, 104°01’14.3 E, 21 Dec. 2012. town, Phar Poon mt., 18°56’11.5” N, 102°20’05.0” E.L. Konstantinov et al., 35P-C-GPS042 (LE), E, 800–900 m, 15 March 2013, N.T. Hiep et al., LA- 42P-C-GPS042, Fig. 4, A (LE). VN 465 (CPC, NHOL, NUOL). LAOS, Vientiane D. signatum Rchb. f. prov., Vang Vieng distr., Nam Pe village, about 12 LAOS, Borikhamxai prov., Khamkeut km to the W of Vang Vieng town, 18°58’41.0” N, distr., Dan Khum Ngan locality, Watershed 102°18’54.1” E, 400 m, 18 March 2013, N.T. Hiep Managing Protected Authority region, 17°57’32” N, et al., LA-VN 586 (CPC, NHOL, NUOL). 105°01’05” E, 538 m, 18 Jan. 2009, S. Bounphanmy et al., LA-VN 21d (CPC, LE). Eria Lindl. D. spatella Rchb. f. E. boniana (Gagnep.) Tang et F.T. Wang LAOS, Borikhamxai prov., Khamkeut distr., LAOS, Vientiane prov., Kasi distr., Thong Phone Sa At village, 18°03’38” N, 105°02’28” E, 635 m, Mout village, 19°26’51.5” N, 102°07’45.8” E, 18 Jan. 2009, S. Bounphanmy et al., LA-VN 38 (CPC). 1550–1650 m, 20 March 2013, N.T. Hiep et al., LA- D. wardianum R. Warner VN 609 (CPC, NHOL, NUOL). LAOS, Vientiane prov., Kasi distr., Thong *E. calcarea V.N. Long et Aver. Mout village, Pudang Ridge, 19°23’33.7” N, LAOS, Vientiane prov., Vang Vieng distr., 102°08’05.7” E, 1950–2000 m, 22 March 2013, L. Phol Xai village, about 8 km to the W of Vang Vieng Averyanov, N.S. Khang, S. Lorphengsy, LA-VN 705 town, Phar Poon mt., 18°56’11.5” N, 102°20’05.0” (CPC, LE, NHOL, NUOL). E, 800–900 m, 15 March 2013, N.T. Hiep et al., LA- *Didymoplexiopsis Seidenf. VN 480 (CPC, NHOL, NUOL). *D. khiriwongensis Seidenf. *E. corneri Rchb. f. LAOS, Attapeu prov., Saysettha distr., Dong LAOS, Vientiane prov., Vang Vieng distr., Am Pham N.P.A., to the E of Paosamphanmixay Na Khun Village, 18°52’28” N, 102°24’21” E, 300 village, 14°48’24.8” N, 107°24’23.2” E, 700–20 m, m, 21 Jan. 2009, O. Souliya et al., LA-VN 64 (CPC). 28 Feb. 2013, L. Averyanov et al., LA-VN 110, Fig. LAOS, Vientiane prov., Kasi distr., Tam Tai village, 4, B (LE). about 10 km to the NW of Kasi town, 19°12’57”

Turczaninowia 2013, 16 (4) : 26–46 37

N, 102°14’59” E, 500–650 m, 24 March 2013, L. Vieng town, Pa Nang Oua mt., 18°55’44.9” N, Averyanov, N.S. Khang, S. Lorphengsy LA-VN 764 102°20’20.1” E, 700–1000 m, 17 March 2013, N.T. (CPC, NHOL, NUOL). Hiep et al., LA-VN 527a (LE). +E. lasiopetala (Willd.) Ormerod E. spirodela Aver. CAMBODIA, Kampot prov., Phnom Bokor LAOS, Attapeu prov., Saysettha distr., Dong N.P., 10°37’92.3 N, 104°01’14.3 E, 21 Dec. 2012, Am Pham N.P.A., to the E of Paosamphanmixay E.L. Konstantinov et al., 39P-C-GPS042, Fig. 4, village, 14°48’24.8” N, 107°24’23.2” E, 700–920 E (LE). LAOS, Borikhamxai prov., Khamkeut m, 28 February 2013, L. Averyanov et al., LA-VN distr., Dan Khum Ngan locality, Watershed 106 (CPC, NHOL, NUOL). LAOS, Kham Mouan Managing Protected Authority region, 17°57’32” N, prov., Bounlapha distr., Thong Sam village, Hinnam 105°01’05” E, 538 m, 18 Jan. 2009, S. Bounphanmy No N.P.A., Pu Pha Song mt. composed by sandstone, et al., LA-VN 13 (CPC). LAOS, Saravan prov., Ta 17°35’13.2” N, 105°47’49.8” E, 1300 m, 9 March Oy distr., to the SW of Sa Mouay town, near Thong 2013, L. Averyanov et al., LA-VN 316 (CPC, NHOL, Hang village, 15°37’30.9” N,, 106°12’26.5” E, NUOL). 650–700 m, 4 March 2013, L. Averyanov et al., LA- *E. thao Gagnep. VN 218 (CPC, NHOL, NUOL). LAOS, Saravan prov., Sa Mouay distr., +E. muscicola (Lindl.) Lindl. to the SEE of Sa Mouay town, elev. from Ta Oy CAMBODIA, Bokor N.P., Bokor mt., 1047 village, 1350 m, 16°08’47.0” N, 106°52’35.4” E, 4 m, 10°38’17” N, 104°00’46” E, 24 Nov. 2012, T. March 2013, L. Averyanov et al., LA-VN 153 (CPC, Maisak et al., 925, Fig. 4, F (LE). NHOL, NUOL). E. paniculata Lindl. E. truncata Lindl. LAOS, Saravan prov., Sa Mouay distr., LAOS, Saravan prov., Sa Mouay distr., to to the SEE of Sa Mouay town, elev. from Ta Oy the SEE of Sa Mouay town, elev. from Ta Oy village, village, 1350 m, 16°08’47.0” N, 106°52’35.4” E, 4 1350 m, 16°08’47.0” N, 106°52’35.4” E, 4 March March 2013, L. Averyanov et al., LA-VN 176 (CPC, 2013, L. Averyanov et al., LA-VN 170 (CPC, LE, NHOL, NUOL). LAOS, Vientiane prov., Kasi distr., NHOL, NUOL). LAOS, Vientiane prov., Kasi distr., Namken village, Phachao mt., 19°18’45.5” N, Thong Mout village, Pudang Ridge, 19°23’33.7” N, 102°22’31.4” E, 1500–1750 m, 24 March 2013, L. 102°08’05.7” E, 1950–2000 m, 22 March 2013, L. Averyanov, N.S. Khang, S. Lorphengsy LA-VN 801 Averyanov, N.S. Khang, S. Lorphengsy, LA-VN 706 (CPC, NHOL, NUOL). (CPC, LE, NHOL, NUOL). E. pannea Lindl. CAMBODIA, Kampot prov., Phnom Bokor Eulophia R. Br. N.P., 10°37’92.3 N, 104°01’14.3 E, 21 Dec. 2012, E. graminea Lindl. E.L. Konstantinov et al., 41P-C-GPS042 (LE). LAOS, Vientiane prov., Vang Vieng distr., LAOS, Borikhamxai prov., Khamkeut distr., Dan vic. of Vang Vieng town, 18°54’58” N, 102°24’52” Khum Ngan locality, Watershed Managing Protected E, 400–450 m, 24 April 2011, L. Averyanov, P.V. The Authority region, 17°57’32” N, 105°01’05” E, 538 CPC 2441 (CPC). m, 18 Jan. 2009, S. Bounphanmy et al., LA-VN 12 (CPC). LAOS, Saravan prov., Sa Mouay distr., to Flickingeria A.D. Hawkes the SEE of Sa Mouay town, elev. from Ta Oy village, *F. angustifolia (Blume) A.D. Hawkes 1350 m, 16°08’47.0” N, 106°52’35.4” E, 4 March LAOS, Saravan prov., Sa Mouay distr., 2013, L. Averyanov et al., LA-VN 191 (CPC, NHOL, to the SEE of Sa Mouay town, elev. from Ta Oy NUOL). LAOS, Saravan prov., Ta Oy distr., to the village, 1350 m, 16°08’47.0” N, 106°52’35.4” E, 4 SW of Sa Mouay town, near Thong Hang village, March 2013, L. Averyanov et al., LA-VN 182, Fig. 4, 15°37’30.9” N, 106°12’26.5” E, 650–700 m, 4 G (CPC, NHOL, NUOL). LAOS, Vientiane prov., March 2013, L. Averyanov et al., LA-VN 216 (CPC, Kasi distr., Tam Tai village, about 10 km to the NW NHOL, NUOL). LAOS, Vientiane prov., Vang of Kasi town, 19°12’57” N, 102°14’59” E, 500–650 Vieng distr., Phol Xai village, about 8 km to the W m, 24 March 2013, L. Averyanov, N.S. Khang, S. of Vang Vieng town, Phar Poon mt., 18°56’11.5” N, Lorphengsy, LA-VN 752 (CPC, NHOL, NUOL). 102°20’05.0” E, 800–900 m, 15 March 2013, N.T. Hiep et al., LA-VN 471 (CPC, NHOL, NUOL). Gastrochilus D. Don E. perpusilla Par. et Rchb. f. *G. calceolaris (Buch.-Ham. ex Sm.) D. Don LAOS, Vientiane prov., Vang Vieng distr., LAOS, Kham Mouan prov., Bounlapha Na Po village, about 12 km to the W of Vang distr., Thong Sam village, Hinnam No N.P.A., Pu

38 Averyanov L.V. New and rare orchids (Orchidaceae) in the flora of Cambodia and Laos

Pha Song mt., 17°35’13.2” N, 105°47’49.8” E, *Lecanorchis Blume 1000–1100 m, 9 March 2013, L. Averyanov et al., *L. vietnamica Aver. LA-VN 303 (CPC, NHOL, NUOL). LAOS, Saravan prov., Sa Mouay distr., *G. pseudodistichus (King et Pantl.) Schltr. to the SEE of Sa Mouay town, elev. from Ta Oy LAOS, Vientiane prov., Kasi distr., Namken village, 1350 m, 16°08’47.0” N, 106°52’35.4” E, 4 village, Phachao mt., 19°18’45.5” N, 102°22’31.4” March 2013, L. Averyanov et al., LA-VN 171 (CPC, E, 1600 m, 24 March 2013, L. Averyanov, N.S. Khang, NHOL, NUOL). S. Lorphengsy, LA-VN 794 (CPC, NHOL, NUOL). Liparis Rich. Goodyera R. Br. *L. balansae Gagnep. *G. fumata Thwaites LAOS, Kham Mouan prov., Bounlapha LAOS, Kham Mouan prov., Bounlapha distr., Thong Sam village, Hinnam No N.P.A., Pu distr., Thong Sam village, Hinnam No N.P.A., Pu Pha Song mt.. 17°35’13.2” N, 105°47’49.8” E, 1425 Pha Song mt., 17°35’13.2” N, 105°47’49.8” E, 1000 m, 9 March 2013, L. Averyanov et al., LA-VN 309 m, 9 March 2013, L. Averyanov et al., LA-VN 301 (LE). (CPC, NHOL, NUOL). L. bootanensis Griff. *G. hispida Lindl. LAOS, Kham Mouan prov., Bounlapha LAOS, Vientiane prov., Kasi distr., Namken distr., Thong Sam village, Hinnam No N.P.A., Pu village, Phachao mt., 19°18’45.5” N, 102°22’31.4” Pha Song mt., 17°35’13.2” N, 105°47’49.8” E, 1200 E, 1500 m, 24 March 2013, L. Averyanov, N.S. m, 9 March 2013, L. Averyanov et al., LA-VN 309 Khang, S. Lorphengsy, LA-VN 797, Fig. 4, H (CPC, (CPC, NHOL, NUOL). NHOL, NUOL). L. nervosa (Thunb.) Lindl. LAOS, Kham Mouan prov., Bounlapha *Herpysma Lindl. distr., Thong Sam village, Hinnam No N.P.A., Pu *H. longicaulis Lindl. Pha Song mt., 17°35’13.2” N, 105°47’49.8” E, 900 LAOS, Vientiane prov., Kasi distr., Namken m, 9 March 2013, L. Averyanov et al., LA-VN 296 village, Phachao mt., around point 19°18’45.5” (CPC, NHOL, NUOL). N, 102°22’31.4” E, 1600 m, 24 March 2013, L. *L. pumila Aver. Averyanov, N.S. Khang, S. Lorphengsy, LA-VN 791 LAOS, Borikhamxai prov., Khamkeut distr., (CPC, LE, NHOL, NUOL). Phone Sa At village, 18°03’38” N, 105°02’28” E, 635 m, 18 Jan. 2009, S. Bounphanmy et al., LA-VN *Hetaeria Blume 34 (CPC). LAOS, Vientiane prov., Kasi distr., Tam *H. affinis (Griff.) Seidenf. et Ormerod Tai village, about 10 km to the NW of Kasi town, LAOS, Vientiane prov., Vang Vieng distr., 19°12’57” N, 102°14’59” E, 500–650 m, 24 March Phol Xai village, about 8 km to the W of Vang Vieng 2013, L. Averyanov, N.S. Khang, S. Lorphengsy LA- town, Phar Poon mt., 18°56’11.5” N, 102°20’05.0” VN 750 (CPC, NHOL, NUOL). E, 300 m, 15 March 2013, N.T. Hiep et al., LA-VN L. viridiflora (Blume) Lindl. 429 (CPC, LE, NHOL, NUOL). LAOS, Vientiane prov., Kasi distr., Thong Mout village, 19°26’51.5” N, 102°07’45.8” E, Holcoglossum Schltr. 1550–1650 m, 20 March 2013, N.T. Hiep et al., LA- *H. subulifolium (Rchb. f.) Christenson VN 607 (CPC, NHOL, NUOL). LAOS, Vientiane LAOS, Kham Mouan prov., Bounlapha prov., Vang Vieng distr., Phol Xai village, about 8 distr., Thong Sam village, Hinnam No N.P.A., Pu km to the W of Vang Vieng town, Phar Poon mt., Pha Song mt., 17°35’13.2” N, 105°47’49.8” E, 1400 18°56’11.5” N, 102°20’05.0” E, 800–900 m, 15 m, 9 March 2013, L. Averyanov et al., LA-VN 260, March 2013, N.T. Hiep et al., LA-VN 479 (CPC, Fig. 4, I (CPC, LE, NHOL, NUOL). NHOL, NUOL).

Kingidium P.F. Hunt Luisia Gaudich. K. deliciosum (Rchb. f.) H.R. Sweet L. filiformis Hook. f. LAOS, Vientiane prov., Vang Vieng distr., LAOS, Attapeu prov., Saysettha distr., Dong Nathong village, about 5 km to the W of Vang Vieng Am Pham N.P.A., to the N of Paosamphanmixay town, 18°55’34.2” N, 102°22’58.4” E, 300 m, 13 village, 14°51’53.4” N, 107°11’38.2” E, 178 m, March 2013, N.T. Hiep et al., LA-VN 351 (CPC, 28 February 2013, L. Averyanov et al., LA-VN 115 NHOL, NUOL). (CPC, LE, NHOL, NUOL).

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Fig. 4. New orchids in the flora of Cambodia and Laos. A – Dendrobium setifolium Ridl. (E.L. Konstantinov et al., 35P-C-GPS042). B – Didymoplexiopsis khiriwongensis Seidenf. (L. Averyanov et al., LA-VN 110). C, D – Eclecticus chungii P. O’Byrne (N.T. Hiep et al., LA-VN 385c). E – Eria lasiopetala (Willd.) Ormerod (E.L. Konstantinov et al., 39P-C-GPS042). F – E. muscicola (Lindl.) Lindl. (T. Maisak et al., 925). G – Flickingeria angustifolia (Blume) A.D. Hawkes (L. Averyanov et al., LA-VN 182). H – Goodyera hispida Lindl. (L. Averyanov, N.S. Khang, S. Lorphengsy, LA-VN 797). I – Holcoglossum subulifolium (Rchb. f.) Christenson (L. Averyanov et al., LA-VN 260). Photos by E. Konstantinov (A, E), Nguyen Sinh Khang (B, G), L. Averyanov (C, D, E, H) and Pham Van The (I).

40 Averyanov L.V. New and rare orchids (Orchidaceae) in the flora of Cambodia and Laos

Micropera Lindl. Averyanov, N.S. Khang, S. Lorphengsy LA-VN 704 *M. poilanei (Guillaumin) Garay (CPC, LE, NHOL, NUOL). LAOS, Vientiane prov., Vang Vieng distr., Nathong village, about 5 km to the W of Vang Vieng Panisea (Lindl.) Lindl. town, Tham Kang mt., 18°55’51.4” N, 102°23’50.6” P. uniflora (Lindl.) Lindl. E, 300–500 m, 14 March 2013, N.T. Hiep et al., LA- CAMBODIA, to the N of Sianukwille VN 377 (CPC, NHOL, NUOL). town, Kaoh Rong Samoloem Khong Island in Siam Gulf, 10°36’45” N, 103°18’38” E, 19 Nov. 2011, Nephelaphyllum Blume T. Maisak, M. Telepova, L. Osinovets, 907 (LE). *N. tenuiflorum Blume LAOS, Borikhamxai prov., Khamkeut distr., Dan LAOS, Saravan prov., Sa Mouay distr., Khum Ngan locality, Watershed Managing Protected to the SEE of Sa Mouay town, elev. from Ta Oy Authority region, 17°57’32” N, 105°01’05” E, 538 village, 1350 m, 16°08’47.0” N, 106°52’35.4” E, 4 m, 18 Jan. 2009, S. Bounphanmy et al., LA-VN 21i March 2013, L. Averyanov et al., LA-VN 168 (CPC, (CPC, LE). LAOS, Vientiane prov., Vang Vieng NHOL, NUOL). distr., Na Khun Village, 18°52’28” N, 102°24’21” E, 400–600 m, 21 Jan. 2009, O. Souliya et al., LA- *Neuwiedia Blume VN 92 (CPC). LAOS, Vientiane prov., Vang Vieng *N. zollingeri Rchb. f. distr., vic. of Vang Vieng town, 18°54’58” N, LAOS, Kham Mouan prov., Bounlapha 102°24’52” E, 400–450 m, 24 April 2011, L. distr., Thong Sam village, Hinnam No N.P.A., Pu Averyanov, P.V. The, CPC 2439 (CPC). LAOS, Pha Song mt., 17°35’13.2” N, 105°47’49.8” E, 500– Vientiane prov., Vang Vieng distr., Phol Xai village, 900 m, 9 March 2013, L. Averyanov et al., LA-VN about 8 km to the W of Vang Vieng town, Phar Poon 297 (CPC, NHOL, NUOL). mt., 18°56’11.5” N, 102°20’05.0” E, 800–900 m, 15 March 2013, N.T. Hiep et al., LA-VN 475 (CPC, Oberonia Lindl. NHOL, NUOL). LAOS, Vientiane prov., Vang *O. cavaleriei Finet Vieng distr., Na Po village, about 12 km to the W of LAOS, Vientiane prov., Kasi distr., Tam Vang Vieng town, Pa Nang Oua mt., 18°55’44.9” N, Tai village, about 10 km to the NW of Kasi town, 102°20’20.1” E, 500–1000 m, 17 March 2013, N.T. 19°12’57” N, 102°14’59” E, 500–650 m, 24 March Hiep et al., LA-VN 515a (CPC, LE, NHOL, NUOL). 2013, L. Averyanov, N.S. Khang, S. Lorphengsy LA- LAOS, Vientiane prov., Kasi distr., Tam Tai village, VN 757 (CPC, NHOL, NUOL). about 10 km to the NW of Kasi town, 19°12’57” O. falcata King et Pantl. N, 102°14’59” E, 500–650 m, 24 March 2013, L. LAOS, Borikhamxai prov., Khamkeut Averyanov, N.S. Khang, S. Lorphengsy, LA-VN 761 distr., Dan Khum Ngan locality, Watershed (CPC, NHOL, NUOL). Managing Protected Authority region, 17°57’32” N, 105°01’05” E, 538 m, 18 Jan. 2009, S. Bounphanmy Paphiopedilum Pfitzer et al., LA-VN 17 (CPC). P. appletonianum (Gower) Rolfe CAMBODIA, Kampot prov., Phnom Bokor Ornithochilus (Lindl.) Benth. N.P. Epiphyte, 10°37’92.3 N, 104°01’14.3 E, 21 O. difformis (Wall. ex Lindl.) Schltr. Dec. 2012, E.L. Konstantinov et al., 50P-C-GPS042 LAOS, Phongsali prov., Muong May distr., (LE). LAOS, Saravan prov., Sa Mouay distr., to the 21°12’32” N, 102°53’44” E, 1250–1400 m, 22 April SEE of Sa Mouay town, elev. from Ta Oy village, 2011, L. Averyanov, P.V. The, CPC 2431 (CPC). 1350 m, 16°08’47.0” N, 106°52’35.4” E, 4 March LAOS, Saravan prov., Sa Mouay distr., to the SEE 2013, L. Averyanov et al., LA-VN 178 (CPC, NHOL, of Sa Mouay town, elev. from Ta Oy village, 1350 NUOL). LAOS, Kham Mouan prov., Bounlapha m, 16°08’47.0” N, 106°52’35.4” E, 4 March 2013, distr., Thong Sam village, Hinnam No N.P.A., Pu L. Averyanov et al., LA-VN 167 (CPC, NHOL, Pha Song mt., 17°35’13.2” N, 105°47’49.8” E, NUOL). 1200–1300 m, 9 March 2013, L. Averyanov et al., LA-VN 305 (CPC, NHOL, NUOL). Otochilus Lindl. *P. canhii Aver. et O. Gruss *O. psudoporrectus Aver. LAOS, Vientiane prov., Kasi distr., Namken LAOS, Vientiane prov., Kasi distr., Thong village, Phachao mt., 19°18’45.5” N, 102°22’31.4” Mout village, Pudang Ridge, 19°23’33.7” N, E, 1600 m, 24 March 2013, L. Averyanov, N.S. 102°08’05.7” E, 1950–2000 m, 22 March 2013, L.

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Khang, S. Lorphengsy, LA-VN 793, Fig. 5, A, B 635 m, 18 Jan. 2009, S. Bounphanmy et al., LA-VN (CPC, LE, NHOL, NUOL). 30 (CPC). LAOS, Saravan prov., Ta Oy distr., to the P. concolor (Lindl.) Pfitzer SW of Sa Mouay town, near Thong Hang village, LAOS, Vientiane prov., Kasi distr., Tam 15°37’30.9” N, 106°12’26.5” E, 650–700 m, 4 Tai village, about 10 km to the NW of Kasi town, March 2013, L. Averyanov et al., LA-VN 221 (CPC, 19°12’57” N, 102°14’59” E, 500–650 m, 24 March NHOL, NUOL). LAOS, Vientiane prov., Vang 2013, L. Averyanov, N.S. Khang, S. Lorphengsy LA- Vieng distr., Phol Xai village, about 8 km to the W VN 743 (CPC, NHOL, NUOL). of Vang Vieng town, Phar Poon mt., 18°56’11.5” P. dianthum Tang et F.T. Wang N, 102°20’05.0” E, 800–900 m, 15 March 2013, LAOS, Vientiane prov., Kasi distr., Namken N.T. Hiep et al., LA-VN 477 (CPC, NHOL, NUOL). village, Phachao mt., 19°18’45.5” N, 102°22’31.4” LAOS, Vientiane prov., Kasi distr., Namken village, E, 1500–1750 m, 24 March 2013, L. Averyanov, N.S. Phachao mt., 19°18’45.5” N, 102°22’31.4” E, Khang, S. Lorphengsy LA-VN 781 (CPC, NHOL, 1500–1750 m, 24 March 2013, L. Averyanov, N.S. NUOL). Khang, S. Lorphengsy LA-VN 775 (CPC, NHOL, NUOL). LAOS, Vientiane prov., Kasi distr., Tam Papilionanthe Schltr. Tai village, about 10 km to the NW of Kasi town, *P. biswasiana (Ghose et Mukerjee) Garay 19°12’57” N, 102°14’59” E, 500–650 m, 24 March LAOS, Vientiane prov., Kasi distr., Namken 2013, L. Averyanov, N.S. Khang, S. Lorphengsy LA- village, Phachao mt., 19°18’45.5” N, 102°22’31.4” VN 763 (CPC, NHOL, NUOL). E, 1500–1750 m, 24 March 2013, L. Averyanov, N.S. P. chinensis Lindl. Khang, S. Lorphengsy LA-VN 799, Fig. 5, C (CPC, LAOS, Saravan prov., Sa Mouay distr., to LE, NHOL, NUOL). the SEE of Sa Mouay town, elev. from Ta Oy village, 1350 m, 16°08’47.0” N, 106°52’35.4” E, 4 March Pelatantheria Ridl. 2013, L. Averyanov et al., LA-VN 184 (CPC, NHOL, P. ctenoglossum Ridl. NUOL). LAOS, Kham Mouan prov., Bounlapha LAOS, Vientiane prov., Kasi distr., Tam distr., Thong Sam village, Hinnam No N.P.A., Pu Tai village, about 10 km to the NW of Kasi town, Pha Song mt., 17°35’13.2” N, 105°47’49.8” E, 1100 19°12’57” N, 102°14’59” E, 500–650 m, 24 March m, 9 March 2013, L. Averyanov et al., LA-VN 318 2013, L. Averyanov, N.S. Khang, S. Lorphengsy LA- (CPC, NHOL, NUOL). VN 748 (CPC, NHOL, NUOL). P. imbricata Lindl. LAOS, Borikhamxai prov., Khamkeut distr., Phaius Lindl. Phone Sa At village, 18°03’38” N, 105°02’28” E, P. flavus (Blume) Lindl. 635 m, 18 Jan. 2009, S. Bounphanmy et al., LA-VN 35 LAOS, Vientiane prov., Vang Vieng distr., (CPC). LAOS, Attapeu prov., Saysettha distr., Dong Na Po village, about 12 km to the W of Vang Am Pham N.P.A., to the N of Paosamphanmixay Vieng town, Pa Nang Oua mt., 18°55’44.9” N, village, 14°51’53.4” N, 107°11’38.2” E, 200 m, 1 102°20’20.1” E, 600–900 m, 17 March 2013, N.T. March 2013, L. Averyanov et al., LA-VN 135 (CPC, Hiep et al., LA-VN 550 (CPC, NHOL, NUOL). NHOL, NUOL). LAOS, Vientiane prov., Vang LAOS, Vientiane prov., Kasi distr., Namken village, Vieng distr., Nathong village, about 5 km to the W Phachao mt., 19°18’45.5” N, 102°22’31.4” E, 1000– of Vang Vieng town, Tham Kang mt., 18°55’51.4” 1600 m, 24 March 2013, L. Averyanov, N.S. Khang, N, 102°23’50.6” E, 300–500 m, 14 March 2013, S. Lorphengsy LA-VN 779 (CPC, NHOL, NUOL). N.T. Hiep et al., LA-VN 389 (CPC, NHOL, NUOL). LAOS, Vientiane prov., Kasi distr., Tam Tai village, Phalaenopsis Blume about 10 km to the NW of Kasi town, 19°12’57” *P. wilsonii Rolfe N, 102°14’59” E, 500–650 m, 24 March 2013, L. LAOS, Vientiane prov., Kasi distr., Thong Averyanov, N.S. Khang, S. Lorphengsy LA-VN 760 Mout village, 19°26’51.5” N, 102°07’45.8” E, (CPC, NHOL, NUOL). 1550–1650 m, 20 March 2013, N.T. Hiep et al., LA- P. pallida Lindl. VN 606, Fig. 5, D (CPC, NHOL, NUOL). LAOS, Vientiane prov., Kasi distr., Namken village, Phachao mt., 19°18’45.5” N, 102°22’31.4” Pholidota Lindl. E, 1500–1750 m, 24 March 2013, L. Averyanov, N.S. P. articulata Lindl. Khang, S. Lorphengsy LA-VN 780 (CPC, NHOL, LAOS, Borikhamxai prov., Khamkeut distr., NUOL). Phone Sa At village, 18°03’38” N, 105°02’28” E,

42 Averyanov L.V. New and rare orchids (Orchidaceae) in the flora of Cambodia and Laos

Plocoglottis Blume 2013, L. Averyanov et al., LA-VN 186 (CPC, NHOL, P. bokorensis (Gagnep.) Seidenf. NUOL). LAOS, Kham Mouan prov., Bounlapha LAOS, Kham Mouan prov., Bounlapha distr., Thong Sam village, Hinnam No N.P.A., Pu distr., Thong Sam village, Hinnam No N.P.A., Pu Pha Song mt., 17°35’13.2” N, 105°47’49.8” E, 1400 Pha Song mt., 17°35’13.2” N, 105°47’49.8” E, 500– m, 9 March 2013, L. Averyanov et al., LA-VN 322 1000 m, 9 March 2013, L. Averyanov et al., LA-VN (CPC, NHOL, NUOL). LAOS, Vientiane prov., 284 (CPC, NHOL, NUOL). Kasi distr., Thong Mout village, 19°24’23.7” N, 102°09’50.2” E, 1760 m, 20 March 2013, N.T. Hiep Podochilus Blume et al., LA-VN 645 (CPC, NHOL, NUOL). *P. oxystophylloides Ormerod LAOS, Vientiane prov., Kasi distr., Namken Smitinandia Holttum village, Phachao mt., 19°18’45.5” N, 102°22’31.4” E, S. micrantha (Lindl.) Holttum 1200–1750 m, 24 March 2013, L. Averyanov, N.S. Khang, LAOS, Borikhamxai prov., Khamkeut S. Lorphengsy LA-VN 783 (CPC, NHOL, NUOL). distr., Dan Khum Ngan locality, Watershed Managing Protected Authority region, 17°57’32” N, Pomatocalpa Breda 105°01’05” E, 538 m, 18 Jan. 2009, S. Bounphanmy P. angustifolium Seidenf. et al., LA-VN 21c (CPC). LAOS, Vientiane prov., LAOS, Vientiane prov., Vang Vieng distr., Vang Vieng distr., Nathong village, about 5 km to the Nathong village, about 5 km to the W of Vang Vieng W of Vang Vieng town, Tham Kang mt., 18°55’51.4” town, Tham Kang mt., 18°55’51.4” N, 102°23’50.6” N, 102°23’50.6” E, 300–400 m, 14 March 2013, E, 300–500 m, 14 March 2013, N.T. Hiep et al., LA- N.T. Hiep et al., LA-VN 423 (CPC, NHOL, NUOL). VN 385 (CPC, LE, NHOL, NUOL). LAOS, Vientiane prov., Vang Vieng distr., Na Khun Village, 18°52’28” N, 102°24’21” E, 400–600 m, Porpax Lindl. 21 Jan. 2009, O. Souliya et al., LA-VN 84 (CPC). +P. elwesii (Rchb. f.) Rolfe CAMBODIA, to the N of Sianukwille town, Spathoglottis Blume Kaoh Rong Samoloem Khong Island in Siam Gulf, S. affinis de Vriese 10°36’104 N 103°18’056 E, 19 Nov. 2011, T. Maisak, LAOS, Phu Khao Khuay N.P., Xang M. Telepova, L. Osinovets, 851, Fig. 5, E (LE). waterfall, Tad Leuk willage, 18°24’05.8 N, 103°03’03.3 E, 12 Nov. 2011, E. Konstantinov et Renanthera Lour. al., 6 (LE). R. coccinea Lour. S. eburnea Gagnep. LAOS, Vientiane prov., Vang Vieng distr., CAMBODIA, Kampot prov., Phnom Bokor vic. of Vang Vieng town, 18°54’58” N, 102°24’52” N.P., 10°37’92.3 N, 104°01’14.3 E, 21 Dec. 2012, E, 400–450 m, 24 April 2011, L. Averyanov, P.V. E.L. Konstantinov et al., 46P-C-GPS042 (LE). The, CPC 2443 (CPC). LAOS, Namnor, 18°01’66.1 N, 104°41’45.8 E, Nov. 2011, E. Konstantinov et al., 2a (LE). Sarcoglyphis Garay S. mirabilis (Rchb. f.) Garay Sunipia Lindl. LAOS, Vientiane prov., Vang Vieng distr., *S. rimannii (Rchb. f.) Seidenf. Nathong village, about 5 km to the W of Vang Vieng LAOS, Vientiane prov., Kasi distr., Namken town, Tham Kang mt., 18°55’51.4” N, 102°23’50.6” village, Phachao mt., 19°18’45.5” N, 102°22’31.4” E, 300–500 m, 14 March 2013, N.T. Hiep et al., LA-VN E, 1500–1750 m, 24 March 2013, L. Averyanov, N.S. 385a (CPC, LE, NHOL, NUOL). LAOS, Vientiane Khang, S. Lorphengsy LA-VN 802, Fig. 5, F (CPC, prov., Vang Vieng distr., Phol Xai village, about 8 km to LE, NHOL, NUOL). the W of Vang Vieng town, Phar Poon mt., 18°56’11.5” S. scariosa Lindl. N, 102°20’05.0” E, 300–500 m, 15 March 2013, N.T. LAOS, Vientiane prov., Kasi distr., Namken Hiep et al., LA-VN 467 (CPC, LE, NHOL, NUOL). village, Phachao mt., 19°18’45.5” N, 102°22’31.4” E, 1500–1750 m, 24 March 2013, L. Averyanov, N.S. Khang, Schoenorchis Blume S. Lorphengsy LA-VN 774 (CPC, NHOL, NUOL). S. gemmata (Lindl.) J.J. Sm. LAOS, Saravan prov., Sa Mouay distr., to Tainia Blume the SEE of Sa Mouay town, elev. from Ta Oy village, T. longiscapa (Seidenf.) J.J. Wood et A.L. 1350 m, 16°08’47.0” N, 106°52’35.4” E, 4 March Lamb

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LAOS, Vientiane prov., Vang Vieng distr., *Trichosma Lindl. Na Po village, about 12 km to the W of Vang *T. simondii Gagnep. Vieng town, Pa Nang Oua mt., 18°55’44.9” N, LAOS, Saravan prov., Sa Mouay distr., 102°20’20.1” E, 600–900 m, 17 March 2013, N.T. to the SEE of Sa Mouay town, elev. from Ta Oy Hiep et al., LA-VN 551 (CPC, NHOL, NUOL). village, 1350 m, 16°08’47.0” N, 106°52’35.4” E, 4 March 2013, L. Averyanov et al., LA-VN 185 (CPC, Thelasis Blume NHOL, NUOL). *T. khasiana Hook. f. *T. suavis Lindl. LAOS, Vientiane prov., Kasi distr., Thong LAOS, Kham Mouan prov., Bounlapha Mout village, 19°26’51.5” N, 102°07’45.8” E, distr., Thong Sam village, Hinnam No N.P.A., Pu 1550–1650 m, 20 March 2013, N.T. Hiep et al., LA- Pha Song mt., 17°35’13.2” N, 105°47’49.8” E, 1200 VN 610 (CPC, NHOL, NUOL). m, 9 March 2013, L. Averyanov et al., LA-VN 310 T. pygmaea (Griff.) Blume (CPC, NHOL, NUOL). LAOS, Vientiane prov., Vang Vieng distr., Phol Xai village, about 8 km to the W of Vang Vieng Trichotosia Blume town, Phar Poon mt., 18°56’11.5” N, 102°20’05.0” T. dasyphylla (Par. et Rchb. f.) Kraenzl. E, 800–900 m, 15 March 2013 Coll.: N.T. Hiep LAOS, Saravan prov., Sa Mouay distr., to et al., LA-VN 473 (CPC, NHOL, NUOL). LAOS, the SEE of Sa Mouay town, elev. from Ta Oy village, Vientiane prov., Vang Vieng distr., Na Khun Village, 1350 m, 16°08’47.0” N, 106°52’35.4” E, 4 March 18°52’28” N, 102°24’21” E, 400–600 m, 21 Jan. 2013, L. Averyanov et al., LA-VN 169 (CPC, NHOL, 2009, O. Souliya et al., LA-VN 87 (CPC). NUOL). LAOS, Vientiane prov., Vang Vieng distr., Phol Xai village, about 8 km to the W of Vang Vieng Thrixspermum Lour. town, Phar Poon mt., 18°56’11.5” N, 102°20’05.0” T. centipeda Lour. E, 800–900 m, 15 March 2013, N.T. Hiep et al., LA- LAOS, Borikhamxai prov., Khamkeut VN 470 (CPC, NHOL, NUOL). distr., Dan Khum Ngan locality, Watershed T. pulvinata (Lindl.) Kraenzl. Managing Protected Authority region, 17°57’32” N, LAOS, Saravan prov., Sa Mouay distr., 105°01’05” E, 538 m, 18 Jan. 2009, S. Bounphanmy to the SEE of Sa Mouay town, elev. from Ta Oy et al., LA-VN 18 (CPC). village, 1350 m, 16°08’47.0” N, 106°52’35.4” E, 4 *T. sutepense (Rolfe ex Downie) Tang et March 2013, L. Averyanov et al., LA-VN 196 (CPC, F.T. Wang NHOL, NUOL). LAOS, Vientiane prov., Kasi distr., Namken village, Phachao mt., 19°18’45.5” N, 102°22’31.4” Tropidia Lindl. E, 1500–1750 m, 24 March 2013, L. Averyanov, N.S. T. angulosa (Lindl.) Blume Khang, S. Lorphengsy LA-VN 788, Fig. 5, G (CPC, LAOS, Vientiane prov., Vang Vieng distr., LE, NHOL, NUOL). Na Khun Village, 18°52’28” N, 102°24’21” E, 300 m, 21 Jan. 2009, O. Souliya et al., LA-VN 67 (CPC). Trias Lindl. LAOS, Kham Mouan prov., Bounlapha distr., Thong T. nasuta (Rchb. f.) Stapf Sam village, Hinnam No N.P.A., Pu Pha Song mt., LAOS, Saravan prov., Ta Oy distr., to the 17°35’13.2” N, 105°47’49.8” E, 400–900 m, 9 March SW of Sa Mouay town, near Thong Hang village, 2013, L. Averyanov et al., LA-VN 295 (CPC, NHOL, 15°37’30.9” N, 106°12’26.5” E, 650–700 m, 4 NUOL). LAOS, Kham Mouan prov., Bounlapha March 2013, L. Averyanov et al., LA-VN 220 (CPC, distr., Nam Boua Sam village, Hinnam No N.P.A., NHOL, NUOL). Pu Pha Song mt., 17°33’43.6” N, 105°45’08.3” E, 230 m, 10 March 2013, L. Averyanov et al., LA- Trichoglottis Blume VN 329 (CPC, NHOL, NUOL). LAOS, Vientiane T. orchidea (J. Koenig) Garay prov., Vang Vieng distr., Nathong village, about 5 LAOS, Vientiane prov., Vang Vieng distr., km to the W of Vang Vieng town, Tham Kang mt., Phol Xai village, about 8 km to the W of Vang Vieng 18°55’51.4” N, 102°23’50.6” E, 300–500 m, 14 town, Phar Poon mt., 18°56’11.5” N, 102°20’05.0” March 2013, N.T. Hiep et al., LA-VN 388 (CPC, E, 800–900 m, 15 March 2013, N.T. Hiep et al., LA- NHOL, NUOL). LAOS, Vientiane prov., Vang VN 466 (CPC, NHOL, NUOL). Vieng distr., Na Po village, about 12 km to the W of Vang Vieng town, Pa Nang Oua mt., 18°55’36.4”

44 Averyanov L.V. New and rare orchids (Orchidaceae) in the flora of Cambodia and Laos

Fig. 5. New orchids in the flora of Cambodia and Laos. A, B – Paphiopedilum canhii Aver. et O. Gruss (L. Averyanov, N.S. Khang, S. Lorphengsy, LA-VN 793). C – Papilionanthe biswasiana (Ghose et Mukerjee) Garay (L. Averyanov, N.S. Khang, S. Lorphengsy LA-VN 799). D – Phalaenopsis wilsonii Rolfe (N.T. Hiep et al., LA-VN 606). E – Porpax elwesii (Rchb. f.) Rolfe (T. Maisak, M. Telepova, L. Osinovets, 851). F – Sunipia rimannii (Rchb. f.) Seidenf. (L. Averyanov, N.S. Khang, S. Lorphengsy LA-VN 802). G – Thrixspermum sutepense (Rolfe ex Downie) Tang et F.T. Wang (L. Averyanov, N.S. Khang, S. Lorphengsy LA-VN 788). H – Zeuxine flava (Wall. ex Lindl.) Trimen (L. Averyanov, N.S. Khang, S. Lorphengsy LA-VN 777). I – Z. nervosa Trimen (L. Averyanov, N.S. Khang, S. Lorphengsy LA-VN 800). Photos by Nguyen Sinh Khang (A, C, F-I), L. Averyanov (B), Pham Van The (D) and T. Maisak (E).

Turczaninowia 2013, 16 (4) : 26–46 45

N, 102°20’44.6” E, 350–450 m, 17 March 2013, *Z. nervosa Trimen N.T. Hiep et al., LA-VN 501 (CPC, NHOL, NUOL). LAOS, Borikhamxai prov., Khamkeut distr., LAOS, Vientiane prov., Kasi distr., Tam Tai village, Phone Sa At village, 18°03’38” N, 105°02’28” E, about 10 km to the NW of Kasi town, 19°12’57” 635 m, 18 Jan. 2009, S. Bounphanmy et al., LA- N, 102°14’59” E, 500–650 m, 24 March 2013, L. VN 27 (CPC). LAOS, Luang Namtha prov., Ban Averyanov, N.S. Khang, S. Lorphengsy LA-VN 741 Nam Ha, 21°00’15.9 N, 101°24’55.4 E, 2 Dec. (CPC, NHOL, NUOL). 2012, E.L. Konstantinov et al., 14ORa-L-GPS018 T. curcugiloides Lindl. (LE). LAOS, Vientiane prov., Kasi distr., Namken LAOS, Kham Mouan prov., Bounlapha village, Phachao mt., 19°18’45.5” N, 102°22’31.4” distr., Thong Sam village, Hinnam No N.P.A., Pu E, 1500–1750 m, 24 March 2013, L. Averyanov, N.S. Pha Song mt., 17°35’13.2” N, 105°47’49.8” E, 700– Khang, S. Lorphengsy LA-VN 800, Fig. 5, I (CPC, 900 m, 9 March 2013, L. Averyanov et al., LA-VN LE, NHOL, NUOL). 294 (CPC, NHOL, NUOL). Acknowledgements. The author cordially Vanda R. Br. thanks the authorities of the Center for Plant V. brunnea Rchb. f. Conservation (Vietnam Union of Science and LAOS, Phongsali prov., Muong May distr., Technology Associations), National University of 21°12’32” N, 102°53’44” E, 1250–1400 m, 22 April Laos and Royal University of Phnom Penh for their 2011, L. Averyanov, P.V. The, CPC 2432 (CPC). key role in organizing the field works. Field and laboratory studies were funded by U.S.A. National Vandopsis Pfitzer Geographic Society, grants “Flora of relict karstic V. gigantea (Lindl.) Pfitzer formation of central Laos (Vientiane province, LAOS, Vientiane prov., Vang Vieng distr., Vang Vieng and Kasi districts)” #2012–2013, Na Khun Village, 18°52’28” N, 102°24’21” E, 300 “Exploration of primary woods along constructed m, 21 Jan. 2009, O. Souliya et al., LA-VN 82 (CPC). highway Hanoi – Ho Chi Minh for their sustainable LAOS, Kham Mouan prov., Bounlapha distr., Thong conservation (in limits of Ha Tinh and Nghe An Sam village, Hinnam No N.P.A., Pu Pha Song mt., provinces of central Vietnam)” #9129–12 and grant 17°35’13.2” N, 105°47’49.8” E, 500–700 m, 9 of the American Orchid Society – “Orchid flora of March 2013, L. Averyanov et al., LA-VN 324 (CPC, relict karstic formation in central Laos (Vientiane NHOL, NUOL). province, Vang Vieng and Kasi districts)”. I Zeuxine Lindl also grateful to Mr. Nguyen Sinh Khang, Dr. E. *Z. flava (Wall. ex Lindl.) Trimen Konstantinov, Mr. Pham Van The and Mrs. T. LAOS, Vientiane prov., Kasi distr., Namken Maisak for their photographs used in the paper and village, Phachao mt., 19°18’45.5” N, 102°22’31.4” Mr. Andre Schuiteman for fruitful discussion on E, 1500 m, 24 March 2013, L. Averyanov, N.S. species identification. I am addressing my particular Khang, S. Lorphengsy LA-VN 777, Fig. 5, H (CPC, thanks to Dr. Thierry Deroin for his careful language LE, NHOL, NUOL). correction and editing this paper. LITERATURE

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