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Les Collections Paleontologiques Du Musee National D'histoire Naturelle De Luxembourg

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Les Collections Paleontologiques Du Musee National D'histoire Naturelle De Luxembourg ISSN 0251 - 2424 MINISTERE DE LA CULTURE TRA VAUX SCIENTIFIQUES DU MUSEE NATIONAL D' HISTOIRE NATURELLE DE LUXEMBOURG 32 Les collections paleontologiques du Musee national d'histoire naturelle de Luxembourg. Fossiles du Trias et du Jurassique par Dominique DELSATE, Chris DUFFIN & Robi WEIS Luxembourg, 1999 Date de publication: 31 decembre 1999 Prix du volu me 32: LUF (=BEF) 550,00 13,63 Les commandes sont a adresser a: Musee national d' histoire naturelle, Bibliotheque/Echange 25, rue Munster, L-2160 Luxembourg Page de couverturc: "Pholidophorus"friedeni sp. nov., Toarcien de Schouweiler ISSN 0251 - 2424 MINISTERE DE LA CULTURE TRAVAUX SCIENTIFIQUES DU MUSEE NATIONAL D'HISTOIRE NATURELLE DE LUXEMBOURG 32 Les collections paleontologiques du Musee national d 'histoire naturelle de Luxembourg. Fossiles du Trias et du Jurassique par Dominique DELSATE, Chris DUFFIN & Robi WEIS Luxembourg, 1999 Trav. sci. Mus. nat. hist. nat. Lux. 32, 1999 Contenu: Delsate D. & C. J. Duffin: A new fish fauna from the Middle Triassic (Upper Muschelkalk) of Moersdorf (Grand Duchy of Luxembourg) ............................................ ... .... p. 5 Delsate D.: Reptiles terrestres (Lepidosauromorpha et Tra­ versodontidae) du Trias superieur de Medemach (G.-D. de Luxembourg) ...................... .. .......................... ... ............. p. 55 Delsate D.: Haasichthys michelsi, nov. gen., nov. sp. , un nouveau Pachycormiforme (Osteichthyes, Actinopterygii) du Toarcien inferieur (Jurassique) luxembourgeois ...... p. 87 Delsate D.: Un Pholidophoridae nouveau (Osteichthyes, Actinopterygii) du Toarcien (Jurassique inferieur) luxem- bourgeois ..... ... .... ............................ ......... ......... ............ p. 141 Weis R.: Die Belernniten der Minette-Formation (ob. Toarcium-ob. Aalenium) Luxemburgs ...................... .. p. 207 3 Trav. sci. Mus. nat. hist. nat. Lux. 32, 1999 A new fish fauna from the Middle Triassic (Upper Muschelkalk) of Moersdorf (Grand Duchy of Luxembourg) by Dominique Delsate1 & Christopher J. Duffin2 1 Musee national d'histoire naturelle de Luxembourg, Departement Paleonto­ logie, 25, Rue Munster L-2160 Luxembourg 2 146, Church Hill Road, Sutton, Surrey SM3 8NF, England Abstract: A new fish fauna is described from the Upper Muschelkalk (Middle Triassic) of Moersdorf in the eastern part of the Grand Duchy of Luxembourg. It contains the chondrichthyans Hybodus plicatilis Agassiz 1843, the morphotypes of Acrodus gaillardoti Agassiz 1837 and Acrodus lateralis Agassiz 1839, Acrodus cfr substriatus (Schmid), Acrodus mutteri sp. nov, Palaeobates angustissimus (Agassiz, 1838), Polyacrodus spp., Lissodus cristatus sp. nov.and Lissodus sp.; a preliminary list of actinopterygians in terms of tooth morphotypes is introduced, mostly indicating the Perleidiformes. Resume: On decrit ici la faune de poissons du Trias moyen, Muschelkalk superieur, de Moersdorf a !'Est du Grand Duche de Luxembourg. Les chondrichthyens sont representes par Hybodus plicatilis Agassiz, 1843, les morphotypes de Acrodus gaillardoti Agassiz 1837 et Acrodus lateralis Agassiz 1839, Acrodus cfr substriatus (Schmid), Acrodus mutteri sp. nov., Palaeobates angustissimus (Agassiz, 1838), Polyacrodus spp., Lissodus cristatus sp. nov.et Lissodus sp. Les morphotypes dentaires des Actinopterygiens sont presentes; ils evoquent principalement les Perleidiformes. 5 1 Introduction 1.1 Geographical and Geological background 1.1.1 Situation Heselbierg Quarry is located on the Luxemburg side of the River Moselle, close to the village of Moersdorf, on road N 19. The quarry is exploited by the Intermoselle society, for road aggregate. Waterlot et al. (1973 p.170-1, figs.84/4, 85) have given a plan and a stratigraphic log of the Quarry. 1.1.2 General geological background of the Luxembourg Triassic Following Bintz, Hary & Muller (1973) and Courel et al. (1984), the Upper Muschelkalk stratigraphy of Luxemburg can be summarised as follows: the (mo) is 60 m thick in the Moselle and Basse-Sure districts, 50 m around Reisdorf and 20 m near Ettelbrlick. The Upper Muschelkalk in Luxemburg consists of a series of platform deposits which are divided into four megasequences.(mol to mo4). The lower part is characterized by Encrinus liliiformis dolomites (mo 1) , formerly known as the "Couches a entroques" forming the three lower sequences (negative sequences grading from argillaceous mudstones into bioclastic and oolitic grainstones). The platform environment is characterised by widespread bioclastic calcarenites, containing terebratuloids and crinoids. Reactivation of the Devonian basement formed a structural high in the Sierck area, and appears to have been the major factor determining the high energy deposition of oolitic sediments there. The upper part of the Upper Muschelkalk is characterised by marls and dolomites beds with rare Ceratites (mo2). In the Sierck anticline, marly beds grade into dolomites. In the last megasequence, the Couches a Ceratites, the influence of Triassic structural highs increases still further: the Sierck area remains the only site of oolitic deposition. This high energy shoal limits a north­ ern mud-carbonate basin with terrigenous influx.(Courel et al. 1984). 6 Fig. 1: Situation map of Moersdorf Quarry. 10 km Fig. 2: A view of the upper part of the quarry. Fig. 3: Hybodus plicatilis, Moersdorf. 7 1.1.3 Local geology The fauna described below was collected from the base of the top bench of the quarry, as it was being worked economically during the 1980's. The unit which was sampled belongs to the 12 m thick Couches a Ceratites, identified as Upper Muschelkalk mo2 on the Luxembourg Geological Survey map. Underlying units consist of the Couches a Entroques layers (mol), some 28 m.thick, which are rich in Encrinus liliiformis columnals, brachiopods, bivalves, and gastropods, with occasional fish and reptile remains (Hary 1965). The base of the quarry exposes 4 m of the Middle Muschelkalk Lingula tenuissima Dolomites (mm2). The whole section consists of a series of alternating Dolomites and Marls. The marls range from a few mm to several metres in thickness. The thick marl beds divide the section into 4 dolomite packs. The conodont fauna has been described by Demonfaucon (1982) and includes the genera Neogondolella and Hindeodella. Ceratites cf subnodosus, and Myophoria sp. have been found by Mr Germain Delsate in the dolomite banks. Lithology: Dolomites with interbedded blue shaly marls. 1.2 Method Some 50 kg of sediment were collected by knife peeling of the laminated dolomites at the interface between the marl and dolomitic beds, from 3 successive cycles. The sediments were disaggregated in 10% H202 at 60°C, left for 24 hours, and then sieved using 0,3 mm and 3 mm meshes. The residue was then sorted under the stereomicroscope. 1.3 Previously described or cited vertebrates from the Heselbierg Quarry The vertebrate fauna as cited in earlier publications is briefly summarised below: Hary 1989: Hybodus sp, Acrodus, Saurichthys sp, Tholodus sp, Gyrolepis sp, "dents de Chondrichthyens" (these specimens actually dermal denticles). The figures also show Perleidiformes cfr Colobodus. 8 Delsate 1992: Hybodus plicatilis, Acrodus gaillardoti, Acrodus lateralis, Acrodus substriatus, Lissodus sp. Delsate 1995, 1997: Hybodus plicatilis, Acrodus cf gaillardoti, Acrodus cf lateralis, Palaeobates angustissimus, Lissodus cf. minimus. 2 Systematic Palaeontology Class Chondrichthyes Huxley 1880 Subclass Elasmobranchii Bonaparte 1838 Cohort Euselachii Hay 1902 Superfamily Hybodontoidea Zangerl 1981 Family Hybodontidae Owen 1846 Genus Hybodus Agassiz 1837 Type species: H. reticulatus Agassiz 1837; Hettangian to Sinemurian, Early Jurassic, Lyme Regis, Dorset, England. 2.1 Hybodus plicatilis Agassiz 1843 1837 Hybodus plicatilis Geinitz (ex Agassiz MS), 22, plate 3 fig. 8. 1837 Hybodus plicatilis Hohard, plate 2 figs. 8-10. 1843 Hybodus plicatilis Agassiz Agassiz, 3, 189, plate 22a figure 1, plate 24 figs. 10, 13 . (?) 1843 Hybodus longiconus Agassiz, 3, 191 , plate 24 figs. 19-21, 23. 1844 Hybodus plicatilis Meyer & Plieninger, 111, plate 12 figs. 51 , 70, 71. 1844 Hybodus longiconus Meyer & Plieninger, 56, plate 12 figs. 54, 56. 1849 Hybodus plicatilis Meyer, 224, plate 28 figs. 35, 36, 40. 1849 Hybodus mougeoti Meyer, 225, plate 28 fig . 37. 1849 Hybodus obliquus Meyer, 227, plate 28 fig. 41. (?) 1849 Hybodus longiconus Meyer, 227, plate 28 fig. 39. 1850 Hybodus plicatilis Naumann, plate 24 fig. 11. 1851 Hybodus plicatilis Bronn, plate 11 fig. 18. 1851 Hybodus plicatilis Meyer, 224, plate 28 fig. 35. 1852 Hybodus plicatilis Gervais, 12, plate 77 fig. 3. 1855 Hybodus plicatilis Giebel, 114, plate 47 figs. 15a, !Sc, 20. 1856 Hybodus plicatilis Giebel, 426, plate 1 fig. 6. 1857 Hybodus plicatilis Chop, 128. 1857 Hybodus plicatilis Geinitz, 22, plate 3 fig. 8. 1861 Hybodus plicatilis Schmid, 18, plate 3 fig. 9. 1861 Hybodus obliquus Schmid, 19, plate 3 figs.1-3. 1861 Hybodus angustus Schmid, 19, plate 3 figs. 10-12. 1861 Hybodus mougeoti Schmid, 19, plate 3 figs. 7-8. 9 1861 Hybodus longiconus Schmid, 19, plate 3 figs. 4-6. 1865 Hybodus plicatilis Eck, 65 . 1865 Hybodus longiconus Eck, 119 (pars). 1870 Hybodus plicatilis Roemer, plate 12 figs. 13, 14. 1883 Hybodus plicatilis Sauvage, 495, plate 12 figs. 6-11. 1885 Hybodus plicatilis Quenstedt, 276 plate 21 figs. 26, 27. 1887 Hybodus plicatilis Zittel, 67, fig. 62a. 1889 Hybodus plicatilis Woodward, 250. 1889 Hybodus plicatilis Jaekel, 305, plate 7 figs. 1-3. 1903 Hybodus plicatilis Frech, 11 plate 1 fig. 2. 1906 Hybodus
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