Crocodile Specialist Group Newsletter 25(1): 5-6]
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8. Archosaur Phylogeny and the Relationships of the Crocodylia
8. Archosaur phylogeny and the relationships of the Crocodylia MICHAEL J. BENTON Department of Geology, The Queen's University of Belfast, Belfast, UK JAMES M. CLARK* Department of Anatomy, University of Chicago, Chicago, Illinois, USA Abstract The Archosauria include the living crocodilians and birds, as well as the fossil dinosaurs, pterosaurs, and basal 'thecodontians'. Cladograms of the basal archosaurs and of the crocodylomorphs are given in this paper. There are three primitive archosaur groups, the Proterosuchidae, the Erythrosuchidae, and the Proterochampsidae, which fall outside the crown-group (crocodilian line plus bird line), and these have been defined as plesions to a restricted Archosauria by Gauthier. The Early Triassic Euparkeria may also fall outside this crown-group, or it may lie on the bird line. The crown-group of archosaurs divides into the Ornithosuchia (the 'bird line': Orn- ithosuchidae, Lagosuchidae, Pterosauria, Dinosauria) and the Croco- dylotarsi nov. (the 'crocodilian line': Phytosauridae, Crocodylo- morpha, Stagonolepididae, Rauisuchidae, and Poposauridae). The latter three families may form a clade (Pseudosuchia s.str.), or the Poposauridae may pair off with Crocodylomorpha. The Crocodylomorpha includes all crocodilians, as well as crocodi- lian-like Triassic and Jurassic terrestrial forms. The Crocodyliformes include the traditional 'Protosuchia', 'Mesosuchia', and Eusuchia, and they are defined by a large number of synapomorphies, particularly of the braincase and occipital regions. The 'protosuchians' (mainly Early *Present address: Department of Zoology, Storer Hall, University of California, Davis, Cali- fornia, USA. The Phylogeny and Classification of the Tetrapods, Volume 1: Amphibians, Reptiles, Birds (ed. M.J. Benton), Systematics Association Special Volume 35A . pp. 295-338. Clarendon Press, Oxford, 1988. -
01 Oliveira & Pinheiro RBP V20 N2 COR.Indd
Rev. bras. paleontol. 20(2):155-162, Maio/Agosto 2017 © 2017 by the Sociedade Brasileira de Paleontologia doi: 10.4072/rbp.2017.2.01 ISOLATED ARCHOSAURIFORM TEETH FROM THE UPPER TRIASSIC CANDELÁRIA SEQUENCE (HYPERODAPEDON ASSEMBLAGE ZONE, SOUTHERN BRAZIL) TIANE MACEDO DE OLIVEIRA & FELIPE L. PINHEIRO Laboratório de Paleobiologia, Universidade Federal do Pampa, Campus São Gabriel, R. Aluízio Barros Macedo, BR 290, km 423, 97300-000, São Gabriel, RS, Brazil. [email protected], [email protected] ABSTRACT – We describe isolated teeth found in the locality “Sítio Piveta” (Hyperodapedon Assemblage Zone, Candelaria Sequence, Upper Triassic of the Paraná Basin). The material consists of five specimens, here classified into three different morphotypes. The morphotype I is characterized by pronounced elongation, rounded base and symmetry between lingual and labial surfaces. The morphotype II presents serrated mesial and distal edges, mesial denticles decreasing in size toward the base, distal denticles present until the base and asymmetry, with a flat lingual side and rounded labial side. The morphotype III, although similar to morphotype II, has a greater inclination of the posterior carinae. The conservative dental morphology in Archosauriformes makes difficult an accurate taxonomic assignment based only on isolated teeth. However, the specimens we present are attributable to “Rauisuchia” (morphotype II and III) and, possibly, Phytosauria (morphotype I). The putative presence of a phytosaur in the Carnian Hyperodapedon Assemblage Zone would have impact in the South American distribution of the group. The taxonomic assignments proposed herein contribute to the faunal composition of the Hyperodapedon Assemblage Zone, a critical unit on the study of the Upper Triassic radiation of archosaurs. -
On the Presence of the Subnarial Foramen in Prestosuchus Chiniquensis (Pseudosuchia: Loricata) with Remarks on Its Phylogenetic Distribution
Anais da Academia Brasileira de Ciências (2016) (Annals of the Brazilian Academy of Sciences) Printed version ISSN 0001-3765 / Online version ISSN 1678-2690 http://dx.doi.org/10.1590/0001-3765201620150456 www.scielo.br/aabc On the presence of the subnarial foramen in Prestosuchus chiniquensis (Pseudosuchia: Loricata) with remarks on its phylogenetic distribution LÚCIO ROBERTO-DA-SILVA1,2, MARCO A.G. FRANÇA3, SÉRGIO F. CABREIRA3, RODRIGO T. MÜLLER1 and SÉRGIO DIAS-DA-SILVA4 ¹Programa de Pós-Graduação em Biodiversidade Animal, Universidade Federal de Santa Maria, Av. Roraima, 1000, Bairro Camobi, 97105-900 Santa Maria, RS, Brasil ²Laboratório de Paleontologia, Universidade Luterana do Brasil, Av. Farroupilha, 8001, Bairro São José, 92425-900 Canoas, RS, Brasil ³Laboratório de Paleontologia e Evolução de Petrolina, Campus de Ciências Agrárias, Universidade Federal do Vale do São Francisco, Rodovia BR 407, Km12, Lote 543, 56300-000 Petrolina, PE, Brasil 4Centro de Apoio à Pesquisa da Quarta Colônia, Universidade Federal de Santa Maria, Rua Maximiliano Vizzotto, 598, 97230-000 São João do Polêsine, RS, Brasil Manuscript received on July 1, 2015; accepted for publication on April 15, 2016 ABSTRACT Many authors have discussed the subnarial foramen in Archosauriformes. Here presence among Archosauriformes, shape, and position of this structure is reported and its phylogenetic importance is investigated. Based on distribution and the phylogenetic tree, it probably arose independently in Erythrosuchus, Herrerasaurus, and Paracrocodylomorpha. In Paracrocodylomorpha the subnarial foramen is oval-shaped, placed in the middle height of the main body of the maxilla, and does not reach the height of ascending process. In basal loricatans from South America (Prestosuchus chiniquensis and Saurosuchus galilei) the subnarial foramen is ‘drop-like’ shaped, the subnarial foramen is located above the middle height of the main body of the maxilla, reaching the height of ascending process, a condition also present in Herrerasaurus ischigualastensis. -
New Insights on Prestosuchus Chiniquensis Huene
New insights on Prestosuchus chiniquensis Huene, 1942 (Pseudosuchia, Loricata) based on new specimens from the “Tree Sanga” Outcrop, Chiniqua´ Region, Rio Grande do Sul, Brazil Marcel B. Lacerda1, Bianca M. Mastrantonio1, Daniel C. Fortier2 and Cesar L. Schultz1 1 Instituto de Geocieˆncias, Laborato´rio de Paleovertebrados, Universidade Federal do Rio Grande do Sul–UFRGS, Porto Alegre, Rio Grande do Sul, Brazil 2 CHNUFPI, Campus Amı´lcar Ferreira Sobral, Universidade Federal do Piauı´, Floriano, Piauı´, Brazil ABSTRACT The ‘rauisuchians’ are a group of Triassic pseudosuchian archosaurs that displayed a near global distribution. Their problematic taxonomic resolution comes from the fact that most taxa are represented only by a few and/or mostly incomplete specimens. In the last few decades, renewed interest in early archosaur evolution has helped to clarify some of these problems, but further studies on the taxonomic and paleobiological aspects are still needed. In the present work, we describe new material attributed to the ‘rauisuchian’ taxon Prestosuchus chiniquensis, of the Dinodontosaurus Assemblage Zone, Middle Triassic (Ladinian) of the Santa Maria Supersequence of southern Brazil, based on a comparative osteologic analysis. Additionally, we present well supported evidence that these represent juvenile forms, due to differences in osteological features (i.e., a subnarial fenestra) that when compared to previously described specimens can be attributed to ontogeny and indicate variation within a single taxon of a problematic but important -
SP's Aviation Cover 8-2019.Indd 1 07/09/19 5:22 PM Design Holistic Directions: Jayant Baranwal Creative Director: Anoop Kamath Designers: Vimlesh Kumar Yadav, Sonu S
SP’s AN SP GUIDE PUBLICATION A-BASED BUYER ONLY) BUYER A-BASED I 100.00 (IND ` aviationSHARP CONTENT FOR A SHARP AUDIENCE www.sps-aviation.com vol 22 ISSUE 8 • 2019 MILITARY • INDIA INDUCTs 8 apACHE GUNSHIPS • FRANCE STRONGLY PITCHEs 36 ADDITIONAL RAFALE LAST WORD: CHANDRAYAAN 2 FAILURES ARE PILLARS OF SUCCESS 50 YEARS OF EMBRAER PAGE 5 YEARS OF RNI NUMBER: DELENG/2008/24199 5050APOLLO 11 EXCELLENCE IS CONSTANT ACTION, NOT A POSITION. At Lufthansa Technik, excellence is part of our corporate DNA. For more than six decades we have been providing the technology that keeps fleets flying and turns a good aircraft into a great one. Today, we are proud that our company’s name is synonymous with high quality and in-depth knowledge. And we continue to shape the aviation industry – by embracing new challenges and exceeding our customers’ expectations. CSABA MÉDER • MAINTENANCE PROJECT LEADER EXCELLENCE IS CONSTANT ACTION, NOT A POSITION. At Lufthansa Technik, excellence is part of our corporate DNA. For more than six decades we have been providing the technology that keeps fleets flying and turns a good aircraft into a great one. Today, we are proud that our company’s name is synonymous with high quality and in-depth knowledge. And we continue to shape the aviation industry – by embracing new challenges and exceeding our customers’ expectations. CSABA MÉDER • MAINTENANCE PROJECT LEADER PUBLISHER AND EDITOR-IN-CHIEF Jayant Baranwal SENIOR EDITOR Air Marshal B.K. Pandey (Retd) TABLE OF CONTENTS DEPUTY MANAGING EDITOR Neetu Dhulia SENIOR TECHNICAL GROUP EDITOR Lt General Naresh Chand (Retd) GROUP EXECUTIVE EDITOR AN SP GUIDE PUBLICATION Vishal Thapar SP’s CONTRIBUTORS ONLY) BUYER A-BASED I India: Group Captain A.K. -
Heptasuchus Clarki, from the ?Mid-Upper Triassic, Southeastern Big Horn Mountains, Central Wyoming (USA)
The osteology and phylogenetic position of the loricatan (Archosauria: Pseudosuchia) Heptasuchus clarki, from the ?Mid-Upper Triassic, southeastern Big Horn Mountains, Central Wyoming (USA) † Sterling J. Nesbitt1, John M. Zawiskie2,3, Robert M. Dawley4 1 Department of Geosciences, Virginia Tech, Blacksburg, VA, USA 2 Cranbrook Institute of Science, Bloomfield Hills, MI, USA 3 Department of Geology, Wayne State University, Detroit, MI, USA 4 Department of Biology, Ursinus College, Collegeville, PA, USA † Deceased author. ABSTRACT Loricatan pseudosuchians (known as “rauisuchians”) typically consist of poorly understood fragmentary remains known worldwide from the Middle Triassic to the end of the Triassic Period. Renewed interest and the discovery of more complete specimens recently revolutionized our understanding of the relationships of archosaurs, the origin of Crocodylomorpha, and the paleobiology of these animals. However, there are still few loricatans known from the Middle to early portion of the Late Triassic and the forms that occur during this time are largely known from southern Pangea or Europe. Heptasuchus clarki was the first formally recognized North American “rauisuchian” and was collected from a poorly sampled and disparately fossiliferous sequence of Triassic strata in North America. Exposed along the trend of the Casper Arch flanking the southeastern Big Horn Mountains, the type locality of Heptasuchus clarki occurs within a sequence of red beds above the Alcova Limestone and Crow Mountain formations within the Chugwater Group. The age of the type locality is poorly constrained to the Middle—early Late Triassic and is Submitted 17 June 2020 Accepted 14 September 2020 likely similar to or just older than that of the Popo Agie Formation assemblage from Published 27 October 2020 the western portion of Wyoming. -
Macroevolutionary Patterns in the Evolutionary Radiation of Archosaurs (Tetrapoda: Diapsida) Stephen L
Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 101, 367–382, 2011 (for 2010) Macroevolutionary patterns in the evolutionary radiation of archosaurs (Tetrapoda: Diapsida) Stephen L. Brusatte1,2, Michael J. Benton3, Graeme T. Lloyd4, Marcello Ruta3 and Steve C. Wang5 1 Division of Paleontology, American Museum of Natural History, Central Park West at 79th Street, New York, NY 10024, USA Email: [email protected] 2 Department of Earth and Environmental Sciences, Columbia University, New York, NY, USA 3 School of Earth Sciences, University of Bristol, Wills Memorial Building, Queens Road, Bristol, BS8 1RJ, UK 4 Department of Palaeontology, Natural History Museum, Cromwell Road, London SW7 5BD, UK 5 Department of Mathematics and Statistics, Swarthmore College, Swarthmore, PA 19081, USA ABSTRACT: The rise of archosaurs during the Triassic and Early Jurassic has been treated as a classic example of an evolutionary radiation in the fossil record. This paper reviews published studies and provides new data on archosaur lineage origination, diversity and lineage evolution, morpho- logical disparity, rates of morphological character change, and faunal abundance during the Triassic–Early Jurassic. The fundamental archosaur lineages originated early in the Triassic, in concert with the highest rates of character change. Disparity and diversity peaked later, during the Norian, but the most significant increase in disparity occurred before maximum diversity. Archo- saurs were rare components of Early–Middle Triassic faunas, but were more abundant in the Late Triassic and pre-eminent globally by the Early Jurassic. The archosaur radiation was a drawn-out event and major components such as diversity and abundance were discordant from each other. -
“Means of Connecting the Contemporary Indo-Fijian Women Towards the Imaginary Homeland.”
“Means of Connecting the Contemporary Indo-Fijian Women Towards the Imaginary Homeland.” Rohini Ronita Lata “Means of Connecting the Contemporary Indo-Fijian Women Towards the Imaginary Homeland.” by Rohini Ronita Lata A supervised research project submitted in partial fulfillment of the requirements for the degree of Master of Arts in Literature Copyright © 2009 by Rohini Ronita Lata School of Language Arts and Media, Faculty of Arts and Law The University of the South Pacific September, 2009 Acknowledgement I am indebted to a number of individuals with out whom this research would have been unattainable. Firstly, my supervisor, Dr. Mohit Prasad for his continuous patience and endurance with my drafts and his facilitation and insights which helped shape this thesis. Secondly, I would like to affectionately thank the Indo-Fijian women of the Samabula area (primary research) who assisted in the research by filling in the surveys and answering the questionnaires. Thirdly, my sincere gratitude to Ms. Shalini Prasad for her assistance in supplying me with the films which I required for this research and providing an overview of the Hindi satellite television’s daily-life drama series and Mr. Richard Ram for assisting in the typing of the script. My heart felt gratitude to Ms. Artika Narayan for her assistance in providing printouts of the research paper for the editing phase, facilitating with the page setup and constructing the graphical representations for the primary research and Ms. Maelin Bhagwan for her efforts in proofreading the research paper before the final submission. My deepest gratitude is reserved for my family and God without whom I would not have been able to ‘decorate in ink.’ Abstract The ‘Old Indo-Fijian Diaspora,’ of the indentured labourers who came to Fiji between 1879 and 1920 has been an important area of scholarship and research. -
ANNUAL REPORT 2018–19 NATIONAL LIBRARY of AUSTRALIA Annual Report 2018–19
2018–19 Annual Report Report Annual NATIONAL LIBRARY OF AUSTRALIA OF AUSTRALIA LIBRARY NATIONAL ANNUAL REPORT 2018–19 NATIONAL LIBRARY OF AUSTRALIA GOVERNANCE AND ACCOUNTABILTY i NATIONAL LIBRARY OF AUSTRALIA Annual Report 2018–19 NATIONAL LIBRARY OF AUSTRALIA 9 August 2019 The Hon. Paul Fletcher MP Minister for Communications, Cyber Safety and the Arts Parliament House CANBERRA ACT 2600 Dear Minister National Library of Australia Annual Report 2018–19 The Council, as the accountable authority of the National Library of Australia, has pleasure in submitting to you for presentation to each House of Parliament its annual report covering the period 1 July 2018 to 30 June 2019. Published by the National Library of Australia The Council approved this report at its meeting in Canberra on 9 August 2019. Parkes Place Canberra ACT 2600 The report is submitted to you in accordance with section 46 of the T 02 6262 1111 Public Governance, Performance and Accountability Act 2013. F 02 6257 1703 National Relay Service 133 677 We commend the annual report to you. nla.gov.au/policy/annual.html Yours sincerely ABN 28 346 858 075 © National Library of Australia 2019 ISSN 0313-1971 (print) 1443-2269 (online) National Library of Australia The Hon. Dr Brett Mason Dr Marie-Louise Ayres Annual report / National Library of Australia.–8th (1967/68)– Chair of Council Director-General Canberra: NLA, 1968––v.; 25 cm. Annual. Continues: National Library of Australia. Council. Annual report of the Council = ISSN 0069-0082. Report year ends 30 June. ISSN 0313-1971 = Annual report–National Library of Australia. -
Triassic Vertebrate Fossils in Arizona
Heckert, A.B., and Lucas, S.G., eds., 2005, Vertebrate Paleontology in Arizona. New Mexico Museum of Natural History and Science Bulletin No. 29. 16 TRIASSIC VERTEBRATE FOSSILS IN ARIZONA ANDREW B. HECKERT1, SPENCER G. LUCAS2 and ADRIAN P. HUNT2 1Department of Geology, Appalachian State University, ASU Box 32067, Boone, NC 28608-2607; [email protected]; 2New Mexico Museum of Natural History & Science, 1801 Mountain Road NW, Albuquerque, NM 87104-1375 Abstract—The Triassic System in Arizona has yielded numerous world-class fossil specimens, includ- ing numerous type specimens. The oldest Triassic vertebrates from Arizona are footprints and (largely) temnospondyl bones from the Nonesian (Early Triassic: Spathian) Wupatki Member of the Moenkopi Formation. The Perovkan (early Anisian) faunas of the Holbrook Member of the Moenkopi Formation are exceptional in that they yield both body- and trace fossils of Middle Triassic vertebrates and are almost certainly the best-known faunas of this age in the Americas. Vertebrate fossils of Late Triassic age in Arizona are overwhelmingly body fossils of temnospondyl amphibians and archosaurian reptiles, with trace fossils largely restricted to coprolites. Late Triassic faunas in Arizona include rich assemblages of Adamanian (Carnian) and Revueltian (early-mid Norian) age, with less noteworthy older (Otischalkian) assemblages. The Adamanian records of Arizona are spectacular, and include the “type” Adamanian assemblage in the Petrified Forest National Park, the world’s most diverse Late Triassic vertebrate fauna (that of the Placerias/Downs’ quarries), and other world-class records such as at Ward’s Terrace, the Blue Hills, and Stinking Springs Mountain. The late Adamanian (Lamyan) assemblage of the Sonsela Member promises to yield new and important information on the Adamanian-Revueltian transition. -
South Asian Primate Network – SAPN Representing the IUCN/SSC Primate Specialist Group in South Asia
South Asian Primate Network – SAPN Representing the IUCN/SSC Primate Specialist Group in South Asia Sponsored by Margot Marsh Biodiversity Fund & Conservation International Primate Action Fund Produced by Zoo Outreach Organization & IUCN/SSC Primate Specialist Group, South Asia March 2017 (Tenth electronic publication) ZOO OUTREACH ORGANIZATION 12, Thiruvannamalai Nagar, Saravanampatti, Coimbatore 641035, Tamil Nadu, India Ph: +91 9385339862 & 9385339863 Email: [email protected] www.zooreach.org, www.zoosprint.zooreach.org, www.southasianprimatenetwork.zooreach.org INTRODUCTION This Directory is a long pending commitment to the South Asian Primate Network, which represents the Primate Specialist Group and is funded by the Primate Action Fund. The objective of the Directory is to assist people who study or otherwise help the primates of South Asia to know about and find one another. You will note just in a very casual read, that most people do not have information under their name, except for their address (and some, not even that). The reason is that they haven't supplied this information as yet. For that reason we have deleted the terminology of "member" from the network and substituted "associate" because there may be people who are hesitant to join something. The Directory is only as useful as the amount and currency of its information. So if you have not contributed your C.V., interests, achievements, hopes, dreams, etc. please do so. If you have corrections which need to be made for your section could you do us a big favour and simply use the copy tool in PDF to copy your whole entry, paste it into a word file and then make your corrections, using our same format. -
Diversificação De Archosauromorpha Após a Extinção Permo-Triássica Survivors: Diversification of Archosauromorpha After Permo-Triassic Extinction
Terræ REVISÃO 10.20396/td.v16i0.8656060 Didatica Sobreviventes: Diversificação de Archosauromorpha após a Extinção Permo-Triássica SURVIVORS: DIVERSIFICATION OF ARCHOSAUROMORPHA AFTER PERMO-TRIASSIC EXTINCTION TIANE MACEDO DE-OLIVEIRA¹, FELIPE LIMA PINHEIRO², LEONARDO KERBER³ 1 - PROGR. PÓS-GRAD. BIODIVERSIDADE ANIMAL, UNIVERSIDADE FEDERAL DE SANTA MARIA (UFSM). SANTA MARIA, RS, BRASIL. 2 - LABORATÓRIO DE PALEOBIOLOGIA, UNIPAMPA, SÃO GABRIEL, RS, BRASIL. 3 - CENTRO DE APOIO À PESQUISA PALEONTOLÓGICA DA QUARTA COLÔNIA (CAPPA-UFSM), SÃO JOÃO DO POLÊSINE, RS, BRASIL. E-MAIL: [email protected], [email protected], [email protected] Abstract: The Archosauromorpha is defined as the clade that comprises all diapsids more closely related Citation/Citação: De-Oliveira, T. M., to the lineage of birds (Ornithodira or Avemetatarsalia) and crocodylians (Pseudosuchia or Crurotarsi) than Pinheiro, F. L., & Kerber, L. (2020). to Lepidosauria. In addition to their ‘basal’ taxa (e.g., Tanystropheidae, Rhynchosauria, and Allokotosau- Sobreviventes: Diversificação de Ar- ria), Archosauromorpha includes the clade Archosauriformes, which, by its turn, includes Archosauria. chosauromorpha após a Extinção During the Triassic, the archosauromorphs became the main group of diapsids to diversify in body size Permo-Triássica. Terræ Didatica, and morphological disparity. The Permo-Triassic mass extinction occurred 252 million years ago and 16, 1-23, e020009. doi: 10.20396/ produced a dramatic change in the composition of the floral and faunal communities at the beginning td.v16i0.8656060 of the Triassic period. The diversification of Archosauromorpha during the Triassic has been the focus of several recent studies, as this clade is a classic example of adaptive radiation. This paper synthesizes Keywords: Diapsida.