GORTANIA. Geologia,GORTAN Paleontologia,IA Paletnologia 34 (2012) U X SS : Geologia, Paleontologia, Paletnologia 34 (2012) 13-36 dine, 10. II.2013 I N 2038-0410

Angelo Mossoni FAMENNIAN (LATE DEVONIAN) CONODONTS Carlo Corradini Monica Pondrelli FROM THE PIZZUL WEST SECTION (CARNIC ALPS, ITALY)

CONODONTI DEL FAMENNIANO (DEVONIANO SUPERIORE) DALLA SEZIONE MONTE PIZZUL OVEST (ALPI CARNICHE, ITALIA)

Riassunto breve - Vengono illustrati conodonti dalla sezione Pizzul Ovest, situata nell’area del Passo del Cason di Lanza/ Monte Zermula nelle Alpi Carniche centrali. La sezione, potente circa 24 metri, è costituita da calcari del Devoniano Superiore. L’associazione comprende 41 taxa di conodonti, che permettono di riconoscere sette biozone del Frasniano e del Famenniano: Upper rhenana, Upper crepida, Uppermost crepida, Lower rhomboidea, Upper rhomboidea, Lower marginifera e Lower expansa. Parole chiave: Conodonti, Biostratigra+ a , Devoniano Superiore, Alpi Carniche.

Abstract - Conodonts from the Pizzul West section are presented.  e section is located in the Cason di Lanza/Mt. Zermula area of the central Carnic Alps and it exposes about twenty-four metres of Upper Devonian limestone.  e forty-one taxa documented allow the discrimination of seven biozones of Frasnian and Famennian: Upper rhenana, Upper crepida, Uppermost crepida, Lower rhomboidea, Upper rhomboidea, Lower marginifera and Lower expansa. Key-words: Conodonts, Biostratigraphy, Upper Devonian, Carnic Alps.

Introduction along the section, one more (PZW Z) have been picked several meters to the south, and its relationship with the ! e Clymeniae limestone have been extensively section is not clear. ! e sample came from an irregular investigated for biostratigraphic purposes in the Carnic bed just below the Carboniferous sediments of the Alps in the last decades. ! e unit crops out widely along Hochwipfel Formation, very distinct in the + eld, being the Carnic Alps, but most of the research have been constituted by irregular gravels and cobbles scattered in concentrated in the central-western part of the chain, a grey micritic cement mainly in Wolayer, Passo di Monte Croce Carnico, Timau-Pramosio areas. From the central-eastern part of the Carnic Alps data available are by far less abundant, Geological settings probably because of the less spectacular outcrops, and/ or more di" cult accessibility of some areas. ! e succession of the Mt. Pizzul area ranges from In this paper we present the conodont fauna and Ordovician to Carboniferous (C' $!*%* et al. biostratigraphy from a section located in the Clymeniae 2012, 2013; P'%! "##* et al. 2011). ! e rocks here Limestone on the western # ank of Mt. Pizzul, in the exposed belong to the Variscan sequences of the central part of the Carnic Alps, just south of Passo Carnic Palaeozoic, that were a< e cted by the Variscan del Cason di Lanza (Fig. 1). From these area only orogeny during the Westphalian and Alpine tectonics, M$%&'%* (1966) + gured a few latest Devonian-early including both extensional and compressional phases, Carboniferous conodonts from some spot localities that involved the whole Carnic area starting from the along the crest of Mt. Zermula massif. Cenozoic (V"%$% *%* 1990a; V"%$% *%* et al. 2009). ! e Pizzul West (PZW) section is located in a First ! e oldest unit cropping in the area is represented World War trench on the western # ank of Mt Pizzul at by the Upper Ordovician “Uqua shales”, consisting of q. 1905, at coordinates 46° 33’ 21,67’’ N, 13° 18’ 18,16’’ highly fossiliferous shales, siltstones, sandstones and E (Fig. 1). About 24 meters of Clymeniae limestones are rare conglomerates. ! e sequence continues with few exposed, even if some tectonic elisions and repetitions meters of nodular limestone (“Uqua limestones”), and a< ected the s e ction. B e side the thirteen samples collected calcareous sandstones (Plöcken Formation). Silurian

13 A. MOSSONI, C. CORRADINI, M. PONDRELLI GORTANIA. Geologia, Paleontologia, Paletnologia 34 (2012)

and drowned (V"%$% *%* et al. 2009 and reference therein). ! e Upper Devonian is almost exclusively represented by “Clymeniae limestones”: pelagic massive and/or nodular limestones, cropping out at Forca di Lanza and on the western # a nk of Mt. Pizzul. ! e sequence of Mt. Pizzul area ends with the Hochwipfel Fm.: gravitative driven accumulation of breccias, conglomerates, sandstones and pelites originated by the Lower Carboniferous transtensional to transpressional tectonics (S$##"$$$ et al. 1980). For a more detailed description of the geology of the Cason di Lanza - Mt. Pizzul area refer to C' $!*%* et al. (2012, 2013).

 e “Clymeniae limestones”

! e “Clymeniae limestones” crops out widely in the Carnic Alps, and have been extensively studied by several authors. ! is informal unit of Frasnian- Tournaisian age is indicated in literature with various names, according to the di< e rent authors: Pal Fm., Gross Pal Fm., Calcari di Pramosio, etc. It consists of grey massive limestone, grey to moderate pink and red very thin to thin (rarely medium) bedded nodular mudstone to wackestone. In the lower part of the units a few levels of breccia occur in some localities. ! e depositional environment is interpreted as pelagic (e.g., S€%#$%‚ 1992) with local gravitative-driven Fig. 1 - Location map. deposits near the base of the unit. - Ubicazione della sezione. ! e fossil content is represented by trilobites, ostracodes, radiolarians and conodonts, and less rocks are poorly exposed and are represented only abundant echinoderms, molluscs, bivalves, brachiopods by a few meters of “Orthoceras limestones” (Alticola and + sh teeth (S€%#$%‚ 1992). Formation) of Pridoli age. A detailed biostratigraphy of the unit have been Compared to the older terms of the sequence, provided by several authors on the basis of the rich Devonian rocks are largely more abundant and conodont associations (for a summary see P" * & di< erentiated. ! e oldest unit is the Rauchkofel Fm., S$##"$$$ 1998a). constituted by dark cephalopod limestone with black shales interbedded, followed by the middle-upper Lochkovian La Valute Fm: a light grey-ochre nodular The Pizzul West section limestone. ! e sequence continues with the Findenig Fm., which consists of nodular purple red mudstones ! e Pizzul West section (Fig. 2) exposes about and wackestones with marly millimetric thick 24 meters of pelagic mudstones-wackestones of intercalations. the Clymeniae limestones (Fig. 3). ! e lower and Starting from around the Lower-Middle Devonian central part of the section (below sample PZW 1) is boundary, the basin started to di< e rentiate: in the tectonically disturbed and is a< ected by folds and/or shallower parts a thick reefal sequence started to faults, as con+ rmed by conodont data (see below for build up, represented in the area by the white cli< s discussion). of Mt. Zermula. In the deeper parts of the basin, now ! r ee di< e rent facies can be distinguished in the represented by the units cropping out in Mt. Pizzul - section: a light grey massive micritic limestone, a Forca di Lanza area, gravitative driven deposits from dark red nodular limestone and a grey-ochre nodular the reef, intecalated with narrow pelagic levels, were limestone (Fig. 3). In general massive grey limestone deposited. ! ese sediments belongs to the Vinz and are more abundant in the lower part, whereas the red Hohe Trieb formations. nodular limestone prevails in the upper part. A few thin During the Frasnian the Carnic basin underwent pelitic levels are present in the section between sample extensional tectonic pulses and the reefal facies collapsed PZW 1 and sample PZW 4.

14 GORTANIA. Geologia, Paleontologia, Paletnologia 34 (2012) FAMENNIAN CONODONTS FROM THE PIZZUL WEST SECTION CARNIC ALPS, ITALY

! e microfacies of the grey limestone consists of ! e grey-ochre nodular limestone consists of a a wackestone with a light grey color and few fossils wackstone-packstone similar to the red one, but without remains scattered in the matrix (mostly ostracods the haematite precipitations that most probably give the and shells); some stylolite structures are also evident red color to the former unit (Fig. 4). (Fig. 4). Beside the samples collected along the section, one ! e red nodular facies consists of a wackstone- more sample (PZW Z) has been picked several meters packstone with nodules up to 1 cm of diameter, to the south, and its relationship with the section is probably due to a synsedimentary diagenesis, with not clear. ! e sample came from an irregular bed just haematite precipitations (Fig. 4). below the Carboniferous sediments of the Hochwipfel ! e fossil contents is higher and includes trilobites, Formation, very distinct in the + eld, being constituted small shells (brachiopods or bivalves), ostracods, a few by irregular gravels and cobbles scattered in a grey- cephalopods and sponge spiculae. brownish micritic cement (Fig. 4).

a b

1 6 2 3 7 5A 4 5 c

d e

Fig. 2 - Views of the Pizzul West section. a) Panoramic view of Mt. Pizzul with indicated in red the position of section; b) general view of the section in the First World War trench; c) the undisturbed part of the section, with location of samples; d) detail of the reddish nodular limestone; e) the irregular level constituted by gravels and cobbles scattered in a grey micritic cement (sample PZW Z). - Vedute della sezione Pizzul Ovest. a) veduta panoramica di M. Pizzul con la sezione indicata in rosso; b) veduta generale della sezione nella trincea della Prima Guerra Mondiale; c) la parte indisturbata della sezione, con ubicazione dei campioni; d) dettaglio del calcare nodulare rosso; e) livello irregolare con litici immersi in un cemento micritico (campione PZW Z)

15 A. MOSSONI, C. CORRADINI, M. PONDRELLI GORTANIA. Geologia, Paleontologia, Paletnologia 34 (2012)

ata ob era d ea cilis ta a ae s er ra k ut na a ra a tu ex oi er ginif t is a ata ob tinodosal ar cu ecti ar lab id ea lab oglab ob o l ern ii a a wols a l a min a subg ul c u s en sis eli . m a prima b ata s c hindewol  ran lt g er ae ip unctata ata er nda g ra p ra g ra a er g er e ut ut ut ut e u pid pp r lo omb tu ar ob e uad lt. a en ami a. m a. sto a. rh a. glab a. j a. ro a. subp r a. cf. a. min a. simpla a. q a. glab a. min a. cr a. glab a. min a. min cr. a cr. cr. olivi cr. P P I Pa. glabr Pa. pe P P glab Po. nodocostaus nod. Po. curvata Anc. l Anc. nodosa Anc. P lyaiol Pa. P P P P P Po. normalis Po. miri  Pa. q uadr. in P t Pa. P P P I P glab Po. P P

LEGEND / LEGENDA Massive grey limestone Calcare massivo grigio Nodular grey limestone Calcare nodulare grigio Nodular red limestone Calcare nodulare rosso Pelitic levels / Livelli pelitici Tectonic contact / Contatto tettonico 1 m

Fig. 3 - Stratigraphic log of the Pizzul West section and conodont distribution - Colonna stratigra ca e distribuzione dei conodonti nella sezione Pizzul Ovest.

16 GORTANIA. Geologia, Paleontologia, Paletnologia 34 (2012) FAMENNIAN CONODONTS FROM THE PIZZUL WEST SECTION CARNIC ALPS, ITALY

Fig. 4 - Microfacies of the Pizzul West section (scale bar 0,5 cm). a) Grey massive limestone with some stylolite structures from the sample PZW 1; b) Red nodular limestone with haematite precipitations and nodules (Sample PZW 5); c) grey-ochre nodular facies of the sample PZW 5A; d) + ne-grained breccia of sample PZW Z. - Microfacies della sezione Pizzul Ovest (scala 0,5 cm). a) Calcare grigio massivo con strutture stilolitiche del campione PZW 1; b) precipitazioni di ematite e noduli millimetrici nella facies nodulare rossa (campione PZW 5); c) facies grigio-ocracea nodulare del campione PZW 5A; d) breccia  ne del campione PZW Z.

Conodont data Palmatolepis is the predominant genus, so the whole Famennian part of the section belongs to the Fourteen samples (Fig. 3), weighting from 1.2 to palmatolepid-bispathodid biofacies of S$%!‚"  2.6 kg, have been collected from the Pizzul West (1976). section (PZW), for a total amount of about 24 Kg of limestone. ! e samples have been solved with conventional Biostratigraphy formic acid technique. All the samples were productive, yielding more than 900 conodonts. ! e ! e conodont zonation scheme followed in this paper state of preservation is good, even if some specimens is the scheme proposed by C' $!*%* (2008), that is a are broken. Conodonts color is black (CAI = 5-5.5). rielaboration of the Late Devonian Standard Conodont ! e abundance is very variable, from a maximum of Zonation (Z*"#" & S$%!‚"  1990) and the Late 154 conodonts/kg in sample PZW 5 to a minimum Devonian-Early Carboniferous Zonation of S$%!‚"  of 0.59 conodonts/kg in sample PZW C; the average et al. (1978). abundance is 25.85 conodont/kg. Sample PZW Z, Seven Biozones has been recognized: collected few meters from the section, has a high - ! e Upper rhenana Zone (sample PZW D) has abundance of about 154 conodont/kg (Tab. I). been discriminated thanks to the joint occurrence Fourty-one taxa, between species and subspecies, of Icriodus alternatus alternatus, Palmatolepis belonging to six genera (Ancyrodella, Bispathodus, rotunda, Ancyrodella lobata, Palmatolepis jamiae and Icriodus, Palmatolepis, P seudopolygnathus, P olygnathus) Palmatolepis lyaiolensis. In fact the + r st two taxa have have been recognized (Fig. 3). their + rst occurrence, while the others became extinct

17 A. MOSSONI, C. CORRADINI, M. PONDRELLI GORTANIA. Geologia, Paleontologia, Paletnologia 34 (2012)

PIZZUL WEST SECTION Stage FAMENNIAN FRASNIAN FAMENNIAN

a ea ea ea era

pid id id id if a o o o re a a s n AL Conodont c gi

an pid pid omb omb omb biozone ar re re xp henana TOT c r c rh rh rh e m

er er er ermost er er er er er pp pp pp pp pp Low U ? U U U Low Low U Low

Sample A B C D 1 2 3 4 5 5A 6 6A 7 Z Ancyrodella curvata 1 1 Ancyrodella lobata 4 4 Ancyrodella nodosa 4 4 Bispathodus stabilis 22 22 Icriodus alternatus alternatus 1 8 9 Icriodus olivierii 6 6 Palmatolepis crepida 1 1 Palmatolepis glabra acuta 2 3 5 Palmatolepis glabra glabra 1 23 5 29 Palmatolepis glabra pectinata 4 3 7 Palmatolepis glabra prima 4 3 1 1 23 26 1 4 63 Palmatolepis gracilis gracilis 1 20 21 Palmatolepis gracilis sigmoidalis 5 5 Palmatolepis jamiae 5 5 Palmatolepis lyaiolensis 4 4 Palmatolepis m. marginifera 1 1 Palmatolepis minuta minuta 7 7 Palmatolepis minuta loba 1 1 Palmatolepis minuta wolskae 2 2 Palmatolepis p. schindewol 4 1 3 2 26 36 Palmatolepis perlobata postera 2 2 Palmatolepis regularis 1 1 Palmatolepis rhomboidea 3 3 5 11 Palmatolepis rotunda 1 1 Palmatolepis rugosa rugosa 2 2 Palmatolepis simpla 2 2 Palmatolepis stoppeli 1 1 Palmatolepis subperlobata 1 1 4 11 17 Palmatolepis tenuipunctata 3 2 5 Pseudopolygnathus controversus 5 5 Pseudopolygnathus irregularis 2 2 Pseudopolygnathus m. marburgensis 2 2 Pseudopolygnathus micropunctatus 2 2 Polygnathus glaber eoglaber 2 2 Polygnathus glaber glaber 6 1 8 1 16 Polygnathus marginvolutus 8 8 Polygnathus miri cus 1 1 Polygnathus n. nodocostatus 3 1 4 8 Polygnathus normalis 1 1 Polygnathus obliquicostatus 7 7 Polygnathus styriacus 50 50 Ramiforms 1 7 2 1 34 4 2 5 127 183 Ancyrodella sp. 5 5 Icriodus sp. 2 2 1 5 10 Palmatolepis sp. 18 1 12 1 1 1 15 5 54 Polygnathus sp. 7 1 8 Pb elements 3 3 Unidenti+ ed 21 58 4 42 23 6 4 117 275 Total 64 5 1 110 20 7 8 92 122 55 3 13 16 401 917 Weight 1,9 1,8 1,7 1,9 2,2 1,7 1,4 1,2 1,2 2,1 2,4 1,9 1,9 2,6 25,9 Abundance 33,7 2,8 0,6 57,9 9,1 4,1 5,9 74,8 100,0 26,2 1,3 6,8 8,4 154,2 485,8

Tab. I - Conodont distribution chart of the Pizzul West section. - Tabella di distribuzione dei conodonti nella sezione Pizzul Ovest.

18 GORTANIA. Geologia, Paleontologia, Paletnologia 34 (2012) FAMENNIAN CONODONTS FROM THE PIZZUL WEST SECTION CARNIC ALPS, ITALY

within this zone (Z*"#" & S$%!‚"  1990; J* & 1993 Ancyrodella curvata B $% '% & M"€# - J* & Z*"#" 1993; O%$$$%'$ & K'%'%'$ 2008). Z*"#" , p. 96, pl. 2, + gs 4-5. - ! e Upper crepida Zone has been discriminated in 1998 Ancyrodella curvata B $% '% & M"€# - S$#- samples PZW 1-3, thanks of the + r st occurrence of the #"$$$  P" *, p. 204, pl. 2.2.1, + g. 1. marker Palmatolepis glabra prima, and of Palmatolepis glabra lepta and Palmatolepis minuta subgracilis. ! e Remarks : Ancyrodella curvata is characterized absence of taxa having a younger + rst occurrence by a strongly pronounced latero-posterior lobe which suggests that also sample PZW B may belong to this can bear a secondary carina and a secondary keel. ! e biozone. anterior lobes are well developed. ! e carina goes all - ! e Uppermost crepida Zone is recognized in along the element and is bounded by two row of nodes, sample PZW 4 by the joint occurrence of the marker one on each side of the carina. ! e platform is covered Palmatolepis glabra pectinata and of Palmatolepis by nodes. It is distinguished from Ancyrodella lobata by tenuipunctata, that has its last appearance datum a secondary carina on the latero-posterior lobe and a within this zone (J* & Z*"#" 1993). Also, Icriodus secondary keel. olivierii enters here at the base of its known range Range: From within the Lower hassi Zone to the (C' $!*%* 2008). end of the linguiformis Zone (J* & Z*"#" 1993). - ! e Lower rhomboidea Zone is discriminated in S tudied material: 1 specimen from sample samples PZW 5-5A by the presence of the marker PZW D. Palmatolepis rhomboidea, and of Palmatolepis minuta wolskae and Palmatolepis subperlobata that have their Ancyrodella lobata B $% '% & M"€#, 1934 last occurrence within this Zone (J* & Z*"#" 1993). (Fig. 5.2) - ! e Upper rhomboidea Zone (Samples PZW 6, 6A) is here recognized by the extinction of Palmatolepis 1934 Ancyrodella lobata n. sp. - B $% '% & M"€#, p. minuta wolskae. 239-240, pl. 19, + g. 14, pl. 21, + gs 22-23. - ! e Lower marginifera Zone (Sample PZW 7) is 1971 Ancyrodella lobata B $% '% & M"€# - S&%#- recognized by the + r st occurrence of the marker &" €*, p. 13, pl. 3, + gs 1-4 only. Palmatolepis marginifera marginifera. ! e presence 1985 Ancyrodella lobata B $% '% & M"€# - K#$- of Palmatolepis stoppeli, which became extinct within " & L$%", p. 923-924, pl. 14, + g. 12, 13, 16, this Zone (J* & Z*"#" 1993) con+ rms the attribution 17. of PZW 7. 1989 Ancyrodella lobata B $% '% & M"€# - J*, pl. 3, - ! e Lower expansa Zone (Sample Z) is discriminated + g . 3. by the joint occurrence of Palmatolepis rugosa 1993 Ancyrodella curvata (B $% '% & Mehl) - J* & rugosa, Polygnathus nodocostatus nodocostatus and Z*"#" , p. 96, pl. 2, + gs 6-10. Polygnathus styriacus. ! e + r st one make its + rst occurrence in this Zone, while the other two became Remarks : Ancyrodella lobata is characterized by a extinct (J* & Z*"#" 1993). platform covered with nodes, with two raws of big nodes on each side. ! e platform is bilaterally asymmetrical with a well-developed lobe-like protrusion on the Sistematic Palaeontology outer side. ! e lobe is covered with random nodes and underneath is evident a secondary keel. It is Synonymy lists are limited to main captions and to distinguished by Ancyrodella curvata by a lobe-like reports from the Carnic Alps. ! e whole conodont protrusion, rather than a distinct latero-posterior lobe collection is housed in the Museum of Palaeontology on the outer platform, and by the lack of the secondary “Domenico Lovisato” of Cagliari University (MDLCA); carina on this lobe-like protrusion. catalog numbers of + gured specimens (Figs 5-6) can be Range: Nearly from the start of the Lower hassi Zone obtained from the + gure captions. into the Upper rhenana Zone (J* & Z*"#" 1993). S tudied material: 4 specimens from sample Genus Ancyrodella U# *€ & B$ #" , 1926 PZW D.

Ancyrodella curvata B $% '% & M"€#, 1934 Ancyrodella nodosa U# *€ & B$ #" , 1926 (Fig. 5.3) (Fig. 5.1)

1934 Ancyrodella curvata n. sp. - B $% '% & M"€#, 1926 Ancyrodella nodosa n. sp. - U# *€ & B$ #" , p. 241, pl. 19, + gs 6, 11. p. 48, pl. 1, + gs 1-13. 1966 Ancyrodella curvata B $% '% & M"€# - 1958 Ancyrodella nodosa U# *€ & B$ #" - Z*"#" , G#"%* $" & K#$" , p. 798, pl. 86, + gs 13-15. p. 44, pl. 11, + g. 1.

19 A. MOSSONI, C. CORRADINI, M. PONDRELLI GORTANIA. Geologia, Paleontologia, Paletnologia 34 (2012)

2 3 4 5

1

7 8 9 6

14 10 11 13 12

17

16 18 15 500 µm 19

20 GORTANIA. Geologia, Paleontologia, Paletnologia 34 (2012) FAMENNIAN CONODONTS FROM THE PIZZUL WEST SECTION CARNIC ALPS, ITALY

1966 Ancyrodella nodosa U# *€ & B$ #" - G#"%*- 1998d Icriodus alternatus alternatus B $% '% & M"€#- $" & K#$" , p. 798-799, pl. 86, + gs 5-12. P" * & S$##"$$$, p. 204, pl. 2.2.1, + gs 4-5. 1993 Ancyrodella nodosa U# *€ & B$ #" - J* & 2003 Icriodus alternatus alternatus B $% '% & M"€#- Z*"#" , p. 96, pl. 2, + gs 11-12; text-+ g. 8, + gs C' $!*%*, p. 92, pl. 2, + gs 9-12. 8-9. 2008 Ancyrodella nodosa U# *€ & B$ #" - O%$- Remarks : Icriodus alternatus alternatus is characte- $$%'$ & K'%'%'$, p. 1080-1081, pl. 26, + gs rized by a thin and elongated platform. It has three rows 11-16. of longitudinal nodes that covered the platform. ! e central row is located anterior to those of the lateral Remarks : Ancyrodella nodosa is characterized by rows. ! e central raw has a cusp at the posterior end. an alate platform with the surface covered by random ! e basal cavity is deep and narrow in the anterior nodes. ! e carina extends beyond the platform in the half of the platform, wider in the posterior third. It is posterior part of the element, due to the constriction distinguished from Icriodus alternatus helmsi, which of the posterior margins of the platform. It is has the posterior cusp aligned with one of the lateral distinguished from Ancyrodella lobata by the absence rows. of the lobe-like protrusion on the outer platform. Range: Upper rhenana Zone to Uppermost crepida Range: Nearly from the start of the Upper hassi Zone (S€#€" 1999). Zone to the end of the linguiformis Zone (J* & Z*"#" , S tudied material: 9 specimens from samples 1993). PZW A, PZW D. S tudied material: 4 specimens from sample PZW D. Icriodus olivierii C' $!*%*, 1998 (Fig. 5.10) Genus Icriodus B $% '% & M"€#, 1938 1970 Icriodus symmetricus B $% '% & M"€# - Icriodus alternatus B $% '% & M"€#, 1934 O#**" *, pl. 14, + g. 9. (Fig. 5.5) 1998 Icriodus olivierii - C' $!*%*, pl. 1.4.1, + g. 8. 2003 Icriodus olivierii C' $!*%* - C' $!*%*, p. 1934 Icriodus alternatus alternatus n. sp. - B $% '% 92-93, pl. 2, + gs 14-21 (cum syn.). & M"€#, p. 225-226, pl. 13, + gs 4-6. 1984 Icriodus alternatus alternatus B $% '% & M"€#- Remarks : ! i s species is characterized by a plat- S$%!‚"  & D "" "%, pl. 2, + gs 5, 11. form thin and elongated, with the longitudinal axis 1993 Icriodus alternatus alternatus B $% '% & M"€# straight or slightly curved. ! ere are three rows of - J* & Z*"#" , p. 55, pl. 5, + gs 5-8; text-+ g. 6, longitudinal nodes. ! e nodes of the middle row + g 2. laterally compressed and longitudinally elongated, in

Fig. 5 - Upper views of P1 elements from PZW section. 1. Ancyrodella nodosa U# *€  B$ #" , 1926 (Sample PZW D); 2. Ancyrodella lobata B $% '%  M"€#, 1934 (Sample PZW D); 3. Ancyrodella curvata B $% '% & M"€#, 1934 (Sample PZW D); 4. Palmatolepis simpla Z*"#" & S$%!‚" , 1990 (Sample PZW D); 5. Icriodus alternatus alternatus B $% '% & M"€#, 1934 (Sample PZW D); 6. Palmatolepis lyaiolensis K€ % $€"$ & K%&*%, 1996 (Sample PZW D); 7. Palmatolepis rotunda Z*"#" & S$%!‚" , 1990 (Sample PZW D); 8. Palmatolepis jamiae Z*"#" & S$%!‚" , 1990 (Sample PZW D); 9. Palmatolepis rhomboidea S$%%"$%%, 1955 (Sample PZW 5); 10. Icriodus olivierii C' $!*%*, 1998 (Sample PZW 4). 11. Palmatolepis minuta loba H"# , 1963 (Sample PZW 2); 12. Palmatolepis regularis C''" , 1931 (Sample PZW 2); 13. Palmatolepis tenuipunctata S$%%"$%%, 1955 (Sample PZW 1); 14. Palmatolepis quadrantinodosalobata S$%%"$%%, 1955 (PZW 4); 15. Palmatolepis glabra glabra U# *€ & B$ #" , 1926 (Sample PZW 5); 16. Palmatolepis glabra prima Z*"#" & H%!!#", 1969 (Sample PZW 5); 17. Palmatolepis subperlobata B $% '%  M"€#, 1934 (Sample PZW 4); 18. Palmatolepis crepida S$%%"$%%, 1955 (Sample PZW 2); 19. Palmatolepis glabra pectinata Z*"#" , 1962 (Sample PZW 4). - Veduta superiore di elementi P1 dalla sezione PZW. 1. Ancyrodella nodosa U€‚ƒ & B€€€, 1926 (Campione PZW D); 2. Ancyrodella lobata B€€€ & Mƒ, 1934 (Campione PZW D); 3. Ancyrodella curvata B€€€ & Mƒ, 1934 (Campione PZW D); 4. Palmatolepis simpla Z€ & S€€, 1990 (Campione PZW D); 5. Icriodus alternatus alternatus B€€€ & Mƒ, 1934 (Campione PZW D); 6. Palmatolepis lyaiolensis Kƒ€€€‚ƒ & K€, 1996 (Campione PZW D); 7. Palmatolepis rotunda Z€ & S€€, 1990 (Campione PZW D); 8. Palmatolepis jamiae Z€ & S€€, 1990 (Campione PZW D); 9. Palmatolepis rhomboidea S€, 1955 (Campione PZW 5); 10. Icriodus olivierii C€€€, 1998 (Campione PZW 4). 11. Palmatolepis minuta loba H€€, 1963 (Campione PZW 2); 12. Palmatolepis regularis C€€€, 1931 (Campione PZW 2); 13. Palmatolepis tenuipunctata S€, 1955 (Campione PZW 1); 14. Palmatolepis quadrantinodosalobata S€, 1955 (PZW 4); 15. Palmatolepis glabra glabra U€‚ƒ & B€€€, 1926 (Campione PZW 5); 16. Palmatolepis glabra prima Z€ & H, 1969 (Campione PZW 5); 17. Palmatolepis subperlobata B€€€ & Mƒ, 1934 (Campione PZW 4); 18. Palmatolepis crepida S€, 1955 (Campione PZW 2); 19. Palmatolepis glabra pectinata Z€, 1962 (Campione PZW 4).

21 A. MOSSONI, C. CORRADINI, M. PONDRELLI GORTANIA. Geologia, Paleontologia, Paletnologia 34 (2012) some cases are almost joined together. ! e basal cavity S tudied material: 5 specimens from samples is deep and narrow in the anterior half of the platform, PZW 4 and PZW 5. wider in the posterior third. Range: From the Upper rhenana Zone to the Palmatolepis glabra glabra U# *€ & B$ #" , 1926 Uppermost crepida Zone (C' $!*%* 2003). (Fig. 5.15) S tudied material: 6 specimens from sample PZW 4. 1926 Palmatolepis glabra n. sp. - U# *€ & B$ #" , p. 51, pl. 9, + g. 20. Genus Palmatolepis U# *€ & B$ #" , 1926 1993 Palmatolepis glabra glabra U# *€ & B$ #" - J* & Z*"#" , p. 60-61, pl. 17, + gs 13-15; text-+ g. Palmatolepis crepida S$%%"$%%, 1955 17, + g. 4. (Fig. 5.18) 2003 Palmatolepis glabra glabra U# *€ & B$ #" - C' $!*%*, p. 79, pl. 4, + gs 1-2. 1955 Palmatolepis crepida n. sp. - S$%%"$%%, p. 134, pl. 6, + g. 21. Remarks : Palmatolepis glabra glabra is characte- 1962 Palmatolepis crepida crepida S$%%"$%% - rized by a narrow and elongated platform. ! e inner Z*"#" , p. 55, pl. 6, + gs 13-19 (no + g. 12). anterior margin joins the blade at a right angle. ! e 1993 Palmatolepis crepida S$%%"$%% - J* & Z*"#" , carina is slightly sigmoidal. It is distinguished from p. 59, pl. 22, + gs 1-7; text-+ g. 13, + g. 4. Palmatolepis glabra prima because of the angle of the insertion of the inner anterior margin into the blade, Remarks : Palmatolepis crepida is characterized and by Palmatolepis glabra distorta by the lack of a by having a drop-shape platform, with a shagreen pronounced parapet. surface. ! e inner anterior margin is convex, the Range: From the Lower rhomboidea Zone to the outer margin is almost straight. ! e carina is strongly Lower marginifera Zone (J* & Z*"#" 2003). curved, the central node is situated in the second half S tudied material: 23 specimens from samples of the element, and the posterior carina is weakly PZW 5 and PZW 5A. pronounced. It is distinguished by Palmatolepis tenuipunctata by the lack of the outer lobe. Palmatolepis glabra pectinata Z*"#" , 1962 Range: From the Lower crepida Zone to the Lower (Fig. 5.19) rhomboidea Zone (J* & Z*"#" 1993). S tudied material: 1 specimen from sample 1962 Palmatolepis glabra pectinata n. sub. sp. - PZW 2. Z*"#" , p. 398-399, pl. 2, + gs 3-5. 1966 Palmatolepis glabra pectinata Z*"#" - G#"%*- Palmatolepis glabra acuta H"# , 1963 $" & K#$" , p. 814, pl. 89, + gs. 1-3, 5, 9, 10; pl. 10, + gs 4-5; pl. 91, + gs 1, 3, 5. 1963 Palmatolepis (Panderolepis) serrata acuta n. sp. 1993 Palmatolepis glabra pectinata Z*"#" - Ji & - H"# , p. 468, pl. 3, + g. 1-4, 6. Z*"#" , p. 61, pl. 16, + gs 5-10, pl. 17, + gs 1-12; 1971 Palmatolepis glabra acuta H"# - S&%#&" €*, text-+ g 17, + gs 7-8. p. 33, pl. 14, + gs 6, 7. 1998b Palmatolepis glabra pectinata Z*"#" - P" * 1990 Palmatolepis glabra acuta H"# - P" * & & S$##"$$$, p. 156, pl. 1.3.1, + gs 1-2. S$##"$$$, p. 60, pl. 1, + gs 4a-b. 1993 Palmatolepis glabra acuta H"# - J* & Z*"#" ; Remarks : ! i s subpecies is distinguished from Pa. pl. 16, + gs 11, text-+ g. 17, + g. 5. glabra prima and Palmatolepis glabra glabra having a long parapet that lies close and parallel to the carina, Remarks : This subspecies of Palmatolepis glabra and from Palmatolepis glabra distorta which is more is characterized by a parapet in the inner anterior sigmoidal and by the lack of the bulge in the posterior margin of the platform, that ends with a thorn-like part of the outer platform. According to J* & Z*"#" projection in the anterior ends. The carina is slightly (1993) there are two morphotypes of this specie that sigmoidal. It is distinguished by Palmatolepis glabra di< ers from the shape of the parapet. glabra by the presence of the thorn-like projection Range: From the Uppermost crepida Zone to the on the inner anterior margin, and by Palmatolepis Upper marginifera Zone (J* & Z*"#" 1993). glabra distorta by the lack of the pronounced S tudied material: 7 specimens from samples parapet. PZW A, PZW 4. Range: From the upper part of the Lower rhomboidea Zone to the base of the Upper marginifera Zone (J* & Palmatolepis glabra prima Z*"#" & H%!!#", 1969 Z*"#" 1993). (Fig. 5.16)

22 GORTANIA. Geologia, Paleontologia, Paletnologia 34 (2012) FAMENNIAN CONODONTS FROM THE PIZZUL WEST SECTION CARNIC ALPS, ITALY

1969 Palmatolepis glabra prima - Z*"#" & H%!!#", Range: From the Upper rhomboidea Zone to the p. 379 (cum syn). Upper praesulcata Zone (J* & Z*"#" 1993). 1970 Palmatolepis glabra prima Z*"#" & H%!!#" S tudied material: 20 specimens from sample - O#**" *, p.100, pl.17, + gs 1-4. PZW Z. 1977 Palmatolepis glabra prima Z*"#" & H%!!#" - Z*"#" in Z*"#" (ed.), p. 309, pl. Palmatolepis- Palmatolepis gracilis sigmoidalis Z*"#" , 1962 7, + g. 4-7 (cum syn.). (Fig. 5.11) 1990 Palmatolepis glabra prima Z*"#" & H%!!#" - P" * & S$##"$$$, p. 61, pl. 2, + gs.1a-b. 1962 Palmatolepis de ectens sigmoidalis n. subsp. - 1993 Palmatolepis glabra pectinata Z*"#" - J* & Z*"#" , p. 56 pl. 3, + gs 24-28. Z*"#" , p. 61, pl. 16, + gs 12-17, pl. 17, text-+ g 1969 Palmatolepis gracilis sigmoidalis Z*"#" - 17, + gs 2, 9, 17. P# " , p. 399, pl. 6, + g. 22. 1998c Palmatolepis glabra prima Z*"#" & H%!!#" 1979 Palmatolepis gracilis sigmoidalis Z*"#" - - P" * & S$##"$$$, p. 156, pl. 1.3.1, + gs. 3-4, S$%!‚"  & Z*"#" ; p. 178, pl. 1, + gs 3-5. 5. 1991 Palmatolepis gracilis sigmoidalis Z*"#" - P" * 2003 Palmatolepis glabra prima Z*"#" & H%!!#" & S$##"$$$, p. 64, pl. 4, + g. 6. - C' $!*%*, p. 79, pl. 4, + gs 3-6. 1993 Palmatolepis gracilis sigmoidalis Z*"#" r - J* & Z*"#" , pl. 5, + gs 1-3; text-+ g. 14, + g. 6. Remarks : Palmatolepis glabra prima is distingui- shed by the other subspecies of Palmatolepis glabra by Remarks : ! i s subspecies di< e rs from the the rounded, bulge-like parapet on the anterior inner other subspecies of Palmatolepis gracilis having a platform, and by Palmatolepis tenuipunctata by the characteristic twisted platform and by the o< set of the lack of the outer lobe. anterior carina. Range: From the Upper crepida Zone to the Upper Range: From within the Upper trachytera Zone to marginifera Zone (J* & Z*"#" 1993). the Upper praesulcata Zone (J* & Z*"#" 1993). S tudied material: 66 specimens from samples S tudied material: 5 specimens from sample PZW B, PZW A, PZW 1, PZW 3, PZW 4, PZW 5, PZW PZW Z. 6, PZW 7. Palmatolepis jamiae Z*"#" & S$%!‚" , 1990 Palmatolepis gracilis gracilis B $% '% & M"€#, 1934 (Fig. 5.8) (Fig. 6.15-16) 1990 Palmatolepis jamiae - Z*"#" & S$%!‚" , p. 1934 Palmatolepis gracilis - B $% '% & M"€#, p. 238, 50-51, pl. 6, + gs 1-3, 9, 10 (only) pl. 18, + g. 8. 1993 Palmatolepis jamiae Z*"#" & S$%!‚"  - J* & 1969 Palmatolepis gracilis gracilis B $% '% & M"€#- Z*"#" , pl. 27, + gs 1-3. P# " , p. 399, pl. 6, + g. 21. 2008 Palmatolepis jamiae Z*"#" & S$%!‚"  - 1977 Palmatolepis gracilis gracilis B $% '% & M"€# - O%$$$%'$ & K'%'%'$, pl. 10, + gs 16-18; Z*"#" in Z*"#" (ed.), p. 315, pl. Palmatolepis- pl. 11, + gs. 1-4, 5?, 6, 7?, 8, 9; pl. 14, + g. 10. 7, + gs 8-10 (cum syn.). 1990 Palmatolepis gracilis gracilis B $% '% & M"€#- Remarks : Palmatolepis jamiae is characterized by P" * & S$##"$$$, p. 61, pl. 2, + g. 2. a shagreen platform that tapers in the anterior part 1991 Palmatolepis gracilis gracilis B $% '% & M"€#- where the two margins join the blade more or less in P" * & S$##"$$$, p. 62, pl. 4, + gs 3-4. the same position. A rounded well pronounced lobe is 1993 Palmatolepis gracilis gracilis B $% '% & M"€#- present just anteriorly of the central node. ! e inner J* & Z*"#" , p. 63, pl. 6, + gs 4-7; text-+ g. 14, posterior margin of the platform is concave. ! e carina + g . 2. is slightly sigmoidal, and well developed posterior of 1998c Palmatolepis gracilis gracilis B $% '% & M"€#- the central node where is composed of three or four C' $!*%*, pl. 1.4.2, + gs 16-17. nodes. ! is species di< er from Palmatolepis foliacea by having a well-developed lobe with two sinuses. Remarks : Palmatolepis gracilis gracilis is characte- Range: From the start of the jamiae Zone to the rized by a small, narrow platform with a raised margin Upper rhenana Zone (Z*"#" & S$%!‚"  1990). rim. ! e element is curved, in some species almost at S tudied material: 5 specimens from sample a right angle. ! e keel underneath the central node is PZW D. twisted around the small basal cavity. It is distinguished from Palmatolepis minuta minuta by the characteristic Palmatolepis lyaiolensis K€ % $€"$ & K%&*%, 1996 twisted keel. (Fig. 5.6)

23 A. MOSSONI, C. CORRADINI, M. PONDRELLI GORTANIA. Geologia, Paleontologia, Paletnologia 34 (2012)

1996 Palmatolepis lyaiolensis - K€ % $€"$ & K%&- Palmatolepis minuta B $% '%  M"€#, 1934 *%, p. 93, pl. 11, + gs 1-2. (Fig. 6.17) 2008 Palmatolepis lyaiolensis K€ % $€"$ & K%&*% - O%$$$%'$ & K'%'%'$, pl. 13, + gs 4-11. 1934 Palmatolepis minuta minuta - B $% '%  M"€#, p. 236, pl. 18, + gs 1, 6-7. Remarks : ! i s species is characterized by having 1962 Palmatolepis minuta minuta B $% '%  M"€#- a broad shagreen platform. ! e inner platform is Z*"#" ; pl. 3, + gs 1-10, text + g. 5 b-n. rounded, while the outer platform is sub-triangular, 1990 Palmatolepis minuta minuta B $% '%  M"€# because of the presence of a poorly di< e rentiated - P" * & S$##"$$$, p. 62, pl. 3, + gs 1, 7. lobe. ! e carina is slightly sigmoidal, and posterior 1993 Palmatolepis minuta minuta B $% '%  M"€#- J* of the central node is composed by one or two nodes. & Z*"#" , pl. 7, + gs 1-19; pl. 9, + gs 8-18; text- Palmatolepis lyaiolensis di< e rs from Palmatolepis hassi + g . 13, + gs 9, 15, 16. in the poorly developed lobe, lacking well pronounced 1998b Palmatolepis minuta minuta B $% '%  M"€#- sinuses. P" * & S$##"$$$, p. 156, pl. 1.3.1, + g. 8. ! i s species is here reported for the + rst time in 1998 Palmatolepis minuta minuta B $% '%  M"€# Europe. - C' $!*%*, pl. 1.4.1, + g . 14. Range: From within the Lower rhenana Zone to the Upper rhenana Zone (O%$$$%'$ & K'%'%'$ Remarks : Palmatolepis minuta minuta is characte- 2008). rized by a lanceolate smooth platform. In some elements Studied material : 4 specimens from sample a weak outer lobe is present. ! e carina is straight or PZW D. weakly curved. ! i s species di< e r from Palmatolepis rhomboidea by the lacking of the bulge in the anterior Palmatolepis marginifera marginifera H"# , 1959 part of the inner platform. (Fig. 6.18) Range: From the Upper triangularis Zone to the Upper trachytera Zone (J* & Z*"#" , 1993). 1959 Palmatolepis quarantinodosa marginifera Z*"#" S tudied material: 7 specimens from sample (sic.) - H"# , p. 649, pl. 5, + gs 22-23. PZW 5. 1973 Palmatolepis marginifera marginifera H"# - S$%!‚"  & Z*"#" ; p. 104, pl. 3, + gs 13-14. Palmatolepis minuta loba H"# , 1963 1977 Palmatolepis marginifera marginifera H"# - (Fig. 5.11) Z*"#" in Z*"#" (ed.), p. 328, pl. Palmatolepis- 7, + g. 17-18; pl. Palmatolepis-8, + gs 1-2 (cum 1963 Palmatolepis (De ectolepis) minuta loba - H"# , syn.). p. 470, pl. 2, + gs 13-14; pl. 3 + g. 12; text + g. 2, + g. 1990 Palmatolepis marginifera marginifera H"# 39. - P" * & S$##"$$$, p. 61, pl. 2, + gs 3-4. 1970 Palmatolepis minuta loba H"# - O#**" *, p. 1993 Palmatolepis marginifera marginifera H"# - J* 107, pl. 20, + g. 5. & Z*"#" , p. 64, pl. 13, + gs 7-10; pl. 14, + gs 1-6; 1993 Palmatolepis minuta loba H"# - J* & Z*"#" , text-+ g. 17, + g. 14. p. 64, pl. 10, + gs 1-16, text-+ g. 13, + g. 11-12. 1998b Palmatolepis marginifera marginifera H"# 2003 Palmatolepis minuta loba H"# - C' $!*%*, - P" * & S$##"$$$, p. 156, pl. 1.3.1, + g. 6. p. 80, pl. 6, + g. 9. 1998 Palmatolepis marginifera marginifera H"# - C' $!*%*, pl. 1.4.1, + g. 14. Remarks : ! i s species is characterized by a 2013 Palmatolepis marginifera marginifera H"# lanceolate platform with a pronounced lobe in the outer - M' '%* et al., + g. 3.2. side. ! e anterior carina is straight and the posterior carina is not well developed. It is distinguished from Remarks : Palmatolepis marginifera marginifera Palmatolepis minuta minuta by the presence of the is characterized by a rounded platform with a well- lobe. developed parapet parallel to the carina. ! e parapet Range: From the Lower crepida Zone to the Lower starts in the inner anterior platform and it extend rhomboidea Zone (J* & Z*"#" 1993). posterior the central node. It is distinguished from S tudied material: 1 specimen from sample Palmatolepis glabra distorta by the length of the parapet PZW 2. and by the shape of the platform. Range: From the base of the Lower marginifera Palmatolepis minuta subgracilis B* €', 1956 Zone into the velifer Zone (J* & Z*"#" 1993). S tudied material: 1 specimen from sample 1956 Palmatolepis minuta subgracilis - B* €', p. 130, PZW 7. pl. 9, + gs 9-12; pl. 10, + g. 13.

24 GORTANIA. Geologia, Paleontologia, Paletnologia 34 (2012) FAMENNIAN CONODONTS FROM THE PIZZUL WEST SECTION CARNIC ALPS, ITALY

1977 Palmatolepis minuta subgracilis B* €' - S tudied material: 2 specimens from sample Z*"#" in Z*"#" (ed.), p. 343, pl. Palmatolepis- PZW Z. 9, + g. 6 (cum syn). 2003 Palmatolepis minuta subgracilis B* €' - C' - Palmatolepis perlobata schindewol M##" , 1956 $!*%*; p. 80, pl. 6, + g. 21. (Fig. 6.20)

Remarks : Palmatolepis minuta subgracilis is 1956 Palmatolepis perlobata schindewol - M##" , p. characterized by a small and slender platform, 27, pl. 8, + gs 22-31; pl. 9, + g. 33. with a small lobe. It is distinguished by the other 1969 Palmatolepis perlobata schindewol M##" - subspecies of Palmatolepis minuta by the very small P# " , p. 399, pl. 5, + gs 1-2, 9. platform. 1970 Palmatolepis perlobata schindewol M##" - Range: From the Upper crepida Zone to the Lower Olivieri, p. 109, pl. 20, + gs 11-14. rhomboidea Zone (Z*"#" 1977). 1977 Palmatolepis perlobata schindewol M##" - S tudied material: 1 specimens from sample Z*"#" in Z*"#" (ed.), p. 361, pl. Palmatolepis- PZW 2. 11, + g. 1-7 (cum syn.). 1990 Palmatolepis perlobata schindewol M##" - Palmatolepis minuta wolskae S&%#&" €*, 1971 P" * & S$##"$$$, + g. 63, pl. 3, + gs 4-5, 8. 1991 Palmatolepis perlobata schindewol M##" - 1971 Palmatolepis minuta wolskae - S&%#&" €*, p. P" * & S$##"$$$, + g. 66, pl. 4, + g. 7. 36, pl. 15, + gs 2, 12-14. 1993 Palmatolepis perlobata schindewol M##" - J* 1993 Palmatolepis minuta wolskae S&%#&" €* - J* & & Z*"#" ; p. 67, pl. 18, + gs 9-15; text-+ g. 15, Z*"#" ; p. 85 pl. 11, + gs 1-11, text-+ g. 13, + gs + g . 3. 7-8. 2003 Palmatolepis perlobata schindewol M##" - 2003 Palmatolepis minuta wolskae S&%#&" €* - C' $!*%*, pl. 7, + gs 1-5. C' $!*%*; p. 80, pl. 6, + gs 7-8. Remarks : Palmatolepis perlobata schindewol is Remarks : ! is species is characterized by a small characterized by an arc-shaped and elongated plat- ovoidal or subtriangular platform, with a small lobe in form. ! ere is a small lobe on the outer platform, and the outer platform. ! e posterior carina is absent. It generally the posterior end is pointed downward. ! e is distinguished from Palmatolepis minuta loba by the surface is smooth or weakly ornamented. ! is species absence of the posterior carina. is distinguished from Palmatolepis perlobata perlobata Range: From the Middle crepida Zone to the Lower by its slender shape, the small lobe and the weak or- rhomboidea Zone (C' $!*%* 2003). namentation. S tudied material: 2 specimens from sample Range: From the Upper crepida Zone to the Upper PZW 5A. expansa Zone (J* & Z*"#" 1993). S tudied material: 24 specimens from samples Palmatolepis perlobata postera Z*"#" , 1960 PZW A, PZW B, PZW 5 and PZW6A. (Fig. 6.2) Palmatolepis quadrantinodosalobata 1960 Palmatolepis perlobata postera n. subsp. - S$%%"$%%, 1955 Z*"#" , p. 27, pl. 8, + gs 22-31; pl. 9, + g. 33. (Fig. 5.14) 1979 Palmatolepis perlobata postera Z*"#" - S$%!‚"  & Z*"#" , p. 180, pl. 2, + gs 1-4. 1955 Palmatolepis quadrantinodosalobata - S$%%"$%%, 1993 Palmatolepis perlobata postera Z*"#" - J* & p. 328, pl. 24, + g. 6. Z*"#" ; text-+ g 15, + g. 10. 1969 Palmatolepis quadrantinodosalobata S$%%"$%% - P# " , p. 399, pl. 6, + gs 13-14. Remarks : Palmatolepis perlobata postera is 1970 Palmatolepis quadrantinodosalobata S$%%"$%% characterized by a broad curved platform covered by - O#**" *, p. 112, pl. 18, + g. 9-11. small nodes. It is distinguished by the other subspecies 1973 Palmatolepis quadrantinodosalobata S$%%"$%% - of Palmatolepis perlobata by the weak lobe on the Z*"#" in Z*"#" (ed.), p. 295, pl. Palmatolepis- outer part of the platform. ! is species was choosen as 4, + gs 6-8 (cum syn.). marker of the postera Zone by Z*"#" & S$%!‚"  1993 Palmatolepis quadrantinodosalobata S$%%"$%% (1984), however in North Gondwana is a very rare - J* & Z*"#" , p. 69, pl. 23, + g. 5-7; text-+ g. 12, taxon (C' $!*%*, 2008). + g . 3, 7-8. Range: From the Lower postera Zone to the Upper 2003 Palmatolepis quadrantinodosalobata S$%%"$%% expansa Zone (J* & Z*"#" 1993). - C' $!*%*, pl. 5, + gs 7-9.

25 A. MOSSONI, C. CORRADINI, M. PONDRELLI GORTANIA. Geologia, Paleontologia, Paletnologia 34 (2012)

1 2 3 4 5

7 6 8 9 10

13 14 15 11 12

16

17 20 18 19 500 µm

26 GORTANIA. Geologia, Paleontologia, Paletnologia 34 (2012) FAMENNIAN CONODONTS FROM THE PIZZUL WEST SECTION CARNIC ALPS, ITALY

Remarks : Palmatolepis quadrantinodosalobata is has a narrow and elongated platform, while morphotype distinguished by a well-developed rounded lobe in the 2 has a broader platform. ! e specimen from Pizzul outer part of the platform and the inner anterior part West section belongs to morphotype 2. However, it covered with aligned or randomly disposed nodes (J* should be pointed out that, since the range of the two & Z*"#" 1993). A few specimens may have a few morphotypes is the same and coincides with the range small nodes on the outer anterior part of the platform. of the species, their utility is questionable. ! i s species Palmatolepis quadrantinodosalobata is distinguished is distinguished by Palmatolepis subperlobata by the from Palmatolepis subperlobata by the presence of the lack of the outer lobe. ornamentation, and by Palmatolepis sandbergi that has Range: From the Upper triangularis Zone to the the whole inner platform covered by nodes Lower rhomboidea Zone (J* & Z*"#" 1993). Range: From the base of the Lower crepida zone into S tudied material: 1 specimen from sample the Lower rhomboidea Zone (J* & Z*"#" 1993). PZW 2. S tudied material: 10 specimens from samples PZW 1, PZW 2, PZW 4. Palmatolepis rhomboidea S$%%"$%%, 1955 (Fig. 5.9) Palmatolepis regularis C''" , 1931 (Fig. 5.12) 1955a Palmatolepis rhomboidea - S$%%"$%%, p. 329, pl. 24, + g. 14. 1931 Palmatolepis regularis n. sp. - C''" , p. 242, pl. 1970 Palmatolepis rhomboidea S$%%"$%% - O#**"- 28, + g. 36. *, p. 114, pl. 16, + gs. 11-14. 1962 Palmatolepis cf. regularis C''" - Z*"#" , p. 1985 Palmatolepis rhomboidea S$%%"$%% - Z*"#" 75-76, pl. 6, + gs 20-24. in Z*"#" (ed.), p. 299, pl. Palmatolepis-1, + g. 1969 Palmatolepis cf. regularis C''" - P# " , p. 6-7 (cum syn.). 399, pl. 5, + gs 3-4. 2003 Palmatolepis rhomboidea S$%%"$%% - C' $- 1993 Palmatolepis cf. regularis C''" - J* & Z*"#" , !*%*, p. 83, pl. 3, + gs 19-21. pl. 21, + gs 6-10; text-+ g. 16, + gs 7, 9. Remarks : Palmatolepis rhomboidea is characterized Remarks : ! is species is characterized by a sha- by a small rhomboidal platform with an evident green, strongly sigmoidal platform lacking the outer bulge in the outer anterior part. It is distinguished by lobe. J* & Z*"#" (1993) proposed two morphotypes Palmatolepis minuta minuta by the bulge in the inner that di< er from the width of the platform. Morphotype 1 anterior platform and the shorter free blade.

Fig. 6 - Upper views of P1 elements from PZW section, unless di< erently stated. 1. Palmatolepis rugosa rugosa B $% '% & M"€#, 1934 (Sample PZW Z); 2. Palmatolepis perlobata postera Z*"#" , 1960 (Sample PZW Z); 3. Polygnathus cf. nodocostatus B $% '% & M"€#, 1934 (Sample PZW Z); 4. Pseudopolygnathus irregularis T $€"#"% & H$ $"%"# , 2011 (Sample PZW Z); 5. Pseudopolygnathus micropunctatus B* €' & Z*"#" , 1956 (Sample PZW Z); 6. Pseudopolygnathus controversus S$%!‚"  & Z*"#" , 1979 (Sample PZW Z); 7. Polygnathus styriacus Z*"#" , 1957 (Sample PZW Z); 8. Bispathodus stabilis (B $% '%  M"€#, 1934) (PZW Z); 9. Polygnathus marginvolutus G"!*€, 1969 (PZW Z); 10. Pseudopolygnathus marburgensis marburgensis B* €' & Z*"#" , 1956 (Sample PZW Z); 11. Palmatolepis gracilis sigmoidalis Z*"#" , 1962 (Sample PZW Z); 12. Polygnathus obliquicostatus Z*"#" , 1962 (Sample PZW Z); 13. Polygnathus glaber glaber U# *€ & B$ #" , 1926 (Sample PZW 5); 14. Polygnathus nodocostatus nodocostatus B $% '% & M"€#, 1934 (Sample PZW 5); 15. Palmatolepis gracilis gracilis B $% '% & M"€#, 1934, lower view (Sample PZW Z); 16. Palmatolepis gracilis gracilis B $% '% & M"€#, 1934 (Sample PZW Z); 17. Palmatolepis minuta minuta B $% '% & M"€#, 1934 (Sample PZW 5A); 18. Palmatolepis marginifera marginifera H"# , 1959 (Sample PZW 7); 19. Palmatolepis stoppeli S$%!‚"  & Z*"#" , 1973 (Sample PZW 7); 20. Palmatolepis perlobata schindewol M%##" , 1956 (Sample PZW 5). - Veduta superiore di elementi P1 dalla sezione PZW, a meno di indicazioni di erenti nella didascalia. 1. Palmatolepis rugosa rugosa B€€€ & Mƒ, 1934 (Campione PZW Z); 2. Palmatolepis perlobata postera Z€, 1960 (Campione PZW Z); 3. Polygnathus cf. nodocostatus B€€€ & Mƒ, 1934 (Campione PZW Z); 4. Pseudopolygnathus irregularis T€ƒ & H€€€, 2011 (Campione PZW Z); 5. Pseudopolygnathus micropunctatus B€ƒ€ & Z€, 1956 (Campione PZW Z); 6. Pseudopolygnathus controversus S€€ & Z€, 1979 (Campione PZW Z); 7. Polygnathus styriacus Z€, 1957 (Campione PZW Z); 8. Bispathodus stabilis (B€€€ & Mƒ, 1934) (PZW Z); 9. Polygnathus marginvolutus G, 1969 (PZW Z); 10. Pseudopolygnathus marburgensis marburgensis B€ƒ€ & Z€, 1956 (Campione PZW Z); 11. Palmatolepis gracilis sigmoidalis Z€, 1962 (Campione PZW Z); 12. Polygnathus obliquicostatus Z€, 1962 (Campione PZW Z); 13. Polygnathus glaber glaber U€‚ƒ & B€€€, 1926 (Campione PZW 5); 14. Polygnathus nodo- costatus nodocostatus B€€€ & Mƒ, 1934 (Campione PZW 5); 15. Palmatolepis gracilis gracilis B€€€ & Mƒ, 1934, veduta inferiore (Campione PZW Z); 16. Palmatolepis gracilis gracilis B€€€ & Mƒ, 1934 (Campione PZW Z); 17. Palmatolepis minuta minuta B€€€ & Mƒ, 1934 (Campione PZW 5A); 18. Palmatolepis marginifera marginifera H€€, 1959 (Campione PZW 7); 19. Palmatolepis stoppeli S€€ & Z€, 1973 (Campione PZW 7); 20. Palmatolepis perlobata schindewol+ M€, 1956 (Campione PZW 5).

27 A. MOSSONI, C. CORRADINI, M. PONDRELLI GORTANIA. Geologia, Paleontologia, Paletnologia 34 (2012)

Range: From the Lower rhomboidea Zone to the Remarks : Palmatolepis simpla is characterized lower part of the Upper marginifera Zone (C' $!*%* by a broad platform, almost rounded in the posterior 2003). part, while in the anterior part became narrow. ! ere S tudied material: 11 specimens from samples is a rounded lobe in the outer part of the anterior PZW A, PZW 5, PZW 5A. platform. ! e anterior outer platform margin is concave. It is distinguished from Palmatolepis proversa Palmatolepis rotunda Z*"#" & S$%!‚" , 1990 by the weaker marginal forti+ cation and for the less (Fig. 5.7) pronounced lobe. ! i s species is here reported for the + rst time from 1990 Palmatolepis rotunda n. sp. - Z*"#" & S$%!- the Carnic Alps. ‚" , p. 62, pl. 10, + gs 1-5. Range: From the Upper hassi Zone to the Upper 1998 Palmatolepis rotunda Z*"#" & S$%!‚"  - rhenana Zone (Z*"#" & S$%!‚"  1990). S$##"$$$  P" *, p. 204, pl. 2.2.1, + g. 12. S tudied material: 2 specimens from sample PZW D. Remarks : Palmatolepis rotunda is characterized by a broad rounded inner posterior platform. ! ere is a well Palmatolepis stoppeli S$%!‚"  & Z*"#" , 1973 developed rounded lobe in the outer platform, with two (Fig. 6.19) evident sinuses. ! e anterior carina is strongly curved, while the posterior carina is not well pronounced. 1960 Palmatolepis sp. - Z*"#" pl. 7 + gs 12-13 Range: From the start of the Upper rhenana Zone to 1977 Palmatolepis stoppeli n. sp. - S$%!‚"  & the top of the linguiformis Zone (Z*"#" & S$%!‚"  Z*"#" , p. 106-107, pl. 3 + gs 1-11, pl. 5, + g. 13. 1990). 1993 Palmatolepis stoppeli S$%!‚"  & Z*"#" - J* S tudied material: 1 specimen from sample & Z*"#" , p. 71, pl. 14, + gs 7-12, text-+ g. 17, PZW D. + g . 12. 1998 Palmatolepis stoppeli S$%!‚"  & Z*"#" - Palmatolepis rugosa rugosa B $% '% & M"€#, 1934 C' $!*%*, pl. 1.4.2, + g. 19 (Fig. 6.1) Remarks : ! i s species is characterized by a broad 1934 Palmatolepis rugosa n.sp. - B $% '% & M"€#, p. sub-ovoidal platform with an evident ramp in the 236, pl. 18, + gs 15, 16, 18, 19. upper part of the inner platform. It is distinguished 1979 Palmatolepis rugosa rugosa B $% '% & M"€# from Palmatolepis quadrantinodosa in exa by the - S$%!‚"  & Z*"#" , p. 180, pl. 2, + gs 1-4. lack of ornamentation on the surface of the platform 1991 Palmatolepis rugosa rugosa B $% '% & M"€# and from Palmatolepis rhomboidea because the latter - P" * & S$##"$$$, p. 66, pl. 4, + gs 8-9. have in the inner platform a small bulge instead of an 1993 Palmatolepis rugosa rugosa B $% '% & M"€# evident ramp. - J* & Z*"#" ; text-+ g 15, + g. 12. Range: Upper rhomboidea Zone to Lower marginifera Zone (J* & Z*"#" 1993). Remarks : ! i s species is characterized by a broad S tudied material: 1 specimen from sample strongly ornamented platform, with a very pronounced PZW 7. outer lobe. ! e carina is strongly curved anterior the central node. It is distinguished from the other Palmatolepis subperlobata B $% '% & M"€#, 1934 subspecies of Palmatolepis rugosa by the pattern of (Fig. 5.17) the ornamentation, that shows a ridge-type nodes in the inner parapet and a series of coarse nodes in the 1934 Palmatolepis subperlobata n. sp. - B $% '%  anterior part of the outer platform. M"€#, p. 235, pl. 18, + gs 11, 21. Range: From the Lower expansa Zone to the Upper 1971 Palmatolepis subperlobata B $% '% & M"€# expansa Zone (J* & Z*"#" , 1993). - S&%#&" €*, p. 40-41, pl. 13, + g. 12. S tudied material: 2 specimens from sample 1993 Palmatolepis subperlobata B $% '% & M"€# PZW Z. - J* & Z*"#" , pl. 20, + gs 3-9; pl. 21, + gs 11-12; text-+ g. 16, + gs 5, 6, 8. 2003 Palmatolepis subperlobata B $% '% & M"€# Palmatolepis simpla Z*"#" & S$%!‚" , 1990 - C' $!*%*, pl. 3, + gs 1-4. (Fig. 5.4) Remarks : Palmatolepis subperlobata is characte- 1990 Palmatolepis simpla n. sp. - Z*"#" & S$%!‚" , rized by a shagreen platform with a well-developed lobe p. 47-48, pl. 4, + gs 9-12. on the outer platform. ! e carina is strongly sigmoidal.

28 GORTANIA. Geologia, Paleontologia, Paletnologia 34 (2012) FAMENNIAN CONODONTS FROM THE PIZZUL WEST SECTION CARNIC ALPS, ITALY

! i s species is distinguished from Palmatolepis 1990 Bispathodus stabilis (B $% '% & M"€#) M1 tenuipunctata which has a relatively narrow, elongated - P" * & S$##"$$$, p. 60, pl. 1, + g. 2. platform and a small outer lobe. 1998c Bispathodus stabilis (B $% '% & M"€#) M1 Range: From the base of the Lower triangularis Zone - P" * & S$##"$$$, p. 177, pl. 1.5.1, figs 5- to the Upper marginifera Zone (C' $!*%* 2003). 6. S tudied material: 15 specimens from samples 2003 Bispathodus stabilis (B $% '% & M"€#) M1 PZW 2, PZW 3, PZW 4 and PZW 5. - C' $!*%*, p. 95, pl. 1, + gs 1-2.

Palmatolepis tenuipunctata S$%%"$%%, 1955 Remarks : ! is element is characterized by a thin (Fig. 5.13) and nearly straight blade, bearing discrete denticles. Close to the posterior end, the denticles are less heigh. 1955b Palmatolepis tenuipunctata - S$%%"$%%, p. 136, ! e re are two morphotypes of Bispathodus stabilis, pl. 6, + g. 22. di< ering by the shape of the basal cavity: in M1 is 1969 Palmatolepis tenuipunctata S$%%"$%% - small and do not reaches the posterior end, while in P# " , p. 399, pl. 5, + g. 21. M2 is wide, slightly asymmetrical, and reaches the 1970 Palmatolepis tenuipunctata S$%%"$%% - O#*- posterior end of the element. In our material both the *" *, p. 117, pl. 18, + gs 1-2. morphotypes are present. 1993 Palmatolepis tenuipunctata S$%%"$%% - J* & Range: From the Upper marginifera Zone through Z*"#" , p. 72, pl. 19, + g. 1-6; text-+ g.16, + g. the Lower Carboniferous (Z*"#" , S$%!‚"  & 2. A% $*% 1974). 2003 Palmatolepis tenuipunctata S$%%"$%% - C' - S tudied material: 22 specimens from sample $!*%*, pl. 3, + gs 11-13. PZW Z.

Remarks : Palmatolepis tenuipunctata is characte- Genus Polygnathus, H*%!", 1879 rized by an elongated platform, with a small lobe in the outer part. ! e blade-carina is slightly sigmoidal. Polygnathus glaber eoglaber J* & Z*"#" , 1993 Underneath the element there is a thin keel that goes all along the platform. It is distinguished from Palmatolepis 1993 Polygnathus eoglaber - J* & Z*"#" , p. 78, pl. 36, subperlobata by the less developed lobe and from + g s 10-15; text-+ g 21, + g. 10. Palmatolepis glabra prima by the presence of the lobe. 2003 Polygnathus glaber eoglaber J* & Z*"#" - Range: From the Upper triangularis Zone to the C' $!*%*, pl. 8, + g. 2. Uppermost crepida Zone (J* & Z*"#" , 1993). S tudied material: 5 specimens from samples Remarks : ! is species is characterized by a small PZW 1 and PZW 4. smooth platform, and by the prolongation of the carina slightly a er the posterior end of the platform, almost to form a small free blade. Range: From the Upper triangularis Zone to the Genus Bispathodus M##" , 1962 Upper rhomboidea Zone (J* & Z*"#" 1993). S tudied material: 2 specimen from sample Bispathodus stabilis (B $% '% & M"€#), 1934 PZW 4. (Fig. 6.8) Polygnathus glaber glaber U# *€ & B$ #" , 1926 1934 Spathodus stabilis - B $% '% & M"€#, p. 188, (Fig. 6.13) pl. 17, + g. 20. 1962 Spathognathodus stabilis (B $% '% & M"€#) 1926 Polygnathus glaber - U# *€ & B$ #" , p. 46, - Z*"#" , p. 110, pl. 13, + gs 4-5, 9-10. pl. 7, + g. 13. 1969b Spathognathodus stabilis (B $% '% & M"€#) 1969a Polygnathus glabra glabra U# *€ & B$ #" - S€%#$%‚, p. 321, pl. 3, + gs 14-15. - S€%#$%‚, p. 295, pl. 2, + g. 14. 1969 Spathognathodus stabilis (B $% '% & M"€#) 1998c Polygnathus glaber glaber U# *€ & B$ #" - P# " , p. 399, pl. 5, + gs 15-16. - C' $!*%*, pl. 1.4.1, + g. 5. 1974 Bispathodus stabilis (B $% '% & M"€#) M1 2003 Polygnathus glaber glaber U# *€ & B$ #" - Z*"#" , S$%!‚"  & A% $*%, p. 103, pl. 3, - C' $!*%*, pl. 8, + gs 3-5. + g s 1-3 1974 Bispathodus stabilis (B $% '% & M"€#) M2 - Remarks : Polygnathus glaber glaber is characte- Z*"#" , S$%!‚"  & A% $*%, p. 103, pl. 3, + g. rized by a small, ovate shagreen platform; a few 2 specimen present slightly raised lateral margins. It is

29 A. MOSSONI, C. CORRADINI, M. PONDRELLI GORTANIA. Geologia, Paleontologia, Paletnologia 34 (2012) di< erent from Polygnathus glaber eoglaber by the lack 1969a Polygnathus nodocostata nodocostata B $% '% of the posterior free blade. & M"€# - S€%#$%‚, p. 295, pl. 2, + g. 12. Range: From the base of the Lower rhomboidea 1970 Polygnathus nodocostatus nodocostatus B $% '% Zone into the Lower trachytera Zone (C' $!*%*, & M"€# - O#**" *, p. 125, pl. 22, + gs 1-5. 2003). 1993 Polygnathus nodocostata nodocostata B $% '% & S tudied material: 17 specimens from samples M"€# - J* & Z*"#" , pl. 34, + gs 13-15; text-+ g. PZW A, PZW 4, PZW 5, PZW 7. 20, + g. 1. 2003 Polygnathus nodocostatus nodocostatus B $% '% Polygnathus marginvolutus G"!*€, 1969 & M"€# - C' $!*%*; pl. 9, + gs 1-2. (Fig. 6.9) Remarks : Polygnathus nodocostatus nodocostatus 1969 Polygnathus marginvolutus - G"!*€, pl. 237, pl. is characterized by a large platform, with a very 5, + gs 2-8. variable outline. ! e platform is totally covered by 1991 Polygnathus marginvolutus G"!*€ - P" * & rows of nodes parallel to the carina. It is distinguished S$##"$$$, p. 237, pl. 6, + gs 1-2. by Polygnathus perplexus by the lack of the collar 1998d Polygnathus marginvolutus G"!*€ - P" * & formed by two asymmetrically developed rostral S$##"$$$, p. 179, pl. 1.5.2, + g. 7. ridges, and by Polygnathus granulosus because the latter has a randomly disposition of the nodes in the Remarks : ! i s species is characterized by a upper surface. subtriangular or heart-shape platform with upturned Range: From the Lower crepida Zone to the Lower margins. ! e anterior margins are o en scalloped. expansa Zone (J* & Z*"#" 1993). ! e posterior part of the platform bear weak ridges S tudied material: 8 specimens from samples which don’t reach the carina, while the anterior part PZW A, PZW 5A and PZW Z. is smooth. Range: From within the Upper trachytera Zone to Polygnathus normalis M*##" & Y'%%%* $, 1947 the Upper expansa Zone (P" * & S$##"$$$ 1991). S tudied material: 8 specimen from sample 1947 Polygnathus normalis n. sp. - M*##" & Y'%%- PZW Z. %* $, p. 515, pl. 74, + gs 4-5. 1966 Polygnathus normalis M*##" & Y'%%%* $ Polygnathus miri cus J* & Z*"#" , 1993 - G#"%* $" & K#$" , p. 829-830, pl. 95, + gs 6, 21-22. 1993 Polygnathus miri cus n. sp. - J* & Z*"#" , pl. 1993 Polygnathus normalis M*##" & Y'%%%* $ - J* 37, + gs 16-21. & Z*"#" , pl. 39, + gs 9-15; text-+ g. 18, + g.14.

Remarks : Polygnathus miri cus is characterized by Remarks : Polygnathus normalis is characterized an asymmetrical and lanceolate platform ornamented by having an asymmetric platform with a posterior with transverse ridges. ! e anterior margin of the plat- margin incurved and a slightly constricted anterior form bears small denticles. ! e carina, generally ex- margin. ! e platform is covered by transversal tended to the posterior tip of the platform, is low and ridges. Some authors consider Polygnathus normalis composed of fused denticles. Polygnathus miri cus is as young synonym of Polygnathus webbi. However distinguished from Polygnathus alatus, Polygnathus we believe that the two forms are di< e rent species webbi and Polygnathus normalis by having an asymmet- because Polygnathus webbi has a strongly constricted rical platform with strong denticulate outer margin. anterior platform and a more expanded posterior ! i s species is here reported for the + rst time from outer platform. the Carnic Alps. Range: From within the Upper rhenana Zone to the Range: From within the Upper rhenana Zone to the Lower postera Zone (J* & Z*"#" , 1993). linguiformis Zone (J* & Z*"#" 1993). S tudied material: 1 specimen from sample S tudied material: 1 specimen from sample PZW D. PZW D. Polygnathus obliquicostatus Z*"#" , 1962 Polygnathus nodocostatus nodocostatus (Fig. 6.12) B $% '% & M"€#, 1934 (Figs 6.3, 6.14) 1962 Polygnathus obliquicostatus n. sp. - Z*"#" , p. 92, pl. 11, + gs 8-12. 1934 Polygnathus nodocostata - B $% '% & M"€#, p. 1970 Polygnathus obliquicostatus Z*"#" - O#**" *, 246, pl. 20, + gs 9-13; pl. 21, + g. 15. p. 128, pl. 23, + gs 4-5.

30 GORTANIA. Geologia, Paleontologia, Paletnologia 34 (2012) FAMENNIAN CONODONTS FROM THE PIZZUL WEST SECTION CARNIC ALPS, ITALY

1993 Polygnathus obliquicostatus Z*"#" - J*& Polygnathus granulosus by the sub-triangular platform Z*"#" , text-+ g. 19, + g. 5. and the downward de# ection of the anterior part of the 1998c Polygnathus obliquicostatus Z*"#" - P" * & platform. S$##"$$$, p. 166, pl. 1.4.2, + gs 10a-b. Range: From the base of the Lower styriacus Zone 1998f Polygnathus obliquicostatus Z*"#" – P" * & (Lower postera Zone) to the Lower expansa Zone (J* & S$##"$$$, p. 226, pl. 2.5.1, + g. 6. Z*"#" 1993) 2003 Polygnathus obliquicostatus Z*"#" - C' $!*%*, S tudied material: 50 specimens from sample p. 112, pl. 10, + gs 3-5. PZW Z.

Remarks : Polygnathus obliquicostatus is characte- Genus Pseudopolygnathus B $% '% & M"€#, 1934 rized by a thin and elongated platform, with the posterior part turned downward. ! e platform bears Pseudopolygnathus controversus oblique transverse ridges that forms an angle of S$%!‚"  & Z*"#" , 1979 about 45° with the carina, more evident posterior of (Fig. 6.6) the carina, where occupy the whole platform. It is distinguished by Polygnathus semicostatus because the 1979 Pseudopolygnathus controversus n. sp. - S$%!‚"  latter has on the inner platform ridges perpendicular & Z*"#" , p. 182, pl.3, + gs 12-17. to the carina, and a generally more developed tongue; 1998f Pseudopolygnathus controversus S$%!‚"  & di< ers from Polygnathus extralobatus in the more thin Z*"#" - P" * & S$##"$$$, p. 226, pl. 2.5.1, and symmetrical platform. + g . 10. Range: From the Lower styriacus Zone to the Lower 2011 Pseudopolygnathus controversus S$%!‚"  & praesulcata Zone (C' $!*%* et al. 2003). Z*"#" - H$ $"%"# p. 510, pl. 62, + gs 10-11. S tudied material: 7 specimens from sample PZW Z. Remarks : Pseudopolygnathus controversus is characterized by an asymmetrical lanceolate platform Polygnathus styriacus Z*"#" , 1957 with an ornamented surface. ! e lenght of the right (Fig. 6.7) side of the platform extends much farther anteriorly than the le side as a row of transverses ridges or 1957 Polygnathus styriacus - Z*"#" , p. 47, pl. 1, + gs nodes. It is distinguished from Pseudopolygnathus 12-13. brevipennatus by the asymmetrical platform. 1979 Polygnathus styriacus Z*"#" - S$%!‚"  & Range: From the Upper styriacus Zone to the Lower Z*"#" , p. 186, pl. 4, + gs 14-18. expansa Zone (S$%!‚"  & Z*"#" 1979). 1993 Polygnathus styriacus Z*"#" - J* & Z*"#" , p. S tudied material: 5 specimens form sample 84, pl. 34, + g. 6-10; text-+ g. 20, + g. 12. PZW Z. 1998d Polygnathus styriacus Z*"#" - P" * & S$##"$$$, p. 179, pl. 1.5.2, + g. 8. Pseudopolygnathus irregularis 1998f Polygnathus styriacus Z*"#" - P" * & T $€"#"% & H$ $"%"# , 2011 S$##"$$$, p. 226, pl. 2.5.1, + g. 8. (Fig. 6.4) 2003 Polygnathus styriacus Z*"#" - C' $!*%*, p. 110, pl. 9, + g. 10. 2011 Pseudopolygnathus irregularis n. sp. - T $€"#"% 2011 Polygnathus styriacus Z*"#" - T $"#"€% & & H$ $"%"# , p. 8, pl. 2, + g. 16-23. H$ $"%"# , p. 12, pl. 1, + gs 10-19. 2013 Pseudopolygnathus irregularis T $€"#"% & 2011 Polygnathus protostyriacus T $"#"€% & H$ $"%"# - M' '%* et al., p. 88, + gs 3.10. H$ $"%"# , p. 12, pl. 1, + gs 3-9. Remarks : ! i s species h a s an asymmetric p l atform, Remarks : Polygnathus styriacus is characterized which extends to the posterior tip of the element. ! e by a small sub-triangular platform covered by weak edges are slightly raised. ! e outline of the platform is nodes irregularly arranged in the posterior part; irregular. ! e surface is covered with distinctive and the anterior part of the platform is not ornamented irregular nodes. It is distinguished from the others and strongly de# e cted downward. T $"#"€% & species of Pseudopolygnathus by the irregular outline H$ $"%"# (2011) introduced a new species and two of the platform and by the ornamentation. morphotypes, previously attributed to Polygnathus Range: From the Upper styriacus Zone to the styriacus. In our opinion they represents variability Lower expansa Zone (T $€"#"% & H$ $"%"# within the population of Polygnathus styriacus. 2011). Polygnathus styriacus is di< e rent from Polygnathus S tudied material: 2 specimens form sample vogesi by the ornamentation pattern; di< e rs from PZW Z.

31 A. MOSSONI, C. CORRADINI, M. PONDRELLI GORTANIA. Geologia, Paleontologia, Paletnologia 34 (2012)

Pseudopolygnathus marburgensis marburgensis specimens show a weak lobe in the inner part of B* €' & Z*"#" , 1956 the platform, but it’s not a distinctive character. ! e (Fig. 6.10) weak ornamentation distinguish Pseudopolygnathus micropunctatus among all the other representative of 1956 Pseudopolygnathus marburgensis n. sp. - B* €' genus Pseudopolygnathus. & Z*"#" , p. 162-163, pl. 11, + gs 9, 11-13. Range: From the Upper trachytera Zone to the - 1979 Pseudopolygnathus marburgensis marburgensis Upper expansa Zone (C' $!*%* 2003). B* €' & Z*"#" - S$%!‚"  & Z*"#" , p. S tudied material: 2 specimens from sample 182, pl. 3, + gs 1-4. PZW Z. 1981 Pseudopolygnathus marburgensis marburgensis - K#$" (in Z*"#" ) 381-382, Pseudopoly- gnathus - pl 1, + gs. 1-7. Conclusions 2011 Pseudopolygnathus marburgensis marburgensis B* €' & Z*"#" - H$ $"%"# p. 512, pl. ! e main results of this study on the Clymeniae 64, + g. 5. limestones in the Pizzul West section can be summarized as follows: Remarks : ! i s subspecies of Pseudopolygnathus 1. Fourty-one conodont taxa, between species marburgensis is characterized by a trilobate platform and subspecies, belonging to six genera and a trilobate basal cavity. ! e upper surface of the (Ancyrodella, Bispathodus, Icriodus, Palmatolepis, platform is strongly ornamented, with some distinctive Pseudopolygnathus, Polygnathus) have been nodes in the anterior part of the platform. ! e outer recognized. lobe has a secondary carina that forms nearly a right 2. ! e following species have been reported for the angle with the main carina, the inner lobe bear large + r st time from the Carnic Alps: Icriodus olivierii crowded nodes or a bifurcate pattern. ! i s subspecies C' $!*%*, 2003, Palmatolepis lyaiolensis J* & of Ps. marburgensis di< ers from Pseudopolygnathus Z*"#" , 1993, Palmatolepis simpla Z*"#" & marburgensis trigonicus because the latter has a cross- S$%!‚" , 1990, Pseudopolygnathus irregularis shape basal cavity instead, while Pseudopolygnathus T $€"#"% & H$ $"%"# , 2011, Polygnathus marburgensis marburgensis has a broad basal cavity. miri cus J* & Z*"#" , 1993. Range: From the Upper styriacus Zone to the 3. Seven conodont biozones, one from the Frasnian Middle expansa Zone (Z*"#" & S$%!‚"  1984). (Upper rhenana) and six from the Famennian (Upper S tudied material: 2 specimens from sample crepida, Uppermost crepida, Lower rhomboidea, PZW Z. Upper rhomboidea, Lower marginifera, Lower expansa) have been discriminated. Pseudopolygnathus micropunctatus B* €' & Z*"#" , 1956 Manuscript received on 19.IX.2013, accepted on 11.XI.2013. (Fig. 6.5) Acknowledgements 1956 Pseudopolygnathus micropunctata - B* €' & Z*"#" , p. 163, pl. 11, + gs 7-8, 10. Angelo Mossoni gratefully acknowledges Sardinia €  1979 Pseudopolygnathus cf. micropunctatus B* ' Regional Government for the + nanancial support of his PhD & Z*"#" - S$%!‚"  & Z*"#" , p. 183, pl. 3, scholarship (P.O.R. Sardegna F.S.E. “Operational Programme + g s 5-7. of the Autonomous Region of Sardinia, European Social Fund 1998f Pseudopolygnathus micropunctatus B* €' & 2007-2013” - Axis IV Human Resources, Objective l.3, Line Z*"#" - P" * & S$##"$$$, p. 226, pl. 2.5.1, of Activity l.3.1). Nicola Carta, Luca Simonetto and Maria + g . 12. Corriga helped in the Field work. 2003 Pseudopolygnathus micropunctatus B* €' & We are deeply grateful to Claudia Spalletta for critical Z*"#" - C' $!*%*, p. 112, pl. 10, + g. 14. revision of the manuscript. ! i s study was supported by R.A.S. (grants LR7/07 - 2010, Resp. C. Corradini). ! is paper 2011 Pseudopolygnathus micropunctatus B* €' & is a contribution to IGCP Project n. 596 “Mid Palaeozoic Z*"#" - H$ $"%"# , p. 510, pl. 63, + gs 1-6, climate and biodiversity”. 8-10. 2013 Pseudopolygnathus micropunctatus B* €' & "#" !  S ##"$$ Z* - M' '%*, C' $ *%* $ $, References p. 88, + g. 3.12. B$ #" , R. 1925. Classi+ cation and stratigraphic use of the Remarks : ! i s species is characterized by a conodonts. Bulletin Geological Society of America 36: 218- lanceolate platform with a shagreen surface. Some 20.

32 GORTANIA. Geologia, Paleontologia, Paletnologia 34 (2012) FAMENNIAN CONODONTS FROM THE PIZZUL WEST SECTION CARNIC ALPS, ITALY

B* €', G. 1956. Oberdevonische Conodonten (to I ä) aus J*, Q. 1989. On the Frasnian conodont biostratigraphy in the dem Rheinischen Schiefergebirge. Notizblatt des Hessisches Guilin Area of Guangxi, South Cina. Courier Forschungs- Landesamt für Bodenforschung 84: 115-37. Institut Senckenberg 117: 303-19. B* €', G., & Z*"#" , W. 1956. Das Alter der “Urfer J*, Q., & W. Z*"#" . 1993. ! e Lali Section: an excellent Schishten” im Marburger Hinterland nach Conodonten. Reference Section for Upper Devonian in South China. Notizblatt des Hessisches Landesamt für Bodenforschung Courier Forschungs-Institut Senckenberg 157: 1-183. 84: 138-69. K€ % €"$ E.N., & A.V. K%&*%. 1996. New late Frasnian B $% '%, E.B., & M.G. M"€#. 1934a. Conodonts from conodonts of the Genus Palmatolepis from the Lyaiol’skaya the Bushberg sandstone and equivalent formations of Formation of southern Timan. Palaeontologicheskyi Missouri. Missouri University Studies 8, n. 4: 265-300. Zhurnal 3: 90-3. B $% '%, E.B., & M.G. M"€#. 1934b. Conodonts from K#$" , G., & H.R. L$%". 1985. Frasnian (Upper the Grassy Creek shale of Missouri. Missouri University Devonian) Conodont sequence at Luscar Mountain and Studies 8 (3): 171-259. Mount Haultain, Alberta Rocky Mountains. Canadian B $% '%, E.B., & M.G. M"€#. 1938. Conodonts from the Society of Petroleum Geology 3: 469-78. lower Mississippian of Missouri. In Stratigraphy and K#$" , G., C. S""$ & W. Z*"#" , cur. 1991. Catalogue palaeontology of the lower Mississippian of Missouri, cur. of Conodonts, volume 5. Stuttgart: E. Schweizerbart’sche E.B. B $% '% et al., Missouri University Studies, 13: 128- Verlagsbuchhandlung, 212 pp. 48. M$%&'%*, M. 1966. Conodonti neodevonici ed eocarboniferi C' $!*%*, C. 1998. Famennian conodonts from two al Monte Zermula (Alpi Carniche). Giornale di Geologia s. sections near Villasalto. Giornale di Geologia 60, special 2, 33: 461-88. issue: 122-35. M*##" , A.K., & W. Y'%%%* $. 1947. Conodonts from the C' $!*%*, C. 2003. Late Devonian (Famennian) conodonts type section of the Sweetland Creek Shale in Iowa. Journal from the Corona Mizziu Sections near Villasalto (Sardinia of Paleontology 21, n. 6: 501-17. Italy). Palaeontographia Italica 89: 65-116. M' '%*, A., C. C' $!*%* & C. S$##"$$$ C. 2013. C' $!*%*, C. 2008. Revision of Famennian-Tournaisian Conodonts from the Monte Taccu section (Famennian- (Late Devonian-Early Carboniferous) conodont biostrati- Tournaisian, Sardinia, Italy). Asociación Paleontológica graphy of Sardinia, Italy. Revue de micro-paléontologie 51: Argentina, Publicación Especial 13: 85-90. 123-32. M##" , K.J. 1956. Zur Kenntnis der Conodonten-Fauna C' $!*%*, C., M.G. C' *$, E. K*!', G. M% *', M. des europaischen Devons, 1; Die Gattung Palmatolepis. P'%! "##*, T.J. S%$$%" , L. S*'%"$$' & C. S$##"$$$. Abhandlunghen and Senckenbergischen Naturforschenden 2013. Cason di Lanza - leggere il passato nelle rocce. Le Gesellscha 494: 1-70. guide del Geoparco della Carnia 1: 48 pp. M##" , K.J. 1962. Zur systematischen Einteilung der C' $!*%*, C., M. P'%! "##*, M.G. C' *$, L. Conodontoporida. Palaontologische Zeitschri 36: 109- S*'%"$$', E. K*!', T.J. S%$$%" , C. S$##"$$$ & N. 17. C$ $$. 2012. Geology and stratigraphy of the Cason di O#**" *, R. 1970. Conodonti e zonatura del Devoniano Lanza area (Mount Zermula, Carnic Alps, Italy). Berichte superiore e riconoscimento del Carbonifero inferiore nei des Institutes für Erdwissenscha en, Karl-Franzens- calcari di Corona Mizziu (Gerrei, Sardegna). Bollettino Universität Graz 17: 83-103. della Società Paleontologica Italiana 8, n. 2: 63-152. C''" , C.L. 1931. New conodonts from the Woodford O%$$$%'$, N.S., & L.I. K'%'%'$. 2008. Frasnian Formation of Oklahoma. Journal of Paleontology 5, n. 3: conodonts from the Eastern Russian Platform. 230-43. Palaeontological Journal 42, n. 10: 997-1166. G"!*€, I. 1969. Karnik Alpler’den Alt Karbonifer’e ait P" *, M.C., & C. S$##"$$$. 1990. Famennian conodonts conodont’lar.  e Mineral Research and Exploration from climenid pelagic limestone, Carnia Alps, Italy. Institut of Turkey, Bulletin 70: 229-42. Palaeontographia Italica 77: 55-83. G#"%* $" , B.F., & G. K#$" . 1966. Upper Devonian P" *, M.C., & C. S$##"$$$. 1991. Famennian conodonts conodonts from the Canning Basin, Western Australia. from Cava Cantoniera and Malpasso sections, Carnic Journal of Paleontology 40, n. 4: 777-842. Alps, Italy. Bollettino della Società Paleontologica Italiana H$ $"%"# , S. 2011. Die globalen Annulata-Events und die 30, n. 1: 47-78. Dasberg-Krise (Famennium, Oberdevon) in Europa und P" *, M.C., & C. S$##"$$$. 1998a. Updating of the Nord-Afrika: hochau# ösende Conodonten-Stratigraphie, conodont biostratigraphy in the Carnic Alps (Italy). In Karbonat-Mikrofazies, Paläoökologie und Paläodiversität. Southern Alps Fiel Trip Guidebook, Ecos VII, cur. M.C. Münstersche Forschungen zur Geologie und Paläontologie P" * & C. S$##"$$$, 116-9. Giornale di Geologia, s. 3, 105: 17-527. 60, Spec. Issue. H"# , J. 1959. Conodonten aus dem Saalfelder Oberdevon P" *, M.C., & C. S$##"$$$. 1998b. ! e Upper marginifera (! uringen). Geologie 8, n. 6: 634-77. Zone (Late Devonian) in the Casera Collinetta di Sotto H"# , J. 1963. Zur “Phylogenese” und Taxionomie von C section (Carnic Alps, Italy). In Southern Alps Fiel Trip Palmatolepis (Conodontida, Oberdevon). Geologie 12, n. Guidebook, Ecos VII, cur. M.C. P" * & C. S$##"$$$, 4: 449-85. 150-7. Giornale di Geologia, s. 3, 60, Spec. Issue. H*%!", G.J. 1879. On conodonts from the Chazy and P" *, M.C., & C. S$##"$$$. 1998c. Late Famennian Cincinnati group of the Cambro-Silurian and from the conodonts from the Casera Collinetta di Sotto B section Hamilton and Genesee shale divisions of the Devonian in (Carnic Alps, Italy). In Southern Alps Fiel Trip Guidebook, Canada and the United States. Geological Society of London Ecos VII, cur. M.C. P" * & C. S$##"$$$, 158-67. Quarterly Journal 35, n. 3: 351-69. Giornale di Geologia, s. 3, 60, Spec. Issue.

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P" *, M.C., & C. S$##"$$$. 1998d. Latest Devonian and Montagne Noire (Southern France). Geologica et Early Carboniferous conodonts from the Casera Collinetta Palaeontologica, SB 3: 1-123. di Sotto A section (Carnic Alps, Italy). In Southern Alps S$##"$$$, C., G.B. V$* & C. V"%$% *%*. 1980. II # ysch Fiel Trip Guidebook, Ecos VII, cur. M.C. P" * & C. ercinico nella geologia dei Monti Paularo e Dimon (Alpi S$##"$$$, 168-81. Giornale di Geologia, s. 3, 60, Spec. Carniche). Memorie della Società Geologica Italiana 20: Issue. 243-65. P" *, M.C., & C. S$##"$$$. 1998e. Middle Famennian S$##"$$$, C., & M.C. P" *. 1998. ! e Frasnian-Famennian conodonts of the Elios section (Carnic Alps, Italy). In boundary at the Pramosio A section (Carnic Alps, Italy). Southern Alps Fiel Trip Guidebook, Ecos VII, cur. M.C. In Southern Alps Fiel Trip Guidebook, Ecos VII, cur. M.C. P" * & C. S$##"$$$, 205-13. Giornale di Geologia, s. P" * & C. S$##"$$$, 198-205. Giornale di Geologia, s. 3, 60, Spec. Issue. 3, 60, Spec. Issue. P" *, M.C., & C. S$##"$$$. 1998f. Late Famennian S&%#&" €*, M. 1971. Upper Devonian conodonts, conodonts of the Malpasso section (Carnic Alps, Italy). stratigraphy and facial development in the Holy Cross In Southern Alps Fiel Trip Guidebook, Ecos VII, cur. M.C. Mts.Acta Geologica Polonica 21, n. 1: 1-130. P" * & C. S$##"$$$, 220-7. Giornale di Geologia, s. 3, T $€"#"%, H., & S. H$ $"%"# . 2011. Neue Conodont 60, Spec. Issue. taxa aus dem höheren Famennium (Oberdevon) des P# " , P. 1969. Conodonten aus dem Devon der Frankenwaldes. Münstersche Forschungen zur Geologie Karnischen Alpen (Findenigkofel, Österreich). Jahrbuch and Paläontologie 105: 1-15. der Geologischen Bundesanstalt, 112: 399-440. U# *€, E.O., & R.S. B$ #" . 1926. A classi+ cation of the P'%! "##*, M., C. C' $!*%*, M.G. C' *$, E. K*!', toothlike fossils, conodonts, with descriptions of American L. S*'%"$$', C. S$##"$$$, T.J. S%$$%" & N. C$ $$. Devonian and Mississippian species. Proceedings of U.S. 2011. Pragian to Famennian depositional evolution of the Natural History Museum 68: 1-63. M. Pizzul area (Carnic Alps, Italy): preliminary results. 78- V"%$% *%*, C. 1990a. Geologia delle Alpi Carniche centro 9. In IGCP 596 - Opening Meeting, Graz, 19-24th September orientali. Udine: Pubblicazioni del Museo Friulano di 2011, cur. T.J. S%$$%" , E. K*!', W.E. P*##" & P. Storia Naturale, 36: 220 pp. K%* €', Berichte des Institutes für Erdwissenscha en, V"%$% *%*, C., cur. 1990b. Field Workshop on Carboniferous Karl-Franzens-Universität Graz, 16. to Permian sequence of the Pramollo-Nassfeld Basin S$%!‚" , C.A. 1976. Conodont biofacies of Late Devonian (Carnic Alps). Guidebook. Udine: Arti Gra+ c he Friulane: Polygnathus styriacus Zone in western United States. 159 pp. In Conodont Palaeoecology, cur. C.R. B$ %" , 171-86. V"%$% *%*, C., S$##"$$$, C., V$*, G.B., P'%! "##*, M., Geological Association of Canada, Special Paper 15. D"# Z'$$', S., F'%$$%$, C., L'%' S$#$!' , G. & G.B. S$%!‚" , C.A., & R. D "" "%. 1984. Late Devonian C$ %##*. 2009. Note Illustrative al Foglio 031 Ampezzo, icriodontid biofacies models and alternate shallow- Carta geologica d’Italia alla scala 1:50.000. ISPRA, Servizio water conodont zonation. In Conodont biofacies and Geologico Italiano: 232 pp. provincialism, cur. D.L. C#$ €, 143-78. Geological Society Z*"#" , W. 1958. Conodonten-feinstratigraphische Unter- of America. Special Paper 196. suchungen an der Grenze Mitteldevon/Oberdevon und S$%!‚" , C.A., W. Z*"#" , K. L"%$" *& & S.M. B *##. in der Adorfstufe. Notizblatt des Hessisches Landesamt für 1978. Phylogeny, speciation and zonation of Siphonodella Bodenforschung 87: 7-77. (Conodonta, Upper Devonian and Lower Carboniferous). Z*"#" , W. 1960. Conodonten aus dem Rheinischen Newsletter on Stratigraphy 7: 102-20. Untedevon (Gedinnium) des Remscheider sattels S$%!‚" , C.A., & W. Z*"#" . 1979. Taxonomy and (Rheinisches Schiefergebirge). Palaontologische Zeitschri biofacies of important conodonts of Late Devonian 34, n. 2: 169-201. styriacus-Zone, United States and Germany. Geologica et Z*"#" , W. 1962. Taxionomie und Phylogenie Ober- Palaeontologica 13: 173-212. devonischer Conodonten und ihre stratigraphische S$%%"$%%, D. 1955a. Beitrag zur Untergliederung des Bedeutung. Abhandlunghen des Hessisches Landesamt für Oberdevons nach Conodonten. Neues Jahrbuch für Bodenforschung 38: 166. Geologie und Paläontologie Abhandlungen, 100, n. 3: 324- Z*"#" , W. 1971. Conodont Stratigraphy of the European 31. Devonian. Geological Society of America Memoir 127, 227- S$%%"$%%, D. 1955b. Oberdevonische Conodonten (to II 84. á). Senckenbergiana Lethaea 26 n. 1-2: 123-56. Z*"#" , W., cur. 1973. Catalogue of Conodonts, volume 1. S€%#$%‚, H.P. 1969a. Das Paläozoikum zwischen Stuttgart: E. Schweizerbart’sche Verlagsbuchhandlung, Bishofalm und Hohem Trieb (Zentrale Karnischen Alpen). 504 pp. Jahrbuch der Geologischen Bundesanstalt 112: 265-320. Z*"#" , W., cur. 1975. Catalogue of Conodonts, volume 2. S€%#$%‚, H.P. 1969b. Conodonten aus dem Oberdevon Stuttgart: E. Schweizerbart’sche Verlagsbuchhandlung, und Unterkarbon des Kronhofgrabens (Karnischen Alpen, 404 pp. Österreich). Jahrbuch der Geologischen Bundesanstalt 112: Z*"#" , W., cur. 1977. Catalogue of Conodonts, volume 3. 321-54. Stuttgart: E. Schweizerbart’sche Verlagsbuchhandlung, S€%#$%‚, H.P. 1992. Stratigraphy, Biogeography and 574 pp. Paleoclimatology of the Alpine Paleozoic and its impli- Z*"#" , W., cur. 1981. Catalogue of Conodonts, volume 4. cations for Plate movements. Jahrbuch der Geologischen Stuttgart: E. Schweizerbart’sche Verlagsbuchhandlung, Bundesanstalt 135: 381-418. 445 pp. S€#€", I. 1999. Conodont multielement reconstruction Z*"#" , W., & J.W. H%!!#". 1969. Die Palmatolepis glabra- from the Early Famennian (Late Devonian) of the Gruppe (Conodonta) nach der Revision der Typen von

34 GORTANIA. Geologia, Paleontologia, Paletnologia 34 (2012) FAMENNIAN CONODONTS FROM THE PIZZUL WEST SECTION CARNIC ALPS, ITALY

Ulrich & Bassler durch J. W. Huddle. Fortschritte in der Geologie der Rheinland und Westfalen 16: 377-86. Z*"#" , W., & S$%!‚" , C.A. 1984. Palmatolepis-based revision of upper part of standard Late Devonian conodont zonation. In Conodont biofacies and provincialism, cur. D.L. C#$ €, Geological Society of America, Special Paper 196: 179-94. Z*"#" , W., & C.A. S$%!‚" . 1990. ! e Late Devonian Standard Conodont Zonation. Courier Forschungs-Institut Senckenberg 121: 1-115. Z*"#" , W., & C.A. S$%!‚" . 1994. Conodont Phylo- genetic-Zone Concept. Newsletter of Stratigraphy 30: 105- 23. Z*"#" , W., C.A. S$%!‚"  & R.L. A% $*%. 1974. Revision of Bispathodus group (Conodonta) in the Upper Devonian and Lower Carboniferous. Geologica et Palaeontologica 8: 97-112.

35 A. MOSSONI, C. CORRADINI, M. PONDRELLI GORTANIA. Geologia, Paleontologia, Paletnologia 34 (2012)

Authors’ addresses - Indirizzi degli Autori: - Angelo M' '%* Dipartimento di Scienze Chimiche e Geologiche Università degli Studi di Cagliari Via Trentino 51, I-09127 CAGLIARI e-mail: [email protected] - Carlo C' $!*%* Dipartimento di Scienze Chimiche e Geologiche Università degli Studi di Cagliari Via Trentino 51, I-09127 CAGLIARI e-mail: [email protected] - Monica P'%! "##* International Research School of Planetary Sciences, Dipartimento di Ingegneria e Geologia, Università D’Annunzio Via Pindaro 42, I-65217 PESCARA e-mail: [email protected]

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