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IDF-Report 95 (2016)
IDF International Dragonfly Fund Report Journal of the International Dragonfly Fund 125 Dejan Kulijer, Iva Miljević & Jelena Jakovljev Contribution of the participants of 4th Balkan Odonatological Meeting to the knowledge of Odonata distribution in Bosnia and Herzegovina Published 26.04.2016 95 ISSN 14353393 The International Dragonfly Fund (IDF) is a scientific society founded in 1996 for the impro vement of odonatological knowledge and the protection of species. Internet: http://www.dragonflyfund.org/ This series intends to publish studies promoted by IDF and to facilitate costefficient and ra pid dissemination of odonatological data.. Editorial Work: Milen Marinov, Geert de Knijf & Martin Schorr Layout: Martin Schorr IDFhome page: Holger Hunger Indexed: Zoological Record, Thomson Reuters, UK Printing: Colour Connection GmbH, Frankfurt Impressum: Publisher: International Dragonfly Fund e.V., Schulstr. 7B, 54314 Zerf, Germany. Email: [email protected] Responsible editor: Martin Schorr Cover picture: Cordulegaster heros Photographer: Falk Petzold Published 26.04.2016 Contribution of the participants of 4th Balkan Odonatological Meeting to the knowledge of Odonata distribution in Bosnia and Herzegovina Dejan Kulijer1, Iva Miljević2 & Jelena Jakovljev3 1National Museum of Bosnia and Herzegovina, Zmaja od Bosne 3, 71000 Sarajevo, Bosnia and Herzegovina. Email: [email protected] 2Center for Environment, Cara Lazara 24, 78 000 Banja Luka, Bosnia and Herzegovina. Email: [email protected] 3Univesity of Natural Resources and Life Sciences, Baumgasse 58/19 1030 Vienna, Austria. Email: [email protected] Abstract As a result of increased interest in dragonflies and close cooperation between odo natologists on the Balkan Peninsula, the Balkan Odonatological Meeting (BOOM) has been established in 2011. -
New Insects from the Earliest Permian of Carrizo Arroyo (New Mexico, USA) Bridging the Gap Between the Carboniferous and Permian Entomofaunas
Insect Systematics & Evolution 48 (2017) 493–511 brill.com/ise New insects from the earliest Permian of Carrizo Arroyo (New Mexico, USA) bridging the gap between the Carboniferous and Permian entomofaunas Jakub Prokopa,* and Jarmila Kukalová-Peckb aDepartment of Zoology, Faculty of Science, Charles University, Viničná 7, CZ-128 43 Praha 2, Czech Republic bEntomology, Canadian Museum of Nature, Ottawa, ON, Canada K1P 6P4 *Corresponding author, e-mail: [email protected] Version of Record, published online 7 April 2017; published in print 1 November 2017 Abstract New insects are described from the early Asselian of the Bursum Formation in Carrizo Arroyo, NM, USA. Carrizoneura carpenteri gen. et sp. nov. (Syntonopteridae) demonstrates traits in hindwing venation to Lithoneura and Syntonoptera, both known from the Moscovian of Illinois. Carrizoneura represents the latest unambiguous record of Syntonopteridae. Martynovia insignis represents the earliest evidence of Mar- tynoviidae. Carrizodiaphanoptera permiana gen. et sp. nov. extends range of Diaphanopteridae previously restricted to Gzhelian. The re-examination of the type speciesDiaphanoptera munieri reveals basally coa- lesced vein MA with stem of R and RP resulting in family diagnosis emendation. Arroyohymen splendens gen. et sp. nov. (Protohymenidae) displays features in venation similar to taxa known from early and late Permian from the USA and Russia. A new palaeodictyopteran wing attributable to Carrizopteryx cf. arroyo (Calvertiellidae) provides data on fore wing venation previously unknown. Thus, all these new discoveries show close relationship between late Pennsylvanian and early Permian entomofaunas. Keywords Ephemeropterida; Diaphanopterodea; Megasecoptera; Palaeodictyoptera; gen. et sp. nov; early Asselian; wing venation Introduction The fossil record of insects from continental deposits near the Carboniferous-Permian boundary is important for correlating insect evolution with changes in climate and in plant ecosystems. -
Dragonf Lies and Damself Lies of Europe
Dragonf lies and Damself lies of Europe A scientific approach to the identification of European Odonata without capture A simple yet detailed guide suitable both for beginners and more expert readers who wish to improve their knowledge of the order Odonata. This book contains images and photographs of all the European species having a stable population, with chapters about their anatomy, biology, behaviour, distribution range and period of flight, plus basic information about the vagrants with only a few sightings reported. On the whole, 143 reported species and over lies of Europe lies and Damself Dragonf 600 photographs are included. Published by WBA Project Srl CARLO GALLIANI, ROBERTO SCHERINI, ALIDA PIGLIA © 2017 Verona - Italy WBA Books ISSN 1973-7815 ISBN 97888903323-6-4 Supporting Institutions CONTENTS Preface 5 © WBA Project - Verona (Italy) Odonates: an introduction to the order 6 WBA HANDBOOKS 7 Dragonflies and Damselflies of Europe Systematics 7 ISSN 1973-7815 Anatomy of Odonates 9 ISBN 97888903323-6-4 Biology 14 Editorial Board: Ludivina Barrientos-Lozano, Ciudad Victoria (Mexico), Achille Casale, Sassari Mating and oviposition 23 (Italy), Mauro Daccordi, Verona (Italy), Pier Mauro Giachino, Torino (Italy), Laura Guidolin, Oviposition 34 Padova (Italy), Roy Kleukers, Leiden (Holland), Bruno Massa, Palermo (Italy), Giovanni Onore, Quito (Ecuador), Giuseppe Bartolomeo Osella, l’Aquila (Italy), Stewart B. Peck, Ottawa (Cana- Predators and preys 41 da), Fidel Alejandro Roig, Mendoza (Argentina), Jose Maria Salgado Costas, Leon (Spain), Fabio Pathogens and parasites 45 Stoch, Roma (Italy), Mauro Tretiach, Trieste (Italy), Dante Vailati, Brescia (Italy). Dichromism, androchromy and secondary homochromy 47 Editor-in-chief: Pier Mauro Giachino Particular situations in the daily life of a dragonfly 48 Managing Editor: Gianfranco Caoduro Warming up the wings 50 Translation: Alida Piglia Text revision: Michael L. -
1 General Introduction
1 General Introduction If the karate-ka (student) shall walk the true path, first he will cast aside all preference. Tatsuo Shimabuku, Grand Master of Isshin-ryu Karate 1.1 The Importance of Insects ~30% of the plants we grow for food and materials. Because of their great numbers and diversity, insects Insects transmit some of these pathogens. While have a considerable impact on human life and indus- weeds can often reduce pest attack, they can also try, particularly away from cities and in the tropics. harbour the pest’s enemies or provide alternative On the positive side they form a large and irreplace- resources for the pest itself. Then in storage, insects, able part of the ecosystem, especially as pollinators mites, rodents and fungi cause a further 30% loss. of fruit and vegetable crops and, of course, many Apart from such biotic damage, severe physical con- wild plants (Section 8.2.1). They also have a place ditions such as drought, storms and flooding cause in soil formation (Section 8.2.4) and are being used additional losses. For example, under ideal field increasingly in ‘greener’ methods of pest control. conditions new wheat varieties (e.g. Agnote and Biological control using insects as predators and Humber) would give yields of ~16 tonnes/ha, but parasites of pest insects has been developed in the produce typically about half this under good hus- West for over a century, and much longer in China. bandry. Pre-harvest destruction due only to insects More recently integrated pest management (IPM) is 10–13% (Pimentel et al., 1984; Thacker, 2002). -
Frank Morton Carpenter (1902-1994): Academic Biography and List of Publications
FRANK MORTON CARPENTER (1902-1994): ACADEMIC BIOGRAPHY AND LIST OF PUBLICATIONS BY DAVID G. FURTH 18 Hamilton Rd., Arlington, MA 02174 The present paper is meant to accompany the preceding one by Elizabeth Brosius, Assistant Editor at the University of Kansas, Paleontological Institute, who was extremely instrumental in aid- ing Prof. Frank Carpenter to finish his Treatise on Invertebrate Paleontology volumes on fossil insects. The Brosius paper is a brief profile taken from her personal interaction with Prof. Carpen- ter as well as numerous interviews about him with his friends, stu- dents, and colleagues. The present paper is intended to be more of an account of Prof. Carpenter's academic background and accom- plishments with the addition of some personal and academic accounts of the author's interaction with Frank Carpenter. Frank Morton Carpenter was born in Boston on 6 September 1902. When he was three years old his family (father Edwin A. and mother Maude Wall) moved from Boston to Revere and at age six his family moved to Melrose where he began to attend Lincoln School the following year. His father worked for the American Express Company but had a strong interest in natural history and taught his elder son (Edwin, four years older than Frank) about the constellations. Edwin later graduated from Harvard, studied astronomy, and became Director of the Astronomical Laboratory at the University of Arizona in Tucson. When Frank Carpenter was a sixth grader at Lincoln School his father encouraged his interest in butterflies and moths. In ninth grade Frank Carpenter began taking out books about insects from the Melrose Public Library. -
The Morphology of the Pterothorax of Ephemeroptera, Odonata
ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Stuttgarter Beiträge Naturkunde Serie A [Biologie] Jahr/Year: 2008 Band/Volume: NS_1_A Autor(en)/Author(s): Willkommen Jana Artikel/Article: The morphology of the pterothorax of Ephemeroptera, Odonata and Plecoptera (Insecta) and the homology of wing base sclerites and flight muscles 203-300 Stuttgarter Beiträge zur Naturkunde A, Neue Serie 1: 203–300; Stuttgart, 30.IV.2008. 203 The morphology of the pterothorax of Ephemeroptera, Odonata and Plecoptera (Insecta) and the homology of wing base sclerites and flight muscles1 JANA WILLKOMMEN Abstract The ability to fly was the decisive factor for the evolutionary success of the most diverse group of insects, the Pterygota. Nevertheless, the ground plan of the functionally important wing base has not been sufficiently clari- fied. The aim of this study is to homologise the wing base sclerites of Ephemeroptera, usually regarded as sister group of the remaining Pterygota, with that of other basal pterygote lineages and to reconstruct the ground plan of the wing base of Pterygota. The pterothoracic musculature of representatives of the three basal lineages of Ptery- gota (Ephemeroptera, Odonata and Neoptera) is also described and discussed. Contrary to previous hypotheses, it is shown that most elements of the neopteran wing base are also present in Ephemeroptera and Odonata. The wing base in the ground plan of Pterygota is presumably composed of three axil- lary sclerites. The proximal median plate is probably also present in the ground plan of Pterygota. The first axillary is provided with two muscles. -
Index to Contents
Index to Contents Author(s) Title Year Vol Pages Holland, Sonia Dragonfly Survey Reports – 1. Gloucestershire 1983 1 (1) 1-3 Butler, Stephen Notes on finding larvae of Somatochlora arctica (Zetterstedt) in N. W. Scotland 1983 1 (1) 4-5 Winsland, David Some observations on Erythromma najas (Hansemann) 1983 1 (1) 6 Merritt, R. Is Sympetrum nigrescens Lucas a good species? 1983 1 (1) 7-8 Vick, G. S. Is Sympetrum nigrescens Lucas a good species? 1983 1 (1) 7-8 Merritt, R. Coenagrion mercuriale (Charpentier) with notes on habitat 1983 1 (1) 9-12 Chelmick, D. G. Observations on the ecology and distribution of Oxygastra curtisii (Dale) 1983 1 (2) 11-14 Khan, R. J. Observations of Wood-mice (Apodemus sylvaticus) and Hobby (Falco subbuteo) feeding on dragonflies 1983 1 (2) 15 Marren, P. R. Scarce Species Status Report 2. A review of Coenagrion hastulatum (Charpentier) in Britain 1983 1 (2) 16-19 Merritt, R. Is Sympetrum nigrescens Lucas a good species? 1983 1 (2) 16-19 Mayo, M. C. A. Coenagrion mercuriale (Charpentier) on the flood plains of the River Itchen and River Test in Hampshire 1983 1 (2) 20-21 Welstead, A. R. Coenagrion mercuriale (Charpentier) on the flood plains of the River Itchen and river Test in Hampshire 1983 1 (2) 20-21 Kemp, R. G. Notes and observations on Gomphus vulgatissimus (Linnaeus) on the river Severn and River Thames 1983 1 (2) 22-25 Vick, G. S. Notes and observations on Gomphus vulgatissimus (Linnaeus) on the river Severn and River Thames 1983 1 (2) 22-25 Corbet, P. -
Journal Vol 26 No 2, October 2010
J. Br. Dragonfly Society, Volume 26 No. 2, October 2010 Journal of the CONTENTS DAVID CHELMICK - Studying British dragonflies in the British Dragonfly Society 1970s: the wilderness years .............................................. 57 Volume 26 Number 2 October 2010 BARRY NATTRESS - Folding wing behaviour in Cordulagaster boltonii (Donovan) ............................................................. 64 DAVID CHELMICK - Species Review 4: The Scarce Emerald Damselfly Lestes dryas Kirby with notes on the family Lestidae in the Western Palearctic ....................................66 JONATHAN. R. DIXON & DOROTHY E. GENNARD - The influence of meteorological conditions on the flight activity of the Blue-tailed Damselfly Ischnura elegans (Vander Linden), the Azure Damselfly Coenagrion puella (Linnaeus) and the Emerald Damselfly Lestes sponsa (Hansemann) ..... .............................................................................................. 83 ADRIAN J. PARR -. Migrant and dispersive dragonflies in Britain during 2009 ............................................................97 PAM TAYLOR & DAVE SMALLSHIRE - A change in status of the Dainty Damselfly Coenagrion scitulum (Rambur) in the United Kingdom ………....................................................107 Corrigendum ..........................................................................i The aims of the British Dragonfly Society (BDS) are to promote and encourage the study and conservation INSTRUCTIONS TO AUTHORS of Odonata and their natural habitats, especially in the -
Diversification of Insects Since the Devonian
www.nature.com/scientificreports OPEN Diversifcation of insects since the Devonian: a new approach based on morphological disparity of Received: 18 September 2017 Accepted: 12 February 2018 mouthparts Published: xx xx xxxx Patricia Nel1,2, Sylvain Bertrand2 & André Nel1 The majority of the analyses of the evolutionary history of the megadiverse class Insecta are based on the documented taxonomic palaeobiodiversity. A diferent approach, poorly investigated, is to focus on morphological disparity, linked to changes in the organisms’ functioning. Here we establish a hierarchy of the great geological epochs based on a new method using Wagner parsimony and a ‘presence/ absence of a morphological type of mouthpart of Hexapoda’ dataset. We showed the absence of major rupture in the evolution of the mouthparts, but six epochs during which numerous innovations and few extinctions happened, i.e., Late Carboniferous, Middle and Late Triassic, ‘Callovian-Oxfordian’, ‘Early’ Cretaceous, and ‘Albian-Cenomanian’. The three crises Permian-Triassic, Triassic-Jurassic, and Cretaceous-Cenozoic had no strong, visible impact on mouthparts types. We particularly emphasize the origination of mouthparts linked to nectarivory during the Cretaceous Terrestrial Revolution. We also underline the origination of mouthparts linked to phytophagy during the Middle and the Late Triassic, correlated to the diversifcation of the gymnosperms, especially in relation to the complex ‘fowers’ producing nectar of the Bennettitales and Gnetales. During their ca. 410 Ma history, hexapods have evolved morphologically to adapt in a continuously changing world, thereby resulting in a unique mega-biodiversity1. Age-old questions2–4 about insects’ macroevolution now- adays receive renewed interest thanks to the remarkable recent improvements in data and methods that allow incorporating full data, phylogenomic trees besides fossil record5–9. -
INDEX for VOLUME 52 (New Names in Boldface)
Journal of the Lepidopterists' Society 52(4). 1998, 388- 401 INDEX FOR VOLUME 52 (new names in boldface) 2-methyloctadecane. 356 Amblyscirtes, 237, 240 fimbriata pallida, 54 Abaeis nicippe, 57 jluonia, 54 Acacia farnesiana, 215 folia, 54 Acanthaceae, 74, 107,215 patriciae, 54 Acer rubrum, 128 raphaeli, 54 Aceraceae, 128 tolteca tolteca, 54 AchaZarus, 236, 240 Ampelocera hottleii, 109 casica,50 Anaea, 239, 242 toxeus,50 aidea, 62 Achlyodes, 240 Anartia, 242 busirus heros, 52 amathea colima, 59 selva , .52 Jatrophe, 25 Acrodipsas illidgei, 139 Jatrophe luteipicta, 59 Acronicta albarufa, 381 Anastnls adaptation, 207 robigus, 52 Adelia triloba, 338 sempiternus sempiternus, 52 Adelotypa eudocia, 66 Anatrytone, 237 Adelpha, 239 mazai,54 basiioicies has i/o ides , 62 Anchistea virginica, 128 celerio diademata , 62 Ancyloxypha, 240 fessonia fessonia , 62 arene,54 iphiclus massilides, 62 Anemeca ehrenbergii, 60 ixia leucas, 62 Annonaceae, 107 leuceria leuceria, 62 Anteos, 241 naxia epiphicla, 62 clorinde nivifera, 57 phylaca phyiaca, 62 maerula lacordairei , 57 serpa massilia, 62 Anteros carausius carausius, 65 Adhemarius gannascus, 110 Anthanassa, 239 ypsilon, 11 0 alexon alexon, 60 Aegiceras corniculatum, 141 ardys anlys, 60 Aellopos pto/yea arrUltor, 60 ceculus, 111 sitalces cortes, 60 clavi pes , III texana texana, 60 fadus , III tulcis, 60 Aeshnidae, 137 Antlwcharis Aethilla lavochrea, 52 cardamines, 156 Agathymus rethon, 55 euphenoid~s, 1.56 Aglais urticae, 156 Anthoptus insignis, 53 Agraulis Antig()nus vanillae, 25 emorsa,52 vanillae incarnata, -
PALAEODICTYOPTERA, MEGASECOPTERA and DIAPHANOPTERODEA (PALEOPTERA)* by JARMILA KUKALOVA-Peck Department of Geology, Carleton University Ottawa, Ontario, Canada
PTERALIA OF THE PALEOZOIC INSECT ORDERS PALAEODICTYOPTERA, MEGASECOPTERA AND DIAPHANOPTERODEA (PALEOPTERA)* BY JARMILA KUKALOVA-PEcK Department of Geology, Carleton University Ottawa, Ontario, Canada For an understanding of insect evolution the structure of the wing base is of major significance. However, the fossil record o pteralia in extinct orders is extremely scanty. This paper is concerned with the wing bases of certain Paleozoic Paleoptera, namely, Palaeodictyop- tera, Megasecoptera and Diaphanopterodea rom the Upper Carboni- ferous (Namurian) of Czechoslovakia, the Upper Carboniferous (Stephanian) of France, and the Lower Permian of Czechoslovakia and Kansas. Independently of the Neoptera, the Diaphanopterodea acquired the ability to flex the wings backwards over the abdomen. In this respect, the order is of special interest, and an attempt is made here to compare the pteralia o the Diaphanopterodea with those of extant Ephemeroptera. Our present knowledge of the wing base in Paleozoic Paleoptera is restricted to the axillary plate o several palaeodictyopteran adults (Kukalova, 96o, 969-7o), and palaeodictyopteran nymphs (Woot- ton, 972; Sharov, 97I) [See figures , 2 and 4]. Recently, the wing base has been described in the Diaphanopterodea, Family E1- moidae (Kukalova-Peck, 974) (/]g. 8). In the present paper, the axillary plates in Martynoviidae and Asthenohymenidae o the Order Diaphanopterodea are included, and for the first time the axillary sclerites in Megasecoptera are described. The interpretation and terminology of the pteralia in extinct Pale- optera are necessarily dependent upon the detailed functional mor- phology of extant Ephemeroptera and Odonata. At the same time, the wing base structures found in extinct orders provide an evolu- tionary view and might be helpful in unraveling the enigmatic archi- *This research has been aided in part by a Publication Grant from Carleton University and in part by a National Science Foundation Grant, GB 39720, F. -
The Occurrence and Diversity of Coal Measure Insects
Journal of the Geological Sociev, London, Vol. 144, 1987, pp. 5W-511, 3 figs. Printed in Northern Ireland The occurrence and diversity of Coal Measure insects E. A. JARZEMBOWSKI Booth Museum of Natural Hktory, Dyke Road, Brighton BN1 5AA, UK Abslrnd Insects are generally considered to be rare in the Upper Carboniferous Coal Measures. However, recent work in the Westphalian D of SW England suggests that many have been overlooked in the past. This is because wings, which are the most characteristic insect fossils, may be mistaken for detached ‘fern’ pinnules, which are much more common. The resemblance may be functional convergence rather than leaf-mimicry. The earliest members of the class Insecta in the strict sense occur in the Upper Carboniferous. Eleven major divisions or orders are represented in the Coal Measures of which only four are still living. Primitively wingless insects (Archaeognatha) are present, relatives of familiar living insects such as the silverfish. There are numerous winged insects, some ofwhich could fold their wings (Neoptera) and others which could not (Palaeoptera). Palaeopterous insects were more diverse than today. They include three extinct orders (Palaeodictyoptera, Megasecoptera, Diaphanopterodea) which were probably plant suckers like present clay bugs. Other extinct palaeopterous insects (order Protodonata) were probably aerial predators like modem dragonflies, and included some of the largest insects of all time (‘giant dragonflies’). By far the most common neopterous insects were cockroaches (Blattodea) which outnumber all other insects in theUpper Carboniferous. This abundance is perhaps less surprising when one considers the general picture of the coal forests as warm, humid, and rich in organic matter.