The Amaryllis
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Guide to the Flora of the Carolinas, Virginia, and Georgia, Working Draft of 17 March 2004 -- LILIACEAE
Guide to the Flora of the Carolinas, Virginia, and Georgia, Working Draft of 17 March 2004 -- LILIACEAE LILIACEAE de Jussieu 1789 (Lily Family) (also see AGAVACEAE, ALLIACEAE, ALSTROEMERIACEAE, AMARYLLIDACEAE, ASPARAGACEAE, COLCHICACEAE, HEMEROCALLIDACEAE, HOSTACEAE, HYACINTHACEAE, HYPOXIDACEAE, MELANTHIACEAE, NARTHECIACEAE, RUSCACEAE, SMILACACEAE, THEMIDACEAE, TOFIELDIACEAE) As here interpreted narrowly, the Liliaceae constitutes about 11 genera and 550 species, of the Northern Hemisphere. There has been much recent investigation and re-interpretation of evidence regarding the upper-level taxonomy of the Liliales, with strong suggestions that the broad Liliaceae recognized by Cronquist (1981) is artificial and polyphyletic. Cronquist (1993) himself concurs, at least to a degree: "we still await a comprehensive reorganization of the lilies into several families more comparable to other recognized families of angiosperms." Dahlgren & Clifford (1982) and Dahlgren, Clifford, & Yeo (1985) synthesized an early phase in the modern revolution of monocot taxonomy. Since then, additional research, especially molecular (Duvall et al. 1993, Chase et al. 1993, Bogler & Simpson 1995, and many others), has strongly validated the general lines (and many details) of Dahlgren's arrangement. The most recent synthesis (Kubitzki 1998a) is followed as the basis for familial and generic taxonomy of the lilies and their relatives (see summary below). References: Angiosperm Phylogeny Group (1998, 2003); Tamura in Kubitzki (1998a). Our “liliaceous” genera (members of orders placed in the Lilianae) are therefore divided as shown below, largely following Kubitzki (1998a) and some more recent molecular analyses. ALISMATALES TOFIELDIACEAE: Pleea, Tofieldia. LILIALES ALSTROEMERIACEAE: Alstroemeria COLCHICACEAE: Colchicum, Uvularia. LILIACEAE: Clintonia, Erythronium, Lilium, Medeola, Prosartes, Streptopus, Tricyrtis, Tulipa. MELANTHIACEAE: Amianthium, Anticlea, Chamaelirium, Helonias, Melanthium, Schoenocaulon, Stenanthium, Veratrum, Toxicoscordion, Trillium, Xerophyllum, Zigadenus. -
Differences in Floral Development Between Lycoris Radiata And
R ESEARCH ARTICLE doi: 10.2306/scienceasia1513-1874.2020.032 Differences in floral development between Lycoris radiata and Lycoris sprengeri a,† b,† c d a a,e, Junhuo Cai , Junjun Fan , Xuying Wei , Donglin Zhang , Jiajia Ren , Lu Zhang ∗ a College of Landscape and Art, Jiangxi Agricultural University, Nanchang 330045 China b College of Horticulture, Jinling Institute of Technology, Nanjing 210037 China c College of Art, Jiangxi Finance and Economics University, Nanchang 330032 China d Department of Horticulture, University of Georgia, Athens, GA 30602 USA e Collaboration Innovation Center of Jiangxi Typical Trees Cultivation and Utilization, Jiangxi Agricultural University, Nanchang 330045 China ∗Corresponding author, e-mail: [email protected], Cai J. and Fan J. contributed equally to this paper. Received 12 Sep 2019 Accepted 21 Mar 2020 ABSTRACT: Lycoris radiata and Lycoris sprengeri are two typical plants of the genus Lycoris. They have different leafing and flowering patterns. To understand the habits of Lycoris species and regulate the flowering period effectively, we compared flower bud development of these two species using phenological, anatomical, and physiological measurements. Our results showed there were significant differences in phenology between the two species. The sprouting stage of L. radiata took place in autumn, the leaf growth period lasted seven months, dormancy lasted approximately 100 days before flowering, and there was no dormancy period after flowering. In comparison, the sprouting period of L. sprengeri was delayed until the spring of the following year, the leaf growth stage lasted approximately 3 months, the flowering stage took place from Jul–Aug, and dormancy lasted 1–2 months before flowering and for another 6 months after flowering. -
Comparative Chloroplast Genomes of Four Lycoris Species (Amaryllidaceae) Provides New Insight Into Interspecific Relationship and Phylogeny
biology Article Comparative Chloroplast Genomes of Four Lycoris Species (Amaryllidaceae) Provides New Insight into Interspecific Relationship and Phylogeny Fengjiao Zhang 1,2, Ning Wang 1,2, Guanghao Cheng 1,2, Xiaochun Shu 1,2, Tao Wang 1,2 , Weibing Zhuang 1,2, Ruisen Lu 1,2,* and Zhong Wang 1,2,* 1 Jiangsu Key Laboratory for the Research and Utilization of Plant Resources, Institute of Botany, Jiangsu Province and Chinese Academy of Sciences (Nanjing Botanical Garden Mem. Sun Yat-Sen), Nanjing 210014, China; [email protected] (F.Z.); [email protected] (N.W.); [email protected] (G.C.); [email protected] (X.S.); [email protected] (T.W.); [email protected] (W.Z.) 2 The Jiangsu Provincial Platform for Conservation and Utilization of Agricultural Germplasm, Institute of Botany, Jiangsu Province and Chinese Academy of Sciences (Nanjing Botanical Garden Mem. Sun Yat-Sen), Nanjing 210014, China * Correspondence: [email protected] (R.L.); [email protected] (Z.W.) Simple Summary: The genus Lycoris (Amaryllidaceae) comprises about 20 species with high orna- mental and medicinal value. However, germplasm identification is still difficult due to frequent interspecific hybridization and intraspecific morphological variation within this genus. Plastid genome sequencing has been proven to be a useful tool to identify closely related species and is widely used in the field of plant evolution and phylogeny. In the present study, we provided four Citation: Zhang, F.; Wang, N.; chloroplast genomes of Lycoris and retrieved seven published species in the genus for comparative Cheng, G.; Shu, X.; Wang, T.; Zhuang, genomics and phylogenetic analyses. All these chloroplast genomes possess the typical quadripartite W.; Lu, R.; Wang, Z. -
Amaryllis – Hardy Scientific Name: Hippeastrum Johnsoni Common
Name: Amaryllis – Hardy Scientific name: Hippeastrum johnsoni Common Names: Cluster Amaryllis, Hurricane Lily, Magic Lily, Spider Lily, Stone Garlic. Life Cycle: Hardy bulb. Height: 12 to 36 inches (30 to 90 cm). Native: Asia. Growing Region: Zones 7 to 10. Flowers: Late summer through to autumn. Flower Details: White, red, pink, orange, yellow. Lily- like. Umbel; four to eight flowers. Foliage: Slender. Long. Grow Outside: Usually grown from bulbs or vegetatively propagated plants as seed grown plants can take up to 12 years to bloom. Bulbs: 3 to 8 inches (8 to 20 cm) depending upon species. End of summer Requirements and care: Full sunlight or partial shade. Good drainage. Acidic to neutral soil. Rich soil, moist soil. Regular watering to maintain soil moisture. Requires a feed every two years; do this during the growing season. Propagate: by planting bulblets once blooming has finished. Source: http://www.plant-biology.com/Lycoris-Hardy-Amaryllis.php http://www.brecksbulbs.ca/product/Hardy-Amaryllis-Mixture/Summer_Bulbs Extension programs service people of all ages regardless of socioeconomic level, race, color, sex, religion, disability, or national origin. The Texas A&M University System, U.S. Department of Agriculture, and the County Commissioners Courts of Texas Cooperating A member of The Texas A&M University System and its statewide Agriculture Program. Common Name: Artemesia - Powis Castle Botanical name: Artemesiax Powis Castle Plant Type: Perennial Light Requirement: High Water Requirement: Low Hardiness/Zone: 4 - 8 Heat/Drought Tolerance: High Height: 3 ft Width/Spacing: 3ft Flower Color: Yellow Blooming Period: Rarely flowers Plant Form or Habit: Evergreen woody perennial, or shrub Foliage Color and Texture: Leaves are finely dissected like filigreed silver lacework. -
Complete Chloroplast Genomes Shed Light on Phylogenetic
www.nature.com/scientificreports OPEN Complete chloroplast genomes shed light on phylogenetic relationships, divergence time, and biogeography of Allioideae (Amaryllidaceae) Ju Namgung1,4, Hoang Dang Khoa Do1,2,4, Changkyun Kim1, Hyeok Jae Choi3 & Joo‑Hwan Kim1* Allioideae includes economically important bulb crops such as garlic, onion, leeks, and some ornamental plants in Amaryllidaceae. Here, we reported the complete chloroplast genome (cpDNA) sequences of 17 species of Allioideae, fve of Amaryllidoideae, and one of Agapanthoideae. These cpDNA sequences represent 80 protein‑coding, 30 tRNA, and four rRNA genes, and range from 151,808 to 159,998 bp in length. Loss and pseudogenization of multiple genes (i.e., rps2, infA, and rpl22) appear to have occurred multiple times during the evolution of Alloideae. Additionally, eight mutation hotspots, including rps15-ycf1, rps16-trnQ-UUG, petG-trnW-CCA , psbA upstream, rpl32- trnL-UAG , ycf1, rpl22, matK, and ndhF, were identifed in the studied Allium species. Additionally, we present the frst phylogenomic analysis among the four tribes of Allioideae based on 74 cpDNA coding regions of 21 species of Allioideae, fve species of Amaryllidoideae, one species of Agapanthoideae, and fve species representing selected members of Asparagales. Our molecular phylogenomic results strongly support the monophyly of Allioideae, which is sister to Amaryllioideae. Within Allioideae, Tulbaghieae was sister to Gilliesieae‑Leucocoryneae whereas Allieae was sister to the clade of Tulbaghieae‑ Gilliesieae‑Leucocoryneae. Molecular dating analyses revealed the crown age of Allioideae in the Eocene (40.1 mya) followed by diferentiation of Allieae in the early Miocene (21.3 mya). The split of Gilliesieae from Leucocoryneae was estimated at 16.5 mya. -
Generic Classification of Amaryllidaceae Tribe Hippeastreae Nicolás García,1 Alan W
TAXON 2019 García & al. • Genera of Hippeastreae SYSTEMATICS AND PHYLOGENY Generic classification of Amaryllidaceae tribe Hippeastreae Nicolás García,1 Alan W. Meerow,2 Silvia Arroyo-Leuenberger,3 Renata S. Oliveira,4 Julie H. Dutilh,4 Pamela S. Soltis5 & Walter S. Judd5 1 Herbario EIF & Laboratorio de Sistemática y Evolución de Plantas, Facultad de Ciencias Forestales y de la Conservación de la Naturaleza, Universidad de Chile, Av. Santa Rosa 11315, La Pintana, Santiago, Chile 2 USDA-ARS-SHRS, National Germplasm Repository, 13601 Old Cutler Rd., Miami, Florida 33158, U.S.A. 3 Instituto de Botánica Darwinion, Labardén 200, CC 22, B1642HYD, San Isidro, Buenos Aires, Argentina 4 Departamento de Biologia Vegetal, Instituto de Biologia, Universidade Estadual de Campinas, Postal Code 6109, 13083-970 Campinas, SP, Brazil 5 Florida Museum of Natural History, University of Florida, Gainesville, Florida 32611, U.S.A. Address for correspondence: Nicolás García, [email protected] DOI https://doi.org/10.1002/tax.12062 Abstract A robust generic classification for Amaryllidaceae has remained elusive mainly due to the lack of unequivocal diagnostic characters, a consequence of highly canalized variation and a deeply reticulated evolutionary history. A consensus classification is pro- posed here, based on recent molecular phylogenetic studies, morphological and cytogenetic variation, and accounting for secondary criteria of classification, such as nomenclatural stability. Using the latest sutribal classification of Hippeastreae (Hippeastrinae and Traubiinae) as a foundation, we propose the recognition of six genera, namely Eremolirion gen. nov., Hippeastrum, Phycella s.l., Rhodolirium s.str., Traubia, and Zephyranthes s.l. A subgeneric classification is suggested for Hippeastrum and Zephyranthes to denote putative subclades. -
Forcing Bulbs “Forcing” – Coaxing, Actually – Is the Term Used To
Forcing Bulbs “Forcing” – coaxing, actually – is the term used to describe the process that stimulates bulbs to bloom out of season. Among the most commonly forced bulb flowers are amaryllis, paper white narcissus, muscari, and hyacinths. Certainly they are the easiest! When selecting bulbs for forcing, look for varieties that are specifically recommended. Spring flowering bulbs usually require a rooting period of about 12 to 15 weeks at temperatures between 41 to 48°F in order to produce a good root system; which is essential if they are to be “forced” into flowering. Storing Bulbs Bare bulbs can be stored for several weeks in your refrigerator prior to potting without damage if they are packed properly. The best way to store them is in a mesh bag or paper bag with holes that permit ventilation. You don’t have to worry about water, as the humidity inside a refrigerator is usually high enough. Check them periodically to make sure they are not molding or drying out. Pot your bulbs right away if you have an appropriate location immediately available, such as an old (functioning) refrigerator, a root cellar or cool basement – or if outdoor temperatures are below 45°F. If, however, your area experiences an “Indian Summer’, your fridge can serve as a suitable rooting area. Vegetable or crisper drawers can be used, but don’t store bulbs in the same drawer you keep ripening fruit or vegetables – these give off ethylene gas which can harm the bulbs. (Also some bulbs are poisonous, so make sure to keep them out of reach of children). -
Introducing Lycoris to U.S. Flower Lovers
PEGGY GREB (D300-1) Introducing Lycoris to Golden-yellow fl owers of Lycoris aurea. U.S. Flower Lovers or more than 20 years ARS horticulturist Mark Roh has been intrigued by the origins and habitats of the exotic and beautiful Lycoris. Though various Lycoris species have been grown as ornamentals in China, F Korea, and Japan for many centuries, only two species are readily available here: L. squamigera and L. radiata. They—and the rarer L. incarnata, L. chejuensis, and L. flavescens—are maintained at the U.S. National Arboretum (USNA), in Washington, D.C., and in Beltsville, Maryland. In 1984, Roh collected several unidentified Lycoris species from Anduck Valley, on Korea’s Jeju Island. This subtropical area hosts about 4,000 species of plants. Then in 1998, more Lycoris species were collected in Japan, Korea, and China. DNA molecular markers and chromosome studies proved that some of the unidentified Lycoris collected from Anduck Valley were L. incarnata, a species previously known to be native only to China. It is possible that this accession was brought from China to Korea by bulb collectors, but no record of that can be found. In the past 6 years, extensive visits to areas near Anduck Valley have been made by Roh or by collaborators Mun Seok Seong, of Jeju-do Agricultural Research and Extension Services, and Yong Bong Park, 12 Agricultural Research/December 2005 387664.indd 12 11/11/05 6:23:47 AM PEGGY GREB (D303-1) of Jeju National University. But they have of drought and waterlogging, as well as failed to locate any more L. -
Amaryllis Handout.Indd
No other indoor fl owering bulb provides such a spectacular display of blooms as the amaryllis – it’s no wonder it is so popular. Amaryllis are prized for their huge 4 to 6 inch Success with wide bold, bell-like fl owers on thick, sturdy stalks that are 30 inches high or taller. Originally from South Africa and South Amaryllis America, today American and Dutch hybrids of amaryllis (Hippeastrum) come in white, various shades of red, scarlet, pink, salmon, orange and white marked with red or pink. Amaryllis is one of the easiest indoor bulbs to grow. They are available to purchase October through December (while supplies last) and can be planted from October until the end of April. For success with your Flowering time begins in early December and amaryllis, we recommend can extend through these fi ne products: the end owf May, depending on when the bulb is planted. Blooms come along rapid- ly after planting, with some amaryllis bulbs fl ower- ing within 21 How to days after planting. Watch- ing the fl ower stalks shoot Shopping List: plant and up and the buds expand and ❑ open will keep the whole family Amaryllis Bulb grow these in suspense. ❑ A Pot With proper care and ❑ Potting Mix spectacular handling, amaryllis bulbs ❑ Indoor Houseplant Food will bloom indoors year ❑ after year. Small Watering Can bulbs ❑ Stakes & Twine (if necessary) 100 Jericho Tpke., Westbury, NY 11590 100 Jericho Tpke., Westbury, NY 11590 516-334-0066 • www.HicksNurseries.com 516-334-0066 • www.HicksNurseries.com 12/17 How to Get Your Amaryllis to Bloom Again • When the fl owers fade cut them from the stem and allow the leaves to grow. -
Taxonomic Novelties in Southern Brazilian Amaryllidaceae – Iv: Hippeastrum Correiense (Bury) Worsley, the Correct Name of the Famous H
BALDUINIA, n. 64, p. 42-58, 04-XI-2018 http://dx.doi.org/10.5902/2358198035738 TAXONOMIC NOVELTIES IN SOUTHERN BRAZILIAN AMARYLLIDACEAE – IV: HIPPEASTRUM CORREIENSE (BURY) WORSLEY, THE CORRECT NAME OF THE FAMOUS H. MORELIANUM LEM.; AND H. VERDIANUM, A NEW SPECIES FROM SANTA CATARINA1 HENRIQUE MALLMANN BÜNEKER2 REGIS EDUARDO BASTIAN3 ABSTRACT In this article, Hippeastrum verdianum, a new species of Amaryllidaceae (Amaryllidoideae, Hippeastreae), which occurs in rocky cliffs in Santa Catarina (Brazil), is described and illustrated. Data are provided on their habitat, ecology and geographical distribution. The new species shows morphological affinity with H. correiense and H. papilio. In order to establish a consistent argumentative basis for the description of the new species, we clarify the taxonomic identity of H. correiense, proposing lectotypes for it as well as other binomials that we consider as synonyms. Keywords: Taxonomy, Monocot, Amaryllidoideae, Hippeastreae, Hippeastrinae, Hippeastrum subgen. Omphalissa RESUMO [Novidades taxonômicas em Amaryllidaceae sul-brasileiras – IV: Hippeastrum correiense (Bury) Worsley, o nome correto do famoso H. morelianum Lem.; e H. verdianum, uma nova espécie para Santa Catarina]. É descrito e ilustrado Hippeastrum verdianum, uma nova espécie de Amaryllidaceae (Amaryllidoideae, Hippeastreae) que ocorre em escarpas rochosas de Santa Catarina (Brasil). São fornecidos dados sobre seu hábitat, ecologia e distribuição geográfica. A nova espécie apresenta afinidade morfológica com H. correiense e H. papilio. -
Amaryllis in Alachua County Fact Sheet # ACFFL1311
UF/IFAS Extension Alachua County 2800 NE 39th Ave. Gainesville, FL 32609 Tel: (352)955-2402 Amaryllis in Alachua County Fact Sheet # ACFFL1311 Taylor Clem, PhD Published Date: January 15, 2019 About the Amaryllis Amaryllis in the Landscape An Amaryllis (Hippeastrum x hybridum) is a beautiful Plant Amaryllis bulbs outside between September and tropical and subtropical flower from South Africa, January in an area with partial sun and well-drained commonly given as gifts around the holidays. Florida’s soils. If you plant the Amaryllis in too deep of shade, warmer temperatures allow us to plant Amaryllis bulbs they will not be as showy in your landscape. within our landscapes once we begin to warm-up in the late winter and early spring. Planting Amaryllis Bulbs in Pots Amaryllis Flower To plant an Amaryllis bulb, select a pot that has a diameter, at minimum, 1.5x the diameter of the bulb. Amaryllis can be top-heavy, so a larger pot helps keep the flower from toppling over. Fill your pot with a well-drained potting media and place the Amaryllis bulb in the center of the pot. To help encourage early flowering of the Amaryllis when planting, keep half or two-thirds of the bulb above the soil line. Make sure you firmly press onto the soil around the bulb to keep it from moving around. After planting the Amaryllis, water thoroughly and place in a bright spot (not direct sun) around your home that stays relatively cool. Now you wait. No watering or fertilizing is required until the flower sprouts. -
Networks in a Large-Scale Phylogenetic Analysis: Reconstructing Evolutionary History of Asparagales (Lilianae) Based on Four Plastid Genes
Networks in a Large-Scale Phylogenetic Analysis: Reconstructing Evolutionary History of Asparagales (Lilianae) Based on Four Plastid Genes Shichao Chen1., Dong-Kap Kim2., Mark W. Chase3, Joo-Hwan Kim4* 1 College of Life Science and Technology, Tongji University, Shanghai, China, 2 Division of Forest Resource Conservation, Korea National Arboretum, Pocheon, Gyeonggi- do, Korea, 3 Jodrell Laboratory, Royal Botanic Gardens, Kew, Richmond, United Kingdom, 4 Department of Life Science, Gachon University, Seongnam, Gyeonggi-do, Korea Abstract Phylogenetic analysis aims to produce a bifurcating tree, which disregards conflicting signals and displays only those that are present in a large proportion of the data. However, any character (or tree) conflict in a dataset allows the exploration of support for various evolutionary hypotheses. Although data-display network approaches exist, biologists cannot easily and routinely use them to compute rooted phylogenetic networks on real datasets containing hundreds of taxa. Here, we constructed an original neighbour-net for a large dataset of Asparagales to highlight the aspects of the resulting network that will be important for interpreting phylogeny. The analyses were largely conducted with new data collected for the same loci as in previous studies, but from different species accessions and greater sampling in many cases than in published analyses. The network tree summarised the majority data pattern in the characters of plastid sequences before tree building, which largely confirmed the currently recognised phylogenetic relationships. Most conflicting signals are at the base of each group along the Asparagales backbone, which helps us to establish the expectancy and advance our understanding of some difficult taxa relationships and their phylogeny.