Butterflies of Reserva Las Gralarias
By Tim Kell and Vicki Liu
© 2012
All butterflies listed have been found and photographed at Reserva Las Gralarias and are ordered taxonomically.
The compilation of this list would not have been possible without the help of Keith Willmott to whom we are very grateful for his assistance in identifying butterflies in the families Nymphalidae, Pieridae, Papilionidae and Riodinidae. Also thanks are extended to Robert Busby for help in identifying butterflies within the Lycaenidae and to Tomasz Pyrcz for confirming some species within the Satyrinae (Nymphalidae).
Family Genus Species Subspecies Author_year_taxon Lycaenidae Contrafacia marmoris H. H. Druce, 1907
Lycaenidae Marachina maraches
Lycaenidae Megathecla cupentus
Lycaenidae Micandra aegides Felder & Felder, 1865
Lycaenidae Rhamma arria
Lycaenidae Thaeides theia
Lycaenidae Theritas monica
Nymphalidae Catonephele chromis chromis Doubleday, 1848 Nymphalidae Epiphile epicaste epicaste
Nymphalidae Epiphile epimenes Fassl, 1912
Nymphalidae Epiphile orea distalis Attal, 1997 Nymphalidae Diaethria clymena bourcieri Guenée, 1872 Nymphalidae Perisama oppelii erebina Oberthür, 1916 Nymphalidae Perisama tryphena insignis
Nymphalidae Perisama yeba ilia Röber, 1916 Nymphalidae Marpesia marcella C. & R. Felder, 1861
Nymphalidae Marpesia zerynthia Hübner, 1823
Nymphalidae Caligo eurilochus
Nymphalidae Caligo oberthurii
Nymphalidae Eryphanis greeneyi
Nymphalidae Fountainea centaurus C. & R. Felder, 1867
Nymphalidae Fountainea nessus Latreille, 1813
Nymphalidae Fountainea ryphea ecuadorialis Johnson & Comstock, 1941 Nymphalidae Memphis beatrix pseudiphis Staudinger, 1887 Nymphalidae Memphis lyceus H. Druce, 1877
Nymphalidae Memphis perenna austrina Comstock, 1961 Nymphalidae Noreppa chromus
Nymphalidae Danaus plexippus
Nymphalidae Dircenna adina steinheili Staudinger, 1884 Nymphalidae Episcada polita ssp. Haensch, 1905 Nymphalidae Hyalenna sulmona tersa
Nymphalidae Pteronymia latilla barilla Haensch, 1903 Nymphalidae Pteronymia veia ssp Willmott & Lamas Nymphalidae Pteronymia zerlina pronuba
Nymphalidae Greta andromica andromica
Nymphalidae Greta ochretis ochretis Haensch, 1903 Nymphalidae Greta polissena polissena Hewitson, 1863 Nymphalidae Ithomia terra
Nymphalidae Pagyris ulla vanilla
Nymphalidae Mechanitis polymnia chimborazona H. Bates, 1864 Nymphalidae Methona confusa
Nymphalidae Patricia dercyllidas dercyllidas Hewitson, 1864 Nymphalidae Oleria padilla padilla Hewitson, 1863 Nymphalidae Oleria quadrata quadrata Haensch, 1903 Nymphalidae Actinote anteas suspecta Jordan, 1913 Nymphalidae Actinote euclia euclia
Nymphalidae Actinote hilaris desmiala
Nymphalidae Actinote ozomene ozomene Hewitson, 1868 Nymphalidae Agraulis vanillae Linnaeus, 1758
Nymphalidae Dione glycera
Nymphalidae Dione juno andicola Bates, 1864 Nymphalidae Dione moneta
Nymphalidae Dryas iulia alcionea Cramer, 1779 Nymphalidae Eueides procula
Nymphalidae Heliconius clysonymus hygiana Hewitson, 1867 Nymphalidae Heliconius sara sprucei
Nymphalidae Podotricha judith mellosa
Nymphalidae Adelpha corcyra collina Fruhstorfer, 1907 Nymphalidae Adelpha cythera
Nymphalidae Adelpha godmani
Nymphalidae Adelpha leucopthalma irminella
Nymphalidae Adelpha lycorias spruceana
Nymphalidae Antirrhea undulata
Nymphalidae Smyrna blomfildia blomfildia Fabricius, 1782 Nymphalidae Anartia amathea amathea Linnaeus, 1758 Nymphalidae Siproeta epaphus
Nymphalidae Siproeta stelenes sophene
Nymphalidae Castilia neria Hewitson, 1869
Nymphalidae Eresia carme lycus Hall, 1928 Nymphalidae Eresia poecilina casiphia
Nymphalidae Tegosa pastazena H.W. Bates, 1864
Nymphalidae Telenassa delphia guajoni Dognin, 1887 Nymphalidae Hypanartia dione dione Latreille, 1813 Nymphalidae Hypanartia lethe Fabricius, 1793
Nymphalidae Hypanartia trimaculata
Nymphalidae Vanessa braziliensis Moore, 1883
Nymphalidae Manataria maculata Hopffer, 1874
Nymphalidae Euptychoides nossis Hewitson, 1862
Nymphalidae Hermeuptychia harmonia Butler, 1877
Nymphalidae Magneuptychia alcinoe C. & R. Felder, 1867
Nymphalidae Pareuptychia occirhoe
Nymphalidae Forsterinaria enjuerma Pena & Lamas, MS
Nymphalidae Forsterinaria pallida aurita Peña & Lamas, 2005 Nymphalidae Parataygetis lineata Godman & Salvin, 1880
Nymphalidae Taygetis puritana Weeks, 1902
Nymphalidae Corades chelonis lactefusa Thieme, 1907 Nymphalidae Corades enyo almo Thieme, 1907 Nymphalidae Corades medeba Hewitson, 1850
Nymphalidae Corades pannonia ploas Thieme, 1907 Nymphalidae Corades violacea
Nymphalidae Daedalma eliza Pyrcz, 2010
Nymphalidae Eretris apuleja subrufescens Grose-Smith & Kirby, 1895 Nymphalidae Eretris calisto calisto C. & R. Felder, 1867 Nymphalidae Lasiophila ciris Thieme, 1907
Nymphalidae Lasiophila confusa Staudinger, 1897
Nymphalidae Lymanopoda albocincta fina Pyrcz Nymphalidae Lymanopoda obsoleta Westwood, 1851
Nymphalidae Manerebia inderena fina Pycrz & Willmott, 2006 Nymphalidae Mygona irmina Doubleday, 1849
Nymphalidae Oxeoschistus puerta isolda Thieme, 1907 Nymphalidae Panyapedaliodes drymaea Hewitson, 1858
Nymphalidae Panyapedaliodes panyasis Hewitson, 1862
Nymphalidae Panyapedaliodes tomentosa Thieme, 1905
Nymphalidae Pedaliodes costipunctata
Nymphalidae Pedaliodes montagna Adams & Bernard, 1981
Nymphalidae Pedaliodes peucestas Hewitson, 1862
Nymphalidae Pedaliodes poesia poema Pyrcz & Viloria, 1998 Nymphalidae Pedaliodes polla Thieme, 1905
Nymphalidae Pedaliodes simpla costipunctata Weymer, 1912 Nymphalidae Pedaliodes sonata
Nymphalidae Pedaliodes zingara Heredia & Viloria, 2004
Nymphalidae Proboscis propyleae
Nymphalidae Pronophila orcus
Nymphalidae Pseudomaniola ilsa
Nymphalidae Steroma bega
Papilionidae Pterourus cacicus
Papilionidae Eurytides servile columbus
Pieridae Eurema xantochlora ectriva Butler, 1873 Pieridae Phoebis neocypris rurina
Pieridae Dismorphia lelex xiomara Lamas, 2004 Pieridae Dismorphia medora
Pieridae Dismorphia zaela zaela Hewitson, 1858 Pieridae Lieinix nemesis nemesis Latreille, 1813 Pieridae Archonias brassolis rosacea Butler, 1873 Pieridae Catasticta chelidonis maja Reissinger, 1972 Pieridae Leodonta chiriquensis intermedia
Pieridae Leptophobia caesia caesia Lucas, 1852 Pieridae Leptophobia eleusis Lucas, 1852
Pieridae Leptophobia helena Lucas, 1852
Pieridae Leptophobia penthica
Pieridae Leptophobia tovaria tovaria C. & R. Felder, 1861 Riodinidae Emesis cypria cilix
Riodinidae Mesosemia mamilia Hewitson, 1870
Riodinidae Mesosemia mancia
Riodinidae Mesosemia species 1
Riodinidae Mesosemia species 2
Riodinidae Teratophthalma nigrita Salazar, 1993
Riodinidae Napaea merula Thieme, 1907
Riodinidae Necyria zaneta Hewitson, 1875
Riodinidae Siseme aristoteles sprucei Bates, 1868