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Butterflies of Reserva Las Gralarias Butterflies of Reserva Las Gralarias By Tim Kell and Vicki Liu © 2012 All butterflies listed have been found and photographed at Reserva Las Gralarias and are ordered taxonomically. The compilation of this list would not have been possible without the help of Keith Willmott to whom we are very grateful for his assistance in identifying butterflies in the families Nymphalidae, Pieridae, Papilionidae and Riodinidae. Also thanks are extended to Robert Busby for help in identifying butterflies within the Lycaenidae and to Tomasz Pyrcz for confirming some species within the Satyrinae (Nymphalidae). Family Genus Species Subspecies Author_year_taxon Lycaenidae Contrafacia marmoris H. H. Druce, 1907 Lycaenidae Marachina maraches Lycaenidae Megathecla cupentus Lycaenidae Micandra aegides Felder & Felder, 1865 Lycaenidae Rhamma arria Lycaenidae Thaeides theia Lycaenidae Theritas monica Nymphalidae Catonephele chromis chromis Doubleday, 1848 Nymphalidae Epiphile epicaste epicaste Nymphalidae Epiphile epimenes Fassl, 1912 Nymphalidae Epiphile orea distalis Attal, 1997 Nymphalidae Diaethria clymena bourcieri Guenée, 1872 Nymphalidae Perisama oppelii erebina Oberthür, 1916 Nymphalidae Perisama tryphena insignis Nymphalidae Perisama yeba ilia Röber, 1916 Nymphalidae Marpesia marcella C. & R. Felder, 1861 Nymphalidae Marpesia zerynthia Hübner, 1823 Nymphalidae Caligo eurilochus Nymphalidae Caligo oberthurii Nymphalidae Eryphanis greeneyi Nymphalidae Fountainea centaurus C. & R. Felder, 1867 Nymphalidae Fountainea nessus Latreille, 1813 Nymphalidae Fountainea ryphea ecuadorialis Johnson & Comstock, 1941 Nymphalidae Memphis beatrix pseudiphis Staudinger, 1887 Nymphalidae Memphis lyceus H. Druce, 1877 Nymphalidae Memphis perenna austrina Comstock, 1961 Nymphalidae Noreppa chromus Nymphalidae Danaus plexippus Nymphalidae Dircenna adina steinheili Staudinger, 1884 Nymphalidae Episcada polita ssp. Haensch, 1905 Nymphalidae Hyalenna sulmona tersa Nymphalidae Pteronymia latilla barilla Haensch, 1903 Nymphalidae Pteronymia veia ssp Willmott & Lamas Nymphalidae Pteronymia zerlina pronuba Nymphalidae Greta andromica andromica Nymphalidae Greta ochretis ochretis Haensch, 1903 Nymphalidae Greta polissena polissena Hewitson, 1863 Nymphalidae Ithomia terra Nymphalidae Pagyris ulla vanilla Nymphalidae Mechanitis polymnia chimborazona H. Bates, 1864 Nymphalidae Methona confusa Nymphalidae Patricia dercyllidas dercyllidas Hewitson, 1864 Nymphalidae Oleria padilla padilla Hewitson, 1863 Nymphalidae Oleria quadrata quadrata Haensch, 1903 Nymphalidae Actinote anteas suspecta Jordan, 1913 Nymphalidae Actinote euclia euclia Nymphalidae Actinote hilaris desmiala Nymphalidae Actinote ozomene ozomene Hewitson, 1868 Nymphalidae Agraulis vanillae Linnaeus, 1758 Nymphalidae Dione glycera Nymphalidae Dione juno andicola Bates, 1864 Nymphalidae Dione moneta Nymphalidae Dryas iulia alcionea Cramer, 1779 Nymphalidae Eueides procula Nymphalidae Heliconius clysonymus hygiana Hewitson, 1867 Nymphalidae Heliconius sara sprucei Nymphalidae Podotricha judith mellosa Nymphalidae Adelpha corcyra collina Fruhstorfer, 1907 Nymphalidae Adelpha cythera Nymphalidae Adelpha godmani Nymphalidae Adelpha leucopthalma irminella Nymphalidae Adelpha lycorias spruceana Nymphalidae Antirrhea undulata Nymphalidae Smyrna blomfildia blomfildia Fabricius, 1782 Nymphalidae Anartia amathea amathea Linnaeus, 1758 Nymphalidae Siproeta epaphus Nymphalidae Siproeta stelenes sophene Nymphalidae Castilia neria Hewitson, 1869 Nymphalidae Eresia carme lycus Hall, 1928 Nymphalidae Eresia poecilina casiphia Nymphalidae Tegosa pastazena H.W. Bates, 1864 Nymphalidae Telenassa delphia guajoni Dognin, 1887 Nymphalidae Hypanartia dione dione Latreille, 1813 Nymphalidae Hypanartia lethe Fabricius, 1793 Nymphalidae Hypanartia trimaculata Nymphalidae Vanessa braziliensis Moore, 1883 Nymphalidae Manataria maculata Hopffer, 1874 Nymphalidae Euptychoides nossis Hewitson, 1862 Nymphalidae Hermeuptychia harmonia Butler, 1877 Nymphalidae Magneuptychia alcinoe C. & R. Felder, 1867 Nymphalidae Pareuptychia occirhoe Nymphalidae Forsterinaria enjuerma Pena & Lamas, MS Nymphalidae Forsterinaria pallida aurita Peña & Lamas, 2005 Nymphalidae Parataygetis lineata Godman & Salvin, 1880 Nymphalidae Taygetis puritana Weeks, 1902 Nymphalidae Corades chelonis lactefusa Thieme, 1907 Nymphalidae Corades enyo almo Thieme, 1907 Nymphalidae Corades medeba Hewitson, 1850 Nymphalidae Corades pannonia ploas Thieme, 1907 Nymphalidae Corades violacea Nymphalidae Daedalma eliza Pyrcz, 2010 Nymphalidae Eretris apuleja subrufescens Grose-Smith & Kirby, 1895 Nymphalidae Eretris calisto calisto C. & R. Felder, 1867 Nymphalidae Lasiophila ciris Thieme, 1907 Nymphalidae Lasiophila confusa Staudinger, 1897 Nymphalidae Lymanopoda albocincta fina Pyrcz Nymphalidae Lymanopoda obsoleta Westwood, 1851 Nymphalidae Manerebia inderena fina Pycrz & Willmott, 2006 Nymphalidae Mygona irmina Doubleday, 1849 Nymphalidae Oxeoschistus puerta isolda Thieme, 1907 Nymphalidae Panyapedaliodes drymaea Hewitson, 1858 Nymphalidae Panyapedaliodes panyasis Hewitson, 1862 Nymphalidae Panyapedaliodes tomentosa Thieme, 1905 Nymphalidae Pedaliodes costipunctata Nymphalidae Pedaliodes montagna Adams & Bernard, 1981 Nymphalidae Pedaliodes peucestas Hewitson, 1862 Nymphalidae Pedaliodes poesia poema Pyrcz & Viloria, 1998 Nymphalidae Pedaliodes polla Thieme, 1905 Nymphalidae Pedaliodes simpla costipunctata Weymer, 1912 Nymphalidae Pedaliodes sonata Nymphalidae Pedaliodes zingara Heredia & Viloria, 2004 Nymphalidae Proboscis propyleae Nymphalidae Pronophila orcus Nymphalidae Pseudomaniola ilsa Nymphalidae Steroma bega Papilionidae Pterourus cacicus Papilionidae Eurytides servile columbus Pieridae Eurema xantochlora ectriva Butler, 1873 Pieridae Phoebis neocypris rurina Pieridae Dismorphia lelex xiomara Lamas, 2004 Pieridae Dismorphia medora Pieridae Dismorphia zaela zaela Hewitson, 1858 Pieridae Lieinix nemesis nemesis Latreille, 1813 Pieridae Archonias brassolis rosacea Butler, 1873 Pieridae Catasticta chelidonis maja Reissinger, 1972 Pieridae Leodonta chiriquensis intermedia Pieridae Leptophobia caesia caesia Lucas, 1852 Pieridae Leptophobia eleusis Lucas, 1852 Pieridae Leptophobia helena Lucas, 1852 Pieridae Leptophobia penthica Pieridae Leptophobia tovaria tovaria C. & R. Felder, 1861 Riodinidae Emesis cypria cilix Riodinidae Mesosemia mamilia Hewitson, 1870 Riodinidae Mesosemia mancia Riodinidae Mesosemia species 1 Riodinidae Mesosemia species 2 Riodinidae Teratophthalma nigrita Salazar, 1993 Riodinidae Napaea merula Thieme, 1907 Riodinidae Necyria zaneta Hewitson, 1875 Riodinidae Siseme aristoteles sprucei Bates, 1868 .
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