MBL ANNUAL REPORT 2013

Marine Biological Laboratory, Woods Hole

AN AFFILIATE OF THE UNIVERSITY OF CHICAGO

2013 MBL ANNUAL REPORT i1 i2 MBL ANNUAL REPORT 2013

CONTENTS

1 MESSAGE FROM THE PRESIDENT & DIRECTOR

2 HIGHLIGHTS

87 FINANCIALSMBL AT-A-GLANCE

11 GIFTS

22 LEADERSHIP

The MBL Annual Report is published by the Marine Biological Laboratory. Although the greatest possible care has been taken in the preparation of this record, the MBL recognizes the possibility of inaccuracies. If any are noted, please accept our apology and advise us of any corrections to be made.

Office of Communications MBL 7 MBL Street Woods Hole, MA 02543 [email protected] 2013 MBL ANNUAL REPORT i3 i4 MBL ANNUAL REPORT 2013 2013 was a historic year for the MBL. chaired by Neil Shubin, the Robert R. Bensley Professor of From the President As the laboratory commemorated its past with a 125th Organismal Biology and Anatomy and Senior Advisor to and Director anniversary celebration, we embarked on the promising future the President and Vice President for Research and National of a full affiliation with a major university. Laboratories at the University of Chicago, was formed in the Fall of 2013 to foster and develop the affiliation through With the support and hard work of the Board, scientists, staff, collaborative research and educational initiatives. These efforts and the greater MBL community, we found a partner in the remain underway and have expanded to include joint scientific University of Chicago, a remarkable institution with strong and operational retreats in both Chicago and Woods Hole. historical and intellectual ties to the MBL that date back to the laboratory’s founding in 1888. At a special meeting of the This spring, the MBL Board of Trustees welcomed Robert J. MBL Corporation in June of 2013, members voted for a change Zimmer, president of the University of Chicago, as the new in the bylaws that made it possible for the MBL to form a chair of the MBL Board. President Zimmer succeeds Jack W. landmark affiliation with the University. Rowe, who led the MBL through an exceptionally successful capital campaign and two significant renovation projects Both UChicago and the MBL have reputations for scientific that transformed Loeb and Rowe laboratories. The MBL has excellence and highly collaborative cultures that draw top benefitted tremendously from Jack Rowe’s leadership and from biologists from around the world. And with our affiliation, the exceptional generosity of the Jack and Valerie Rowe family, a shared mission of leadership and innovation in scientific for which we are very grateful. research and education was strengthened. As the MBL looks toward its next 125 years, we are energized In the months following the affiliation’s official start on July 1, by the opportunity to craft a vision and strategy that will make 2013, MBL scientists and staff worked diligently with University the most of our affiliation with the University of Chicago and colleagues with the essential task of implementing the new further distinguish the quality and impact of our research and partnership through both operational and scientific planning. educational programs. There is enormous room for discovery before us as we work to ensure that the future of the MBL is as MBL’s resident faculty members met regularly to discuss ways bright, if not brighter, than its remarkable past. to enhance interactions among programs within the MBL, and with faculty and programs at the University of Chicago. A committee comprised of the heads of MBL’s administrative units was convened in the summer of 2013, and focused on Joan Ruderman operational implementation. A University faculty committee, MBL President and Director

2013 MBL ANNUAL REPORT 1 Senator Warren Visits the MBL Highlights U.S. Senator Elizabeth Warren (D-Massachusetts) toured the MBL accompanied by MBL President and Director Joan Ruderman, University of Chicago President Robert J. Zimmer, Argonne National Laboratory Director Eric Isaacs, and other members of the institutions’ leadership teams. The MBL’s affiliation with the University of Chicago, Senator Warren said, “is a great partnership; great for Massachusetts and for [the MBL].” Her visit, she said, was a “reminder of why we are investing in research … Woods Hole is the place the world is looking to for the best research and latest ideas.”

Life in the “Plastisphere” of the MBL Celebrates 125th Anniversary University of Chicago and MBL Form Landmark Affiliation World’s Oceans The MBL commemorated its Plastics are the most abundant form of In July 2013 the University of Chicago and the MBL 125th anniversary with a series of marine debris, with global production special events, including Discovery strengthened their combined eminence and innovation in rising and documented impacts in Day, a free family science fair and some marine environments. Associate biological research and education by forming an affiliation anniversary party. In conjunction with Scientist Linda Amaral Zettler and the celebration, the MBL Community that builds upon their shared values and historical ties. Both collaborators published a study Archives Project collected documenting the microbial community institutions have reputations for scientific excellence, highly community photos, memorabilia, associated with plastic marine debris and remembrances for the MBL collaborative cultures that draw top biologists from around at several locations in the North Archives and History of MBL website, Atlantic. They uncovered a diverse the world, and programs that will benefit from the affiliation’s developed by the MBLWHOI Library, multitude of microbes, a community EOL Informatics Group, and Arizona combination of strengths. now called the “Plastisphere.” They State University. showed that plastisphere communities are distinct from surrounding surface water, implying that plastic serves as a novel ecological habitat in the open ocean.

2 MBL ANNUAL REPORT 2013 University of Chicago, MBL Launch Research Awards, Scholarship Programs The MBL and UChicago introduced the Frank R. Lillie Research Innovation Awards. The competitive, multi-year grant program, one of the first results of the UChicago-MBL affiliation, supports innovative, interdisciplinary research that will lead to novel ideas and biological discovery. UChicago also initiated a full-tuition, merit based, renewable scholarship for undergraduate study at the University to be awarded annually to the child of a year-round MBL employee who is accepted for enrollment at the University of Chicago.

Long-Term Tundra Warming Study New Sculptures Grace Provides Insight into Ecosystem’s Waterfront Park Resiliency MBL’s Waterfront Park was enhanced University of California Santa with the installation of two new Barbara Ph.D. student Seeta Sistla, sculptures. “Flukes,” a cast-bronze a graduate of the Brown-MBL, abstract form inspired by the shape program, compared plant species and of a whale’s tail, was generously total carbon and nitrogen storage in donated to the MBL by the artist, warmed and control tundra plots. Gordon Gund. A life-sized bronze These plots were established by statue of environmentalist and writer Ecosystems Center Senior Scientist Rachel Carson was also unveiled. Gus Shaver more than 20 years The statue commemorates Carson’s ago as part of the Arctic Long-Term ties to Woods Hole and Cape Cod Ecological Research Project. Sistla and is modeled on a photograph found that warming increased plant taken of the scientist seated on a biomass and woody dominance, Woods Hole dock, notebook in hand, indirectly increased winter soil looking out at the sea. The statue is temperature and suppressed surface- the culmination of the work of the soil-decomposer activity, but did not Falmouth, MA-based Rachel Carson change total soil carbon or nitrogen Statue Committee. stocks and thereby increased ecosystem carbon storage. This is an important prediction that indicates carbon loss could be partially offset by greater storage in vegetation as the tundra warms.

2013 MBL ANNUAL REPORT 3 Jellyfish Built for Efficiency, Genes Linked to Human Neurological to shallow dry soil delivers sips of Not Speed Disorders Found in Lamprey Genome sustaining water during drought Jellyfish may not be fast, but they are A consortium of scientists including to surface soil microbes that make one of the most efficient swimmers Jennifer Morgan of the Eugene Bell nitrogen available to plants. Working on the planet, reported MBL Center for Regenerative Biology in Utah’s sagebrush steppe, the team Whitman Investigators Brad Gemmell and Tissue Engineering published monitored this “hydraulic lift” of and John Costello of Providence the genome of the sea lamprey, a soil water and discovered that the College and Sean Colin of Roger vertebrate fish used by Morgan to mechanism stimulated microbial Williams University. The key to study the molecular and cellular basis activity and more than doubled the the energy savings is a previously of spinal cord regeneration. Morgan plants’ uptake of nitrogen from the overlooked part of the way jellies and MBL Whitman Investigators surrounding surface soil at exactly move, a mechanism scientists Ona Bloom and Joseph Buxbaum the time they were flowering and call “passive energy.” During the identified several genes linked to setting seed. Greater understanding relaxation phase, after the jellyfish human neurological disorders, of hydraulic lift could aid scientists contracts, the scientists including Alzheimer’s disease, and growers in finding ways to discovered that another vortex ring in Parkinson’s disease, and spinal protect crops from the effects of the water gives the animal a second cord injury. prolonged drought. locomotive boost. The discovery may explain why jellyfish can “bloom” and New Program Helps Scientists Build overrun an ecosystem, outcompeting a SUCCESSful Career much swifter predators like fish. The MBL launched SUCCESS (Shaping and Understanding Career Choices Microbial Diversity Course Named in Education, Science and Self), a “Milestones in Microbiology” Site new career development program The MBL Microbial Diversity Course for young scientists. The program was honored as the 2013 “Milestones features a series of weekly workshops in Microbiology Site” by the led by panels of established scientists American Society for Microbiology. from institutions across the United The designation recognizes places States. Topics address everything where major developments in from choosing a career, to honing microbiology occurred and/or where presentation skills, to obtaining outstanding microbiologists made funding and finding mentors. seminal discoveries. The course, initiated in 1971, has shaped the Staying Alive in the High and Dry careers of generations of outstanding Ecosystems Center Senior Scientist microbiologists, and continues to be Zoe Cardon and colleagues showed a premier site for advanced training how the movement of water from at the leading edge of microbiological deep moist soil, through plant roots, investigation.

4 MBL ANNUAL REPORT 2013 Shedding Light on Cell Structures Regenerative Biology and Tissue Engineering. Bilogan conducted Cellular Dynamics Program scientists her doctoral research in the lab of Rudolf Oldenbourg, Michael Shribak, Associate Scientist Marko Horb. and Shalin Mehta developed the Bilogan’s research focused on optical setup, image acquisition, embryonic development, in particular and processing algorithms needed development of the pancreas. to measure the polarization Understanding how the pancreas sensitive absorption of pigments develops is vital to finding treatments New Insight on Cuttlefish Decision- in photoreceptors and in other for a range of pancreatic diseases, Making During Camouflage light absorbing cell structures. including diabetes and pancreatic In related work, members of the A study from Kimberly Ulmer and cancer. Oldenbourg laboratory Roger Hanlon in the Program in created OpenPolScope.org for users Sensory Physiology and Behavior and developers of polarized light Impacts of Beach Oiling on Coastal and their colleagues showed that microscopy techniques. The open- Microbes when cuttlefish change their skin access website is a platform for patterns for camouflage purposes, The beach ecosystems along the the collection and dissemination of they respond to vertical visual northern Gulf Coast underwent a knowledge about the technology, cues in their environment more major disturbance in the months its applications, and its further strongly than to horizontal cues. following Deepwater Horizon oil spill. development. Many prior experiments had shown Bay Paul Center scientists published the influence of two-dimensional a study of the changes in microbial substrates, such as sand and gravel community composition from oil habitats, yet many marine habitats Study Examines Effects of Cropland hydrocarbon contaminated sands at have three-dimensional structures, Intensification on Amazon seven coastal sites that experienced such as rocks and coral, among Watershed a range of oiling. They documented which cuttlefish camouflage from Ecosystems Center Senior Scientists that, in addition to increases in oil predators. Such choices highlight the Christopher Neill and Linda Deegan hydrocarbon-degrading bacteria, selective decision-making that occurs and colleagues study ways that there were community-wide impacts in cuttlefish as they determine their clearing of Amazon forest for including increased variability of core, camouflage body patterns. resident, and transient members of soybean cropland influences runoff to streams, water quality and stream the bacterial community in coastal Bell Center Student Awarded Ph.D. sands. These changes could affect habitats. The work also indicates the biological and chemical stability Brown-MBL Program student that cleared and intensively farmed of coastal ecosystems. Cassandra Bilogan successfully regions of the Amazon are near a defended her Ph.D. dissertation, point where clearing for cropland is making her the first Brown-MBL reducing rainfall by lowering the student to complete her degree amount of water recycled to the in the Eugene Bell Center for atmosphere.

2013 MBL ANNUAL REPORT 5 Rudolf Oldenbourg Appointed jaw identity in mammals and bony Director of Cellular Dynamics fishes—is actually a primitive, shared Program feature of the jaw and gill arches MBL Senior Scientist Rudolf of jawed vertebrates. The evidence Oldenbourg was named director supports the hypothesis made by of the Cellular Dynamics Program, anatomist Karl Gegenbauer more whose research focuses on than a century ago and demonstrates developing new techniques and that the structures were primitively technologies for the study of living specified by a common Dlx blueprint. cells. Oldenbourg has pioneered advances in transmitted light Embryology Course Turns 120 microscopy and invented the The MBL’s Embryology course LC-PolScope, a polarized light celebrated its 120th session in 2013. microscope enhanced by the The course was founded in 1893 use of liquid crystal technology, by University of electronic imaging, and digital image Chicago scientists How to Survive Without Sex processing. The LC-PolScope is now Charles Otis Bdelloid rotifers, a group of more widely used in clinical and research Whitman (then the 400 small aquatic invertebrate laboratories around the world and MBL Director) species, are considered an has found particular relevance in and Frank Rattray “evolutionary scandal” due to their fertility clinics, where it can be used Lillie. Considered successful evolution apparently noninvasively to visualize meiotic the premier without sex. An international spindles in human eggs. course on animal collaboration of scientists including developmental the Bay Paul Center laboratories Fish Study Probes Evolution of biology, the course of Irina Arkhipova and David Mark Vertebrate Jaw brings together Welch published the first genome The emergence of the biting jaw is leading faculty sequence of a bdelloid rotifer. Their a major novelty in the evolution of and students in an analysis is shedding new light on the vertebrates. A classical and widely intensive research evolutionary significance of sex, and cited hypothesis that the jaw evolved experience that suggests that the homogenizing and from the gill skeleton remained explores the latest paradigms, diversifying roles of sex may have unsupported by palaeontological problems, and technologies in been replaced in bdelloids by gene evidence or genetic data. Using modern developmental biology. conversion and horizontal shark, skate, and paddlefish embryos, To celebrate the course’s legacy, a gene transfer. Whitman Investigator Andrew commemorative symposium was Gillis of Dalhousie University and held at which former directors, his colleagues showed that nested faculty, and students discussed expression of the Dlx genes—the “Dlx their research, and considered the code” that specifies upper and lower course’s role in shaping the practice of science.

6 MBL ANNUAL REPORT 2013

The Marine Biological Laboratory (MBL) is dedicated to scientific discovery and improving the human condition through research and MBL At-A-Glance* education in biology, biomedicine, and environmental science. Founded in Woods Hole, Massachusetts, in 1888, the MBL is a private, nonprofit institution and an affiliate of the University of Chicago.

LEADERSHIP Bell Center for Regenerative Biology & Tissue Encyclopedia of Life (EOL): The MBL is a founding member Chairman of the Board of Trustees: Robert J. Zimmer Engineering: Cell, developmental and regenerative of the Encyclopedia of Life, a global consortium working President and Director: Joan V. Ruderman biology; tissue engineering and regeneration to provide free, open access to curated digital knowledge Speaker of the MBL Society: Colleen M. Cavanaugh about life on Earth, with a special focus on more than Cellular Dynamics Program: Microscopy, imaging and 200,000 marine organisms. The software programming team SCIENTISTS & AFFILIATED STAFF cellular dynamics; cellular physiology and biocurrents; responsible for building EOL is based at the MBL. The MBL has approximately 250 year-round employees, biosensors about half of which are scientists and science support SCIENTIFIC CONFERENCES staff. The staff is joined each year by more than 300 Program in Sensory Physiology & Behavior: Sensory The MBL annually hosts scientific meetings, conferences, visiting scientists, summer staff, and research associates biology (especially vision, balance, hearing); cephalopod and departmental retreats, accommodating more than from hundreds of institutions around the world, as well camouflage; animal behavior 2,600 participants from around the world. as a large number of faculty and students participating in MBL courses (see below). Whitman Center for Visiting Research: Collaborative RESOURCES research by leading scientists from around the world The MBLWHOI Library houses one of the world’s foremost AWARD-WINNING SCIENCE print and electronic literature collections in the biological, Among the scientists with a significant affiliation with EDUCATIONAL PROGRAMS biomedical, ecological, and oceanographic sciences; and the MBL (scientists, course faculty and students) are 55 MBL Courses: 25 courses providing advanced, conducts digitization and informatics projects. Nobel Prize winners (since 1929); 124 Howard Hughes laboratory-based research training in fields such as Medical Institute investigators, early career scientists, cellular physiology, embryology, neurobiology, and The W. M. Keck Ecological and Evolutionary Genetics international researchers, and professors (since 1960); microbiology, including (in 2013) 557 students from 416 Facility is a high-throughput DNA template production and 229 Members of the National Academy of Sciences institutions and 45 countries; and 845 faculty, staff, and sequencing facility in the MBL’s Josephine Bay Paul Center. (since 1960); and 198 Members of the American lecturers from 276 institutions and 17 countries. Academy of Arts and Sciences (since 1960). The Marine Resources Department Educational programs also include undergraduate is an advanced facility for maintaining, culturing, and AFFILIATION WITH THE UNIVERSITY OF CHICAGO education and independent research, graduate training, providing aquatic organisms essential to biological, The MBL and the University of Chicago formed the Brown-MBL Graduate Program in Biological and biomedical, and ecological research. an affiliation on July 1, 2013, that enhances both Environmental Sciences, postdoctoral training, and institutions’ missions of leadership and innovation in community continuing education programs. NationalXenopus Resource breeds and maintains scientific research and education. The affiliation builds Xenopus (frog) genetic stocks; and provides training on shared values and historical ties between Chicago and OTHER PARTNERSHIPS in Xenopus husbandry, cell biology, imaging, genetics, the MBL, which was led by University of Chicago faculty The Brown-MBL Partnership fosters research transgenesis, and genomics. members for the first four decades of its existence. collaborations between scientists from the MBL and Brown University, and includes undergraduate training, FINANCES MAJOR PROGRAMS & RESEARCH AREAS a joint graduate program, graduate training grants, and 2014 Operating Budget: $47.4 million (40% government Ecosystems Center: Ecosystems processes, collaborative research programs. grants and contracts; 11% nonfederal grants and contracts; biogeochemistry, and climate change 8% contributions; 7% endowment earnings; 34% other) The Woods Hole Consortium is a research and Endowment: $76.0 million(as of Dec. 31, 2013) Bay Paul Center for Comparative Molecular Biology educational alliance between the MBL, the Woods Net Assets: $166.7 million (as of Dec. 31, 2013) & Evolution: Microbial ecology; genetics and Hole Oceanographic Institution, and the Woods Hole metagenomics; molecular evolution Research Center.

* as of August 2014 2013 MBL ANNUAL REPORT 7 2013 brought several significant financial changes to the MBL, Report of the in addition to transitioning to the new affiliation with the University of Chicago, which Treasurer became effective July 1, 2013. The University has committed to substantial financial support to the MBL for the coming years, and in 2013 $6.0 million in cash was received as income for operations. MBL realized a $2.0 million loss from operating activities, after including the University support, largely driven by decreases in federal revenue as available grant streams continue to tighten.

The affiliation with the University necessitated a review of the fair market value of MBL’s assets and liabilities, as the MBL results will be consolidated into the University’s financial statements. The result of the review was a revaluation of MBL’s real estate assets, and a $50.0 million increase in land value was recorded on MBL’s balance sheet in 2013. Net assets as of December 31, 2013 were $166.7 million, up from $109.7 million as of December 31, 2012. MBL’s endowment value rose to $76.0 million at the end of 2013, up from $69.4 million at the end of 2012.

MBL has been working closely with the University on intensive strategic planning activities, both scientific and operational, since July 2013. Revenues and expenditures of each major business unit are being reviewed in detail, and an operational modeling exercise has begun to consider new activities and their revenue impacts on the institution. Partnerships between staff and researchers at both institutions have developed,

integrating activities and contributing to initial programmatic successes and operational The financial statements of the improvements. While there is much work left to be done, leadership of both the MBL and Marine Biological Laboratory for the fiscal year ending the University are optimistic and enthusiastic about the potential for mutual benefit, and December 31, 2013, were audited the future growth of the MBL. by Grant Thornton LLP.

Complete financial statements are available on request from:

— Mary S. Harrington, Treasurer Financial Services MBL 7 MBL Street Woods Hole, MA 02543

8 MBL ANNUAL REPORT 2013 Statement of Financial Position December 31, 2013 Statement of Activities: Year ended December 31, 2013

Temporarily Permanently Unrestricted Restricted Restricted Total ASSETS Operating support and revenues: Cash and cash equivalents $ 734,067 Federal grants $ 18,883,765 $ - $ - $ 18,883,765 Cash held for others 302,915 Non-Federal grants and contracts 4,913,541 1,136,000 - 6,049,541 Accounts receivable, net of allowance for doubtful accounts Income from affiliate 6,040,000 - - 6,040,000 Fees for conferences and services 4,436,109 - - 4,436,109 of $35,000 567,396 Contributions 2,155,322 1,089,088 295,058 3,539,468 Contributions receivable, net 1,765,351 Investment earnings used for operations 387,920 2,569,511 - 2,957,431 Receivables due for costs incurred on grants and contracts 3,195,727 Laboratory rentals 1,011,158 - - 1,011,158 Inventory and prepaid expenses 786,214 Tuition, net 936,324 - - 936,324 Investments, at fair value 80,846,876 Other revenue 615,635 - - 615,635 Charitable remainder trusts 480,797 Investment loss (6,301) - - (6,301) Plant assets, net 115,192,315 Net assets released from restrictions and reclassifications 5,749,042 (5,781,543) 32,501 - Other assets 1,343,419 Total operating support Total assets $ 205,215,077 and revenues 45,122,515 ( 986,944) 327,559 44,463,130

Expenses: LIABILITIES AND NET ASSETS Research 27,612,268 - - 27,612,268 Education 10,000,661 - - 10,000,661 Visiting research 4,161,126 - - 4,161,126 Accounts payable and accrued expenses $ 3,133,860 Conferences, housing and dining 3,419,631 - - 3,419,631 Future program resources 302,915 Other expenses 1,954,660 - - 1,954,660 Deferred income 2,029,653 Annuities and unitrusts payable 422,372 Total expenses 47,148,346 - - 47,148,346 Interest rate swap 3,049,573 Long-term debt 29,615,000 Change in net assets before non-operating activities ( 2,025,831) ( 986,944) 327,559 (2,685,216) Total liabilities 38,553,373 Non-operating revenue (expense): Change in value of interest rate swap 3,171,103 - - 3,171,103 Commitments and contingencies Present value adjustment to annuities (17,903) - - (17,903) Post retirement - related charges other Net assets: than periodic pension costs 638,409 - - 638,409 Unrestricted 93,244,744 Contributions for investment in plant 19,024 - - 19,024 Temporarily restricted 21,900,525 Change in net assets from Permanently restricted 51,516,435 non-operating activities 3,810,633 - - 3,810,633

Total net assets 166,661,704 Investment income and gains (losses): Net investment income and gains 1,109,848 7,650,791 - 8,760,639 Total liabilities and net assets $ 205,215,077 Less: investment earnings used for operations (387,920) (2,569,511) - (2,957,431) Recovery of deficiencies in historical values 299,729 ( 299,729) - -

Investment income and gains 1,021,657 4,781,551 - 5,803,208

Changes in net assets 2,806,459 3,794,607 327,559 6,928,625

Net assets, December 31, 2012 40,438,285 18,105,918 51,188,876 109,733,079

Fair value adjustment due to Affiliation 50,000,000 - - 50,000,000

Net assets, December 31, 2013 $ 93,244,744 $ 21,900,525 $ 51,516,435 $ 166,661,704

2013 MBL ANNUAL REPORT 9 I am pleased to report that in 2013, we raised $6,794,685 in new Report of the private support for MBL’s research and education programs. Development This is a strong accomplishment, given that 2013 was the first year following the very Committee successful Catalyst Campaign.

Funds raised in 2013 include $1.47 million in Annual and unrestricted funds that were immediately available to meet operational needs, and a $1.25 million grant from the Alfred P. Sloan Foundation in support of The Bay Paul Center’s Deep Carbon Project, which seeks to generate new insights about the abundance and diversity of subsurface marine microorganisms.

2013 proved to be a landmark year for the MBL in many ways. As the 125th anniversary of the MBL’s founding, it was a time of reflection about where we’ve been, and for looking ahead to the next generation of MBL science. It was also the start of the MBL’s affiliation with the University of Chicago, and the beginning of a new era in the lab’s history.

Thank you to all our Trustees, Overseers, scientists, alumni, friends, staff, and extraordinary donors! On behalf of the Development Committee of the Board of Trustees, I offer my gratitude and thanks. Your generosity and belief in our work is an inspiration!

Thank you.

— Jeffrey Pierce, Chair

10 MBL ANNUAL REPORT 2013 Major Gifts THE G. UNGER VETLESEN $10,000-$49,999 FOUNDATION Anonymous (2) The MBL gratefully acknowledges the $350,000 for support of the Bay Paul The Agouron Institute generous contributions and pledges of the Center and the Marine Resources The American Society for Cell Biology individuals, foundations, and companies American Society for Reproductive Center. Gifts who supported our programs and Medicine operations in 2013. Margaret C. and Francis P. Bowles SIMONS FOUNDATION Worthington and Dorothy Campbell $250,000 in support of the Microbial $1,000,000-$2,000,000 Carolyn Cohen Diversity course. The Company of Biologists Limited ALFRED P. SLOAN FOUNDATION Sally Cross $1,256,000 grant for support of the Deep GEORGE AND KATHY PUTNAM Saint Michael and All Angels Carbon Project; Mitchell Sogin, Principal $129,788 in support of the Strategic Episcopal Church Investigator. Planning Fund and the Annual Fund. Fund for Science Howard Hughes Medical Institute $100,000-$999,999 Barbara Woll Jones MILLICENT BELL Ethan and Lisa Lerner BURROUGHS WELLCOME FUND $125,000 in support of the Strategic Marianne F. Milkman $820,000 in support of the Frontiers in Planning Fund and the Annual Fund. Gordon and Betty Moore Foundation Reproduction and Embryology courses Richard S. and Susan G. Morse and the Physiology Alumni Fund. DAVID AND SUSAN HIBBITT The Morton Cure Paralysis Fund $102,348 in support of the Strategic Nikon Instruments, Inc. Saul Pannell and Sally Currier CHARLES M. AND PHYLLIS ROSENTHAL Planning Fund and the Annual Fund. William Townsend Porter Foundation $550,000 in support of the Ecosystems Society for Developmental Biology, Inc. Recruitment Fund for Postdoctoral Ivan and Virginia Valiela Scientists and the Annual Fund. Restricted Gifts Waksman Foundation for Microbiology $50,000-$99,999 Wellcome Trust THE BAY AND PAUL FOUNDATIONS James F. Case $500,000 for support of the Bay Paul Patrick Gage and Irina Wilson $5,000-$9,999 Donald R. Aukamp Center for Molecular Evolution’s Jeffrey Gardner William and Susan Cobb expanding investigation of human Shinya and Sylvia Inoué Kurt J. and Rhoda S. Isselbacher Peter and Ginny Foreman microbial biome. Joan C. Miller Tom and Maryann Jalkut Donald and Patricia Mintmire George Frederick Jewett Foundation East Joan V. and Gerald S. Ruderman THE GRASS FOUNDATION Jeffrey and Margaret Pierce Edward D. and Nancy M. Salmon $450,000 in support of the Neural Systems Promega Corporation Walter J. and Marjorie B. Salmon and Behavior and Neurobiology courses. George and Kathy Putnam Woods Hole Foundation, Inc. Patricia and Charles Robertson ESTATE OF LAURA HUNTER COLWIN Ann E. Stuart and John W. Moore $401,549 for discretionary use.

2013 MBL ANNUAL REPORT 11 $1,000-$4,999 Other Gifts Jason Hoobler Cynthia L. Perreault-Micale Gifts in Kind American Society of Naturalists Jennifer M. Huettner Bertha Person Porter W. Anderson, Jr. Anonymous (1) Patricia A. Hunt Niles and Mary Peterson Paul Besse Gloria S. Borgese Robert S. Adelstein Hunter College Leslie and Donna Pinsof Brenda J. Boleyn Paul N. Chervin American Society for Microbiology David Isenberg and Paula Blumenthal Harmony Glazebrook Pittet Fine Science Tools Bradley and Katie Cohen Tammy S. and Douglas Amon Kenneth Jackson James D. Potter Barbara Woll Jones Marion Cohen Clay M. Armstrong and Joseph Johnston Robert A. Prendergast Kerry Walton Organization for Computational Clara Franzini-Armstrong Freda Kaminer Dixon and Rita Purcell World Precision Instruments Neurosciences Samuel C. and Elizabeth F. Armstrong Frank S. and Nancy Karger Agnes Jancso Radek Carl Zeiss Microscopy, LLC The Cooperative Bank of Cape Cod Carol Arnosti and Andreas P. Teske Richard F. Karger Jack A. Rall Clive R. Bagshaw In 2013, the MBL continued to benefit David and Karen Davis Doreen R. Kelly Shirley Raps from major multi-year gifts pledged in Stephen and Lois Eisen Brenda J. Bodian John Kendrick-Jones Alfred G. and Sarah C. Redfield Giovanna Breu previous years from the following: Christopher Foreman Thomas C. and Mary Ann Kenny Thomas and Sally Reeve Anonymous (1) Prosser Gifford Scott G. and Penelope P. Chrysler Marc W. and Phyllis Kirschner Joe M. Regenstein David Clarke Burroughs Wellcome Fund Mary L. Goldman and Debra Weiner Kathiann Kowalski William S. and Catherine A. Reznikoff The Ellison Medical Foundation Robert D. and Anne E. Goldman Sarah Cohen George M. and Sylvia T. Langford Fred and Gloria Rhoades Seymour S. Cohen The Grass Foundation Piroska Hardwicke Leonard and Ruth Ann Laster Rex R. Robison Howard Hughes Medical Institute J. Woodland Hastings Edward and Carolyn Conrad William J. Lehman Michael G. Ryan and Linda A. Joyce Arthur Roger Cooke Gordon and Betty Moore Foundation Carmela J. Huettner Sherwin Lehrer Joseph W. and Jean M. Sanger George and Kathy Putnam Hugh E. and Frances M. Huxley Jon C. Cooper Leslie Ann Leinwand Susan E. Seaman Roger W. Craig Alfred P. Sloan Foundation Hyockman Kwon Jack and Francine Levin Timothy and Kathleen Short The G. Unger Vetlesen Foundation Hans Laufer John and Gwen Daniels Birgit R. and Werner R. Loewenstein Rolly L. Simpson Paul B. Lazarow Elizabeth M. Davis Thomas E. Lovejoy Klara Sirokman Lhing-Yew Li Carolyn and Anatol Eberhard James and Christine Lynch Avril V. Somlyo Anna Lysakowski Sharyn A. Endow John A. Mankovich and Ellen Guthrie Walter F. Stafford, III Eve Marder Andrew and Amy Feinberg Mary Bates McIntyre Alan B. and Sala Steinbach William N. and Marianne W. Mebane Sarah Q. Foster Anne Messer StemCell Technologies Inc. John Mills Anne Freeman Steven Miano Ete Z. Szuts and Susan Oleszko-Szuts Pentair Aquatic Eco-Systems Christopher J. Freeman Mid-American Water of Wauconda, Edwin and Heather Taylor Sidney K. Pierce, Jr. Friends of The Falmouth Public Inc. Saul Teichberg Thomas D. and Patricia S. Pollard Library Charles and Judith Miller Daniel C. and Patricia A. Thums Rhys E. Probyn Chandler M. Fulton and Elaine Lai David A. and Virginia I. Miller Margaret A. Titus The Rockefeller University Press James N. and Nancy Galloway Russell Miller David Trentham Robert D. Schatz Ann M. Gardener Ronald A. Milligan Volker and Mary Ulbrich Sara Schupf Sallyann Garner Milwaukee Valve Kensal and Barbara van Holde Alexandra E. Shea Mark Gasarch James Morrell Peter and Marian Vibert Society of General Physiologists Yale Goldman Pamela Nelson and Leonard and Eve Warren Bruce and Mary Goodman Raymond E. Stephens Christopher Olmsted Cory A. Waters Andrew and Ursula Szent-Györgyi Laura Grosch and Herb Jackson Kenneth W. and Ann W. Nickerson Edward and Dia Weil Ronald D. Vale and Karen Dell Stephen and Dayna Harris Amy Nobu Michael Wheeler Washington & Jefferson College Peter Hartline and Rebecca Kucera Alexander Notopoulos and Clare M. Wilber Robert and Patricia Waterston James and Martha Hartmann Alexis Anderson John and Patty Wineman Alfred M. and Naida D. Weisberg Ellen Hertzmark Joy Olotu Ann S. Wolff John W. Weisel Sarah E. Hitchcock-DeGregori Elizabeth O’Neall-Hennessey Ralph G. Yount John and Olivann Hobbie Michelle Peckham and Richard Holt Helena Aegerter

12 MBL ANNUAL REPORT 2013 Gifts to Named Fellowships, Diane and Norman Bernstein The Fischbach Research Awards Shinya and Sylvia Inoué Fund for S. O. Mast Endowed Scholarship Scholarships, and Other Endowment Fund for the Marine Simons Foundation Cellular Dynamics Imaging Fred and Gloria Rhoades Resources Center Lawrence B. Cohen Funds Joshua and Lisa Bernstein Frontiers in Reproduction Scholarship Dan Kiehart and Patricia Lee Saylor Microbial Diversity Course Fellowships, scholarships and Norman and Diane Bernstein Fund Edward D. and Nancy M. Salmon Endowment other funds enable the MBL to American Society for Reproductive Caroline S. Harwood and Thomas A. and Gloria S. Borgese Rare stay competitive by attracting and Medicine Irvin Isenberg Lecture E. P. Greenberg Book and Journal Fund supporting top researchers and Burroughs Wellcome Fund David Isenberg and Paula Gloria S. Borgese Roger D. Milkman Endowed Lecture students. We gratefully acknowledge Patricia A. Hunt Blumenthal Rolly L. Simpson Freda Kaminer in Microbial Diversity donors to the following funds: Patricia A. Case Endowed Scholarship Marianne F. Milkman AFAR Aging Research Program James F. Case General Education Endowment Holger and Friederun Jannasch James A. and Faith Miller Research Fellowship Anonymous (1) Scholarship in Microbial Diversity William D. Cohen Endowed Fund for Awards American Federation for Aging Ann Sutherland and Raymond Keller Carol Arnosti and Andreas P. Teske Undergraduate Education David A. and Virginia I. Miller Research Everett and Anne Bergman Albert and Ellen Grass Faculty Grant Benjamin Kaminer Scholarship Mountain Memorial Fund American Society for Cell Biology Marion Cohen Program Freda Kaminer Brenda J. Bodian Summer Research Awards Sarah Cohen The Grass Foundation Bertha Person The American Society for Cell Cornell Research Development Award Neural Systems and Behavior Biology Daniel S. and Edith T. Grosch Masakazu Konishi Endowed Molly N. Cornell Scholarship Fund Lectureship in Neural Systems and Course Scholarship Fund James and Cynthia Angstadt Associates Scholarship Institutional Laura Grosch and Herb Jackson Behavior Janis C. Weeks and Fund Norman and Sally Cross Scientific Marc F. Schmidt Aquaculture Endowment William M. Roberts Tammy S. and Douglas Amon Aline D. Gross Scholarship Sally Cross Lucy B. Lemann Institutional Fund Samuel C. and Elizabeth F. Armstrong Freda Kaminer Alfred M. and Naida D. Weisberg Mary L. Goldman and Debra Weiner Nikon Research Awards Friends of The Falmouth Public Bernard Davis Scholarship Fund Nikon Instruments, Inc. Library Porter W. Anderson, Jr. Hartline MacNichol Reasearch Edwin S. Linton Scholarship Fund John and Olivann Hobbie Elizabeth M. Davis Awards Washington & Jefferson College Physiology Course Endowment Hunter College Peter Hartline and Rebecca Kucera Thomas D. and Patricia S. Pollard Barbara Woll Jones Eugene Floyd DuBois Memorial Fund Logan Science Journalism Program Thomas C. and Mary Ann Kenny Harmony Glazebrook Pittet Tay Hayashi Lectureship in Cell Giovanna Breu George and Kathy Putnam Biodiversity Informatics Fund Leonard and Ruth Ann Laster Physiology Howard Hughes Medical Institute George and Kathy Putnam James and Christine Lynch Embryology Fund Freda Kaminer Kathiann Kowalski Joan C. Miller The Company of Biologists Limited Waksman Foundation for Hibbitt Endowed Education Fund Abigail Salyers Endowed Russell Miller Society for Developmental Biology, Inc. Microbiology David and Susan Hibbitt Scholarship in Microbial Diversity Niles and Mary Peterson Herman Epstein Endowed Memorial American Society for Microbiology Shirley Raps Marine Biological Laboratory Lectureship The John E. Hobbie Fund for Science General Endowment Edward and Carolyn Conrad Susan E. Seaman Freda Kaminer Frank C. and Sheila G. Carotenuto Mary Bates McIntyre Ann M. Gardener Daniel C. and Patricia A. Thums George D. and Bernice Pappas David Clarke Jeffrey Gardner Clare M. Wilber Bertha Person Thomas E. Lovejoy Joe Martinez, Jr. and James Townsel Lhing-Yew Li Michael G. Ryan and Linda A. Joyce Endowed Lectureship (SPINES) Frederik B. and Betsy G. Bang John A. Mankovich and Ellen Guthrie Falmouth Forum Endowment Fund Anne Etgen Research Awards Charles and Judith Miller Anonymous (1) Robert Huettner Rare Book Endowed Paul A. Gray Jack and Francine Levin William S. and Catherine A. Reznikoff Fund Joe L. Martinez and Kimberly Falmouth Forum Fund Carmela J. Huettner Smith-Martinez The Cooperative Bank of Cape Cod Jennifer M. Huettner Prosser Gifford Doreen R. Kelly

2013 MBL ANNUAL REPORT 13 Sheldon J. Segal Friday Evening Semester in Environmental Science Leslie Ann Leinwand John J. Valois Marine Resources Gifts to the MBL Lecture Fund General Endowment Birgit R. and Werner R. Loewenstein Center Endowment Stanley Mierzwa Margaret C. and Francis P. Bowles Eve Marder Anonymous (2) Annual Fund Harriet F. Segal The Clowes Fund, Inc. Anne Messer Duncan P. and Dorothy D. Aspinwall Through their annual unrestricted Laura Segal and Matthew Petrie Ronald A. Milligan Freda Kaminer gifts, annual fund donors enable Society of General Physiologist Kenneth W. and Ann W. Nickerson Jahn Pothier Semester in Environmental Scholarships the MBL to cover costs not met by Elizabeth O’Neall-Hennessey grants or fees, seed key strategic Science Fund Society of General Physiologists Michelle Peckham and Helena Aegerter Stanley W. Watson Education Fund initiatives, and provide important Bradley and Katie Cohen Cynthia L. Perreault-Micale Anonymous John and Gwen Daniels Burr and Susie Steinbach Research support to its scientists and Awards James D. Potter Whitman Center General Endowment Stephen and Lois Eisen Robert A. Prendergast programs. Andrew and Amy Feinberg Volker and Mary Ulbrich Richard L. and Alice M. Chappell Dixon and Rita Purcell Robert D. and Anne E. Goldman Christopher Foreman The Chairman’s Circle Jane Lazarow Stetten Scholarship Agnes Jancso Radek Hans Laufer Peter and Ginny Foreman Fund Jack A. Rall Bryan D. and Elaine K. Noe $25,000 or more Sarah Q. Foster Paul B. Lazarow Alfred G. and Sarah C. Redfield Anonymous (1) Anne Freeman Joshua J. Zimmerberg and Joe M. Regenstein Millicent Bell Ann E. Stuart and John W. Moore Teresa L. Z. Jones Christopher J. Freeman Edward D. and Nancy M. Salmon Fund Patrick Gage and Irina Wilson Sallyann Garner Joseph W. and Jean M. Sanger Walter L. Wilson Endowed Ann E. Stuart and John W. Moore David and Susan Hibbitt Mark Gasarch Klara Sirokman Scholarship Fund Bruce and Mary Goodman Ethan and Lisa Lerner Andrew Szent-Györgyi Endowed Avril V. Somlyo Paul N. Chervin Stephen and Dayna Harris Edward and Linda Owens Lecture in Physiology Walter F. Stafford, III James and Martha Hartmann Young Scholars Program Saul Pannell and Sally Currier Robert S. Adelstein Alan B. and Sala Steinbach Ellen Hertzmark David and Karen Davis Jeffrey and Margaret Pierce Clay M. Armstrong and Raymond E. Stephens Kenneth Jackson George and Kathy Putnam Clara Franzini-Armstrong Andrew and Ursula Szent-Györgyi Frank S. and Nancy Karger Patricia and Charles Robertson Clive R. Bagshaw Ete Z. Szuts and Susan Oleszko-Szuts Richard F. Karger Carolyn Cohen Edwin and Heather Taylor Charles M. and Phyllis Rosenthal Steven Miano Seymour S. Cohen Saul Teichberg John and Valerie Rowe Mid-American Water of Wauconda, Arthur Roger Cooke Margaret A. Titus Inc. Jon C. Cooper David Trentham The Director’s Circle John Mills Roger W. Craig Kensal and Barbara van Holde $10,000 - $24,999 Milwaukee Valve Carolyn and Anatol Eberhard Peter and Marian Vibert Donald and Patricia Mintmire Anonymous (3) Sharyn A. Endow Cory A. Waters James Morrell Alison and Robert Ament Chandler M. Fulton and Elaine Lai Robert and Patricia Waterston Amy Nobu Porter W. Anderson, Jr. Yale Goldman John W. Weisel Alexander Notopoulos and Norman and Diane Bernstein Piroska Hardwicke Ralph G. Yount Alexis Anderson Marianne and Eric Billings J. Woodland Hastings Leslie and Donna Pinsof Eva Szent-Györgyi Scholarship Fund Martha W. and William C. Cox Sarah E. Hitchcock-DeGregori Thomas and Sally Reeve Freda Kaminer John E. and Judith F. Dowling Hugh E. and Frances M. Huxley Robert D. Schatz Andrew and Ursula Szent-Györgyi Shinya and Sylvia Inoué Gerald D. and Ruth L. Fischbach Sara Schupf John Kendrick-Jones Craig and Nancy Gibson Alexandra E. Shea William Townsend Porter Scholarship Marc W. and Phyllis Kirschner William I. and Lauren Huyett Timothy and Kathleen Short William Townsend Porter Foundation Hyockman Kwon Richard S. and Susan G. Morse Edward and Dia Weil George M. and Sylvia T. Langford Robert A. Prendergast Michael Wheeler J.P. and Madeleine Trinkaus Endowed William J. Lehman Elise and Stanley Rose John and Patty Wineman Scholarship Sherwin Lehrer Ann S. Wolff Ann Sutherland and Raymond Keller Vincent J. Ryan and Carla E. Meyer Walter J. and Marjorie B. Salmon

14 MBL ANNUAL REPORT 2013 James and Mary Sharp Kenneth H. Foreman and The Whitman Society Alfred L. and Joan H. Goldberg Bertha Person William T. Speck and Evelyn Lipper Anne E. Giblin Timothy H. and Mary Helen M. Frank Press Jan and Marica Vilcek Charles M. Ganson LIFE MEMBERS Goldsmith Ronald J. Przybylski Dyann and Peter Wirth Philip Grant Frank M. and Julie S. Child Susan P. and Thomas Goux Michael and Diane Rabinowitz Mary Greer Margaret Clowes Katherine Graubard and Edward Rastetter and Karen The President’s Club Caroline Harwood and J. Woodland Hastings William H. Calvin Hendrickson $5,000 - $9,999 E. Peter Greenberg William Jeffery and Meredith William H. and Margaret J. Greer Peter W. and Elizabeth Renaghan Anonymous (1) Pamela C. and Gregory J. Hinkle Yeager-Jeffery GreyBird Ventures Lawrence C. and Victoria H. Rome Hans Laufer Linda L. and Stephen A. Greyser Donald R. Aukamp Gary and Susan Jacobson Jack and Rosalyn Rosenbluth Susan Harding Hawkins Edward D. and Nancy M. Salmon Joshua and Lisa Bernstein Barbara Woll Jones $1,000 - $2,499 Robert and Margot Haselkorn Audrey M. Schneiderman George H. Billings Saul and Gitta Kurlat Louise Adler Diane E. Heck Harriet F. Segal Francis P. and Margaret C. Bowles Deena and Alan Laties Garland E. Allen John G. Hildebrand and Gail D. Burd Shawmut Design and Construction Ed and Amy Brakeman Anna L. and Tom Lawson James L. and Helene M. Anderson John and Olivann Hobbie Osamu and Akemi Shimomura Mary and Jonathan Conrad Jennifer Letitia Peter B. and Margaret T. Armstrong Joseph F. Hoffman and Corinne Steel Michael Fenlon and Linda Sallop Joseph Levine and Samuel C. and Elizabeth F. Armstrong Elena Citkowitz Marjorie E. Steinberg Pamela and David Follett Steven A. Cadwell Elizabeth Heald Arthur Carmela J. Huettner Raymond E. Stephens Sibyl R. Golden William W. and Noelle Locke Robert W. and Jean Ashton Shinya and Sylvia Inoué Edwin and Heather Taylor Robert and Trudy Gottesman George and Harmon Logan David Baltimore and Alice S. Huang Kurt J. and Rhoda S. Isselbacher Edward and Louise Tsoi Thomas F. and Virginia C. Gregg Nawrie Meigs-Brown and Patricia Ann Barlow Freda Kaminer Leonard and Eve Warren Laurie J. Landeau and David Brown Bruce A. Beal and A. Sidney and Alice Knowles Alfred M. and Joyce V. Zeien Robert J. Maze Alice Netter Francis Cunningham Hans and Donna Kornberg H.F. and Marguerite Lenfest R. Dana Ono and Anne Wagner Jonathan and Elizabeth Billings Stephen M. and Cynthia R. Krane Monique Liuzzi Sautter Arthur B. Pardee and Max M. Burger Edward A. and Kathryn F. Kravitz Walter E. and Shirley A. Massey Ann B. Goodman Colleen M. Cavanaugh and Ezra and Aimlee D. Laderman Jean H. Pierce Gregory Weyerhaeuser Piasecki Philip M. Gschwend Leonard and Ruth Ann Laster Richard L. and Alice M. Chappell Burton J. and Ann K. Lee Joan V. and Gerald S. Ruderman Robert W. and Carroll C. D. Pierce James M. and Ruth C. Clark Rodolfo R. and Gillian K. Llinás Robert Samuelson and John S. Price Alexander W. Clowes and Werner R. and Birgit R. Loewenstein Kristin Campbell John and Joan B. Ripple Susan E. Detweiler Laszlo Lorand Bonnie W. and Stephen A. Simon Harris and Jeanne Ripps Mary Clutter Michele Lorand and Paul Ringel Carolyn and John Stimpson Cecily Cannan Selby Beth Colt and P.K. Simonds William K. and Winnie C. Mackey John J. and Frederica W. Valois Dusan Stefoski and Craig Savage Molly N. Cornell Jerry and Lalise Melillo Earl H. Weidner Ann E. Stuart and John W. Moore Thomas S. and Geraldine Crane Matthew Meselson and Gerald and Ann Weissmann Gerard L. and Mary Swope Stephen and Beth Crocker Jeanne Guillemin-Meselson John F. Swope Eric H. Davidson Jason and Jennifer Meyers The Lillie Society Annette Williamson and Paul J. and Maria S. De Weer Gerrish H. Milliken $2,500 - $4,999 Michael McColm Leyla deToledo-Morrell Timothy Mitchison and David and Sandra Bakalar Kathrin R. Winkler and Barbara E. Ehrlich Christine Marie Field John and Lisa Batter Angus Campbell Herman N. Eisen and Merle Mizell George and Harriet Berkowitz Nancy Wolf and Aric Greenfield Natalie Aronson James F. and Lisa M. Mooney Eloise E. Clark Barbara Wu and Eric Larson Martha S. Ferguson Hiroko Naka Bernice Cramer and Paul Friedman Ruth E. Fye Peter A. Nickerson Sally Cross David C. and Patricia S. Gadsby O’Herron Family Foundation James N. and Nancy Galloway James L. Olds and Robin E. Buckley Richard B. and Marcia H. DeWolfe Sallie A. Giffen Thoru and Judith Pederson Joseph and Sarah Dowling

2013 MBL ANNUAL REPORT 15 $500 - $999 Ted Begenisich and Sherrill Spires Arthur B. and Roberdeau C. DuBois Stephen C. Harrison Stefan J. Medwasowski Blaise Aguera Arcas and Marlene and George Belfort Quan-Yang Duh and Ann Comer Jonathan and Dorothy Hasson Wilhelm and Nonie Merck Adrienne L. Fairhall Thomas L. and Mary Jo Benjamin Laurel A. Eckhardt J. Woodland Hastings Melanie and Klein Merriman Kerry S. Bloom and Elaine Yeh Michael V. L. and R. Suzanne Bennett John E. and Carol Edwards Marian B. and Edmund Hazzard Nancy S. and Richard Milburn Darryl A. and Janet Buckingham Francisco Bezanilla Frank Egloff, II Edward S. and Mary G. Heard Ricardo and Mela Miledi Harry F. Conner and Stanley J. Bielen Paul Elias and Marie Lossky Edward E. Heller David A. and Virginia I. Miller Carol E. Scott-Conner Stephen H. and C. Lynn Bishop Gordon and Joyce M. Ellis Robert and Taska Hener Ronald B. Moir and Toni Czekanski Susan Dillmann Thomas P. and Jane W. Bleck Doris W. Epstein Peter K. and Margaret H. Hepler Robert and Maura Morey Carol Bernstein Eckstein Dieter and Gigi Blennemann Thomas E. Evans Howard H. Hiatt Jennifer R. Morgan and Ian G. Davison David and Lois Epel David A. and Marla Bodznick Robert R. and Ann Everett Catherine E. Hibbitt and Tim Rockwell Isabelle Mort David and Doris L. Fausch Thomas C. and Elizabeth D. Bolton Donald and Jo Faber Richard W. and Susan D. Hill Day O. and Kathie C. Mount Joseph G. Gall and Diane M. Dwyer Barbara C. and John F. Boyer Patricia M. Failla Edward G. Horn Phil Nehro John P. and Mary E. Harrington Eleanor Bronson-Hodge Robert E. and Joanne Fallon Ronald R. Hoy and Margaret C. Nelson Ann Little Newbury Emily Ann Coplan Harrison Jennifer Byrnes Brower Barry William Festoff Nicholas A. and Laura M. Ingoglia Peter M. and Virginia Nicholas Buzz and Janis Harvey Donald D. and Linda W. Brown Harvey M. and Susan Fishman Laurinda A. Jaffe and Mark Terasaki Santo V. Nicosia William A. Haskins M. Kathryn S. and Thomas A. Brown Franklin W. Fort Ernest G. and Pauline B. Jaworski Lee A. Niswander Gary G. and Nancy Hayward Melinda Browne Elizabeth Frank Yishi Jin Michael and Carol Oberdorfer Robert E. and Marjorie J. Mainer Ann C. Bucklin and Peter Wiebe Alfred Franzblau Elizabeth Ann Jonas and Thomas D. Eisen Shinpei and Catherine B. Ohki Richard Mains Theo C. Bufferd Arthur and Linda Freeman Robert K. Josephson Janice and Albert Olszowka Craig C. and Darlene M. Malbon Michael and Susan Burke Paul J. and Patricia S. Freyheit Michael G. Kauffman Jeffery and Valerie Oppenheim Edwin McCleskey Harold L. and Joan N. Burstyn Robert A. and Jessica R. Frosch Richard E. and Cynthia Kendall George O’Toole and Deborah A. Hogan Dennis E. Meiss Captain’s Fund Paul A. Fuchs Thomas C. and Mary Ann Kenny Lorrence J. and Beverly C. Otter Abigail Norman Margaret W. and Samuel B. Carr John J. Funkhouser Louis M. and Brenda Kerr David and Lois Parker, Jr. Rudolf and Nannette D. Oldenbourg Donald L.D. Caspar Elinor R. Gabriel Betsy King Robert H. and Pamela D. Pelletreau Keith Purpura Connie Cepko Harold and Ruth S. Gainer John M. and Louise Kingsbury Nancy Pendleton William N. Ross and Nechama Lasser-Ross Albert Chang David Gallagher Peter N. Kivy and Joan Pearlman Matthew A. Person and Jill Erickson Edward S. and Susie W. Rowland John and Sandra Checklick Frank P. and Joyce Gallagher Paul W. and Virginia Knaplund Carey R. and Ryan Petrie Howard K. and Ethel L. Schachman Sallie Chisholm Stephen E. Gellis Susan Kohn Flynn Picardal Laura and Jamey Shachoy Ellen S. and George B. Citron Thomas Gewecke John and Susan Krezoski Helen and David Piwnica-Worms Harriet S. and Howard E. Shapiro Ann P. and James M. Cleary Barbara B. Glade Andrew M. Kropinski and George H. and Louann S. Plough John and Candace Simourian Laurence P. and Katharine B. Cloud Amy S. Gladfelter Peggy A. Pritchard Jeanne S. Poindexter Roger D. and Carol J. Sloboda Carolyn Cohen David L. Glanzman George M. and Sylvia T. Langford Jessica K. Polka John and Evelyn Steele John J. and Joan M. Collier Bert Gold Jeffry B. Lansman Harvey B. and Bette S. Pollard Edward and Anne Stimpson Marc D. Coltrera and Paul K. Goldsmith Louis S. and Camilla B. Larrey Douglas Powers Richard and Dorothy Verney Anne L. Buchinski Charlotte O. Goodwin Matt K. Lee Donald L. Price Rita R. and Jack H. Colwell Marion K. Gordon Vanessa LeFevre Kathleen M. Regan and Jeffrey Leshin $100 - $499 Nathaniel S. and Catherine E. Coolidge Judith P. and J. Frederick Grassle David P. Lenzi Robert V. Rice Donald A. and Barbara R. Abt Sallie Riggs Corgan Michael J. and Rebecca H. Greenberg Jack and Francine Levin Frederick R. and Kathryn G. Rickles Douglas G. and Kathleen Alexander Patrica Cowan Frederic and Jocelyn Greenman Raymond J. Lipicky Barbara Roberts Peggy and Fred Alsup Marilyn E. and J. Sterling Crandall Huson and Edith Gregory Philip and Martha Long Philip S. and Anne Robertson Michael and Marie Angelini Charles M. Crane and Wendy Breuer Lawrence Greksa and Jill Korbin Joanna E. Lowell James Robinson Ellen P. and Richard Armstrong Karen Crawford Newton H. and Evelyn Gresser Victoria H. and Francis C. Lowell Michael Rosen Jelle Atema Joan and Jerry Cross Alexandre Grigorovitch Louise M. Luckenbill John D. and Martha A. Ross David S. and Nancy C. Babin Prince S. Crowell, III Albert and Sondra Grossman Robert P. Malchow Norman B. Rushforth John M. and Carol T. Baitsell Jiamin Cui Gregg Gundersen James Marcello Joseph W. and Jean M. Sanger Hope Baker Elizabeth M. Davis Zach W. Hall and Julia A. Giacobassi Junko Munakata Marr Jay Schippers Michael J. Baltzley Nigel W. and Leila Daw Suzanne Hammett Julian B. and Priscilla K. Marsh Thomas J. Schnitzer John E and Helen V. Barnes Joseph P. Day Tom and Lisa Hammett Lowell V. and Ann J. Martin Nan Schow Harold and Marion Barr Martha Bridge Denckla Clifford V. Harding and Richard B. McElvein Lawrence M. Schwartz and Carol Bigelow David C. Beebe Ellen P. Donaldson Mina K. Chung Robert McIntire Robert and Carole Seidler Deborah Y. Beers-Jones Dorothy L. Drummey Glenn W. Harrington Jane A. McLaughlin Deborah G. Senft

16 MBL ANNUAL REPORT 2013 Joan Sharp Charles R. and Ellen G. Wyttenbach Muriel Gould Mary Elizabeth Rice 2013 MBL 125th Anniversary Jerry W. Shay Charles Yanofsky William Grifferi Viviana I. Risca Richard and Elizabeth Shriner Isabel and Richard Yoder Lawrence I. and Esta S. Grossman Ruth L. Saz Concert Alan R. and Phyllis J. Silver Abigail Young Ronald J. Haley Mary L. Scalera The MBL wishes to thank Patrick Eric J. Simon Harold H. Zakon and M. Lynne McAnelly David S. Hays Mary E. Schiffman Gage and Irina Wilson for their Rolly L. Simpson Michael J. and Naomi Zigmond Rodney Hinkle and Kirstin A. Moritz John F. and Diane S. Schoenfelder generosity in presenting the 2013 Roxanna S. and Ronald J. Smolowitz Kenneth H. and Patricia E. Zimble Lawrence and Layna Hobbie School of the Holy Child Anniversary concert, with students Julian Sosnik Bruce and Elizabeth I. Zimmerli Ann Snow Hobbs Franklin Sher John W. and Cathleen Speer Jeffrey A. and Johanna A. Zinn Betsy and John Honey Matthew Simon and faculty of the renowned Curtis Cynthia and Peter Starosta Jennifer M. Huettner Stephanie M. Sit Institute of Music performing on Barbara A. Stay Up to $100 Carolyn L. Johnson Sven and Anna Spoerri stage as Curtis On Tour. We are also Lucy Steere Anonymous (2) Dean and Mary Johnston John J. Stegeman deeply grateful to the following Alan B. and Sala Steinbach Aggregate George and Sally Judd Walter Sullivan sponsors, hosts, friends, patrons, William K. and Jane B. Stephenson Helen Ahearn Leonard K. and Sheila F. Kaczmarek Frank and Loretta Tang and many others who made the Ann Stern Elizabeth I. Arnold Edna S. Kaneshiro Belle K. Taylor 2013 MBL Anniversary concert a Judith G. Stetson Mike and Elana Arnold Doreen R. Kelly The Auction Company, Inc. great success, raising more than Elijah W. Stommel and Jasmin Bihler Viviana M. Berthoud Patricia E. Keoughan Susan and Robert Veeder $34,000 for the MBL Annual Fund. Gary Stormo Olive C. Beverly Mark D. Kirk Irving and Vida Wagner Dorothy A. Stracher Thomas and Janet Brady Jane and Daniel MacNeil Kathleen C. Wagner Albert J. Stunkard and Margaret S. Maurin Charles and Helen Bresnahan Katherine MacQueen Mary J. Walsh PRESENTING SPONSORS Charles H. and Donna J. Sullivan Judy A. Bruguiere Patrick J. and Lorraine Mahoney Cory A. Waters Patrick Gage and William C. and Karen A. Summers Slade M. Burton Charles K. and Susanne J. Mann George Weinbaum Margaret W. Taft Catherine Emily Carr Dawn Morin Marick Phyllis R. Wendorff Irina Wilson Gage Marvin L. and Betsy C. Tanzer Chappel Hill-Chauncy Hall Joseph C. and Connie Martyna David Alan A. Winspear Martha E. Tarafa John M. and Theresa M. Clarkin William and Kathleen McGoey Ashley V. Wires CONCERT SPONSORS Peter Tassia and Maija Lutz Janet Clay Joseph and Darlene McGuire Urszula Wires Alan M. and Deena Gu Laties D. Lansing and M. Margaret Taylor Mort and Susan Cohan Gary McHatton Alexander and Judith S. Ziss Catherine D. Thaler Katherine Coleman Carol McKeon Ronald Zweig and Christina Rawley CONCERT HOSTS William R. Thomas Concord Christian Academy Cornelia Hanna McMurtrie Anonymous W. Nicholas and Joan I. Thorndike Priscilla N. and Alan Conger Samuel McMurtrie, Jr. John S. Price Mary and James Thorne Lou-Anne Conroy Margaret A. Mills-Michael PrintSynergy Solutions Emil D. Tietje, Jr. Gail Cooper Stephen A. and Marjorie H. Moore Laurence C. Torsher Margaret S. Cooper Thomas C. and Elizabeth D. Moseley Shawmut Design and Construction Noah and Janet B. Totten Mary Couch Christoph Mueller Dyann and Peter K. Wirth Barbara S. Trela Dorothy I. Crossley Mary U. Musacchia and James E. Faber Natalie Trousof Paul Cruikshank Iris Nelson-Schwartz CONCERT FRIENDS Tom Tullius Robert A. and Donna Curtis Brian and Mary Nickerson David and Sandra Bakalar Kenyon S. and Joan Tweedell Syrel Dawson Eliot Nierman Barbo’s Furniture Michael and Frances W. Tytell Frank Egloff Northeast Unlimted Tours Norman and Diane Bernstein Volker and Mary Ulbrich Lynne Eichenfield Charles and Patricia P. Parmenter Eric and Marianne Billings Richard and Susan Vallee Falmouth Newcomers Club Leonard M. Passano James M. and Ruth C. Clark Robin A. Wallace Ruth Alice Fitz Joyce S. Pendery Kerry Walton John H. Ford Ronald J. and Dawn G. Pfohl Flagship Roofing & Sheetmetal Gary Ward and Zail Berry Alvin K. Fossner William G. Phillips Co., Inc. Samuel and Anne Ward Peter Franklin Daniel A. and Linda K. Pollen Robert and Margot Haselkorn Andrew Weissmann Larry Jay Friedman Catherine Prendergast Gerard L. and Mary Swope Robert H. and Marily Werner Michael and Bridget Gabriel Anne T. Pressman Tsoi Kobus & Associates, Inc. Roland L. and Nancy Wigley Lisa Gallagher and Gregory Ruffa Julian B. Prosser Clare M. Wilber Patricia E. Garrett and Oliver H. Woshinsky Cynthia and Russell D. Rankin PATRONS OF THE CONCERT Clayton A. and Arlene Wiley C. Vernon Gaw Fred J. and Catherine E. Ravens Elizabeth W. Brine Gerrard Wilson Sarah C. Goodrich Virginia R. Reynolds Annie Chalkley

2013 MBL ANNUAL REPORT 17 Gloria Cormack Kuhns Gifts to the MBL Alumni Fund John Cunniff Christine L. Howe Matthew McFarlane and Elizabeth M. Davis Mary Ellen M. Cunningham Deborah Hursh and Mark Mortin Jennifer Nadeau James N. and Nancy Galloway MBL alumni and faculty from 37 Gregory Alan Dasch Haruhiko Itagaki Everett Mendelsohn Prosser Gifford states and two countries made Jennine Dawicki McKenna Daniel E. James Lynn Miller Jonathan D. Gitlin and gifts to the Alumni Fund to support Charles Derby Nancy A. Johnson Tetsuto Miyashita Patricia Hodgman students in the 2013 MBL Discovery Bruce A. Diner Rena T. Jones Marius R. Moran Mary L. Goldman Courses. Long Ding Caramai N. Kamei Yasuhiro Morita John and Olivann Hobbie Thelma D. and Jonathan S. Dixon Gordon I. Kaye David E. Naylor Barbara Woll Jones Anonymous (5) Thuy A. Doan Robert G. Kemp Ralph F. Nelson Monique Liuzzi Sautter Robert F. Aarstad Tabitha Doniach R. Emmet and Gail Kenney Jeanne M. Nerbonne George and Harmon Logan Benjamin S. Abrams Laneta Dorflinger Clifford H. Kern, III Jeniel E. Nett Werner R. and Birgit R. Loewenstein C. Ronald Anderson David D. Dow Lynda A. Kiefer Susan Offner Jerry and Lalise Melillo Robert Angerer David and Sherri Durica Jennifer M. King Stephen H. Ogilvy Matthew Meselson and Anthony Apicella Marcia Edwards Marc W. and Phyllis Kirschner Jennifer B. Palenchar Jeanne Guillemin-Meselson Erin and Barry Aprison Judith S. Eisen Alan Klotz Louis Pierro Robert and Maura Morey Irina Arkhipova Stanley Goodman Elfbaum Collin Knauss Sara C. Pimental Saul Pannell and Sally Currier Oge Arum Helen Mittlacher Erickson James J. Knierim Mark Plummer Arthur B. Pardee and Katherine Bachman and Ali Eroglu Paul M. and Carol L. Knopf William K. Plunkett Ann B. Goodman Robert Fleming Alan Scott Fanning Robert E. Knowlton William F. Pomputius, III John and Joan B. Ripple Robert Barker and Karl W. Flessa and Mari N. Jensen Birgit Kovacs James D. Potter John and Valerie Rowe Marla Schay Baker Kenneth Freedman and George H. Kowallis George Radcliffe Joan V. and Gerald S. Ruderman Norman Bauman Steven Weiss Keith G. Kozminski Jane E. Radix Walter J. and Marjorie B. Salmon Elizabeth Kujawinski Behn Hugo D. and Anita R. Freudenthal Susan A. Laessig Karen Russell Randall Gaius and Eleanore Shaver Edward Joseph Behrman Anne E. Fry Deborah A. Lans Jean F. and Ronald R. Regal David and Marilyn Sheprow Jerry A. Bell and Mary Ann Stepp Yoshiko Fujita Men-Jean Lee and Vincent Rehder Richard S. Taylor Gerald Bergtrom Paul E. Gallant Giuseppe Del Priore Marian C. Rice W. B. Mason Edward M. Blumenthal Maureen A. Gannon Ardean Leith Mary Esther Rice Gerald and Ann Weissmann Elizabeth A. Booth and Mark Gaynor Ellen K. LeMosy Austen F. Riggs James A. and Carol Wyban Dustin L. Dovala Susan Gerbi-McIlwain Steven William L’Hernault Viviana I. Risca Andrew W. Zimmerman and John and Renee Breznak Helen W. Gjessing Anne M. Linton Ellen Adeline Robey Susan L. Connors Andrew Brittingham Michele Gallati Glasgow Jennifer Lippincott-Schwartz Ramona M. Rodriguiz Burt V. Bronk James A. Glazier William Longabaugh Richard E. Rohr Robert H. Broyles Holly V. Goodson-Hildreth Howard W. Lowy Allen Rosenspire Amy K. Butler Gary Gorbsky Jane Lubchenco and Bruce Menge Laurens N. Ruben Otger Campas Joel S. Gordon William B. Ludington John G. Rutherford, Jr. Richard Chadwick Martin Gorovsky Gary E. Lyons Carol Ann Ryder Chenbei Chanag David M. Grossman John L. Magnani Jan S. Ryerse A. Kent Christensen Janet Haas Phillip B. and Yvonne R. Maples Scott Reid Saleska George A. Clark E. Janet Hager Deborah Margules Robert J. Saqueton Susan D. Clements-Rabatin John E. Hall Eugene M. Martin Paul R. Schloerb John R. and Annette W. Coleman William Allen Held Sandra K. Masur Loren J. Schmidt Stephen D. Collins Gordon E. Hering Allen Wray Mathies, Jr. Joseph H. Schneider Edwin L. Cooper Scott B. Herrick Mats & Associates, LLC. Andrew Schutrumpf Jeffrey T. Corwin Ralph Hillman Joan Mattson David R. Sherwood Pamela A. Coutchie Raymond W. Holton Anne E. McBride David Sibley Nessly C. and Susan W. Craig Seymour and Victoria E. Holtzman Edwin McCleskey Philip J. Skehan, Jr. Christopher S. Cronan Timothy E. Holy James Slock

18 MBL ANNUAL REPORT 2013 Samuel J. Sober Memorial and Tribute Gifts Abigail Salyers Endowed Fund in Whitman Center General Mark S. Mooseker Alan M. Sooho Microbial Diversity Endowment Fund Christine L. Howe Allan C. Spradling These donors have chosen to American Society for Microbiology, Hans Laufer in memory of: Joel P. Stafstrom support the MBL as a special way to Edward Conrad, Ann M. Gardener, Eli K. Dabora Jeffrey Pierce Robert E. Steele remember or honor a relative Jeffrey Gardner, Lhing-Yew Li, John Lionel Roseff Suzanne Hammett Joan A. and Thomas A. Steitz or friend. A. Mankovich and Ellen Guthrie, Tom and Lisa Hammett Kelly J. Suter Charles and Judith Miller and Annual and Alumni Funds Jonathan and Dorothy Hasson Associates Scholarship Fund Harold E. Sweetman William S. and Catherine A. Reznikoff John and Olivann Hobbie, Hunter Gifts made in honor of: Steven J. Zottoli Ben G. Szaro in memory of: College, Barbara Woll Jones, Joan Jason and Jennifer Meyers Saul Teichberg Abigail Salyers Maggie Carmody C. Miller, Russell Miller, Shirley Raps Holly Thompson Joan Sharp Gifts made in memory of: and Susan E. Seaman in memory of: Jeanine A. Ursitti Rare Books Fund Alice M. Chappell Miranda Van Gelder Pamela Nelson and Christopher James J. Cimino Robert B. Barlow Olmsted in memory of: Stanley J. Bielen Marion K. Gordon Susan Volman Tammy S. and Douglas Amon, Leslie B. Vosshall Leonard Nelson Michael and Carol Oberdorfer Samuel C. and Elizabeth F. Marcia Donovan Matthew K. Waldor Armstrong, Friends of The Falmouth Semester in Environmental Science Dorothy A. Stracher Lester Barth Brant G. Wang Public Library, Barbara Woll Jones, John and Gwen Daniels in honor of: Iris Nelson-Schwartz Yihong Wang Thomas C. and Mary Ann Kenny, William C. Daniels Philip B. Dunham Margery Ward Leonard and Ruth Ann Laster, James Andrew and Amy Feinberg and Leslie B. Vosshall Lucena Barth Joseph T. Warden and Christine Lynch, Niles and Mary Milwaukee Valve in honor of: Iris Nelson-Schwartz Jeffrey M. Welch Peterson, Daniel C. and Patricia Kenneth H. Foreman Frank Egloff Harold Bancroft White A. Thums and Clare M. Wilber in Frank Egloff, II Elaine K. Bernstein Eric Wieschaus and Trudi Schupbach memory of: Anne Freeman in honor of: Carol Bernstein Eckstein Pamela C. Hinkle Benjamin Douglas Williams Cynthia C. Smith Christopher J. Freeman Helene D. Winstanley Christopher J. Freeman in memory Rodney Hinkle and Jim Burton George Wittenberg and Thomas A. and Gloria S. Borgese of: Kirstin A. Moritz Slade M. Burton Alyson H. Bonavoglia Rare Book and Journal Fund Beth Chenicek John E. Hobbie Alice M. Chappell Robert G. Wolk Gloria S. Borgese in memory of: Lawrence and Layna Hobbie Jeffery and Valerie Oppenheim Lily L. Wong Thomas A. Borgese Bradley and Katie Cohen, Stephen and Lois Eisen, Bruce and Mary Susan Wray Edna Kean Shinya Inoué Robert A. Cowan Goodman, Frank S. and Nancy Steven Zeichner Elizabeth I. Arnold Patrica Cowan Richard E. Zigmond Robert Huettner Rare Book Karger, Richard F. Karger, Mid- Endowed Fund Andrew W. Zimmerman and American Water of Wauconda, Inc., Edward A. Kravitz Christopher Dillmann Carmela J. Huettner in honor of and Susan L. Connors John M. Mills, Donald and Patricia Jeffrey T. Corwin Susan Dillmann in memory of: Mintmire, Leslie and Donna Pinsof, Robert Huettner Edward and Dia Weil, John and William B. Kristan Eric C. Dillmann Patty Wineman and Ann S. Wolff in George Wittenberg and Susan Dillmann Jennifer M. Huettner in memory of: memory of: Alyson H. Bonavoglia Robert Huettner Jeanette Fossner Harold E. Foreman Sandra Magowan Alvin K. Fossner Benjamin Kaminer Scholarship Fund Sustainable Aquaculture Fund Lynne Eichenfield Bertha Person in honor of: Scott G. and Penelope P. Chrysler in Mordecai L. Gabriel Freda Kaminer Joe L. Martinez honor of: Elinor R. Gabriel Ramona M. Rodriquiz William Mebane David L. Garbers John W. Moore Carey R. and Ryan Petrie Jennifer R. Morgan and Ian G. Davison

2013 MBL ANNUAL REPORT 19 Margaret C. Gildea Frank Morrell New Century Society Paul B. Lazarow Frank Egloff, II Thomas P. and Jane W. Bleck Edward R. and Gloria A. Leadbetter The New Century Society recognizes Marian E. LeFevre and honors those who have made Richard W. Glade Eugene P. Odum Howard W. Lowy bequests and other deferred gift Barbara B. Glade Douglas G. and Gordon MacIver and arrangements that will benefit the Kathleen Alexander Eleanor MacIver Rose Grant MBL in the future. We thank all the Robert E. and Marjorie J. Mainer Philip Grant members of the New Century Society Abigail Salyers Melanie and Klein Merriman Irina Arkhipova for their generosity. Robert Hill John W. Moore and Ann E. Stuart Arthur B. Pardee and Nancy S. and Richard Milburn Arthur K. Saz Anonymous (6) Ann B. Goodman Ruth L. Saz Peter B. and Margaret T. Armstrong Richard A. Huettner Patricia Ann Barlow Jeanne S. Poindexter Jennifer M. Huettner Sheldon J. Segal Joan R. Berman Thomas D. and Patricia S. Pollard Harold and Marion Barr Robert A. Prendergast Robert Huettner Margaret C. and Francis P. Bowles Ronald Calabrese and Ronald J. Przybylski Jennifer M. Huettner Stephen Simon Christine Cozzens Michael and Diane Rabinowitz David and Susan Hibbitt Hugh E. Huxley Graciela C. Candelas Michael G. Ryan and Linda A. Joyce Stefan J. Medwasowski Adolph S. Cavallo Cecily Cannan Selby Cynthia C. Smith Enid K. Sichel Susan and Robert Veeder Paul N. Chervin Holger W. Jannasch Julie S. and Frank M. Child Philip J. Skehan, Jr. Lynda A. Kiefer Malcolm S. Steinberg Lawrence B. Cohen Roger D. and Carol J. Sloboda John L. Magnani Seymour S. Cohen Marjorie E. Steinberg Diane Jaworski Marjorie E. Steinberg John E. and Judith F. Dowling John Tochko and Ernest G. and Pauline B. Jaworski Joseph and Sarah Dowling Christina Myles-Tochko Alfred Stracher John J. and Frederica W. Valois James W. Lash Paul R. Dupee, Jr. George and Sally Judd Susan Volman Joel S. Gordon Barbara E. Ehrlich Frederick R. and David and Lois Epel Byron and Joyce Waksman Esther M. Lederberg Kathryn G. Rickles Martha S. Ferguson Earl H. Weidner Matthew Simon Gerald D. and Ruth L. Fischbach Clare M. Wilber Cornelius F. Strittmatter Yvonne R. Fuortes John S. Willis Robert H. Broyles Paul G. LeFevre Bruce and Barbara Furie Vanessa LeFevre Robert Weale Susan Gerbi-McIlwain Bequests Realized in 2013 Anthony Liuzzi Harris and Jeanne Ripps Sol H. Goodgal Worthington Campbell, Jr. Monique Liuzzi Sautter Philip Grant James F. Case Lillian Wendorff J. Woodland Hastings Monica Riley Joyce Lorand Phyllis R. Wendorff Gertrude W. Hinsch Laszlo Lorand Joseph Hoffman and Elena Citkowitz Charles G. Wilber Michele A. Lorand and Paul Ringel M Howard and Frances F. Jacobson Clare M. Wilber Daniel and Jean Johnston Luigi Mastroianni Elif Yavuz Sally S. and Ramsey E. Joslin Santo V. Nicosia Emily Ann Coplan Harrison Jane C. Kaltenbach-Townsend Darcy B. Kelley Harold M. Mills Donald J. Zinn Richard G. Kessel Margaret A. Mills-Michael Jeffrey A. and Johanna A. Zinn Andrew M. Kropinski and Alberto Monroy Peggy A. Pritchard Ronald J. and Dawn G. Pfohl Hans Laufer

20 MBL ANNUAL REPORT 2013 Corporate Lending Program Dagan Corporation Life Technologies - a Thermo Fisher QIMAGING Digitimer Ltd. Company Quest Scientific Instruments Inc. The quality and success of the MBL Drummond Scientific Co., Inc. Lonza Walkersville Inc. educational program is maintained Luigs & Neumann Rapp OptoElectronic GmbH through research equipment, Electron Microscopy Sciences Lumen Dynamics Group Research Precision Instruments Co., reagents, and computers valued at EMD Millipore / CellASIC Inc. approximately $27 million loaned by: Eppendorf North America Mathworks Roche Diagnostics Corp. MatTek Corporation AD Instruments FEI Company – TILL Photonics MBF Bioscience SAGE Publications Adams & List Associates FHC, Inc. Micro Video Instruments Inc. Scientifica Ltd. Agilent Technologies, Inc. Fine Science Tools Mightex Systems Semrock, Inc. - a unit of IDEX A-M Systems Fisher Scientific Miltenyi Biotec Inc Corporation Advanced Microscopy Group – Fluidity Software, Inc. Millipore Corporation Siskiyou Corporation a Thermo Fisher Company Molecular Devices, LLC Solamere Technology Group Inc. Andor Technology GE Healthcare Molecular Probes / Life Technologies Soma Scientific Instruments, Inc. Applied Biosystems – a Thermo GeneTools LLC - a Thermo Fisher Company Spectra Physics, A Division of Fisher Company Grass Technologies / Astro-Med, Inc. MyNeuroLab / Leica Biosystems Inc. Newport Corp. Applied Precision / GE Healthcare Growth Curves USA Spectral Applied Research Inc. Applied Scientific Instrumentation, Nanodrop / Thermo Fisher Scientific SPOT Imaging Solutions Hamamatsu Corporation Inc. NanoSight USA West Stoelting Co. / ANY-maze Hamilton Thorne, Inc. Aquaneering Narishige USA, Inc. Sutter Instrument Company HANNA Instruments ATCC National Instruments Harvard Apparatus, Inc. AutoMate Scientific, Inc. Neuralynx Inc. Technical Manufacturing Corporation HEKA Instruments, Inc. Axiom Optics Nevtek Tecniplast Histochemical Society New Era Pump Systems Inc. Ted Pella, Inc. BD Biosciences Horiba Jobin Yvon Inc. Newport Corporation The Jackson Laboratory Beckman/ Coulter, Inc. Flow Thermo Fisher Inteligent Imaging Innovations Inc. Nikon, Inc. Cytometry Thorlabs Imaging Systems INTRACEL Limited Nikon Confocal Systems Beckman / Coutler Inc. Titertek Instruments, Inc. Invitrogen Corp. / Life Technologies – Noldus Information Technology Tokai Hit Bioptechs a Thermo Fisher Company Olympus America, Inc. – Scientific Toohey Company Bio-Rad Laboratories Jackson Immunoresearch Equipment Group Tucker-Davis Technologies BioTek Instruments, Inc. Laboratories, Inc. Omega Optical, Inc. Bitplane Inc. Unisense A/S Jelight Company, Inc. BioVision Technologies Inc. Parker Hannifin Corp. Journal of Histochemistry and Boston Electronic Corporation Pathology Devices, Inc. Varian / Agilent Technologies Cytochemistry Brownlee Precision Company PC’s for Everyone Veeco Instruments GmbH Vibratome / Leica Biosystems, Inc. Bruker - NANO Keyence Corporation of America PCO-TECH Inc. (Cooke Corporation) Viewpoint Life Sciences Kramer Scientific Pearson / Brock Education Cambridge Electronic Design Ltd. Pentair Vutara Carl Zeiss Microscopy, LLC Leica Microsystems Inc. PerkinElmer Life and Analytical VWR International Cell Signaling Technology Inc. Leica Microsystems Biosystems Sciences Charles River Laboratories, Inc. Warner Instruments Division Photometrics Chroma Technology Corporation WaveMetrics, Inc. Leica Microsystems Confocal Photron USA, Inc. Coherent Wescor, Inc. – an ELITech Group Systems North America Prairie Technologies, Inc. Conoptics Inc. USA Company Leica Microsystems Nanotechnology Prior Scientific, Inc. Coriell Instute for Medical Research World Precision Instruments, Inc. Division PROMEGA CrystaLaser LI-COR Biosciences, Inc Protech International, Inc. Yokagawa Corporation of America

2013 MBL ANNUAL REPORT 21 CHAIRMAN OF THE BOARD OF TRUSTEES R. Dana Ono, VMM Capital Partners, LLC Leadership John W. Rowe, Columbia University Saul J. Pannell, Wellington Management Company, LLC Jeffrey H. Pierce, Pierce Aluminum Co. As of December 31, 2013 VICE-CHAIR OF THE BOARD OF TRUSTEES Robert A. Prendergast, Falmouth, MA William I. Huyett, McKinsey & Co., Inc. George Putnam, New Generation Advisors, LLC Patricia K. Robertson, Rockland, DE SPEAKER OF THE MBL SOCIETY Elise Rose, Falmouth, MA Colleen M. Cavanaugh, Harvard University Charles M. Rosenthal, First Manhattan Company John W. Rowe, Columbia University PRESIDENT AND DIRECTOR Walter J. Salmon, Harvard Business School Joan V. Ruderman, Marine Biological Laboratory James A. Sharp, Carl Zeiss Microscopy, LLC Dyann F. Wirth, Harvard Global Health Institute TREASURER Maureen Nunez, MBL TRUSTEES EMERITI DIRECTOR EMERITI Frederick Bay Gary G. Borisy SECRETARY Norman Bernstein John E. Burris David Fithian, The University of Chicago Margaret C. Bowles Paul R. Gross Alexander W. Clowes William T. Speck Seymour S. Cohen BOARD OF TRUSTEES Mary B. Conrad PRESIDENT EMERITUS Robert H. Ament, Esq., Ament Law Firm, LLC Eric H. Davidson John E. Dowling Millicent Bell, Boston, MA Gerald D. Fischbach Rita R. Colwell, University of Maryland Robert D. Goldman Thomas S. Crane, Esq., Mintz, Levin, Cohn, Ferris, Glovsky Mary Greer and Popeo, PC Ruth Hubbard Paul R. Dupee, Haddeo Partners, LLP Kurt J. Isselbacher Gerald R. Fink, Whitehead Institute, Cambridge, MA M Howard Jacobson L. Patrick Gage, enGage Biotech Consulting George M. Langford James N. Galloway. University of Virginia Burton J. Lee David A. Greene, The University of Chicago Irwin B. Levitan Caroline S. Harwood, University of Washington Paul A. Marks Robert E. Mainer Robert Haselkorn, The University of Chicago Jean H. Pierce H. David Hibbitt, Bristol, RI Thomas D. Pollard William I. Huyett, McKinsey & Co., Inc. Frank Press Eric D. Isaacs, Argonne National Laboratory Vincent J. Ryan Ethan A. Lerner, Massachusetts General Hospital John W. Saunders George W. Logan, Charlottesville, VA David Sheprow Walter G. Massey, School of the Art Institute of Chicago Gerald Weissmann John W. McCarter, Jr., Northfield, IL Christopher M. Weld Ambrose K. Monell, Palm Beach, FL Annette L. Williamson Richard S. Morse, Jr., Woods Hole, MA Alfred M. Zeien

22 MBL ANNUAL REPORT 2013 BOARD OF OVERSEERS Karen Tierney, Wellesley, MA Linda Deegan, Senior Scientist Donald R. Aukamp, Woods Hole, MA Jan T. Vilcek, The Vilcek Foundation, Inc. Anne Giblin, Senior Scientist Nazneen Aziz, Lexington, MA Nancy G. Wolf, University Hospitals of Cleveland Edward Rastetter, Senior Scientist David Bakalar, Chestnut Hill, MA Barbara Wu, Wilmette, IL Gaius Shaver, Senior Scientist Richard M. Barno, Weston, MA James H. Wyche, Howard University Joseph Vallino, Senior Scientist Timothy A. Barrows, Matrix Partners William Zammer, Cape Cod Restaurants Jianwu Tang, Assistant Scientist John F. Batter, WilmerHale Nathan Wilson, Director, EOL Biodiversity Informatics Stephen J. Bergman, Newton, MA ASSOCIATES EXECUTIVE BOARD Eugene Bell Center for Regenerative Biology and Tissue George Berkowitz, Legal Sea Foods Thomas F. Gregg, President Engineering Joshua B. Bernstein, Bernstein Management Corporation Huson R. Gregory, Vice President Jonathan Gitlin, Director**, Senior Scientist George H. Billings, Falmouth, MA Michael Fenlon, Secretary Joshua Hamilton, Acting Director** Marianne Billings, Potomac, MD Robert C. King, Treasurer Marko Horb, Associate Scientist, Director National Xenopus Resource Ed Brakeman, Bain Capital LLC Donald R. Aukamp Jennifer Morgan, Associate Scientist Beth Colt, Woods Hole Inn Syrel Dawson Mark Messerli, Assistant Scientist Bernice Cramer, Newton, MA Paul J. De Weer Joel Smith, Assistant Scientist Sally Cross, Falmouth, MA Joe A. DeGiorgis Michael J. Fishbein Diarmaid Douglas-Hamilton, Hamilton Thorne Inc. Cellular Dynamics Program Barbara W. Jones Joseph L. Dowling, The Rhode Island Eye Institute Joshua Hamilton, Acting Director** Hans Kornberg Pamela L. Follett, Lewis Rhodes Lab Rudolf Oldenbourg, Director**, Senior Scientist Ruth Ann Laster Jean-François Formela, Atlas Ventures Shinya Inoué, Distinguished Scientist Jeffrey O. Oppenheim Craig B. Gibson, Lawrence General Hospital Osamu Shimomura, Distinguished Scientist Marjorie B. Salmon Prosser Gifford, Woods Hole, MA Jonathan Gitlin, Senior Scientist Catherine M. Golden, New York, NY ex-officio members Mark Messerli, Assistant Scientist Gary Jacobson, Westwood Pembroke Health System Joan V. Ruderman, President and Director Michael Shribak, Associate Scientist Thomas P. Jalkut, Nutter McClennen & Fish LLP John W. Rowe, Chairman of the Board of Trustees Tomomi Tani, Associate Scientist Barbara W. Jones, Barbara Woll Jones Designer/Builder Pamela Clapp Hinkle, Director of Development and Saul Kurlat, Cambridge, MA Program in Sensory Physiology and Behavior External Relations Deena G. Laties, Philadelphia, PA Roger Hanlon, Director, Senior Scientist Anna L. Lawson, Daleville, VA SCIENTIFIC LEADERSHIP Stephen Highstein, Senior Scientist Jennifer O. Letitia, Westport, CT Alan Kuzirian, Associate Scientist Joseph S. Levine, Concord, MA The Josephine Bay Paul Center for Comparative Molecular Richard Lipkin, Commerce Health Ventures Biology and Evolution SENIOR STAFF William W. Locke, Cataumet, MA Mitchell Sogin, Director**, Senior Scientist Joan Ruderman, President and Director William K. Mackey, Mackey & Foster, PA David Mark Welch, Director**, Associate Scientist Jerry Abarbanel, Interim Director of Human Resources** Dale Mathias, Washington, DC Mark Alliegro, Senior Scientist Richard Cutler, Director Facilities, Services and Projects** Thomas McMurray, Jackson, WY Joshua W. Hamilton, Senior Scientist** Andrea Early, Director of Communications** Nawrie Meigs-Brown, Woods Hole, MA Anton Post, Senior Scientist Jonathan Gitlin, Deputy Director of Research and Programs** Edward P. Owens, Norwell, MA Linda Amaral Zettler, Associate Scientist Susan Goux, Director of Human Resources** Robert W. Pierce, Pierce Aluminum Co. Irina Arkhipova, Associate Scientist Joshua Hamilton, Chief Academic and Scientific Officer** John Ripple, Virdante Pharmaceuticals Julie Huber, Associate Scientist John Harrington, Director Facilities, Services and Projects Robert Samuelson, Monitor Group LLC Sheri Simmons, Assistant Scientist** Pamela Clapp Hinkle, Director of Development & External Relations Cecily C. Selby, New York, NY Shelly Kaplan, Interim Director of Facilities** Bonnie W. Simon, Washington, DC Ecosystems Center Maureen Nunez, Chief Financial Officer Carolyn Stimpson, Wachusett Mountain Christopher Neill, Director, Senior Scientist William Reznikoff, Director of Education Ann E. Sutherland, University of Virginia John Hobbie, Distinguished Scientist Cathy Schmidt, Interim Director of Human Resources** Gerard L. Swope, Woods Hole, MA Jerry Melillo, Distinguished Scientist Zoe Cardon, Senior Scientist John F. Swope, Sheehan, Phinney, Bass and Green **Staff who joined or left the MBL during 2013.

2013 MBL ANNUAL REPORT 23 Photos

Cover: MBL’s R/V Gemma (Daniel Cojanu)

Inside front cover: MBL Campus on Eel Pond, Woods Hole, MA (Tom Kleindinst)

P1: MBL President and Director Joan Ruderman (Bachrach Studios)

P2: clockwise: The University of Chicago and the Marine Biological Laboratory sign an agreement to affiliate (Bruce Gilbert); Massachusetts Senator Elizabeth Warren visits the MBL (Tom Kleindinst)

P3: clockwise: Arctic LTER greenhouses in peak autumn. (Sadie Iverson); The University of Chicago campus (UChicago); Flukes statue (MBL Communications Office); Rachel Carson statue (Tom Kleindinst)

P4: clockwise: Moon jellyfish (Brad Gemmell); Field experiments were conducted in a remote area near Utah’s Bear Lake where the ecosystem’s productivity is limited by both water and nitrogen availability. (Zoe Cardon); MBL’s Microbial Diversity course (Tom Kleindinst)

P5: clockwise: Heavy oiling of Bay Jimmy, Plaquemines Parish, Louisiana (Wikimedia); cuttlefish (C. Vallino); Rain over a soybean field at Tanguro Ranch in the Upper Xingu watershed, Mato Grosso, Brazil. (Christopher Neill); Polarization sensitive absorption of a Siemens star pattern etched into a thin metal film (Shalin Mehta)

P6: clockwise: Bdelloid rotiferAdineta vaga, birefringence image, polarized light microscope. (M. Shribak and I. Arkhipova); cuttlefish embryo (Nipam Patel)

P8: Collosphaera huxleyi microbe (David Patterson, ICoMM)

P 10: Jeff Pierce (Dan Cutrona)

24 MBL ANNUAL REPORT 2013