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 (34@8J35602"$* -.#'$+" ",,# -#90 :#$ -#!34<3J@602$#.#'$+" ",,# -# .#'$+" ", -# .- $( 90      ( 34@8J (34@80      - + -#/J&)#$34562,.#:#$ -#!34<390 : -/ # .#'J:"  #$B#*"( " # #*$"2 4 /: & 34A39  #+&)-'",  '#."  EJ '#."  D# $ - $##'# #-"+" ## &)"# 0     "-"!+#VJ ' ." #  ! . *20.# &#(*&#-9 -#*0 $0:# - >50343A290      &)#$ 3456J #-"+" ##J $' .   ."  7  &+$#  !  45>   3V  24.05.1955; stream at 1150 m, on the path from Kalavrita to Khelmos Mt., 27.05.1955, 6♀; :#$ -#!34<3J Central Greece: Fthiotida, Spercheios, Rachi Tymphrestou, 1200 m, 9.4.1967, 1150 m, 9.4.1967, 1♀; Spercheios at Ag. Georgios, bridge of the route Karpenisi-Lamia, 10.4.1967, 1♂, 1♀, 4♂ ex., 4♀ ex., 1 larva; Asopos $#-0" Spercheios, route Lamia -Athens, 100 m, 10. 4.1967, 3 ex. ♂, 2 ex. ♀; Evritania, Acheloos, Rachi Tymphrestou and Karpenisi, 1000 m, 9.4.1967, 7♂ ex., 6♀ ex.; Euboea: Dirphys, Steni stream, 550 m, 1–2 km upstream of Steni, 35080346< 3VE '/34 >>N <5>M800 m, 27.10.1972, 1♀; Central Greece: Eptalofos, Parnassus Mt, 22°29'E, 38°36'N, 720 m, 1.5.1976, 1♀; 29.5.1976, 1♀; Peloponnese:

 9 a‚‚‡h‘h*%%%*'+1%2 .     About 60 miles west of Tripoli, 29.04.1971, 1♀; Euboea: N Amarynthos, 4M700 m, 8.5.1971, 1♂, 2♀; Stropones, @7 57N 7A 76N <@>M45> @7M24.05.1973, leg. Aspock, Rausch, Ressl, 1♂, 2♀; &$ !@>33 J#-"+" ##J Arcadia, Vytina, stream SW of the city, 3739.031’ N, 2210.156’ E, 960 m, 06.04.2009, 3♂, 8♀, 4 larvae; Arcadia, Magouliana, stream SE of the village, 3739.425’ N, 2208.730’ E, 1035 m, 06.04.2009, 4♂, 7♀, 4 larvae; Arcadia, #$ B * #"."$ $#  ./ $+$ * 7<@30@64R @35A0 >50>80@>>4 4W 4V 5 larvae; Messenia, Agioi Apostoloi, stream E of the village, 3704.158’ N, 2147.275’ E, 415 m, 05.04.2009, 1♀; ,&$#$3A "+#',#.-"' -# ..  ,$"##-"+" ##0

                                                              O"  P$    % &     $       '    &                        (       '    &   )         '   *  

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kontschani

             Nemoura taurica Nemoura uncinata Nemoura Nemoura marginata Nemoura Nemoura flaviscapa Nemoura Nemoura cinerea cinerea cinerea Nemoura turcica cinerea Nemoura Nemoura apollo Nemoura Amphinemura quadrangularis Amphinemura sperchiana Amphinemura aetolica Nemoura caligula Nemoura "&#! arcadia Amphinemura triangularis Amphinemura           Brachyptera phthiotica Brachyptera Brachyptera transcaucasica kykladica turcica transcaucasica Brachyptera Brachyptera helenica Brachyptera Brachyptera

Brachyptera beali cretica cretica beali Brachyptera graeca Brachyptera    "#$! beali beali Brachyptera

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   Protonemura intricata taygetiana intricata Protonemura pseudointricata intricata Protonemura Protonemura malickyi mattheyi Protonemura lagrecai Protonemura oitica Protonemura rauschi Protonemura graeca praecox Protonemura &#! aegaeica Leuctra bronislawi Leuctra candiae Leuctra cretica Leuctra fusca Leuctra            androsiana Protonemura aroania Protonemura bithynica bacuriana Protonemura autumnalis Protonemura beaumonti Protonemura cressa Protonemura intricata intricata Protonemura

 ! "#  a‚‚‡h‘h*%%%*'+1%2.    9 

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macedonica

floridus

   Capnioneura balkanica Capnioneura #"!! ventralis Besdolus Perlodes microcephalus Perlodes floridus peloponnesiacus floridus Perlodes Leuctra minoica Leuctra Leuctra marani Leuctra metsovonica Leuctra moreae Leuctra olympia Leuctra rhodoica Leuctra  !Zwicknia bifrons balcanica schilleri Capnopsis           Leuctra kykladica Leuctra major Leuctra Leuctra inermis Leuctra Leuctra hirsute Leuctra  graeca Leuctra hippopus Leuctra

 9 a‚‚‡h‘h*%%%*'+1%2 .    

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   Siphonoperla neglecta neglecta Siphonoperla Siphonoperla graeca Chloroperla tripunctata Perla pallida Perla ("#"#!Chloroperla russevi           Helenoperla malickyi Marthamea vitripennis** Perla marginata Eoperla ochracea* Eoperla Isoperla tripartita recta tripartita Isoperla #! Dinocras megacephala Isoperla grammatica Isoperla ikariae Isoperla lesbica Isoperla obliqua Isoperla tripartite tripartita Isoperla  nigrita mirabilis Bulgaroperla chius Isoperla              !  !  "  #      $

 ! "#  a‚‚‡h‘h*%%%*'+1%2.    9         ) .#$ *#",-"' ) ,$"-"- .$(#$""  #)$""/0 D"--#'" #$ *#.,$"3>"3@<> #$#  )& .  "' #0###$*# '#+#$".-" *0 "+&- " && --!'" #.",)"##$*#. .&- .-- &$#- $( #0  #!+#-"' -!2 ! . *9$#,#$"&$/ #",#"&,"$B# 0" #)"$.#$", $##'# . # #+&)-' ",  '#."  0     '"--#'#. "& ", #(*#-Q  2[ &#(*&#-9  " +$") )-! $#,#$$ *"+$## . !#$$"$!", $##'#0#$#.#'$+" ", )!&)#$234679$#,#$"  :#$ -#! 34<30# # ++-#,"$#.  ",  2- +-#/ 34>39  . "( -.$#'"$. $#/ " ,$" $##'#0

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&)#$3467J@@802.#'$+" ", -# .,# -# $#.#'$+" ", 90 :#$ -#!34<3J7602"$* -.#'$+" ", -# .,# -#90 '/@>38J3402'- $,, '" ",!+# &90       - + -#/J&)#$34562,.#:#$ -#!34<390      ( J&)#$3467 34682,.#:#$ -#!34<390 : -/ # .#'J:"  #$B#'"( " # #*$"2 4 /: & 34A39  #+&)-'",  '#."  EJ+$&  '#."  D# $ - $##'# #-"+" ## &)"# 0    "-"!+#WJ 35>> -#*0C$0:"$' 3AM@50>5034562:90     &)#$3456J#-"+" ##J$#  3@>> "*  !" !*#" @>0>503455  2♀; Epirus: 2 streams on road to Metsovon, between 1600M1700 m, 09.06.1955, 3♀; &)#$3467JD# $ - $##'#J Oiti Mt., 1500 m, 18–25.6.1956, 2♀, Prioni, Olympus, 1000 m, 03–13.06.1956; Peloponnese: Voïdia, 1500—1927 m, 2♀, 23.05.1929, leg. M. Beier (repeated in Aubert 1964); Taygetos, 20.5.1955, 2♀, leg. J. Aubert; Epirus: Col .# #"("  >40>603455  7V  -#*0 =0 &)#$E :#$ -#! 34<3J D# $ - $##'#J C". +#$'#"  *0 Georgios, bridge of the route Karpenisi-Lamia, 10.04.1967, 1♀; Evrytania, Acheloos, Rachi Tymphrestou and  $+#  3>>> >40>80346< 7W#10 @V#10&)"# J$+! # $#  55> 3M@/&+$# ",#   15.04.1967, 2♂, 1♀; '/ 34>>   13.05.1976, large creek and "Rieselquelle", 1♂, 2♀, 1 larva; &$ !@>33 J #-"+" ##J $## " +#',#. -"' -# ..  ,$"# !*#" .",$" $ " 0       )"  #$#  .)$""/ . )## '"--#'#.,$"--!"* "&   -"' -#0  " )& .  ." -! &$#- $( ##$#,"& ."*##$)"# #$ - .,&--! '"-"$#. .&-&**# *  $$"##$*# '#+#$".0   # .   ",     2- +-#/  34>@9   '"$$#'#. "    )! &)#$ 2346790 ##

 9 a‚‚‡h‘h*%%%*'+1%2 .     $#'"$.  "#(#$  $#,#$ "    2,.# :#$ -#! 34<39 . ,"$     " $#'"$. $# / "  ,$" $##'#0

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&)#$3456J34302"$* -.#'$+" ", -# .,# -#90 :#$ -#!34<3J7502'"+-##  $!.#'$+" ", -# .,# -# .#'$+" ",- $( 90     &)#$ 3456J:#$ -#!34<32+#-- *90  : -/  # .#'J :"    #$B#'"(  :&-* $  ""("  " # #*$"  #+&)-' ",  '#."  EJ+$&  '#."  D# $ - $##'# # -! #-"+" ##0     +$&J"-"!+#W  --"!+#V + $ !+#@W 73VJ' +&$#. $##$#  .$(&-#", #+$# .#", #'/#"(" )### 36>> .3<>> Central Greece: Oiti Mt., 1500 m, 18–25.6.1956, 2♀; Peloponnese: Stymphalia Lake, 16–19.5, 1♂; :#$ -#!34<3JC". +#$'#"  '!+$#"& 3@>> 4080346< 6 larvae; 1150 m, 9.4.1967, 2♂, 4♀, 2 larvae; Spercheios at Ag. Georgios, bridge of the route Karpenisi-Lamia, 10.4.1967, 1♂, 1 ex.♂; Evrytania, Acheloos, Rachi Tymphrestou and Karpenisi, 1000 m, 9.4.1967, 1♀ ex., 4 - $( #E '/ 34Larissa, Ossa Mt., 22 8>N  74 5>N 4>> 3<0>5034<7 -#*0+"'/  &'  #- 3W0  C"& . ".#$ #-!* -&.#.&$ *+$ * .# $-!&#$0  :$ '=""234<3)9 -"$#+"$#.#,# -#)&& '#$  !0

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:#$ -#!34<3J7A02"$* -.#'$+" ", -# .,# -#90 : -/ # .#'J-)  2&$ !@>><9EJD# $ - $##'# #-"+" ## &)"# 0     "-"!+# W  + $ !+# 4VJ D# $ - $##'#J +#$'#"  ' !+$#"&  335>   >40>80346<2DD:90      :#$ -#! 34<3J +#$'#"  ' !+$#"&  3@>>   >40>80346<  7 - $( #E Spercheios at Ag. Georgios, bridge of the route Karpenisi-Lamia, 10.04.1967, 4♂ ex., 1♀ ex., 1 larva; Evrytania, Acheloos, Rachi Tymphrestou and Karpenisi, 1000 m, 09.04.1967, 21♂ ex., 28♀ ex., 3 larvae; '/ 34Parnassos, Quelle 2232'E, 3835'N, 1300 m, 29.05.1976; Euboea: Stropones, 23°53'E, 38°36'N, 720–950 m, 23–24.05.1974, 3♀; &$ ! @>33 J#-"+" ##J$' . 

 ! "#  a‚‚‡h‘h*%%%*'+1%2.    9  Tetrazi Mts, Agia Theodora, stream and karst springs, 3721.269’ N, 2158.782’ E, 490 m, 05.04.2009, 1♂; further " "+#',#.-"' -# .$#'"$. ,$"# !*#" .$"  0    )  ", / $ +$ * . '"-. #$ $#   -&.# )###  84> " 36>> 0  "''&$ && --! -" &)#$  #(#$ )& . 0   :#$ -#!234<39$#+"$#.#- $( #& '#$  ! ." -!$#+"$#.+#'# '"--#'#. #"-"!+# + $ !+#0

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&)#$3456J@>>02"$* -.#'$+" ", -# .,# -#90 :#$ -#!34<3J8@02'"+ $"     90 '/34 @806034552 90      &)#$ 3467J D# $ - $##'#J C".  "$*"+" "  "&    A>>   @6M @40503456  3V0 ="" 34<>J #-"+" ##J #*  +-#"  @@ 3>N  7A 05'N, 1500m, 26.5.1969, 1♂; Pendayi, @@ >5N 7A 35'N, 900m, 4.6.1969, 1♂; Killini, 22 @8N 7< 58'N, 1500m, 28.5.1969, 2♀;   :#$ -#! 234<39 "#.       '"&-. )# &)+#'# ",      2 9 '/ 234

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&)#$3456J34702"$* -.#'$+" ", -# ,# -# .- $( 90 #' *#$34<5J3@702'"+-##  $!.#'$+" ", -# .,# -#90         &)#$ 3456J&)#$34560 : -/ # .#'J:&-* $ #$) EJ+$& D# $ - $##'# #-"+" ##0    "-"!+#W  --"!+#V + $ !+#AW 36VJ#-"+" ##J$' .  *"&-  ."  (#$  @705034552 90      &)#$ 3456J +$&J $#    36>>  $" . " #"("  40603455  6 - $( # 2]9E D# $ - $##'#J #*"  ",  $ "J    (#$  "$ .# ", $" . " $ (  65>  50603455  @ - $( # 2]9E Peloponnese: Kakary, Anogia, 300m, 22.5.1955, 1♂, 5♀, 1 larva; part of a channel 600m, streams at 1200m and 1500m, Anogia route to Taygetos 20–21.5.1955, 7♀, 4 larvae (?); Arcadia, Landona, Magouliana, 1100m, 23.5.1955, 9♂, 17♀, 18 larvae, Ladon and tributaries; Vytina, 950m, 24.5.1955, 6♂, 8♀, 18 larvae; Creek behind #"#- - ($ <>> @60503455 3- $( E'  $#  35>>"  !" - ($ #-" @<0503455  5♂, 6♀; &)#$ 3467J Peloponnese: Laconia, Marathea, 100 m, near Gythion, 21–22.5.1955, 1♀; ="" 34<>J D# $ - $##'#J ($!    -- /"&.  @3 5>N  7A 50'N, 1000m, 7.6.1969, 1♂; #' *#$ 34<5J #-"+" ##J$' .  C"-"  @3 88N  7< 45'N, 700m, 25.5.1974, 4♂, 14♀; Parnon Mt, 22 73N  7< 3>>  13.5.1976, 3♂, 5♀, spring 1♂, 1♀; &$ ! @>33 J #-"+" ##J $' .  !  $#  I ", # '!  3739.031’ N 2210.156’ E, 960 m, 06.04.2009, 3♂; Arcadia, Magouliana, stream SE of the village, 3739.425’ N, 2208.730’ E, 1035 m, 06.04.2009, 2♂; Arcadia, Tetrazi Mts, Agia Theodora, stream and karst springs, 3721.269’ N, 2158.782’ E, 490 m, 05.04.2009, 5♂, 4♀; Messenia, Agioi Apostoloi, stream E of the village, 3704.158’ N, @38<0@<5R 835 >50>80@>>4 

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 '/3438)9E   '#."  2-!+&9 # -! #-"+" ## &)"# 0     "-"!+# WJ  '#."  J "& .# ","&  -!+&    $  @@/ # #+"/ $  22°26'E, 40°00N', 800m, 27.10.1972 (CPZ); paratypes 1♂, 2♀: Pertuli, the main brook of the valley (CPZ); paratypes 2♂: Platamon, 23.10.1972; paratypes 14♂, 4♀: Peloponnese: Kefalarion, 700m, 18.10.1972 (CPZ); paratype ♀: Evrostini, 750 m (CPZ); paratypes 5♂, 2♀: same locality but 1200m (CPZ).       $."  '/ 3447J # -!J #-"  )"(# "$ $  74 @7N  @7 >3N  A7>  @304M13.10.1980, 1♂, 1♀; Euboea, above Stropones, 38°36'N, 23°53'E, 700m, 10M12.10.1980, 2♂, 2♀; Ano Steni, 7A 75N @7 57N 55> 3>M3@03>034A> @3W 338)90

    > 0#$-%%

I# - # $''+#'#  / " ,$" --: -/ '"& $#E  +$&   '#."   D# $ - $##'# # -! #-"+" ## &)"# 0     "#-!347@J Central Greece, Parnassos, 18.04.1966, 1♂, Kruper 1869 leg.;&)#$3456J Epirus: stream at 1600m on road to Metsovo, 07.06.1955, 1♀; &)#$3467JD# $ - $##'#J"&  35>>  3AM25.5.1956, 1♀; '/34> >50>5034<6  1♂, 2♀; Taygetos Mt, stream and springs, 3♂, 6♀; Nemea, 3♂, 3♀ (Wittmer, Mus Basel); Euboea: Monokaria, 1♂; #$ ".#C*&#$" C"'#@>> 7< 83N530>` @@ @AN3405`0  "#-!2347@9 "#. # *-# -# I,$"# $ " ++# $")# # #$ - +#'# 0:#$ -#!234<39$#+"$#.+#'# ,$"#+#$'#":   C".  $##'#& '#$  !0 '/ 2340

     . 7  5 #$%23

: -/ # .#'J:"   .#$B#*"( :&-* $ " # #*$"  #+&)-'", '#."   "  E   '#."  D# $ - $##'# # -! #-"+" ##0

 ! "#  a‚‚‡h‘h*%%%*'+1%2.    9       '/34A

     > 0#$-%%

&$"+# +#'#  #: -/ / " ,$"D$"  2"+Q 6(#'@>>4 9 :"   . #$B#*"( :&-* $ " # #*$"  #+&)-'", '#."  E +$&  '#."  #-"+" ##0     &)#$3456J '#."  J-!+&  ($"-" *" *"" !" A5> 350603455  1♂; Prionia, 1100m, 15.6.1955, 7♂, 14♀, 7 larvae; Epirus: Metsovitiko, road to Metsovon, 600m, 8.6.1955, 1 larva 2*$"&+   ?); Peloponnese: stream in 1200m, from Anogia way to Taygetos, 21.5.1955, 3♂, 10♀; stream 3 km upstream of Vytina, 1050m, 24.5.1955, 8♂, 4♀, 5 larvae; '/ 34

      5)*+#$%%9

 $." '/3447J8@<02"$* -.#'$+" ", -# .,# -#90  .#'" D!'- .#2 1"90    "-"!+#W + $ !+#@VJ 1"J )"(#"$"  7< >AR @5 7@R 66> @6M@A03>034A> 2D D90

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 &$"+#  +#'#    # : -/  / "  ,$" D$"   2"+Q 6  (#' @>>4 9  :"   . #$B#*"(  :&-* $  2:$ '  ="" 34<69  " # #*$"  #+&)-' ",  '#."   -"(#  E   '#."  D# $ - $##'# # -! #-"+" ## &)"# 0      '/34R 5>> @604034A>  2♂; Central Greece: Pelion Mt, E Chania, 39°25’N, 23°06’E, 930m, 21.9–13.10.1980, 2♀; Pelion, above Portaria, 39°23’N, 23°01’E, 830m, 13.10.1980, 1♂, 1♀; Euboea: Ano Steni, 38°35’N, 23°53’E, 550m, 10–12.10.1980, 1♂, 3V0  &)#$234569+&)-#.$#'"$.",# - $( #",N!+#    &)#$"$$ :$ '/N ,$"+$& .# -!0$") )-!)"$#,#$"#$ 0

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&)#$3456J@>802"$* -.#'$+" ", -# .,# -#90  : -/ "G++#    +#'#  / "  ,$" -)   2&$ ! @>><9   -! 2 $."  '/ 34479  #+&)-'", '#."  E +$&  '#."  #-"+" ##0

 9 a‚‚‡h‘h*%%%*'+1%2 .        "-"!+#W  --"!+#V + $ !+#@4W 3AVJ+$&J$# " $" ."#"(" 36>>M 3<>> > 7<:83N530>`  @@:@AN3405`E &$ ! @>33 J #-"+" ##J $' .  #$ B   *  #"."$  $#  . / $ +$ *  7<@30@64R @35A0 >50>80@>>4 3V0   '/234><90&)"&$#'"$.,$" -! $#) #." ,# -# 2 $." '/344790

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  $." '/3447J8@402"$* -.#'$+" ", -# .,# -#90  .#'" D$##0    "-"!+#W + $ !+#7W 37R @5 57R 8<> @50@034A@ (CPZ); paratypes 2♂, 6♀, 1 larva: E of Ag. Ioannis, 35°03’N, 25°53’E, 450m, 16.12.1981 (CPZ, CIP).  .&-"''&$.&$ *## $#'"-.# "  ( *-" *##$*# '#+#$".0

     )*+#$%,-

 '/34>3J@3@02.#'$+" ", -#90  .#'" D# $ - $##'# #-"+" ##0     Holotype ♀: Peloponnese: Evrostini, 22°24'E, 38°04'N, 750m (CPZ); paratypes 21♀: Evrostini, 22°23'E, 38°03'N, 1200m (CPZ); paratypes 3♀: Kefalarion, 22°31'E, 37°54'N, 700m, 18.10.1972, 3♀ 2D 90       $."  '/ 3447J #-"+" ##J $#  -" * $" . " #* +-#"  7A >4R  22°14’E, 130m, 22.4.–21.5.1984, 1♀; #$ " .# C*&#$"   C"'# @>>3J D# $ - $##'#J  $ "  +$ * Kastalla, Delfi, 20.5.1985, 1♂, 1♀, PN leg.; "('@>3@JD# $ - $##'#J"$#*" -&  *" #"$*" +#$'" (#$I",#(-- *# 7A 5>05P @3 56R8@0A0>50@>330  .&-#$#'"--#'#.  ! .'")#$  .'  * $ #$#1# .#.##$*# '#+#$".0

      #$%12

&)#$3456J@>@02"$* -.#'$+" ", -# ,# -# .- $( 90  : -/  # .#'J :"   . #$B#*"(  " # #*$"  #+&)-' ",  '#."  E   '#."  2-!+&9 D# $ - $##'# #-"+" ##0    "-"!+#V + $ !+#3V 3+ $ #W- $( J '#."  J-!+&  ($"-" *" $"   33>> 3505034552 90     :#$ -#!34<3JD# $ - $##'#JC".  '!+$#"& 335> 4080346<  1♀; '/34>  26.5.1974, 1♀; Platamonas, 1♀; Central Greece: Pendayi, 1000 m, 1♀; #$ " .# C*&#$"   C"'# @>>3J Peloponnese: stream near Kaloskopi, 21.5.1985, 1♀, (P. Nicolai leg.); River Ladon, Planiteron, 650–1000m, 7< 56_ @@ >4_ @0802000, 2♀, RF & JMTF leg; #$ ".#C*&#$" C"'#@>>  7<:83N530>`  @@:@AN3405`E &$ ! @>33 J #-"+" ##J$' .  *"&- $# ",#(-- *# 7<7408@5R @@>A0<7>R 3>75 >60>80@>>4 7V0

 ! "#  a‚‚‡h‘h*%%%*'+1%2.    9      5)*+#$%%9

 $." '/3447J8@502"$* -.#'$+" ", -# .,# -#90  .#'" ".#' ##0    "-"!+#W + $ !+#@W AVJ".#' ##J ".#J+ *# 76 35_ @A ><_ 6> 8M 6033034A>2D D90

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"      +  !#$%,-

 %   (  : &    /  34<5J #$ " .# C*&#$"   C"'# @>>< 2,.# &$ !@>38)90 %  ( ( : &  /34<5J&$ !@>33 2,.#&$ !@>38)90 : -/ # .#'J #+&)-'", '#."  E #-"+" ##0     #$ ".#C*&#$" C"'#@>>070@>>5 #-"+" ## $*"-. #, -"($"  stream exiting from Kefalovriso cave, 820m, 37º41΄51.0΄΄N, 22º28΄19.5΄΄E, V. Sbordoni leg.; same locality, 4♂, AV 3<0@0@>>< #$ ".#C*&#$" C"'##BB-#*0E&$ !@>33 J#-"+" ##J$' . ! $#  SW of the city, 3739.031’ N, 2210.156’ E, 960 m, 06.04.2009, 2♂, 5♀; Arcadia, Magouliana, stream SE of the village, 3739.425’ N, 2208.730’ E, 1035 m, 06.04.2009, 1♂, 5♀; Arcadia, Tetrazi Mts, Agia Theodora, stream and / $+$ * 7<@30@64R @35A0 >50>80@>>4 8V0  '#."   .&- $# '(#,$"C#)$& $!"+$- #$# ##-"+" # +#'# #$# ' &*  $' .+$-0--/ " +"+&- " "''&$ +#$ # $#  .+$ *"&-#0

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       #$-99(

       '#J&)#$3467  '/34Thessaloniki, 3–8.6, 1♀; Mount Oiti: 1500m, 18–25.5.1956, 2♂, 2♀; :#$ -#!34<3J+#$'#" at Ag. Georgios, bridge on the route Karpenisi-Lamia, 10.4.1967, 1♀; '/ 34

      > !5? #&3$&

"(' " @>3@2 J"('@>3@9J3A702"$* -.#'$+" ", -# ,# -# - $(  . #**90  : -/  # .#'J -)   2"('   @>3@9  " # #*$" 2"('   @>3@9E   '#."  D# $ - $##'# # -!0     $ !+#VJI# '#."  J $#(# $#*" -&  # # #/" (#$ 8> >8R@<04R  E21°12΄06.0”, 973 m, (2006/114), 14.05.2006, DL-KJ-MD (HNHM); paratype ♀: Eleftherohori, Venetikos River, 24.04.1989, HB-SI, (PMSL); paratypes 1♀, 1 exuviae: same locality, N40°03΄06.6”, E21°28΄50.2”, 475 m, (2006/ 111), 13.05.2006, DL-KJ-MD (HNHM); paratypes 1♂, 9 exuviae: Kipourio, Venetikos River, 24.04.1989, HB-SI (PMSL); paratypes 1♂, 1 larva: Zakas, Venetikos River, N40°02΄19.3”, E21°17΄19.2”, 699 m, (2006/112), 14.05.2006, DL-KJ-MD (HNHM); paratypes 6♂, 6♀, 9 exuviae: Thessaly, Trikala regional unit, Ambelia, Kalambaka, 20.04.1989, HB-SI (PMSL, CGV); paratypes 1♂, 7 exuviae: Kastania, Kalambaka, 20.04.1989, HB- SI (PMSL); paratypes 2♀: Kato Palagokaria, 19.04.1989, HB-SI (PMSL); paratypes 3♀: Longiai, 19.04.1989, HB- SI (PMSL); paratype ♂: Karditsa regional unit, Sarantaporo, small river S of the village, N39°09΄13.9”, E21°49΄50.7”, 700 m, (2011/39), 08.05.2011, KJ-MD-SzT-UZs (HNHM); paratypes 1♂, 1♀: Central Greece, "$#*" -&  *" #"$*" +#$'" (#$I",#(-- *# 7A 5<_>>05P @3 56_8@0

Phocis regional unit, Vardousia Mts, Stromi, 26.04.1990, HB-SI (PMSL); paratypes 4♂, 2♀: Giona Mts, Lefkaditi, NW of Amfissa, 15.04.1987, Drosopoulos (PMSL90 +#'# #$#,"& . .,,#$# !+#",$(#$)### 75 .4<5 , ,-" #(#  # -" -&.#" !&)$ #0 $( ##$#,"& .& .#$- $*#" # #, #'&$$# 0#-,#'!'-### ")#" #G!# $ "#$   +#'#0.&-"''&$$#.)### .G $' .# .", !  # $" # .(#*# " '-"#"$(#$) /0&#".# ) +$#,#$# '# +#'#'"" -! "' #.  !"#$" #,-#0#"+#'#$'-"' -!#+#$'" (#$ *" #"$*"2"('@>3@90

        > !5? #&3$&

"(' " @>3@2 J"('@>3@9J3AA02"$* -.#'$+" ", -# ,# -# - $(  . #**90       2 )&$ 3A8@9J#$ ".#C*&#$" C"'#@>>32,.#"('@>3@90  .#'" #-"+" ##0    "-"!+#V + $ !+#6W @>#1&( #J#-"+" ##J' # $#*" -&  $ " 6

 ! "#  a‚‚‡h‘h*%%%*'+1%2.    9  km W Agridi, 19.04.1990, HB-SI (PMSL, MM, HNHM); paratypes 1♂, 1♀: Ano Vlassia, Upstream of Selinous River, 37°57’N, 21°47’E, 1000 m, (GR 59), 25.05.1992, BC-GR (CGV); paratype ♀: Kalavrita, Vouraikos River, @70>5034A5   2 90      #$ " .# C*&#$"   C"'# @>>3J #-"+" ##J (#$ "&$ /"   - ($  @705034A5 @V -#*02" -!" #+#'# .#* #.  + $ !+#90

        )*+#$%,-

 '/34 @<05034<52D 90

#   )*+#$%,-

 '/34M35>  3AM 20.5.1975 (CPZ); paratypes 2♂, 2♀: S Kurunia 25°55'E, 38°34'N, 350m, 20.5.1975 (CPZ); paratype ♂: 5 km NE Pirama, 25°54'E, 38°3l'N, 170m, 20.5.1975 (CPZ); paratypes 16♂, 9♀: SE Kaminia 38°33'N, 25°56E', 470m, 18.5.1975 (CPZ); paratypes 2♂: 2 km N Fita, 26°00'E, 38°32'N, 510m (CPZ); paratypes 2♂, 4♀: Wikion (CPZ).      '/34M35> 3AM@>05034<5E !- $( #J&$&  @5 55N 7A 78N  75> @>05034<5E- $( #J   7A 77N @5 56N 8<> 3A05034<5E3#1&( #JI/" 0

#      #$,2$(

 %    'J# 3A670  I# - # $''+#'# / " ,$" --#: -/ '"& $#)&D$"  2&$ !@>33)9E   '#."  #-"+" ##2]90     #  3A67J 3@ +#'#   -#*0 $+#$  #-"+" ##2]9E &$ ! @>33)J  '#."  J $#(# J -#,#$""$  # #/" (#$  8>>7R>606  @3@AR5>0@  8<5   370>50@>>6  -#*0 0  !  =0 Kontschán, D. Murányi: 1♂; Zakas, Venetikos River, 4002’19.3N, 2117’19.2”E, 700m, 14.05.2006, leg. L. Dányi, =0" ' 0&$ !J3W 8- $( #   $#("& $#'"$. ",  +#'# ,$" $##'# $#,#$ "-! " #  #$+$# "  ", &#& $ #'&   '# 23A839  # . ", &#& $ #'#  2".  3<63  ,.# '/ 34

#    )*+#$%,-

 '/34M65> @6 @>N 7< 74N  @@05034<72+'/  &'  #-92D 90

 9 a‚‚‡h‘h*%%%*'+1%2 .     #    )*+#$%,-

 '/343N  280m (CPZ); paratypes 8♂6V - $( #J#"'"$" @6 34N 74 01'N, 250m, 25.5.1975 (CPZ); paratypes 7♂ 7VJ/$  @/#)" @6 34N 74 03'N, 370m, 25.5.1975 (CPZ); paratypes 6♂, 7♀: Ipsilometopo, 2 km E, #)" @6 36N 74 19'N, 420m, 27.5.1975 (CPZ); paratypes 10♂, 5♀, 1 larva: Lepetimnos (CPZ); paratypes 1♂, 3V - $( #J$*# " I#)" @5 36N 74 22'N, 370m, 28.5.1975 (CPZ); paratypes 5♂, 1♀: Chidira, 26 >@N  74 13'N, 210m, 28.5.1975 (CPZ); paratypes 31♂, 19♀: Plomari, 26 @6N 7A 54N 33> 7305034<52D 90      '/343N @A>E- $( #J#"'"$" @6 34N 74 >3N @5> @505034<5E 3- $( J#+# "E- $( #J$*# " I#)" @5 36N 74 @@N 7<> @A05034<50

#     )*+#$%,-

 '/340N2"$* -.#'$+" ", -#90 &$ !@>33)J7@02'"+-##  $!.#'$+" ", -# #-#( " "+#'# &90  )       * '/34N  7A >8N  @@   @@05034<52D 90      '/34N 7A >8N @@ m, 22.5.1975 (same locality and data as the types); Paradision, 1♂, 1♀, Larvae, 450 m, 29.5.1973, 1♂, 7♀ 2+'/  &'  #-9E$,  "# @7 53N 7A 34'N, 520m, 24.5.1974, 1♂, 1♀, Larvae; Andros, Apikia, 29.5.1973, 3♂, Larvae (Malicky); Central Greece: Pendayi, 3♂, 4♀, Larvae; Eptalofos, 7♂, 4♀, 4 Ex; Lilaia, @@ 7>N 7A 37'N, 300m, 29.–30.5.l976, 1♀, 50 larvae, incl. Karia, 30.4.–1.5.1976, 2♂, 2♀; Peloponnese: Agios   @@ 533)J'   +$#,#'&$#J "-  $ " #- "&$#  3>>> @50>50344@ -#*00 -'/! .#0 0 " J @W@V0 #/ " -"' -# $#&)"  #$#  .+$ *"&-#0.&- '(#  ! .&& --!  '""  .(#$#'"& !"," #,-#0

#         #$%1'

&)#$3456J@>602"$* -.#'$+" ", -# .,# -#",) 90 :#$ -#!34<3J8602'"+-##  $!.#'$+" ",#- $( 90 '/34>3J6402.#'$+" ",#**90 &$ !@>33)J@702'"+-##  $!.#'$+" ", -# ,# -# - $(  .#**90  )   &)#$ 3456J&)#$3456 &)#$3467 =""34<>0  : -/   . D# $ - &$"+#  +#'#  / "  ,$" -- : -/  '"& $#E   +$&   '#."  D# $ - $##'# #-"+" ## &)"# 0    2",) &)#$ 34569 "-"!+#W  --"!+#V + $ !+#5W > @305034552 90      &)#$3456J Peloponnese: Kakary, Anogia, 300m, 22.5.1955, 4♂, 2♀, 21 larvae; stream in 1200m, Anogia route to Taygetos, 21.5.1955, 6♂, 8♀, 16 larvae; Landonas, Magouliana, 1100m, 23.5.1955, 3♂, 6♀, 7 larvae; Ladon and tributaries, Vytina, 100m, 24.5.1955, 3♂, 23 larvae; Vouraikos, Kalavrita, 700m, 26.5.1955, 4 larvae; Epirus: stream on road to Metsovon, 1600–1700m, 9.6.1955, 6♂, 11♀, 15 larvae; Makedonia: Mounts Neretschka, road from Argos to Vevi, Km 39, 900m, 11.6.1955, 1♂; Olympos, Mavroneri, on

 ! "#  a‚‚‡h‘h*%%%*'+1%2.    9  road from Katerini to Elassona, Km 29, 600m, 16.6.1955, 1♂; &)#$3467J Euboea: Steni, 800 m, 4–11.4, 1♂; Peloponnese: Mistras, near Sparti, 28.4.1956, 1♀; =""34<>J# . ! @@ >5N 7A 35'N, 900m, 4.6.1969, 2♂; :#$ -#!34<3JD# $ - $##'#J+#$'#" *0 #"$*" )$.*#",#$"&# $+# G  3>080346< @ larvae; Euboea: Ilia stream, small village by the sea 8 km east of Aedipsos, 3.4.1967, 1♀, 6 larvae. Kireus, 1.5 km )#,"$#$"/"+" $"&##.+"MD -'E$+! # $#  55> 3M@/&+$# ",#  35080346< 3 - $( E --$# 7M3>. 37080346< 3- $( E '/34>3J#-"+" ##J-"  ( $+"- @605034A5  " "& #-"0 @@05034A5E&$ !@>33)JC-"$ +$#,#'&$#J".#$ #$ " ,"$#$# 7/I", the village, 4047’16.5N, 2113’26.7E, 1315 m, 15.05.2006, leg. L. Dányi, J. Kontschán, D. Murányi: 1♂, 1 exuviae; "  +$#,#'&$#J-# 5/I",#(-- *# 37>> >40>60344@ -#*00 #$#'/# .#0 0 " J7W7V0

#        )*+#$%,-

 '/3402"$* -.#'$+" ", -#9  .#'" $##'#J"$#*# - .0    "-"!+#W + $ !+#WJ#)"J</",-" $ @6 @6N7A 54N 33> 7305034<52D 9E + $ !+#@WJ3/I",#* -"'"$ @6 @3N 7A 01'N, 280 m, 24.5.1975 (CPZ); paratype ♂: same locality, 31.5.1975 (CPZ); paratypes 2♂: Lepetimnos, 25 36N 74 22'N, 330–420 m, 27–28.5.1975 (CPZ); paratypes 3♂: #"'"$"  @6 34N  74 01'N, 250 m, 25.5.1975 (CPZ); paratypes 2♂: Ambeliko, 26 3AN  74 >8N  78>   @505034<52D 90      '/ 34M8@>  27–28.5.1975; 3♀, larvae: Neochorion, 26 34N 74 01'N, 250 m, 25.5.1975; 2♀: Ambeliko, 26 3AN 74 >8N  78> @505034<50

< 

$     >+#$%3,(

- +-#/34><0   : -/   . D# $ - &$"+#  +#'#0   # : -/  / "  ,$" :"   . #$B#*"( :&-* $ D$"  ""(" " # #*$"  #+&)-'", '#."  #$) -"(#  E +$&  D# $ - $##'# #-"+" ## &)"# 0    ! !+#7VJ $##'# -#*0$+#$290     &)#$3467JD# $ - $##'#J "$*"+" " #,""","&  A>> @6M 28.6.1956, 4♀; '/ 3433 J#-"+" ##J$' . -- /"  "$$&  "&" (#$ 7<7@07@0 >60>80@>>4  @W 3- $( 0     # "$* - .#'$+"  # !+# #$#  .' #. " )# .#+"#.   :#$-  2- +-#/ 34><90 "#(#$  " ", # $## +#'#  $#   # 0# +#'#  )# $ * D"!+# - )#-   .#* #.  -#'"!+# )! '/   34A> )& " ,"$ --! +&)-#.0 !+# +$") )-! )#-" * " #  # +#'#     +&)-#.     )!# 23A6790+  - $( -$#'"$.+&)-#."&+#','.# ,' " )! &)#$234569,$"#"(" !$#,#$"+#'#0

 9 a‚‚‡h‘h*%%%*'+1%2 .     %     > #$--1(

 - +-#/34@3J5402"$* -.#'$+" ",,# -#",   90    - +-#/ 34@3J- +-#/34@30  D$'&G#.#$$ #  +#'#    # : -/    / "  ,$" -)   #+&)-' ",  '#."  E +$&  '#."  #-"+" ## "  - .2D"$,&90    2",  - +-#/ 34@39 ! !+#VVJ#-"+" ##J - ($! "$# -#*0"-B 290      &)#$ 3456J Peloponnese: Evrotas, Sparti, 200m, 19.5.1955, 1♂, 1♀, 4 larvae; "&$ /"  -"$"&2#* +-#" 9 5>> @30603455E+$&J+" "" $" ."#"(" 5>> A0603455  3♂, 1♀, 4 larvae; Macedonia: Aliakmonas, between Neapolis and Argos, 580m, 10.6.1955, 13 larvae; '/ 34)& ","$ --!+&)-#.0

&      !#$%%,

(#'344VJ+$&J3>/"&$" 74 38N@>` @> 8@N>>` @>>  26.5.1994, leg. Sivec, I. & Horvat, B. (PMSL, CPZ); paratypes 2♂, 12♀, 36 larvae: same locality, 17.5.1994 (PMSL); paratypes 3♂, 2♀: same locality, 21.5.1987, leg. Malicky, H. (PMSL); paratypes 1♂, 1♀: stream N $' *#-" 74 > 74N A> @>05034A< -#*0 -'/! 0290

'      . #$-9%(

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&)#$3456J@3>02! " !!",   ' #& . 34@690 :#$ -#!34<3J5302'"+-##  $!.#'$+" ",- $( .$#* $.",   !   ! " !!90 '/34>@9E  +$&  '#."  D# $ - $##'# # -! #-"+" ## &)"# "$#*# - .0     - +-#/34@7JParnassos, 1♀; Kalavryta, Morea, leg. Holtz, 3♀;  (34@8J '#."  J "&#1 '-"' -!2+")-!C 9 343A -#*0 (#E#$ 50343AE.# 2".# 9 <0343A -#*0#$*#  D .#- E&)#$3456J '#."  J"& #$#'/ $" .$*""#( /G@8 4>> 330603455 <- $( #E/G 39, 1♂, 7 larvae; Olympus, Mavrolongos, Lithochoron, 300m, 16.6.1955, 2 larvae; Agios Dionysos, 850m, 15.6.1955, 1 larva, 1 ex.; Prionia, 13.6.1955, 1100m, 3♀, 6 larvae; Mavroneri, on road from Katerini to Elassona, km-29, 600m, 16.6.1955, 1♂, 12 larvae; km-40, 900m, 16.6.1955, 2 larvae; Thesally: Kalambaka, 300m, 10.6.1955, 1♂; on road to Metsovo in 650–750m, 6.6.1955, 7 larvae, 7 exuvies; Parnassos: Main river, N side of

 ! "#  a‚‚‡h‘h*%%%*'+1%2.    9  the road to Gravia, 650m, 5.6.1955, 1 larva; Peloponnese: Kakary, Anogia, 300m, 22.5.1955, 1♂, 20 larvae; Landonas, Magouliana, 1100m, 23.5.1955, 1♂; Vytina, 950m, 24.5.1955, 2♂, 2♀, 32 larvae; Vouraikos, Zalorou 2#* +-#" 9  5>>   @30603455 8 - $( #E "&$ /"  - ($ <>> @60503455  3- $( E$# )# . # "#-  - ($ <>> @60503455 @- $( #E&)#$3467JD# $ - $##'#J "$*"+" " )# # 0 A>> m, 26–28.1956, 1♂;=""34<>J#-"+" ##J--  @@ @8N 7< 58'N, 1500m, 28.5.1969, 1♀; '/34>@9 #"''&$$# '#",      $##'#  "-/#-! . $#+"$#.$#'"$.$#,#$" !",#   !+### &"$.#'$)#.0"#(#$ #-# )"(#$#'"$.      " *  --+#'#  ##.")#$##1  #.,"$.##$ " 0

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:#$ -#!34<3J5>02'"+-##  $!.#'$+" ",- $( 90 '/34>@J@702$#.#'$+" ",#** #$#'" ",,"&$"$+"!+#90        '"# #& .J&)#$34672,.#:#$ -#!34<390     - +-#/ 34>>@0 D &'  G "- +#'# .+#$#. "D# $ -&$"+#0 #: -/  +#'#  )## $#'# -!'" ,$#.,$":"   .#$B#*"( :&-* $ D$"  " # #*$"  #+&)-'", '#."   -"(#  2"+Q 6(#'@>>4) (#' $/@>>@9E +$&  '#."  D# $ - $##'# # -!  &)"# 0      &)#$ 3467J Macedonia: Prioni, 1000 m, Olympus, 3–13.6.1956, 1♂ brachypter, 5♀; :#$ -#!34<3JD# $ - $##'#JC". "$$#   '!+$#"& 335> 4080346< <- $( #E+#$'#" *0 #"$*" )$.*#",#$"&# $+# G  3>080346< 8- $( E '/34>@J!+#7#**J +$& #* #$#$ @@05034AA 0(#' 0 '. E+$& !  (#$ "&-"+"&- @305034AA 0(#' 0 '. E!+#8#**JD# $ - $##'# -  " -#+ @5060344> 0(#'E"('@>3@JD# $ - $##'#J "$#*" -&  *" #"$*" +#$'" (#$I",#(-- *# 7A 5>05P @3 56R8@0A0>50@>330  ''"$. *"(#' $/2@>>@9 #: -/  +"+&- "  $#+$") )-! "'" +#','   .#'$)#.,$"#D &' &0#!$#+"$#. $##/+#'# "$#+$## O!+#7 .O!+#8P0

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 9 a‚‚‡h‘h*%%%*'+1%2 .     Kalavrita, 700m, 26.5.55, 20♂, 11♀; Epirus: Metsovitiko, on road to Metsovon, 600m, 8.6.1955, 1 male, 2 ,# -#  $#   365>  " $" ."#"("  40603455  3,# -#E:#$ -#! 34<3J ($  J '#-""  '!+$#"& . $+#  3>>> 4080346< 7- $( #E+#$'#" *0 #"$*" )$.*#",#$"&#  $+# G   3>080346<  6 - $( #E &)"# J -  $#    -- (-- *# )! # #  A / #  ", #.+"  7080346<  3 - $( #E $#&  305 / )#,"$# $"'"+"  $"&# #.+" G D -'0  -- $#  7M3>  .  37080346< 3- $( E$+! $)& $!",# $#  55> 35080346< 3- $( E '/34N 7A 42'N, 250 m, 5♂, 2♀; 1.5.1975, 2♀, 2 - $( #E" "/ $ @7 38N 7A 5 

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