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Two New Stegosaur Specimens from the Upper Jurassic Morrison Formation of Montana, USA
Editors' choice Two new stegosaur specimens from the Upper Jurassic Morrison Formation of Montana, USA D. CARY WOODRUFF, DAVID TREXLER, and SUSANNAH C.R. MAIDMENT Woodruff, D.C., Trexler, D., and Maidment, S.C.R. 2019. Two new stegosaur specimens from the Upper Jurassic Morrison Formation of Montana, USA. Acta Palaeontologica Polonica 64 (3): 461–480. Two partial skeletons from Montana represent the northernmost occurrences of Stegosauria within North America. One of these specimens represents the northernmost dinosaur fossil ever recovered from the Morrison Formation. Consisting of fragmentary cranial and postcranial remains, these specimens are contributing to our knowledge of the record and distribution of dinosaurs within the Morrison Formation from Montana. While the stegosaurs of the Morrison Formation consist of Alcovasaurus, Hesperosaurus, and Stegosaurus, the only positively identified stegosaur from Montana thus far is Hesperosaurus. Unfortunately, neither of these new specimens exhibit diagnostic autapomorphies. Nonetheless, these specimens are important data points due to their geographic significance, and some aspects of their morphologies are striking. In one specimen, the teeth express a high degree of wear usually unobserved within this clade—potentially illuminating the progression of the chewing motion in derived stegosaurs. Other morphologies, though not histologically examined in this analysis, have the potential to be important indicators for maturational inferences. In suite with other specimens from the northern extent of the formation, these specimens contribute to the ongoing discussion that body size may be latitudinally significant for stegosaurs—an intriguing geographical hypothesis which further emphasizes that size is not an undeviating proxy for maturity in dinosaurs. Key words: Dinosauria, Thyreophora, Stegosauria, Jurassic, Morrison Formation, USA, Montana. -
Stegosaurus Scelidosaurus Huayangosaurus Cheeks: No
Huayangosaurus Scelidosaurus Stegosaurus Cheeks: No reptile has ever had a ‘buccinator’ muscle Answer: highly flexible tongue Brains 0.001% of stegosaur body weight Compared to 1.8% in humans (1000x larger per unit body weight!) Brains Brains Locomotion Graviportal Locomotion Elephantine hind feet (weight-bearing) Shin bones fused with astragalus/ calcaneum Femur: Long compared to humerus Columnar Facultative Tripodality? Stocky forelimbs- could be used for turning/posturing (Bakker) Dermal Armour? Pattern of plates and spines is species-specific Plates paired or staggered (Stegosaurus) Plates were probably not for defense... not tough enough Rotation? Surface markings => symmetrical. Rotation unlikely Potential uses: Thermoregulation? Warm up (ectotherms), Cool down (endotherms) Signaling? positioned for maximal lateral visibility Sexual Selection Mate Recognition Grooves for blood vessels Sexual dimorphism Differences between males and females of the same species **New finding** published in 2015 Stegosaurus Morrison formation, Colorado Dinosaur Sex Figuring out how Stegosaurus even could have mated is a prickly subject. Females were just as well-armored as males, and it is unlikely that males mounted the females from the back. A different technique was necessary. Perhaps they angled so that they faced belly to belly, some have guessed, or maybe, as suggested by Timothy Isles in a recent paper, males faced away from standing females and backed up (a rather tricky maneuver!). The simplest technique yet proposed is that the female lay down on her side and the male approached standing up, thereby avoiding all those plates and spikes. However the Stegosaurus pair accomplished the feat, though, it was most likely brief—only as long as was needed for the exchange of genetic material. -
A Phylogenetic Analysis of the Basal Ornithischia (Reptilia, Dinosauria)
A PHYLOGENETIC ANALYSIS OF THE BASAL ORNITHISCHIA (REPTILIA, DINOSAURIA) Marc Richard Spencer A Thesis Submitted to the Graduate College of Bowling Green State University in partial fulfillment of the requirements of the degree of MASTER OF SCIENCE December 2007 Committee: Margaret M. Yacobucci, Advisor Don C. Steinker Daniel M. Pavuk © 2007 Marc Richard Spencer All Rights Reserved iii ABSTRACT Margaret M. Yacobucci, Advisor The placement of Lesothosaurus diagnosticus and the Heterodontosauridae within the Ornithischia has been problematic. Historically, Lesothosaurus has been regarded as a basal ornithischian dinosaur, the sister taxon to the Genasauria. Recent phylogenetic analyses, however, have placed Lesothosaurus as a more derived ornithischian within the Genasauria. The Fabrosauridae, of which Lesothosaurus was considered a member, has never been phylogenetically corroborated and has been considered a paraphyletic assemblage. Prior to recent phylogenetic analyses, the problematic Heterodontosauridae was placed within the Ornithopoda as the sister taxon to the Euornithopoda. The heterodontosaurids have also been considered as the basal member of the Cerapoda (Ornithopoda + Marginocephalia), the sister taxon to the Marginocephalia, and as the sister taxon to the Genasauria. To reevaluate the placement of these taxa, along with other basal ornithischians and more derived subclades, a phylogenetic analysis of 19 taxonomic units, including two outgroup taxa, was performed. Analysis of 97 characters and their associated character states culled, modified, and/or rescored from published literature based on published descriptions, produced four most parsimonious trees. Consistency and retention indices were calculated and a bootstrap analysis was performed to determine the relative support for the resultant phylogeny. The Ornithischia was recovered with Pisanosaurus as its basalmost member. -
Stegosaurian Footprints from the Morrison Formation of Utah and Their Implications for Interpreting Other Ornithischian Tracks Gerard D
Stegosaurian footprints from the Morrison Formation of Utah and their implications for interpreting other ornithischian tracks Gerard D. Gierliński and Karol Sabath Polish Geological Institute, Rakowiecka 4, 00-975 Warsaw, Poland. e-mail: [email protected] ABSTRACT - The supposed stegosaurian track Deltapodus Whyte & Romano, 1994 (Middle Jurassic of England) is sauro- pod-like, elongate and plantigrade, but many blunt-toed, digitigrade, large ornithopod-like footprints (including pedal print cast associated with the manus of Stegopodus Lockley & Hunt, 1998) from the Upper Jurassic of Utah, better fit the stego- saurian foot pattern. The Morrison Formation of Utah yielded other tracks fitting the dryomorph (camptosaur) foot pattern (Dinehichnus Lockley et al., 1998) much better than Stegopodus. If the Stegopodus pedal specimen (we propose to shift the emphasis from the manus to the pes in the revised diagnosis of this ichnotaxon) and similar ichnites are proper stegosaur foot- prints, Deltapodus must have been left by another thyreophoran trackmaker. Other Deltapodus-like (possibly ankylosaurian) tracks include Navahopus Baird,1980 and Apulosauripus Nicosia et al., 1999. Heel-dominated, short-toed forms within the Navahopus-Deltapodus-Apulosauripus plexus differ from the gracile, relatively long-toed Tetrapodosaurus Sternberg, 1932, traditionally regarded as an ankylosaurian track. Thus, the original interpretation of the latter as a ceratopsian track might be correct, supporting early (Aptian) appearance of ceratopsians in North America. Isolated pedal ichnites from the Morrison Formation (with a single tentatively associated manus print, and another one from Poland) and the only known trackways with similar footprints (Upper Jurassic of Asturias, Spain) imply bipedal gait of their trackmakers. Thus, problems with stegosaur tracks possibly stem from the expectation of their quadrupedality. -
The Systematic Position of the Enigmatic Thyreophoran Dinosaur Paranthodon Africanus, and the Use of Basal Exemplifiers in Phyl
1 The systematic position of the enigmatic thyreophoran dinosaur Paranthodon africanus, 2 and the use of basal exemplifiers in phylogenetic analysis 3 4 Thomas J. Raven1,2 ,3 and Susannah C. R. Maidment2 ,3 5 61Department of Earth Science & Engineering, Imperial College London, UK 72School of Environment & Technology, University of Brighton, UK 8 3Department of Earth Sciences, Natural History Museum, London, UK 9 10Corresponding author: Thomas J. Raven 11 12Email address: [email protected] 13 14 15 16 17 18 19 20 21ABSTRACT 22 23The first African dinosaur to be discovered, Paranthodon africanus was found in 1845 in the 24Lower Cretaceous of South Africa. Taxonomically assigned to numerous groups since discovery, 25in 1981 it was described as a stegosaur, a group of armoured ornithischian dinosaurs 26characterised by bizarre plates and spines extending from the neck to the tail. This assignment 27that has been subsequently accepted. The type material consists of a premaxilla, maxilla, a nasal, 28and a vertebra, and contains no synapomorphies of Stegosauria. Several features of the maxilla 29and dentition are reminiscent of Ankylosauria, the sister-taxon to Stegosauria, and the premaxilla 30appears superficially similar to that of some ornithopods. The vertebral material has never been 31described, and since the last description of the specimen, there have been numerous discoveries 32of thyreophoran material potentially pertinent to establishing the taxonomic assignment of the 33specimen. An investigation of the taxonomic and systematic position of Paranthodon is therefore 34warranted. This study provides a detailed re-description, including the first description of the 35vertebra. Numerous phylogenetic analyses demonstrate that the systematic position of 36Paranthodon is highly labile and subject to change depending on which exemplifier for the clade 37Stegosauria is used. -
Síntesis Del Registro Fósil De Dinosaurios Tireóforos En Gondwana
ISSN 2469-0228 www.peapaleontologica.org.ar SÍNTESIS DEL REGISTRO FÓSIL DE DINOSAURIOS TIREÓFOROS EN GONDWANA XABIER PEREDA-SUBERBIOLA 1 IGNACIO DÍAZ-MARTÍNEZ 2 LEONARDO SALGADO 2 SILVINA DE VALAIS 2 1Universidad del País Vasco/Euskal Herriko Unibertsitatea, Facultad de Ciencia y Tecnología, Departamento de Estratigrafía y Paleontología, Apartado 644, 48080 Bilbao, España. 2CONICET - Instituto de Investigación en Paleobiología y Geología, Universidad Nacional de Río Negro, Av. General Roca 1242, 8332 General Roca, Río Negro, Ar gentina. Recibido: 21 de Julio 2015 - Aceptado: 26 de Agosto de 2015 Para citar este artículo: Xabier Pereda-Suberbiola, Ignacio Díaz-Martínez, Leonardo Salgado y Silvina De Valais (2015). Síntesis del registro fósil de dinosaurios tireóforos en Gondwana . En: M. Fernández y Y. Herrera (Eds.) Reptiles Extintos - Volumen en Homenaje a Zulma Gasparini . Publicación Electrónica de la Asociación Paleon - tológica Argentina 15(1): 90–107. Link a este artículo: http://dx.doi.org/ 10.5710/PEAPA.21.07.2015.101 DESPLAZARSE HACIA ABAJO PARA ACCEDER AL ARTÍCULO Asociación Paleontológica Argentina Maipú 645 1º piso, C1006ACG, Buenos Aires República Argentina Tel/Fax (54-11) 4326-7563 Web: www.apaleontologica.org.ar Otros artículos en Publicación Electrónica de la APA 15(1): de la Fuente & Sterli Paulina Carabajal Pol & Leardi ESTADO DEL CONOCIMIENTO DE GUIA PARA EL ESTUDIO DE LA DIVERSITY PATTERNS OF LAS TORTUGAS EXTINTAS DEL NEUROANATOMÍA DE DINOSAURIOS NOTOSUCHIA (CROCODYLIFORMES, TERRITORIO ARGENTINO: UNA SAURISCHIA, CON ENFASIS EN MESOEUCROCODYLIA) DURING PERSPECTIVA HISTÓRICA. FORMAS SUDAMERICANAS. THE CRETACEOUS OF GONDWANA. Año 2015 - Volumen 15(1): 90-107 VOLUMEN TEMÁTICO ISSN 2469-0228 SÍNTESIS DEL REGISTRO FÓSIL DE DINOSAURIOS TIREÓFOROS EN GONDWANA XABIER PEREDA-SUBERBIOLA 1, IGNACIO DÍAZ-MARTÍNEZ 2, LEONARDO SALGADO 2 Y SILVINA DE VALAIS 2 1Universidad del País Vasco/Euskal Herriko Unibertsitatea, Facultad de Ciencia y Tecnología, Departamento de Estratigrafía y Paleontología, Apartado 644, 48080 Bilbao, España. -
The Fighting Pair
THE FIGHTING PAIR ALLOSAURUS VS STEGOSAURUS — Allosaurus “jimmadsoni” and Hesperosaurus (Stegosaurus) mjosi Upper Jurassic Period, Kimmeridgian Stage, 155 million years old Morrison Formation Dana Quarry, Ten Sleep, Washakie County, Wyoming, USA. THE ALLOSAURUS The Official State Fossil of Utah, the Allosaurus was a large theropod carnosaur of the “bird-hipped” Saurischia order that flourished primarily in North America during the Upper Jurassic Period, 155-145 million years In the spring of 2007, at the newly-investigated Dana Quarry in the Morrison Formation of Wyoming, the team from ago. Long recognized in popular culture, it bears the distinction of being Dinosauria International LLC made an exciting discovery: the beautifully preserved femur of the giant carnivorous one of the first dinosaurs to be depicted on the silver screen, the apex Allosaur. As they kept digging, their excitement grew greater; next came toe bones, leg bones, ribs, vertebrae and predator of the 1912 novel and 1925 cinema adaptation of Conan Doyle’s finally a skull: complete, undistorted and, remarkably, with full dentition. It was an incredible find; one of the most The Lost World. classic dinosaurs, virtually complete, articulated and in beautiful condition. But that was not all. When the team got The Allosaurus possessed a large head on a short neck, a broad rib-cage the field jackets back to the preparation lab, they discovered another leg bone beneath the Allosaurus skull… There creating a barrel chest, small three-fingered forelimbs, large powerful hind limbs with hoof-like feet, and a long heavy tail to act as a counter-balance. was another dinosaur in the 150 million year-old rock. -
Remains from Jurassic-Cretaceous Transition Beds of Valencia Province (Southwestern Iberian Range, Spain)
ISSN (print): 1698-6180. ISSN (online): 1886-7995 www.ucm.es/info/estratig/journal.htm Journal of Iberian Geology 36 (2) 2010: 243-252 doi:10.5209/rev_JIGE.2010.v36.n2.10 New stegosaurian (Ornithischia, Thyreophora) remains from Jurassic-Cretaceous transition beds of Valencia province (Southwestern Iberian Range, Spain). Nuevos restos de estegosaurios (Ornithischia, Thyreophora) del tránsito Jurásico-Cretácico de la provincia de Valencia (Cordillera Ibérica Suroccidental, España) J. Company1, X. Pereda Suberbiola2, J.I. Ruiz-Omeñaca3,4 1Departamento de Ingeniería del Terreno, Universidad Politécnica de Valencia, 46022 Valencia, Spain. [email protected] 2Universidad del País Vasco/Euskal Herriko Unibertsitatea, Facultad de Ciencia y Tecnología, Departamento de Estratigrafía y Paleontología, Apartado 644, 48080 Bilbao, Spain. [email protected] 3Museo del Jurásico de Asturias (MUJA), 33328 Colunga, Spain 4Grupo Aragosaurus-IUCA (www.aragosaurus.com), Area de Paleontología, Facultad de Ciencias, Universidad de Zaragoza, 50009 Zaragoza, Spain. [email protected] Received: 20/11/09 / Accepted: 30/06/10 Abstract New stegosaurian remains have been recently recovered from the Jurassic-Cretaceous transition sandstones of the Villar del Arzobispo Formation (Tithonian-Berriasian) in the Valencia province, eastern Spain. Specimens consist of two partially articulated (or closely associated) postcranial skeletons. The Baldovar specimen is composed of appendicular bones (scapula, femur) and two pairs of dermal tail spines, two of them articulated with two distal caudal vertebrae. The second specimen, unearthed in the vicinity of La Yesa village, consists of dorsal vertebrae and ribs, fragments of caudal centra and an incomplete femur. The new specimens are tentatively referred to the clade Dacentrurinae and may belong to Dacentrurus on the basis of features observed on the dorsal vertebrae and caudal dermal spines. -
Two New Stegosaur Specimens from the Upper Jurassic Morrison Formation of Montana, USA
Two new stegosaur specimens from the Upper Jurassic Morrison Formation of Montana, USA D. Cary Woodruff, David Trexler, and Susannah C.R. Maidment Acta Palaeontologica Polonica 64 (3), 2019: 461-480 doi:https://doi.org/10.4202/app.00585.2018 Two partial skeletons from Montana represent the northernmost occurrences of Stegosauria within North America. One of these specimens represents the northernmost dinosaur fossil ever recovered from the Morrison Formation. Consisting of fragmentary cranial and postcranial remains, these specimens are contributing to our knowledge of the record and distribution of dinosaurs within the Morrison Formation from Montana. While the stegosaurs of the Morrison Formation consist of Alcovasaurus, Hesperosaurus, and Stegosaurus , the only positively identified stegosaur from Montana thus far is Hesperosaurus. Unfortunately, neither of these new specimens exhibit diagnostic autapomorphies. Nonetheless, these specimens are important data points due to their geographic significance, and some aspects of their morphologies are striking. In one specimen, the teeth express a high degree of wear usually unobserved within this clade—potentially illuminating the progression of the chewing motion in derived stegosaurs. Other morphologies, though not histologically examined in this analysis, have the potential to be important indicators for maturational inferences. In suite with other specimens from the northern extent of the formation, these specimens contribute to the ongoing discussion that body size may be latitudinally -
Stegosaurus Chirality R
Stegosaurus chirality R. P. Cameron1,*, J. A. Cameron1, and S. M. Barnett1 1School of Physics and Astronomy, University of Glasgow, Glasgow G12 8QQ, United Kingdom. *[email protected] Introduction Most living things appear rather symmetrical: the external human form, for example, has a plane of mirror symmetry to good approximation. Snails, flounders, narwhals, crossbills, fiddler crabs and twining vines are members of the short but fascinating list of living things known instead to defy mirror symmetry by exhibiting exterior chirality1. The study of this symmetry breaking lies at the cutting edge of developmental and evolutionary biology2,3. We have recently extended the aforementioned list4 by adding one of the most recognisable genera of dinosaurs: Stegosaurus5. Here we summarise our research to date into Stegosaurus chirality. Figure 1. The arrangement of Stegosaurus's plates differs from its mirror image and is therefore chiral1, as highlighted here for the largest plate in particular of the Stegosaurus stenops holotype USNM 4939: this specimen was of the (R) rather than the (L) form. Adapted from 8. The currently favoured arrangement of Stegosaurus's plates was put forward by Lucas6-8 and sees them mounted in two staggered rows along the animal’s back. There are, in fact, two conceivable forms consistent with this basic description: if the largest plate in particular tilts to the right we have an (R) Stegosaurus, if it tilts to the left we have an (L) Stegosaurus instead4. That Stegosaurus exhibited exterior chirality is beyond reasonable doubt: many of the plates are manifestly chiral by themselves and no two plates of the same size and shape have been found for an individual8-10. -
Body-Size Evolution in the Dinosauria
8 Body-Size Evolution in the Dinosauria Matthew T. Carrano Introduction The evolution of body size and its influence on organismal biology have received scientific attention since the earliest decades of evolutionary study (e.g., Cope, 1887, 1896; Thompson, 1917). Both paleontologists and neontologists have attempted to determine correlations between body size and numerous aspects of life history, with the ultimate goal of docu- menting both the predictive and causal connections involved (LaBarbera, 1986, 1989). These studies have generated an appreciation for the thor- oughgoing interrelationships between body size and nearly every sig- nificant facet of organismal biology, including metabolism (Lindstedt & Calder, 1981; Schmidt-Nielsen, 1984; McNab, 1989), population ecology (Damuth, 1981; Juanes, 1986; Gittleman & Purvis, 1998), locomotion (Mc- Mahon, 1975; Biewener, 1989; Alexander, 1996), and reproduction (Alex- ander, 1996). An enduring focus of these studies has been Cope’s Rule, the notion that body size tends to increase over time within lineages (Kurtén, 1953; Stanley, 1973; Polly, 1998). Such an observation has been made regarding many different clades but has been examined specifically in only a few (MacFadden, 1986; Arnold et al., 1995; Jablonski, 1996, 1997; Trammer & Kaim, 1997, 1999; Alroy, 1998). The discordant results of such analyses have underscored two points: (1) Cope’s Rule does not apply universally to all groups; and (2) even when present, size increases in different clades may reflect very different underlying processes. Thus, the question, “does Cope’s Rule exist?” is better parsed into two questions: “to which groups does Cope’s Rule apply?” and “what process is responsible for it in each?” Several recent works (McShea, 1994, 2000; Jablonski, 1997; Alroy, 1998, 2000a, 2000b) have begun to address these more specific questions, attempting to quantify patterns of body-size evolution in a phylogenetic (rather than strictly temporal) context, as well as developing methods for interpreting the resultant patterns. -
A New Phylogeny of Stegosauria (Dinosauria, Ornithischia)
Raven, T. J. and Maidment, S. C. R. In press. Palaeontology A new phylogeny of Stegosauria (Dinosauria, Ornithischia) THOMAS J. RAVEN1 AND SUSANNAH C. R. MAIDMENT2* 1Department of Earth Science and Engineering, Imperial College London, South Kensington Campus, London SW7 2AZ, United Kingdom, [email protected]; 2School of Environment and Technology, University of Brighton, Lewes Road, Brighton BN2 4GJ, United Kingdom, [email protected] * Corresponding author 1 Raven, T. J. and Maidment, S. C. R. In press. Palaeontology ABSTRACT The stegosaurs are some of the most easily recognisable dinosaurs, but they are surprisingly rare as fossils. Consequently much remains unknown about their palaeobiology, and every new stegosaurian find contributes to understanding the evolution of the clade. Since the last attempt to examine the evolutionary relationships of Stegosauria, new specimens have come to light, including the most complete individual of Stegosaurus ever found, new taxa have been described and, perhaps most importantly, new methods for analysis of cladistic datasets have been produced. In the light of these new data and technological advances, the phylogenetic relationships of the stegosaurs and basal armoured dinosaurs are investigated. The inclusion of continuous data results in much better resolution than was previously obtained, and the resulting single most parsimonious tree supports re-erection of the genera Miragaia and Hesperosaurus, which had previously been synonymized with Dacentrurus and Stegosaurus respectively. The recently described genus Alcovasaurus is resolved as a basal thyreophoran, but this is likely to be due to a combination of a very high degree of missing data and the questionable ontogenetic stage of the specimen.