Butterflies of Chizami Naro & Sondhi

:ŽƵƌŶĂůŽĨdŚƌĞĂƚĞŶĞĚdĂǆĂͮǁǁǁ͘ƚŚƌĞĂƚĞŶĞĚƚĂǆĂ͘ŽƌŐͮϭϮĞĐĞŵďĞƌϮϬϭϰͮϲ;ϭϯͿ͗ϲϱϵϯʹϲϲϯϰ

çããÙ¥½®Ý;>Ö®ÊÖãÙͿÊ¥«®þÃ®͕W«»®ÝãÙ®ã͕ DÊÄÊ¦ÙÖ« E¦½Ä͕/Ä® dƐŚĞƚƐŚŽůŽEĂƌŽϭΘ^ĂŶũĂǇ^ŽŶĚŚŝϮ ISSN 0974-7907 (Online) /^^EϬϵϳϰͲϳϴϵϯ;WƌŝŶƚͿ 1 North East Network, P.O. Chizami, Phek District, Nagaland 797102, India 2 dŝƚůŝdƌƵƐƚ͕ϰϵZĂũƉƵƌZŽĂĚŶĐůĂǀĞ͕ŚŽƌĂŶ<ŚĂƐ͕ĞŚƌĂĚƵŶ͕hƩĂƌĂŬŚĂŶĚϮϰϴϬϬϭ͕/ŶĚŝĂ KWE^^ 1 [email protected], 2 [email protected] (corresponding author)

ďƐƚƌĂĐƚ͗dŚĞďƵƩĞƌŇŝĞƐŽĨŚŝǌĂŵŝ͕WŚĞŬŝƐƚƌŝĐƚŽĨEĂŐĂůĂŶĚ͕/ŶĚŝĂǁĞƌĞĚŽĐƵŵĞŶƚĞĚŽǀĞƌĂƉĞƌŝŽĚŽĨƚŚƌĞĞǇĞĂƌƐ;ϮϬϭϭʹϮϬϭϰͿ͘dŚŝƐ ƌĞƉƌĞƐĞŶƚƐƚŚĞĮƌƐƚůŝƐƚŽĨďƵƩĞƌŇǇƌĞĐŽƌĚƐĨƌŽŵEĂŐĂůĂŶĚĂŌĞƌƉƵďůŝĐĂƟŽŶƐďǇdǇƚůĞƌďĞƚǁĞĞŶϭϵϭϭĂŶĚϭϵϭϰ͘dǁŽͲŚƵŶĚƌĞĚͲĂŶĚͲƚǁĞůǀĞ ;ϮϭϮͿƐƉĞĐŝĞƐǁĞƌĞƌĞĐŽƌĚĞĚŽĨǁŚŝĐŚϯϰƐƉĞĐŝĞƐĂƌĞůĞŐĂůůǇƉƌŽƚĞĐƚĞĚƵŶĚĞƌǀĂƌŝŽƵƐƐĐŚĞĚƵůĞƐŽĨƚŚĞtŝůĚůŝĨĞ;WƌŽƚĞĐƟŽŶͿĐƚ͕ϭϵϳϮ͘ ŵŽŶŐƐƚƚŚĞŵĂŶǇƐŝŐŶŝĮĐĂŶƚƌĞĐŽƌĚƐĚƵƌŝŶŐƚŚĞƐƵƌǀĞǇ͕ƚǁŽƐƉĞĐŝĞƐ͕ƚŚĞdĂǁŶǇŵƉĞƌŽƌChitoria ulupi ulupiĂŶĚƚŚĞŽŵŝĐKĂŬďůƵĞ Arhopala comicaǁĞƌĞƌĞĐŽƌĚĞĚĨƌŽŵ/ŶĚŝĂĂŌĞƌĂƉĞƌŝŽĚŽĨϭϬϬǇĞĂƌƐ͘

<ĞǇǁŽƌĚƐ͗ƵƩĞƌŇǇĚŝǀĞƌƐŝƚǇ͕ĐŚĞĐŬůŝƐƚ͕ŚŝǌĂŵŝ͕Arhopala comica͕ŝŶǀĞŶƚŽƌǇ͕>ĞƉŝĚŽƉƚĞƌĂ͕EĂŐĂůĂŶĚ͕ƌĞĐŽƌĚ͕Chitoria ulupi.

Junonia almana almana Peacock Pansy

NOT DATA LEAST NEAR CRITICALLY EXTINCT VULNERABLE ENDANGERED EXTINCT EVALUATED DEFICIENT CONCERN THREATENED ENDANGERED IN THE WILD NE DD LC NT VU EN CR EW EX

K/͗ŚƩƉ͗ͬͬĚǆ͘ĚŽŝ͘ŽƌŐͬϭϬ͘ϭϭϲϬϵͬ:Ždd͘Žϯϵϵϱ͘ϲϱϵϯͲϲϯϰͮŽŽĂŶŬ͗ƵƌŶ͗ůƐŝĚ͗ǌŽŽďĂŶŬ͘ŽƌŐ͗ƉƵď͗ϵϮ&ϱϳϲͲϯͲϰϮϮͲϰϯͲϴϭϳϮϯ&ϬϬ

ĚŝƚŽƌ͗ŶŽŶǇŵŝƚǇƌĞƋƵĞƐƚĞĚ͘ ĂƚĞŽĨƉƵďůŝĐĂƟŽŶ͗ϭϮĞĐĞŵďĞƌϮϬϭϰ;ŽŶůŝŶĞΘƉƌŝŶƚͿ

DĂŶƵƐĐƌŝƉƚĚĞƚĂŝůƐ͗DƐηŽϯϵϵϱͮZĞĐĞŝǀĞĚϭϮƉƌŝůϮϬϭϰͮ&ŝŶĂůƌĞĐĞŝǀĞĚϭϲKĐƚŽďĞƌϮϬϭϰͮ&ŝŶĂůůǇĂĐĐĞƉƚĞĚϮϴEŽǀĞŵďĞƌϮϬϭϰ

ŝƚĂƟŽŶ͗EĂƌŽ͕d͘Θ^͘^ŽŶĚŚŝ(2014). ƵƩĞƌŇŝĞƐ;>ĞƉŝĚŽƉƚĞƌĂͿŽĨŚŝǌĂŵŝ͕WŚĞŬŝƐƚƌŝĐƚ͕EĂŐĂůĂŶĚ͕/ŶĚŝĂ. Journal of Threatened Taxaϲ;ϭϯͿ͗ϲϱϵϯʹϲϲϯϰ; ŚƩƉ͗ͬͬ Ěǆ͘ĚŽŝ͘ŽƌŐͬϭϬ͘ϭϭϲϬϵͬ:Ždd͘Žϯϵϵϱ͘ϲϱϵϯͲϲϯϰ

ŽƉǇƌŝŐŚƚ͗ ©EĂƌŽΘ^ŽŶĚŚŝϮϬϭϰ͘ƌĞĂƟǀĞŽŵŵŽŶƐƩƌŝďƵƟŽŶϰ͘Ϭ/ŶƚĞƌŶĂƟŽŶĂů>ŝĐĞŶƐĞ͘:ŽddĂůůŽǁƐƵŶƌĞƐƚƌŝĐƚĞĚƵƐĞŽĨƚŚŝƐĂƌƟĐůĞŝŶĂŶǇŵĞĚŝƵŵ͕ƌĞƉƌŽĚƵĐ- ƟŽŶĂŶĚĚŝƐƚƌŝďƵƟŽŶďǇƉƌŽǀŝĚŝŶŐĂĚĞƋƵĂƚĞĐƌĞĚŝƚƚŽƚŚĞĂƵƚŚŽƌƐĂŶĚƚŚĞƐŽƵƌĐĞŽĨƉƵďůŝĐĂƟŽŶ͘

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ŽŵƉĞƟŶŐ/ŶƚĞƌĞƐƚ͗dŚĞĂƵƚŚŽƌƐĚĞĐůĂƌĞŶŽĐŽŵƉĞƟŶŐŝŶƚĞƌĞƐƚƐ͘

ƵƚŚŽƌŽŶƚƌŝďƵƟŽŶƐ͗dEƐƵƌǀĞǇĞĚďƵƩĞƌŇŝĞƐŝŶƚŚĞĂƌĞĂĨƌŽŵϮϬϭϭƚŽϮϬϭϰ͘dEĂŶĚŵĞŵďĞƌƐŽĨƚŚĞ,'ƚŽŽŬŵŽƐƚŽĨƚŚĞƉŚŽƚŽŐƌĂƉŚƐŽĨƚŚŝƐƉĂƉĞƌ͘^^ŵĂĚĞ ĮǀĞǀŝƐŝƚƐƚŽŚŝǌĂŵŝĂŶĚƐƵƌǀĞǇĞĚďƵƩĞƌŇŝĞƐĚƵƌŝŶŐŚŝƐǀŝƐŝƚƐ͘dEŝĚĞŶƟĮĞĚŵĂŶǇŽĨƚŚĞƐƉĞĐŝĞƐ͕ĂŶĚ^^ĐŽŶĮƌŵĞĚƚŚĞŝƌŝĚĞŶƟƟĞƐ͘^^ĐŽŶĚƵĐƚĞĚƚŚĞůŝƚĞƌĂƚƵƌĞ ƌĞǀŝĞǁ͘dŚĞƉĂƉĞƌǁĂƐǁƌŝƩĞŶũŽŝŶƚůǇǁŝƚŚďŽƚŚĂƵƚŚŽƌƐĐŽŶƚƌŝďƵƟŶŐƚŚĞŝƌƌĞĐŽƌĚƐ͘dEƉƌĞƉĂƌĞĚƚŚĞƉůĂƚĞƐ͘

ƵƚŚŽƌĞƚĂŝůƐ͗ dÝ«ãÝ«Ê½ÊEÙÊ;ůŽͿŝƐĂǇŽƵŶŐŶĂƚƵƌĞĐŽŶƐĞƌǀĂƟŽŶŝƐƚďĂƐĞĚŝŶŚŝǌĂŵŝ͕WŚĞŬŝƐƚƌŝĐƚ͕EĂŐĂůĂŶĚĂŶĚĐƵƌƌĞŶƚůǇǁŽƌŬŝŶŐǁŝƚŚĂŶE'K͕EŽƌƚŚĂƐƚ EĞƚǁŽƌŬ;EEͿ͕ŝŶŝƚƐEĂƚƵƌĞŽŶƐĞƌǀĂƟŽŶĚƵĐĂƟŽŶWƌŽŐƌĂŵŵĞǁŚŝĐŚŝŶǀŽůǀĞƐĐŽŶƐƚĂŶƚĞŶŐĂŐĞŵĞŶƚǁŝƚŚǇŽƵƚŚƐĂŶĚĐŚŝůĚƌĞŶƚŽǁĂƌĚƐĐƌĞĂƟŶŐĂĐŽŶƐĐŝŽƵƐŶĞƐƐ ĨŽƌĐŽŶƐĞƌǀĂƟŽŶ͘ůŽŚĂƐĂŶĞŶƚŚƵƐŝĂƐƟĐŝŶƚĞƌĞƐƚŝŶƐƚƵĚǇŝŶŐďƵƩĞƌŇŝĞƐĂŶĚŚĂƐďĞĞŶŝĚĞŶƟĨǇŝŶŐďƵƩĞƌŇŝĞƐŝŶĂŶĚĂƌŽƵŶĚŚŝǌĂŵŝƐŝŶĐĞϮϬϭϭ͖ŚĞĂůƐŽŚĂƐŬĞĞŶ interest in nature photography and videography. ^Ä¹ù^ÊÄ«®͕&ŽƵŶĚĞƌdƌƵƐƚĞĞdŝƚůŝdƌƵƐƚ͕ŝƐĂŶĂƚƵƌĂůŝƐƚǁŝƚŚĂŶŝŶƚĞƌĞƐƚŝŶ>ĞƉŝĚŽƉƚĞƌĂ͕ŚĞƌƉĞƚŽĨĂƵŶĂĂŶĚ ĂǀŝĨĂƵŶĂ͘,ĞƐƵƉƉŽƌƚƐĐŽŶƐĞƌǀĂƟŽŶƌĞƐĞĂƌĐŚĂŶĚĂĐƟŽŶĂŶĚĐŽŶƐĞƌǀĂƟŽŶĞĚƵĐĂƟŽŶƉƌŽŐƌĂŵƐŝŶĞĂƐƚĞƌŶĂŶĚǁĞƐƚĞƌŶ,ŝŵĂůĂǇĂ͘

ĐŬŶŽǁůĞĚŐĞŵĞŶƚƐ͗dEƚŚĂŶŬƐƚŚĞEŽƌƚŚĂƐƚEĞƚǁŽƌŬĨŽƌƚŚĞŽƉƉŽƌƚƵŶŝƚǇƚŚĞǇŚĂǀĞŐŝǀĞŶŚŝŵƚŽŐĞƚŝŶǀŽůǀĞĚŝŶŶĂƚƵƌĞĐŽŶƐĞƌǀĂƟŽŶĞĚƵĐĂƟŽŶƉƌŽŐƌĂŵŵĞ ĂŶĚŝŶŵĂŶǇŽƚŚĞƌǁĂǇƐ͘^^ĂĐŬŶŽǁůĞĚŐĞƐƚŚĞƐƵƉƉŽƌƚŐŝǀĞŶƚŽŚŝŵďǇEŽƌƚŚĂƐƚEĞƚǁŽƌŬĚƵƌŝŶŐŚŝƐǀĂƌŝŽƵƐǀŝƐŝƚƐƚŽEĂŐĂůĂŶĚ͘dŚĞĂƵƚŚŽƌƐǁŽƵůĚůŝŬĞƚŽƚŚĂŶŬ ƚŚĞƌĞǀŝĞǁĞƌƐĨŽƌƚŚĞŝƌĨĞĞĚďĂĐŬĂŶĚĐŽŵŵĞŶƚƐ͘

6593 Butterflies of Chizami Naro & Sondhi

NCN records in this paper are through photographs taken by members of the HGEC. The hill state of Nagaland has a geographical area of 16,579km2 of which 30% is under forest with 88% of this being private forests. Nagaland falls in S the eastern Himalaya region of the Himalaya Global Biodiversity Hotspot, one of 34 global biodiversity The authors conducted butterfly surveys in the hotspots. Nagaland is home to a variety of forest types area over a period of three years. The butterfly study (altitudes vary from 184–3,841 m) varying from northern was mainly conducted in the following areas (Fig. 2). tropical wet evergreen forest, northern tropical semi- (i) NEN compound (25024’0’’N & 94024’E, 1440m); (ii) evergreen forest, northern sub-tropical broad-leaved In and around Chizami Village; (iii) The road leading to wet hill forest (500–1,800 m), northern sub-tropical Thetsumi Village approximately 5km away from Chizami pine forest (1,000–1,500 m), northern montane wet Village; (iv) The Ewelu path up to approximately 3km temperate forest (2,000–2,500 m) and temperate from Chizami Village; (v) Erule River bank - a mini hydro forest (above 2,500m) (http://ibcn.in/wp-content/ project area 10km away from Chizami Village. Areas uploads/2011/12/33-793_812- Nagaland.pdf). that were primarily visited are listed below along with Chizami is located in the Phek District in southeastern photographs of their habitats (Appendix 1). Nagaland. Phek District, covering an area of 2,026km2 More than 50% of the butterflies seen were in the NEN is bordered by Myanmar on the east and the state compound. TN lives in Chizami, and opportunistically of Manipur on the south. To its north are Kiphire and recorded and photographed butterflies in the area Zunheboto districts, while Kohima District is to its west. beginning 2010. SS made five visits to Chizami in July The village of Chizami is 88km due east of Kohima (Fig. 2011, February 2012, October 2012, January 2013 1). It covers an area of 85,242ha approximately and the altitudinal range varies from 981–2000 m. The forest type at Chizami is mainly sub-tropical pine forest and some northern sub-tropical broad-leaved wet hill forest. The Chakhesang tribe inhabits Chizami. Chizami village shares its boundary with Pholami villages on the north; Khomi, Sumi and Losami Village on the northeast; Lai Village of Manipur State on the east, Thetsumi Village on the south; Enhulumi Village on the west and Porba and Sakraba villages on the northwest. The Kuwari River is to the east and the Tuphale River to the west of Chizami. North East Network (NEN) (www.northeastnetwork. org) is a women’s rights organization working in the northeastern region of India with a focus on women’s rights. NEN has also been working on the issue of natural resource management in the area. As part of the Natural Resource Management activity, wildlife and biodiversity conservation education programme was launched in 2010 and the Hoolock Gibbon Eco Club was formed. The Hoolock Gibbon Eco Club (HGEC) is a children’s eco club consisting of 40 members from the villages of Chizami and Enhulumi between the age group of 10–14 years. The HGEC is focused on conservation education, conservation research and conservation action. As part of the HGEC’s activities, primarily Tshetsholo Naro (TN), Kewekhrozo Thopi, NEN and Sanjay Sondhi (SS), Titli Trust (www.titlitrust.com), introduced the children to butterfly watching and identification. Quite a few of the iure a of Naaand he isri Chizami

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

iure a of Chizami and surroundin areas NN omound n and around Chizami iae hesumi oad eu ah rue ier and March 2013 during which he surveyed butterflies older literature. Amongst the earliest publications on along with TN. The butterflies were photographed and butterflies from the Naga Hills was the publication by identified using existing literature (Marshall 1882–90; Elwes on the collection of butterflies by Mr. W. Doherty Bingham 1905–07; Swinhoe 1912–13; Evans 1932, during two expeditions in 1889 and 1890 (Elwes 1891). 1949; Talbot 1939, 1947; Wynter-Blyth 1957; Eliot 1969; In 1889, Doherty spent the rainy season collecting in the Corbet et. al 1992; Haribal 1992; Smith 1993; Smith Naga Hills from June to September. Subsequent to Mr. 1994; Kunte 2000; Kinyon 2004; Kehimkar 2008; Singh Doherty’s butterfly collection in the Naga Hills, Major 2011; Sondhi et. al 2013; Sondhi & Kunte 2014). Cryptic H.C. Tytler spent many years collecting and reporting species were netted and their external features relevant on the butterflies of the Naga Hills (Tytler 1911, 1912). for identification were observed and photographed. Tytler’s collection covered the lower, medium and higher Wherever possible, the identification of the individuals altitudes of the Naga Hills covering areas between 200 were done up to the subspecies level. In numerous and 2,100 m. Later, Tytler, who had by then become a species, there are different common names in use. Lieutenant Colonel, was posted in Imphal in Manipur and Alternate common names, if in use, are mentioned in further collected butterflies from Manipur and Naga hills brackets. (Tytler 1914, 1915a,b). In this paper, we will be referring to only these species recorded from the Naga Hills. Amongst the significant records during the survey, SS N SCSSN two species, the Tawny Emperor Chitoria ulupi ulupi and the Comic Oakblue Arhopala comica have been recorded Two-hundred-and-twelve (212) species were from India after 100 years, having last been recorded recorded during the survey. The family Nymphalidae during surveys by Tytler (1915a,b). The Comic Oakblue with 102 species representing 48% of the species seen Arhopala comica has previously only been known from a was the most abundant family followed by Lycaenidae few specimens collected in Manipur, hence this record with 36 species (17%) and Hesperiidae with 30 species extends its range marginally to Nagaland. There are (14%). Of the 212 species recorded in Chizami, 34 numerous others species and subspecies for which there species are legally protected under various schedules of exist no or very few recent published records from India the Wildlife (Protection) Act, 1972 (Table 1). such as Assam Darter Ochlodes siva siva, Purple Swift Surveys and collections of butterflies from the Naga Caltoris tulsi, Ruddy Pied Flat Pseudocoladenia fatua, Hills are scanty and Varshney’s bibliography (Varshney Khaki Silverline Spindasis rukmini, Hedge Cupid Bothrinia 2008, 2009) was consulted along with SS’s perusal of chennelli, Blue Duke Euthalia durga splendens, and

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 6595 Butterflies of Chizami Naro & Sondhi

ae Seies reorded from Chizami oered in shedues should be noted that while this subspecies is the one that is most commonly recorded in northeastern India, Shedue Nmhaidae Aemona amathusia species of this genera are best separated by examining aenidae Calinaga brahma brahma male genitalia. Rare. Reported from the Naga Hills Arhopala comica Euthalia franciae franciae Spindasis rukmini Euthalia aconthea garuda (Elwes 1892). Several individuals collected at Kohima Faunis eumeus assama (Tytler 1912). Nmhaidae Hestina persimilis persimilis Bassarona durga splendens Parthenos sylvia gambrisius Chitoria ulupi ulupi Kallima knyvettii Sae Hasora anura anura de Niie Dilipia morgiana Lethe sinorix sinorix Mimathyma chevana chevana Libythea lepita lepita Recorded at NEN compound in November 2011. Sephisa Chandra Chandra Melanitis zitenius zitenius Very rare. Mycalesis mestra sadona Shedue Symbrenthia niphanda niphanda hieanded Hasora taminatus bhavara aenidae ieridae Anthene lycaenina lycambes Appias albina darada ruhsorfer Arhopala ganesa watsoni Bothrinia chennelli Shedue Recorded from the NEN compound in the month of Euchrysops cnejus cnejus eseriidae July 2012. Rare. Flos areste Hyarotis adrastus praba Heliophorus moorei tytleri Jamides elpis pseudoelpis Nmhaidae ami eseriidae Sufami eseriinae Lampides boeticus Euploea mulciber mulciber Buesotted Sru oer Aeromachus kali de Poritia hewitsoni hewitsoni Euthalia lubentina lubentina Niie ieridae Appias galba Recorded on the Chizami road leading to Enhulumi Village in May 2012. Rare. Reported to be common in the Naga Hills (Elwes 1892; Tytler 1912).

Green Duke Euthalia sahadeva nadaka. These species eined Sru oer Aeromachus stigmata obsoleta are discussed in more detail in the notes below. oore The present work is significant because it reports Recorded from the NEN compound and the Erule numerous species not recorded by Tytler and Elwes River at Chizami during the post-monsoon season. during their extensive collections of butterflies in the Uncommon. Reported from the Naga Hills (Elwes 1892; Naga Hills. Equally significant, there are more than a Tytler 1912). hundred species that were collected by them that were not recorded at Chizami, though the survey area and ores oer Astictopterus jama olivascens oore elevations covered during their surveys were far more extensive. Recorded from the Erule River and the Thetsumi trail at Chizami between March and July in the pre-monsoon NS N BS C season. Common. Reported from the Naga Hills (Elwes 1892; Tytler 1912). ami eseriidae Sufami Coeiadinae Bron Badamia exclamationis ariius ure Si Caltoris tulsi de Niie Recorded from the NEN compound in June 2012 and Recorded from the NEN compound in late November July 2013 at the same location. Rare. Reported to be not 2011. Few recent published records from India. Rare. uncommon at Kohima (Tytler 1912). Sraih Si Parnara f guttatus Bremer & re reen e Bibasis vasutana oore Recorded from the NEN compound in October Recorded in the flower garden at NEN compound in 2011. Rare. Reported from the Naga Hills (Elwes 1892). August 2011 and October 2012. Members of this genus Reported to be common at Kohima (Tytler 1912). are best separated by male genitalia; hence this has only been provisionally identified. Common. Reported from ndian in Choaspes benjaminii japonica the Naga Hills (Elwes 1892). Reported to be not common urra at Kohima (Tytler 1912). Recorded from near the stream on the Thetsumi roadside at Chizami in July and November 2011. It

6596 Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

Sma Branded Si Pelopidas mathias mathias ndian am Bo Suastus gremius gremius ariius ariius Recorded from the NEN compound in the flower Single record from Erule River bank at Chizami in garden in August 2011. Uncommon. August 2013. Very rare in the survey area.

are Branded Si Pelopidas sinensis aie Siim am edee Erionota torus ans Recorded from all locations at Chizami. Separated Recorded from the NEN compound in the month from similar-looking E. thrax by the rounded FW apex of February 2012. Reported from the Naga Hills (Elwes and convex termen, though it is pertinent to note that 1892). Reported to be common at Kohima (Tytler 1912). this species is best separated from E. thrax by male genitalia. Mostly seen during the pre-monsoon season. eo So Si Polytremis eltola eltola eison An individual entered the training room at night on two consecutive nights in July 2011, evidently attracted Recorded from all locations at Chizami. Mostly seen to light. Common. Erionota thrax reported from the during the pre- and post- monsoon seasons. Common. Naga Hills with individuals captured at Kohima inside a Reported to be common at Kohima (Tytler 1912). bungalow, apparently attracted to light, just as we had observed (Tytler 1912). However, Evans described E. ar s Potanthus s torus as a separate species from E. thrax only in 1941, Numerous records from the Chizami area in the pre- hence Tytler’s records could have been either of these and post-monsoon seasons. Numerous species reported two species. from the Naga Hills (Elwes 1892). Baeined Branded edee Matapa cf. sasivarna ssam arer Ochlodes siva siva oore oore Two individuals sighted by TN. Seen 400m away from Recorded on the Chizami road leading to Thetsumi NEN office (West) in May 2010 and at the NEN compound in October. The upperside of this individual was in May 2014. The first individual was a female, lacking not observed. This species could be M. cresta, M. the UPF brand. However, UPH discal spots were as purpurascens or M. sasivarna. In the field, these species opaque as the opaque spots in space 1b UPF and not are best separated by a combination of the male UPF hyaline as are the other spots UPF, separating it from brands and wing markings and coloration. While in M. Ochlodes brahma (Evans 1949). In addition, the spot in cresta and M. purpurascens the male brand is black and 1b in O. brahma is much smaller. Rare. These are the inconspicuous, in M. sasivarna, it is grey and conspicuous. first records of this species from Nagaland, having been However, on the basis of the absence of paler UNF apex recorded from Khasi Hills in Meghalaya, Mussoorie in and UNH base, present in M. cresta, and the metallic- Uttarakhand, Assam and north Myanmar (Evans 1949). green scales at HW base and wing coloration, this is These sightings are also the first published record of this provisionally identified as M. sasivarna (de Jong 1983; species from India in many decades. Evans 1949). Very rare.

ih Sra e Pithauria stramineipennis esried emon Notocrypta curvifascia curvifascia stramineipennis oodason & de Niie eder & eder Single record from Erule River bank at Chizami in Recorded from all locations at Chizami. Mostly seen August 2013. SS had observed this species in the foothills in the post-monsoon season. Common. Reported to be from other parts of northeastern India, while it has been common at Kohima (Tytler 1911). recorded up to 640m in Nepal (Smith 1994), hence this record from 700m is unusual. Very rare in the survey Sotted emon Notocrypta feisthamelii alysos area. Reported from the Naga Hills at low elevations in oore March and October (Tytler 1912). Recorded on the Chizami to Awelu trail near the stream in the pre-monsoon season. Uncommon. ued e Sebastonyma dolopia eison Reported to be common at Kohima (Tytler 1912). Recorded from the NEN compound in October 2012. Rare in the survey area. Reported from the Naga Hills (Tytler 1912).

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

Common Banded emon Notocrypta paralysos an ne Ctenoptilum vasava vasava oore asawa ruhsrofer Single record from the NEN compound in July 2011. Recorded from the Erule River at Chizami in March Rare in the survey area. 2012. Rare. A single male collected from Nichuguard in the Naga Hills (Tytler 1912). ree itter Hyarotis adrastus praba oore Recorded on the Chizami to Erule trail in the pre- Chesnu ne Odontoptilum angulata angulata monsoon season. Uncommon. This subspecies is legally eder protected under Schedule IV of the Wildlife (Protection) Recorded from the Erule River at Chizami in March Act, 1972. Reported from the Naga Hills (Tytler 1912). 2013. Rare in the survey area. Reported to be common from the Naga Hills at low elevations (Tytler 1912). ami eseriidae Sufami rinae Common Sotted a Celaenorrhinus leucocera ami aenidae Sufami Curenae oar Brih Suneam Curetis bulis bulis esood Worn individuals were recorded from the NEN Recorded two individuals at the NEN compound and compound in October 2012. Uncommon. Reported on the Erule trail at Chizami in October 2012. Rare in the from the Naga Hills in September (Tytler 1912). survey area. Reported from the Naga Hills (Elwes 1892; Tytler 1912). udd ied a Pseudocoladenia cf. fatua ans ami aenidae Sufami aeninae Recorded from the Thetsumi roadside near the stream oden Sahire Heliophorus brahma brahma and at the NEN compound in May and June. Rare. This oore species most closely resembles P. fatua with the spot in Recorded from all locations at Chizami, throughout two extended to beyond origin of vein 3, UPF, and being the year, though less common in the winter months. brighter, similar to the West Himalayan species P. fatih Very common. Reported to be common in the Naga Hills and its wings being more produced than P. festa which (Elwes 1892; Tytler 1912). also occurs at similar altitudes (Evans 1949). The similar looking P. dan is more of a low elevation species. It is Naa Sahire Heliophorus kohimensis kohimensis pertinent to note that while Evans (1949) had treated er fatua and festa as a subspecies of Pseudocoladenia dan, Recorded from only two locations at Chizami. On subsequent work has elevated both these subspecies to the Ezilu road to Enhulumi and around NEN compound. species level (Huang & Xue, 2004). However, P. fatua Both males and females seen at NEN compound. Seen and P. festa are best separated by differences in male from May to October. Uncommon. Described as a new genitalia, hence this species is provisionally identified. species by Tytler from the Naga Hills (1911). Reported as Coladenia faith from Kohima in the Naga Hills in September and October between 1,800 and 2,130 reen Sahire zure Sahire Heliophorus m (Tytler 1915b), and possibly represents the species P. moorei tytleri ie fatua as described by Evans (1949). Recorded from the Government High school area and NEN compound. Very common. This species is legally Sotted Sno a Tagiades menaka menaka protected under Schedule II of the Wildlife (Protection) oore Act, 1972. Reported as very abundant in the Naga Hills Recorded from the NEN compound in October 2011. (Elwes 1892). Tytler (1911) reports it to be the most Rare. Reported from the Naga Hills (Elwes 1892; Tytler common of the genus in the Naga Hills. 1912). ami aenidae Sufami oommanae Sotted neCaprona agama agama oore oined Ciiae Bue Anthene lycaenina lycambes Recorded from the Erule River at Chizami in March eison 2012. Rare. A single record from the Erule River at Chizami in July 2012. Seen mud-puddling at the riverside. Very rare in the survey area. This species is legally protected under Schedule II of the Wildlife (Protection) Act, 1972.

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

Reported from the Naga Hills (Elwes 1892; Tytler 1912). ram Bue Euchrysops cnejus cnejus ariius A single record from the NEN compound at Chizami Common ede Bue Acytolepis puspa gisca in May 2012. Very rare in the survey area. This species ruhsorfer is legally protected under Schedule II of the Wildlife Recorded from all locations at Chizami. Very (Protection) Act, 1972. common. Reported to be very common from the Naga Hills (Elwes 1892; Tytler 1912). ar Ceruean Jamides bochus bochus So Recorded from all locations at Chizami. Mostly seen in ain ede Bue Celastrina lavendularis limbata May. Very common. Reported from the Naga Hills (Elwes oore 1892; Tytler 1912). A single record by SS in July 2011. Very rare in the survey area. Reported from the Naga Hills (Elwes 1892). Ceruean Jamides seies in S A butterfly was observed by TN at Chizami in ae ede Bue Udara dilecta dilecta oore November 2012, which looked like a Jamides in DSF and Recorded from the NEN compound and Erule River could not be identified to the species level. The upperside at Chizami. Only two individuals seen in October 2011. markings were not observed in the field. This individual Rare in the survey area. Reported from the Naga Hills looked superficially like J. celeno in DSF, but the UNH (Elwes 1892). discal markings were narrower and broken, somewhat like a Nacaduba species, compared with individuals ede Cuid Bothrinia chennelli de Niie observed by SS in other areas of the Himalayas. Recorded from the Erule trail at Chizami. Seen in October 2011 (TN) and March 2013 (SS). Rare. This isenin Ceruean Jamides elpis pseudoelpis species is legally protected under Schedule II of the Buer Wildlife (Protection) Act, 1972. Reported from the Naga Recorded at Chizami near the village (east of Hills (Elwes 1892). Reported to be not common by Tytler Chizami). A single record in the month of January 2011. (1911). Very rare in the survey area. This species is legally protected under Schedule II of the Wildlife (Protection) Chamans Cuid Everes argiades diporides Act, 1972. Reported to be common from the Naga Hills Chaman (Elwes 1892; Tytler 1912). Single record from the NEN compound at Chizami in the month of October 2011. Very rare in the survey area. ea Bue Lampides boeticus innaeus Reported from the Naga Hills (Elwes 1892; Tytler 1912). Recorded from all locations at Chizami. Most common in the pre-monsoon season. Visits flowers. ndian Cuid Everes lacturnus assamica er Very common. This species is legally protected under Recorded from the Erule River and the NEN Schedule II of the Wildlife (Protection) Act, 1972. compound in the pre- and post-monsoon seasons. Rare Reported to be very common from the Naga Hills (Tytler in the survey area. Reported from the Naga Hills (Tytler 1912). 1912). era Bue Leptotes plinius plinius ariius Sier oremeno Catochrysops panormus Recorded from all locations at Chizami in April and exiguus isan May. Common. A single individual collected at Kohima Recorded from two locations at Chizami; the NEN (Tytler 1912). compound and on the Erule trail. Seen mostly during summer. Rare in the survey area. Reported from the in ineue Petrelaea dana de Niie Naga Hills (Tytler 1912). A single record from the Erule riverside at Chizami in October 2011. Very rare in the survey area. Reported oremeno Catochrysops strabo strabo ariius from the Naga Hills (Tytler 1912). Recorded from the NEN compound. Two individuals Common ineue Prosotas nora ardates oore seen. Rare in the survey area. Reported from the Naga Hills (Elwes 1892; Tytler 1912). Mostly recorded from the Erule riverside, mud-

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 6599 Butterflies of Chizami Naro & Sondhi puddling in groups in the summer. Very common. in Manipur. This species is legally protected under Reported sparingly from 600m altitude in the Naga Hills Schedule I of the Wildlife (Protection) Act, 1972. Rare. (Tytler 1912). aiess Bushue Arhopala ganesa watsoni ans ar rass Bue Zizeeria karsandra oore Recorded from the NEN compound. Mostly seen A single record from the Thetsumi roadside at during summer. Common. Reported from the Naga Hills Chizami in January 2013. Very rare in the survey area. (Tytler 1912). This species is legally protected under Schedule II of the Wildlife (Protection) Act, 1972. ami aenidae Sufami oriinae Common em Poritia hewitsoni hewitsoni oore eodis aue Arhopala singla de Niie Recorded at Government High School area, Tahulu Single record from the New Town area at Chizami in trail at Chizami. Recorded two individuals at different September 2013. Very rare in the survey area. Reported locations in the month of June 2012. This subspecies from the Naga Hills (Elwes 1892; Tytler 1912). is legally protected under Schedule II of the Wildlife (Protection) Act, 1972. Rare. A few individuals collected aiess ushue Flos areste eison from Kohima (Tytler 1912). Recorded from the NEN compound and the Erule trail at Chizami in November 2011 and January 2013. ami aenidae Sufami heinae Four individuals sighted. Rare. This species is legally eaf Bue Amblypodia anita dina ruhsrofer protected under Schedule II of the Wildlife (Protection) This species was recorded on the Erule trail and mostly Act, 1972. seen mud-puddling near the riverside. Subspecies level identity could not be confirmed; this could also be the Common inse Catapaecilma major major rue ssp. gigantea, known from Manipur. Rare. Recorded only once at the Erule riverbank at Chizami Biso oa Ancema ctesia ctesia eison in March 2013. Very rare in the survey area. Reported A single record from the NEN compound in from the Naga Hills (Tytler 1912). September 2013. Very rare in the survey area. Reported to be common at 1,500m in the Naga Hills (Tytler 1912). Siersrea Bue Iraota timoleon timoleon So odered aue Arhopala bazalus teetsa de Recorded two individuals from the NEN compound Niie in May 2013. Very rare in the survey area. Reported A single record from NEN compound in November from the Naga Hills in May (Tytler 1915b). 2013. Very rare in the survey area. Reported from the Naga Hills (Elwes 1892). Common ash Rapala nissa nissoides Sinhoe Comi aue Arhopala comica de Niie Recorded from the NEN compound and the Erule Two records by TN from the NEN compound on 1 trail in the pre-monsoon season from February to July. April 2013 and 12 April 2014. Very rare. This Arhopala Rare. Reported from the Naga Hills (Elwes 1892) where species is easily told by the markings on the underside it is not as common as in the Khasi Hills. Tytler (1911) as well as the shape of the hindwing tornus and its tail reports it to be very common in Kohima throughout the (de Nicéville, 1900). The type locality of this species is year. Bhamo in Myanmar while it was recorded as Surendra learmondi (=Arhopala comica) from Loimwe in the hai Sierine Spindasis rukmini de Niie Southern Shan States (Tytler, 1940). From India, two Recorded from the North East Network office females of this species were recorded from Sebong, compound. Single sighting in October 2012. Very rare. Manipur in March (Tytler 1915b; Evans 1957). Hence this This species is legally protected under Schedule I of record of Arhopala comica marginally extends the range the Wildlife (Protection) Act, 1972. A single individual of this species into Nagaland, and is the first record from collected from the Naga Hills (Tytler 1912). India in a hundred years, since Tytler’s (1915b) collection

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

Cu Sierine Spindasis syama peguanus oore Couresan Euripus nyctelius nyctelius oueda Recorded from the Government High school area and Recorded from Khili and inside the village at Chizami in on the Erule trail at Chizami in the pre-monsoon season. September and October 2011 respectively. Uncommon. Rare in the survey area. Reported from the Naga Hills Reported to be common in the Naga Hills (Tytler 1911). (Elwes 1892; Tytler 1912). asha Herona marathus marathus oueda Common aia Bue Surendra quercetorum A single record from Sitikho at Chizami in October quercetorum oore 2011. Rare. Recorded from many different locations at Chizami through most of the year. Very common. Reported Cire Hestinalis nama nama oueda from the Naga Hills (Elwes 1892; Tytler 1912). Recorded from many different locations at Chizami in the pre- and post-monsoon seasons. Very common. ami Nmhaidae Sufami aurinae Reported to be very common in the Naga Hills (Tytler ure meror Apatura ambica ambica oar 1911). Recorded from the Thetsumi trail and Erule riverside Siren Hestina persimilis persimilisesood at Chizami. Seen sitting in groups, mud-puddling. Two individuals recorded from NEN compound and Common. Reported to be common in the Naga foothills Chizami village in Aug 2011 and July 2013 respectively. up to 1,370m (Tytler 1911). This species is legally protected under Schedule II of the Wildlife (Protection) Act, 1972. Rare. A single individual an meror Chitoria ulupi ulupi oher recorded in October in the Naga Hills (Tytler 1911). Recorded by TN from the NEN compound on 14 June 2013 at 12:57hr. Seen sitting on human clothes. Ba rine Rohana parisatis parisatis esood This species is a powerful flier. There have been no published records of this species from India in more Recorded from the Erule riverside at Chizami in than a hundred years, having been last reported from October 2012. Seen sitting on moist stones and mud- India in 1915 (Tytler 1915a). Very rare. The subspecies puddling. Common. Males reported to be exceedingly florenciae, which is now synonymized with subspecies common in the Naga Hills (Tytler 1911). ulupi, is legally protected under Schedule I of the Wildlife (Protection) Act, 1972. Reported to be rare in the Naga asern Courer Sephisa chandra chandra oore Hills (Tytler 1911, 1915a). Tytler (1911) initially believed this to be a separate species from C. ulupi but later Recorded from two locations at Chizami-the stated (Tytler 1915a) that it would turn out to be a race Thetsumi trail and the Erule riverside in October 2011 of the same species. and October 2012. Rare. This subspecies is legally protected under Schedule I of the Wildlife (Protection) oden meror Dilipia morgiana esood Act, 1972. Numerous records from the Naga Hills (Tytler Recorded from the Thetsumi trail and the Erule 1911). riverside at Chizami in October 2012 and August 2013 respectively. Common. Seen visiting an animal shed. ami Nmhaidae Sufami Biidinae This species is legally protected under Schedule I of the ned Casor Ariadne ariadne pallidior ruhsorfer Wildlife (Protection) Act, 1972. Reported to be rather rare in the Naga hills (Tytler 1911). Recorded by SS from Chizami in October 2012. Rare in the survey area. Serean meror Mimathyma chevana chevana oore Common Casor Ariadne merione tapestrina oore Only two individuals recorded from the NEN compound and Erule riverside at Chizami in October A single record from Etsoshulu at Chizami in October 2012. Rare. This subspecies is legally protected under 2012. Very rare in the survey area. Only one individual Schedule I of the Wildlife (Protection) Act, 1972. A single collected from the Naga Hills (Tytler 1911). individual recorded from the Naga Hills (Tytler 1911).

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

ami Nmhaidae Sufami Charainae Rare in the survey area. Reported to be common near Common Naa Charaxes athamas athamas rur Kohima from August to October (Tytler 1911). Recorded from the Erule riverside at Chizami. Chooae ier Parantica melaneus plataniston Seen mud-puddling during the summer. Uncommon. ruhsorfer Reported to be common from the Naga Hills up to Recorded from many different locations at Chizami. 1,500m (Tytler 1911). Common. Reported to be not common; collected near Kohima from August to October (Tytler 1911). rea Naa Charaxes eudamippus eudamippus oueda Chesnu ier Parantica sita sitaoar Recorded two individuals from NEN compound and Recorded from many different locations at Chizami. Erule riverside at Chizami in August 2011 and August Common. Reported to be common near Kohima from 2013. Rare. Reported to be not uncommon in the Naga August to October (Tytler 1911). foothills (Tytler 1911). ar Bue ier Tirumala septentrionis septentrionis ami Nmhaidae Sufami Cresnae Buer Common a Cyrestis thyodamas thyodamas Recorded from all locations at Chizami between Boisdua April and October. Common. Reported to be common; Recorded by SS in January 2013, but no photographic collected near Kohima from August to October and in evidence. Rare in the survey area. Only a single individual the foothills in February and March (Tytler 1911). collected, not common in the Naga Hills (Tytler 1911). ami Nmhaidae Sufami eioniinae ami Nmhaidae Sufami anainae eo Coser Acraea issoria issoria ner ain ier Danaus chrysippus chrysippus innaeus Recorded from many different locations at Chizami. Mostly seen during the summer. Very common. Records by SS in flight at Chizami in July 2011 and Reported to be very common from the Naga Hills in the October 2012. Outside of Chizami, recorded by TN post-monsoon season (Tytler 1911). at Pungro, Kiphire District, Nagaland in the month of October 2013 where he sighted 2–3 individuals. Rare are Siersrie Argynnis childreni childreni Gray in the survey area. A few individuals reported from the foothills of the Naga Hills (Tytler 1911). Recorded from all locations at Chizami between March and October. Visits flowers. Very common. Sried ier Common ier Danaus genutia Reported to be fairly common from the Naga Hills (Tytler genutia Cramer 1911). Recorded from all locations at Chizami. Mostly seen during summer. Common. Reported from the Naga roia riar ndian riar Argynnis Hills from the foothills up to 1,500m throughout the year hyperbius hyperbius innaeus (Tytler 1911). Recorded from all locations at Chizami. Both males and females (less common) seen between March and Sried Bue Cro Euploea mulciber mulciber October. Common. Reported to be very common from Cramer the Naga Hills (Tytler 1911). Males and females recorded from the NEN compound and the Thetsumi trail at Chizami in the pre- ed aein Cethosia biblis tisamena ruhsorfer and post-monsoon seasons. Uncommon. This species is legally protected under the Schedule IV of the Wildlife Recorded from all locations at Chizami between (Protection) Act, 1972. Reported as very common in the March and October. Very common. Reported to be Naga Hills (Tytler 1911). exceedingly common from the Naga Hills (Tytler 1911).

ass ier Parantica aglea melanoides oore eoard aein Cethosia cyane cyane rur Recorded from many different locations at Chizami. Single record on the road linking Kuwari and Chizami. Mostly seen visiting flowers during summer. Uncommon.

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

Reported to be not common from the Naga Hills (Tytler ami Nmhaidae Sufami imenidinae 1911). imaaan Serean Athyma opalina orientalis es are eoman Cirrochroa aoris aoris oueda Several individuals recorded from two locations at the Thetsumi roadside at Chizami between March and Recorded from many different locations at Chizami October. Uncommon. Reported to be not uncommon between March and October. Seen visiting animal from the Naga Hills (Elwes 1891; Tytler 1911). droppings and flowers. Common. Reported to be not very common from the Naga Hills (Tytler 1911). Common Serean Athyma perius perius innaeus Common eoman Cirrochroa tyche mithila oore Recorded from all locations at Chizami between March and October. Common. Reported to be very Single record from the NEN compound in November common from the Naga Hills (Tytler 1911). 2011. Rare in the survey area. Reported to be not uncommon from the Naga Hills (Tytler 1911). rane Sa Serean Athyma cama cama oore us Cupha erymanthis lotis Suzer Recorded from the NEN compound in October 2012. Single record from Kuwari at Chizami in November Both male and female seen. Rare. Reported to be not 2013. Very rare in the survey area. Reported to be uncommon from the Naga Hills (Tytler 1911). rather scarce from the Naga Hills (Tytler 1911). Sa Serean Athyma selenophora selenophora Common eoard Phalanta phalantha phalantha oar rur Two records from the NEN compound and the Erule Recorded from the NEN compound and the Erule trail at Chizami in October 2011. Rare. Reported to be riverside at Chizami. Two records from these two very common from the Naga Hills (Tytler 1911). locations, seen during the pre-monsoon season. Rare in the survey area. Reported to be common from the Naga rea Calinaga brahma brahma Buer Hills (Tytler 1911). Recorded from the NEN compound. Two individuals seen at the same location in May 2011 and March 2012. aran Vagrans egista sinha oar Rare. This species is legally protected under Schedule II Recorded from the Erule riverside at Chizami in the of the Wildlife (Protection) Act, 1972. A few individuals summer months. Seen visiting animal droppings and collected in April and May, near Kohima (Tytler 1911). mud-puddling. Rare in the survey area. Reported to be common at low elevations from the Naga Hills (Tytler Bue ue Bassarona durga splendens er 1911). Recorded from Kuwari at Chizami in November 2013. Seen mud-puddling at the riverside. This is the Cruiser Vindula erota erota ariius rare subspecies splendens in which the blue discal band Recorded from Chizami. Both males and females forms lunules throughout its length from tornus to costa (less common) seen between March and October. Seen described by Tytler (1915a) from the Naga Hills. Very visiting flowers. Uncommon. Males reported to be fairly rare. This species is legally protected under Schedule I of common from the Naga Hills (Tytler 1911). the Wildlife (Protection) Act, 1972. Reported from near Imphal in Manipur (Tytler 1915a). ami Nmhaidae Sufami iheinae Common Bea Libythea lepita lepita oore renh ue Euthalia franciae franciae eder & Two individuals recorded from the NEN compound eder and the Erule River at Chizami in October 2012. Rare in Mostly recorded from the Thetsumi trail at Chizami. the survey area. This species is legally protected under Common between July and October. This species is Schedule II of the Wildlife (Protection) Act, 1972. A few legally protected under Schedule II of the Wildlife individuals collected from the Naga Hills (Tytler 1911). (Protection) Act, 1972. Reported from the Naga Hills (Elwes 1891). It is reported to be not uncommon in August to October between 1,500–1,600 m in the Naga

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

Hills (Tytler 1911). http://www.ifoundbutterflies.org/2770-Neptis/Neptis- ananta wherein some individuals of N. namba have paler Bronze ue Euthalia nara nagaensis er orange markings. While this individual was worn, and A single record on the village footpath leading to its cilia not clearly visible, the UNF/H bluish tinge on the the NEN office in September 2012. Very rare. Reported markings below, so typical of namba was clearly visible from the Naga Hills (Elwes 1891; Tytler 1911). hence we are inclined to provisionally identify this as N. namba. Neptis namba and Neptis ananta have often reen ue Euthalia sahadeva nadaka ruhsorfer been considered as a single species and forms/subspecies of the latter. While Eliot suggested these were separate A male recorded basking high up on a tree in July species, he also stated “dissections of the male genitalia 2011, below the NEN compound. Females recorded in have not helped greatly to establish whether there are the summer in the NEN compound. Common. Reported one, two or three species in the complex.” Clearly more to be not uncommon from the Naga Hills (Elwes 1891; work on the phylogeny and ecology of these species are Tytler 1911; Tytler 1915a). necessary to establish their true status. Neptis namba was described by Tytler (1915a) as a new species on the Common Baron Euthalia aconthea garuda oore basis of his collection from the Naga Hills. Tytler (1915a) states that N. ananta is mostly found between 1,500– Recorded from many different locations at Chizami- 2,100 m, while N. namba is mostly found in the foothills. both males and females seen. Recorded in the pre- and However, SS’s records from the NE India show that while post-monsoon seasons. Common. This species is legally N. ananta is generally found at higher altitudes, both protected under Schedule II of the Wildlife (Protection) species have been recorded from the foothills at least Act, 1972. Reported to be not common from the Naga up to 600m. However this record, if correct, extends its Hills (Tytler 1911). altitudinal range even further.

aud Baron Euthalia lubentina lubentina Cramer Broadanded Saier Neptis sankara amba oore A single record from the Chizami Village in March Two individuals recorded from NEN compound in 2012. Very rare in the survey area. This species is legally June 2012 and August 2012. Rare. protected under Schedule IV of the Wildlife (Protection) Act, 1972. Only a few individuals collected in the Naga aass Saier Neptis sappho astola oore Hills (Tytler 1911). Recorded from all locations at Chizami between March and October. Very common. Reported to be very hieeded Bue Baron Euthalia phemius common from the Naga Hills (Tytler 1911). phemius oueda A single record of a male from the NEN compound in Commander Moduza procris procris Cramer October 2011. Very rare in the survey area. Reported A single record in flight by TN from the Erule River at from the Naga Hills (Tytler 1911). Chizami in 2012. Very rare in the survey area.

eo Saier Neptis ananta ochracea ans Cier Parthenos sylvia gambrisius ariius Two individuals recorded from the Thopfu trail and A single record from the NEN compound in November the NEN compound in Sep 2013. Rare in the survey 2011. Very rare in the survey area. This species is legally area. Reported to be not uncommon in the Naga Hills protected under Schedule II of the Wildlife (Protection) from July to October (Tytler 1911). Act, 1972. Reported to be fairly common at low elevations in the Naga Hills (Tytler 1911). Nama Saier Neptisf namba namba er A single unconfirmed record from the NEN compound ami Nmhaidae Sufami Nmhainae in Oct 2011. Generally N. namba has brighter orange ndian oroiseshe Aglais caschmirensis aesis markings compared with N. ananta (Eliot, 1969), but ruhsorfer this is not always true based on SS’s experience in the Recorded from many different locations at Chizami field as well as images available online http://www. in the pre- and post-monsoon seasons. Common. This ifoundbutterflies.org/2773-Neptis/Neptis-namba and species has extended its range eastwards over time from

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi the western and central Himalaya and has established 2012, Chizami. Rare in the survey area. Reported to be a known population in western Arunachal Pradesh and fairly common from the low elevations of Naga Hills Nagaland (Naro 2012). (Tytler 1911).

ono Araschnia prorsoides dohertyi Banhard Chooae ans Junonia iphita iphita Cramer Recorded only from Thetsumi trail near the Tsoyiphe Recorded from many different locations at Chizami. stream in July 2011 and from Ezilu in March 2011 at Seen in both pre- and post-monsoon seasons. Very Chizami. Some authors treat Araschnia dohertyi as a common. Reported to be very common from the Naga separate species. Rare. Reported to be abundant from Hills (Tytler 1911). the Naga Hills between 1,800–2,400 m (Elwes 1891) who states that its larval food plant is a nettle. Tytler (1911) emon ans Junonia lemonias lemonias innaeus states that it is fairly common post-monsoon. Recorded from NEN compound between January uumn eaf Doleschallia bisaltide indica oore and July. Uncommon. Reported to be very common from the low elevations of Naga Hills and not collected Recorded from the NEN compound and the Erule trail above 1,200 m (Tytler 1911). at Chizami, seen during summer season. Uncommon. Common at low elevations but scarce near Kohima Bue ans Junonia orithya ocyale ner (Tytler 1911). Recorded from many different locations at Chizami in February, July and October. Very common. Reported to rane aeaf Kallima inachus inachus Boisdua be very common from the Naga Hills (Tytler 1911). Recorded from the Erule riverside at Chizami ndian Bue dmira Kaniska canace canace between February and July. Common. Reported to be innaeus fairly common from the Naga Hills (Elwes 1891; Tytler Recorded from many different locations at Chizami. 1911). Common. Reported to be common from the Naga Hills (Tytler 1911). Sare Bue aeaf Kallima knyvettii de Niie Sotted eser Symbrenthia hypselis cotanda Recorded from the Thetsumi trail at Chizami in July oore 2011. Rare. This species is legally protected under A single record from the NEN compound in August Schedule II of the Wildlife (Protection) Act, 1972. 2011. Rare. Reported to be rather common from the Reported to be “not uncommon” from the Naga Hills Naga Hills (Tytler 1911). (Elwes 1891). Tytler (1911) reports it to be rather rare from the Naga Hills. Common eser Symbrenthia lilaea khasiana oore rea fl Hypolimnas bolina jacintha rur Recorded from the NEN compound, the Government high school area and many other different parts of Recorded from NEN compound and Erule riverside Chizami through most of the year. Common. Reported at Chizami. Uncommon. Reported to be common at low to be common from the Naga Hills (Tytler 1911). elevations from the Naga Hills (Tytler 1911). Bueaied eser Symbrenthia niphanda niphanda eao ans Junonia almana almana innaeus oore A single record from the NEN compound in June Recorded from many different locations at Chizami 2013. Very rare. This species is legally protected during the summer. Uncommon. Reported to be not under Schedule II of the Wildlife (Protection) Act, 1972. uncommon from the Naga Hills (Tytler 1911). Reported to be rare in the Naga Hills with only three individuals being collected in two years in August and eo ans Junonia hierta hierta ariius October (Tytler 1911). Recorded only one individual at Erule River in October

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

ained ad Vanessa cardui cardui innaeus une or Thaumantis diores diores oueda Recorded from different locations of Chizami during the summer season. Common. Reported to be very A single record from Chizami Village late evening in common from the Naga Hills (Tytler 1911). November 2012. Very rare in the survey area. Reported from the Naga Hills (Tytler 1911). ndian ed dmira Vanessa indica indica ers Callerebia orixa oore Recorded from many different locations at Chizami Recorded from the NEN compound. A few worn and during the pre- and post- monsoon seasons. Common. poorly marked individuals were recorded and could not Reported to be common from the Naga Hills (Tytler be identified to the species level with certainty, though 1911). it is likely that these were C. orixa. Rare in the survey area. Several specimens collected from the Naga Hills ami Nmhaidae Sufami seuderoinae and reported to be common post-monsoon (Elwes 1891; Consae Dichorragia nesimachus nesimachus Tytler 1911). ore A single record from below the NEN compound in Common amflElymnias hypermnestra undularis August 2011. Very rare in the survey area. rur Recorded from Erule and Kuwari at Chizami during a Pseudergolis wedah wedah oar the summer months. Common. Reported to be common Recorded from Thetsumi and Erule trails at Chizami in the Naga Hills (Tytler 1911). mostly post- monsoon from July onwards. Often seen mud-puddling. Common. Reported to be not uncommon Sotted amfl Elymnias malelas malelas from the Naga Hills (Tytler 1911). eison Recorded from many different parts of Chizami’s oina Stibochiona nicea nicea ra forests. Common. Records from the Thopfu and Tsetsumi trails at Chizami in February 2010 and July 2011. Rare in the Banded reeron Lethe confusa confusa survey area. Reported to be not uncommon in the Naga uriiius Hills (Tytler 1911). Recorded below the NEN compound in Mar 2013. Rare. Reported to be common from Kohima (Tytler ami Nmhaidae Sufami Sarinae 1911). eo rad Aemona amathusia eison Recorded from the NEN compound area at Chizami Common reeron Lethe rohria rohria ariius in the summer. Uncommon. This species is legally protected under Schedule II of the Wildlife (Protection) Recorded from many different locations at Chizami Act, 1972. Reported to be very rare in the Naga Hills between March and October. Common. Reported to be (Tytler 1911). common from Kohima (Tytler 1911).

Common uer Discophora sondaica zal Sraihanded reeron Lethe verma sintica esood ruhsorfer A single record from the NEN compound in May 2012. Recorded from many different locations of Chizami Very rare in the survey area. Reported to be common forests between March and October. Common. Seen from low elevations in the Naga Hills (Tytler 1911). visiting animal sheds. Reported to be very common at altitudes between 1,500–2,000 m in the Naga Hills are aun Faunis eumeus assama esood (Tytler 1911). A single record from the Tahulu trail at Chizami in ned ed oreser Lethe chandica chandica September 2011. Very rare in the survey area. This oore species is legally protected under Schedule II of the Recorded from inside the village and from the NEN Wildlife (Protection) Act, 1972. compound in July 2011 and March 2013. Rare.

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

Common ed oreser Lethe mekara mekara ar enin Bron Melanitis phedima bela oore oore A single record from the NEN compound at Chizami Recorded from the NEN compound in July 2011 and in October 2012. Very rare in the survey area. October 2012. Uncommon. Reported to be common in the Naga Hills (Tytler 1911). aied ed oreser Lethe sinorix sinorix eison rea enin Bron Melanitis zitenius zitenius Recorded from many different locations at Chizami ers sporadically throughout the year including the winter A few records of individuals identified in hand in months. Common. This subspecies is legally protected DSF in forest below NEN compound in Oct 2012. Rare. under Schedule II of the Wildlife (Protection) Act, 1972. This species is legally protected under Schedule II of the Reported from the Naga Hills (Elwes 1891; Tytler 1911). Wildlife (Protection) Act, 1972.

Bamoo oreser Lethe f kansa oore hieine Bushron Heteropsis malsara oore Two sightings of this butterfly in the same location- just outside NEN compound , near a clump of bamboo Recorded from many different locations of the in January 2013. UNH ocellus in four on an even curve forested areas at Chizami and at the NEN compound in hence seems to be Lethe kansa in its dry season form. July 2011 and October 2012. This species was earlier However, the upperside could not be seen, and hence considered to be part of the Genus Mycalesis but recent this identification is only provisional. Rare. Reported phylogenetic studies have re-classified them in Genus from the Naga foothills from May to August (Tytler Heteropsis (Kodandaramaiah et al. 2010). Common. 1911). Only a single record from the Naga Hills (Tytler 1911).

ae oreser Lethe latiaris latiariseison iaine Bushron Mycalesis francisca sanatana Recorded from the NEN compound at Chizami and oore the forest trail below NEN compound in April 2011, Recorded from many different locations at Chizami in October 2012 and August 2013. Rare. Reported to be July and October. Common. Seen mud-puddling at the not uncommon between 1,500–2,000 m, September riverside occasionally, but seen mostly on the footpaths and October (Tytler 1911). leading to the fields and inside the bushes. Reported from the Naga Hills (Elwes 1891, Tytler 1911). Common oodron Lethe sidonis eison eha Bushron Mycalesis lepcha kohimensis Two individuals recorded from the NEN compound in er the summer months. Rare in the survey area. Reported Recorded from many different locations of the from the Naga Hills (Elwes 1891). Reported to be very forested areas at Chizami. Common. Dry season form common at altitudes between 1,500–2,000 m in the reported in March and April and wet season form in Naga Hills (Tytler 1911). August and October from Kohima (Tytler 1911; Tytler 1914). iafor Lethe f sura oueda A single unconfirmed record on the Thopfu trail hieeded Bushron Mycalesis mestra sadona at Chizami in November 2012. The butterfly was er photographed sitting on the upper surface of a leaf, Recorded from two locations at Chizami—the NEN high up in the canopy, and its upperside could not be compound and at Awu in October 2012. Rare in the observed. Reported to be very common in the Naga Hills survey area. This species is legally protected under (Tytler 1911). Schedule II of the Wildlife (Protection) Act, 1972.

Common enin Bron Melanitis leda leda eined arinh Neope pulaha pulaha oore innaeus Recorded from different locations at Chizami in the Recorded from the NEN compound, near the pre- and post- monsoon seasons. Common. Reported to kitchen. Two individuals recorded at the same location be fairly common in the Naga Hills (Tytler 1911). in September 2011 and May 2013. Rare. Reported from

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi the Naga Hills (Elwes 1891). aied a Graphium agamemnon agamemnon innaeus ier Bron Orinoma damaris damaris ra Recorded by Kewekhrozo Thopi on the village Two individuals recorded from different locations at approach road leading to Chizami Village near NEN area Chizami in October 2012. Rare. Reported at 1,500m in and by SS in October 2012. Rare in the survey area. September and October and reported to be not common Reported to be common from the Naga Hills (Tytler in the Naga Hills (Tytler 1911). 1912).

Common ierin Ypthima baldus baldus ass Bueotte Graphium cloanthus cloanthus ariius esood Recorded from all locations at Chizami mostly Recorded from the NEN compound. Often seen between March and July. Common. Reported to be visiting flowers during the summer. Rare. Reported to common from the Naga Hills (Tytler 1911). be not common from the Naga Hills (Tytler 1912).

Near hreerin Ypthima cf. newara newara Common BueotteGraphium sarpedon sarpedon oore innaeus Recorded from all locations at Chizami area in July Recorded from all locations at Chizami. Mostly seen and October. Common. All the individuals recorded had visiting flowers and mud-puddling on the riverbank. a larger sub-apical ocellus UNH and lacked UNF/H sub- Uncommon. Reported to be common from the Naga marginal dark lines, than individuals recorded elsewhere Hills (Tytler 1912). in northeastern India by SS. It is difficult to separate Y. newara and Y. confusa without examining male genitalia Common ime Papilio clytia clytia innaeus hence only provisionally identified. Form dissimillis recorded from the Erule riverbank at Chizami. Seen mud-puddling. Rare. Form dissimilis imaaan ierin Ypthima sakra sakra oore reported to be common from the Naga Hills (Tytler 1912). Recorded from all locations at Chizami in the pre- and post-monsoon seasons. Common. Reported from esser ime Papilio epycides epycides eison the Naga Hills (Elwes 1891; Tytler 1911). A single record from the NEN compound in April Ypthima s 2014. Very rare. At least two separate individuals, which did not match any of the large-sized fiverings:Ypthima savara, Ypthima ed een Papilio helenus helenus innaeus methora, Ypthima dohertyi or Ypthima persimilis. Both Recorded from all locations at Chizami between these individuals had wingspans > 45 mm, were heavily March and October. Mostly seen visiting flowers. Locally striated on both wings UNF/H. UNF: in addition to large common. Reported to be common from the Naga Hills sub-apical doubled-pupilled ocellus, and additional (Tytler 1912). ocellus in two. UNH: with five mostly equal-sized ocelli, with pupils. Tornal ocelli merged. Seen at Chizami in the eo een Papilio nephelus chaon esood forested areas in Oct 2012. Recorded from the Erule riverbank at Chizami in ami aiionidae Sufami aiioninae August 2013. Rare. Reported at low elevations from the reen raonai Lamproptera meges indistincta Naga Hills (Tytler 1912). er Recorded from all locations at Chizami. Mostly rea ormon Papilio memnon agenor innaeus seen during the summer between March and July. Often seen mud-puddling. Common. Reported tobe Recorded from the Erule riverbank and other not uncommon between 1,200–1,500 m from April to forested areas at Chizami in July and October. Seen mud- October (Tytler 1912). puddling, Forms agenor and distantianus seen. Rare. Reported “sparingly” from the Naga Hills (Tytler 1912).

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

Common ormon Papilio polytes romulus Cramer (Tytler 1912). Recorded from the Awelu trail at Chizami. Rare in Birdin s Troides s the survey area. Reported to be common from the Naga A birdwing seen in flight by SS though species level Hills (Tytler 1912). identification was not possible.

edreas Papilio alcmenor alcmenor eder & ami ieridae Sufami Coiadinae eder hreeso rass eo Eurema blanda silhetana Recorded from the Erule riverbank at Chizami. Seen aae mud-puddling. Rare. Reported from the Naga Hills Recorded from all locations at Chizami. Common. (Tytler 1912). Reported to be not common from August to September at Kohima (Tytler 1912). Sane Papilio protenor euprotenor ruhsorfer Sma rass eo Eurema brigitta rubella Recorded from all locations at Chizami. Seen mostly aae in the months of March and July. Twenty-five individuals Recorded from all locations at Chizami. Common. were recorded in March alone. Very common. Reported Reported to be not very common from the Naga Hills to be common from the Naga Hills (Tytler 1912). (Tytler 1912).

ime Butterfl Papilio demoleus demoleus Common rass eo Eurema hecabe hecabe innaeus innaeus Recorded from all locations at Chizami mostly during Recorded from all locations at Chizami. Common. the summer season. Common. Reported from the Naga Reported to be very common from the Naga Hills (Tytler Hills (Tytler 1912). 1912).

Bue eao Papilo arcturus arcturus esood Soess rass eo Eurema laeta sikkima oore Recorded from the NEN compound in October 2012. Recorded from many different locations at Chizami. Rare. Reported to be not uncommon from the Naga Hills Common. Reported to be rather common from Kohima (Tytler 1912). (Tytler 1912).

Common eao Papilio bianor ganesa ami ieridae Sufami ierinae oueda Common aross Appias albina darada eder & Recorded from numerous locations at Chizami. eder Common. Reported rather commonly from the Naga Recorded from the NEN compound. Male and female Hills (Tytler 1912). seen. Rare in the survey area. This subspecies is legally protected under Schedule II of the Wildlife (Protection) aris eao Papilio paris paris innaeus Act, 1972. Reported to be rare from the Naga Hills Recorded from all locations at Chizami. Seen mostly (Tytler 1912). visiting flowers in gardens. Common. Reported tobe very common from the Naga Hills (Tytler 1912). rane aross Appias galba aae Two individuals recorded from Chizami Village. Seen Common Bain Atrophaneura varuna astorion mud-puddling. Rare. This subspecies is legally protected esood under Schedule IV of the Wildlife (Protection) Act, 1972. Recorded from the NEN compound. Rare. Collected Reported to be not uncommon from the Naga Hills “sparingly” in the Naga Hills (Tytler 1912). (Tytler 1912).

Common indmi Byasa polyeuctes polyeuctes So un Appias lalage lalage oueda oueda Recorded from many different locations at Chizami. Recorded from many different locations at Chizami. Common. Reported to be rather common from the Common. Collected “commonly” from the Naga Hills Naga Hills (Tytler 1912).

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

esser u Cepora nadina nadina uas ae anderer Pareronia avatar oore Recorded from two locations—the NEN compound Recorded from different locations of Chizami. and Erule riverbank at Chizami. Rare in the survey area. Common. A few individuals reported from the Naga Hills A few individuals recorded from the Naga Hills (Tytler (Tytler 1912). 1912). are Caae hie Pieris brassicae nepalensis Common miran Catopsilia pomona pomona oueda ariius Recorded from all locations at Chizami through most Recorded from the NEN compound. Uncommon. of the year. Very common. Reported to be very scarce Reported to be not uncommon from the Naga Hills from the Naga Hills (Tytler 1912). (Tytler 1912). ndian Caae hie Pieris canidia indica ans otted miran Catopsilia pyranthe pyranthe innaeus Recorded from all locations at Chizami through most Recorded from the NEN compound. Uncommon. of the year. Very common. Reported to be abundant throughout the year from the Naga Hills (Tytler 1912). edreas ezee Delias acalis pyramus aae Bah hie Pontia daplidice moorei er Recorded from the NEN compound and the Erule Recorded from many different parts of Chizami. riverbank at Chizami in October 2012. Rare. Common. This species has extended its range eastwards in recent times from the western and central Himalaya i ezee Delias belladonna lugens ordan to many locations in eastern Himalaya including Assam, Recorded from many different parts at Chizami. Arunachal Pradesh, Nagaland and Manipur (Naro & Common. Recorded as common between 1,200–2,100 Sondhi 2013; Singh & Gogoi 2013). m in August and September in the Naga Hills (Tytler 1912). reeneined hie Pieris melete ajaka oore edso ezee Delias descombesi descombesi Recorded from all locations at Chizami through Boisdua most of the year. Very common. Reported to be rather A single record from Chizami Village in November uncommon from the Naga Hills (Tytler 1912). 2012. Very rare. A few individuals reported from the Naga Hills (Tytler 1912). Sotted Saooh Prioneris thestylis thestylis oueda edase ezee Delias pasithoe pasithoe Recorded from many different parts at Chizami. innaeus Female recorded feeding on flower nectar in July 2011 Recorded from the NEN compound and the Erule in the NEN office complex. Males common, females riverbank at Chizami during summer. Rare. rare. Reported to be common from the Naga Hills (Tytler 1912). rea rane Hebomoia glaucippe glaucippe innaeus ami iodinidae Sufami Nemeoiinae Recorded from many different locations of Chizami. ar ud Abisara fylla esood Common. Reported to be common from low elevations Recorded from all locations at Chizami. Seen up to 600m in the Naga Hills (Tytler 1912). throughout the year. Very common. Reported to be abundant and common from the Naga Hills (Elwes 1891; eo rane Ixias pyrene familiaris Buer Tytler 1911). Recorded from many different locations at Chizami Sried unh Dodona adonira adonira eison between March and October. Common. Reported to be common from the Naga Hills (Tytler 1912). A single record from Thetsumi roadside near the stream in February 2013. Rare. Reported to be “not common” from the Naga Hills (Elwes 1891; Tytler 1911).

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

unhineo Zemeros flegyas fleygas Cramer Naro Sighting ofAglais cashmirensis aesis Fruhstorfer, 1912 (Nymphalidae) from Nagaland, India. Journal of Threatened Taxa Recorded from all locations at Chizami. Seen 4(4): 2534–2535; http://dx.doi.org/10.11609/JoTT.o3019.2534-5 throughout the year. Very common. Reported to be Naro & S Sondhi Sightings of Bath White Pontia daplidice very common from the Naga Hills (Tytler 1911). moorei Röber, 1907 (Lepidoptera: Pieridae: Pierinae: Pierini) from Arunachal Pradesh and Nagaland, India. Journal of Threatened Taxa 5(7): 4122-4124; http://dx.doi.org/10.11609/JoTT.o3308.4122-4 Sinh Butterflies of India. Om Books International, 183pp. NCS Sinh & ooi A new range record for the Bath White butterfly Pontia daplidice for north-east India. Bionotes 15(2): 59. Smih C Illustrated Checklist of Nepal’s Butterflies. T.C. Binham C The Fauna of British India including Ceylone Majupuria (Ed.). Rohit Kumar, Gwalior. M.P., India, 127pp. and Burma: Butterflies. Vols. I-II. Taylor and Francis Ltd., London. Smih C Butterflies of Nepal. Revised Edition, Tecpress Service Core S & endeur The Butterflies of the Malay L.P., Bangkok, Thailand, 368pp. Peninsula. 4th Edition revised by J.N. Eliot. Malayan Nature Society, Sondhi S & une Butterflies and Moths of Pakke Tiger Kuala Lumpur, 595pp. Reserve. Titli Trust (Dehradun) and Indian Foundation for Butterflies de on Revision of the oriental Genus Matapa Moore (Bengaluru), vi+202pp. (Lepidoptera: Hesperiidae) with discussion of its phylogeny and Sondhi S une aear oaear & oear geographic history. Zoologische Mededelingen 57: 21. Butterflies of the Garo Hills. Samrakshan Trust (New Delhi), Titli de Niie C On new and little-known Lepidoptera from Trust (Dehradun), and Indian Foundation for Butterflies (Bengaluru), the Oriental Region. Journal of Bombay Natural History Society xvi+200pp. 13(1): 157–175. Sinhoe C Lepidoptera Indica. Vol. X. Rhopalocera. io N An Analysis of the Eurasian and Australian Neptini Family Hesperiidae. Reeve & Co, London, 361pp+78pl. (Lepidoptera: Nymphalidae). Bulletin of the British Museum ao The Fauna of British India, including Ceylone (Natural History) Entomology. Supplement 15. and Burma: Butterflies. Vol.. 1 Taylor and Francis, London, es On butterflies collected by Mr. W. Doherty in the 600pp+3pl. Naga and Karen Hills and in Perak. Proceedings of the Zoological ao The Fauna of British India, including Ceylone and Society of London 1891: 249–289. Burma: Butterflies. Vol. 2. Taylor and Francis, London, 506pp+2pl. es On butterflies collected by Mr. W. Doherty in the er C Notes on butterflies from the Naga Hills - Part II. Naga and Karen Hills and in Perak. Proceedings of the Zoological Journal of Bombay Natural History Society 21: 588–606. Society of London 1892: 617–664. er C Notes on butterflies from the Naga Hills - Part I. ans The Identification of Indian Butterflies. 2nd Revised Journal of Bombay Natural History Society 21: 48–65. Edition. Bombay Natural History Society, Bombay, x+454pp+32pl. er C Notes on some new and interesting butterflies ans A catalogue of the Hesperiidae from Europe, from Manipur and the Naga Hills. Part - I. Journal of Bombay Natural Asia and Australia in the British Museum (Natural History). British History Society 23: 216–229. Museum (Natural History), London, 502pp. er C a Notes on some new and interesting butterflies ans A revision of the Arhopala group of Oriental from Manipur and the Naga Hills. Part - II. Journal of Bombay Lycaenidae (Lepidoptera: Rhopalocera). Bulletin of the British Natural History Society 23: 502–515. Museum (Natural History), London Entomology Volume 5: 85–141. er C Notes on some new and interesting butterflies aria The Butterflies of The Sikkim Himalaya and Their from Manipur and the Naga Hills. Part - III. Journal of Bombay Natural History. Sikkim Nature Conservation Foundation, Gangtok, Natural History Society 24: 119–155. 217pp. er C Notes on Some New and Interesting Butterflies uan & ue The Chinese Pseudocoladenia skippers. chiefly from Burma. Part - 2. Journal of Bombay Natural History Neue Entomologische Nachrichten 57: 161–170. Society 42: 109–123. ehimar The Book of the Indian Butterflies. Bombay Natural arshne a First Bibliography of the Butterflies of Indian History Society and Oxford University Press, Oxford, xvi+497pp. Region. Bionotes 10(2): 43–46. inon S Illustrated Checklist for the Butterflies of Myanmar. arshne First Bibliography of the Butterflies of Indian Smithsonian Institution, 197pp. Region. Bionotes 10(3): 81–86. odandaramaiah C ees C er orres aranh arshne First Bibliography of the Butterflies of Indian & N aher Phylogenetics and biogeography of a Region. Bionotes 10(4): 143–146. spectacular Old World radiation of butterflies: the subtribe arshne a First Bibliography of the Butterflies of Indian Mycalesina (Lepidoptera: Nymphalidae: Satyrini). BMC Evolutionary Region. Bionotes 11(1): 30–33. Biology 10: 172. arshne First Bibliography of the Butterflies of Indian une Butterflies of Peninsular India. Universities Press Region. Bionotes 11(2): 47–50. (Hyderabad) and Indian Academy of Sciences (Bangalore), 254pp. arshne First Bibliography of the Butterflies of Indian arsha & de Niie The Butterflies of India, Region. Bionotes 11(3): 86–89. Burmah and Ceylon. Vol. I. The Calcutta Central Press Co., Calcutta, arshne d First Bibliography of the Butterflies of Indian 398pp Region. Bionotes 11(4): 119–121. arsha & de Niie The Butterflies of India, nerBh Butterflies of The Indian Region. Bombay Burmah and Ceylon. Vol. II.The Calcutta Central Press Co., Calcutta, Natural History Society, Bombay, xx+523pp+72pl. 370pp. arsha & de Niie The Butterflies of India, Burmah and Ceylon. Vol. III. The Calcutta Central Press Co., Calcutta, 552pp

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

(©Tshetsholo Naro NEN) (© Sanjay Sondhi) (© Kewekhrozo Thopi NEN) (© Kewekhrozo

mae Bron Badamia exclamationis mae ndian in Choaspes benjaminii japonica

mae reen e Bibasis vasutana (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae Sae Hasora anura anura mae Buesotted Sru oer Aeromachus kali

mae hieanded Hasora taminatus bhavara (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae eined Sru oer

Aeromachus stigmata obsoleta NEN) Naro (©Tshetsholo mae ure Si Caltoris tulsi (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae ores oer Astictopterus jama olivascens (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae Sraih Si Parnara f guttatus NEN) Naro (©Tshetsholo mae are Banded Si Pelopidas sinensis (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae Sma Banded Si Pelopidas mathias mathias

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

© Kewekhrozo Thopi NEN (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae ih Sra e Pithauria (©Tshetsholo Naro NEN) Naro (©Tshetsholo stramineipennis stramineipennis mae ued e Sebastonyma dolopia mae ssam arer Ochlodes siva siva (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae Baeined edee Matapa f sasivarna

mae Siim am edee Erionota torus (© Sanjay Sondhi) mae ndian am Bo Suastus gremius gremius (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae esried emon Notocrypta curvifascia curvifascia Thopi NEN) (© Kewekhrozo mae Common Banded emon mae Sotted emon Notocrypta Notocry pta paralysos asawa feisthamelii alysos

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

(©Tshetsholo Naro NEN)

(©Tshetsholo Naro NEN) mae Common Sotted a mae udd ied a Celaenorrhinus leucoce ra Pseudocoladenia f fatua (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae ree itter Hyarotis adrastus praba

(©Tshetsholo Naro NEN) (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae Sotted ne Caprona agama agama mae an ne Ctenoptilum vasava vasava (©Tshetsholo Naro NEN) mae Sotted Sno a Tagiades menaka menaka (©Tshetsholo Naro NEN) Naro (©Tshetsholo

(© Kewekhrozo Thopi NEN) (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae Chesnu ne Odontoptilum mae Brih Suneam Curetis bulis bulis angulata angulata mae Brih Suneam Curetis bulis bulis (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae Naa Sahire Heliophorus mae oden Sahire Heliophorus kohimensis kohimensis brahma brahma mae Na a Sahire Heliophorus kohimensis kohimensis

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae reen Sahire Heliophorus moorei tytleri mae oined Ciiae Bue Anthene

(©Tshetsholo Naro NEN) Naro (©Tshetsholo lycaenina lycambes

mae Naa Sahire femae Heliophorus kohimensis kohimensis (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae ain ede Bue Celastrina lavendularis limbata

mae ae ede Bue Udara dilecta mae Common ede Bue Acytolepis dilecta puspa gisca

mae Bothrinia chennellii ede Cuid (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae ede Cuid Bothrinia chennellii

mae ede Cuid Bothrinia chennellii (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae Chamans Cuid Everes argiades diporides

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi ©Tshetsholo Naro NEN) Naro ( ©Tshetsholo (© Kewekhrozo Thopi NEN) (© Kewekhrozo Thopi NEN) (© Kewekhrozo

Image 41. Indian Cupid Everes lacturnus Image 42. Silver Forget-me-not assamica Image 41. Indian Cupid Everes lacturnus assamica Catochrysops panormus exiguus ©Tshetsholo Naro NEN) Naro ( ©Tshetsholo ©Tshetsholo Naro NEN) Naro ( ©Tshetsholo ©Tshetsholo Naro NEN) Naro ( ©Tshetsholo

Image 43. Forget-me-not Catochrysops strabo strabo Image 44. Gram Blue female Euchrysops cnejus cnejus Image 45. Dark Cerulean Jamides bochus bochus ©Tshetsholo Naro NEN) Naro ( ©Tshetsholo ©Tshetsholo Naro NEN) Naro ( ©Tshetsholo ©Tshetsholo Naro NEN) Naro ( ©Tshetsholo

Image 47. Glistening Cerulean Jamides elpis Image 46. Cerulean Jamides sp. pseudoelpis Image 48. Pea Blue Lampides boeticus

6616 Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo © Kewekhrozo Thopi NEN © Kewekhrozo mae era Bue Leptotes plinius plinius mae in ineue Petrelaea dana

mae Common ineue Prosotas nora ardates (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae eaf Bue Amblypodia anita dina

mae ar rass Bue Zizeeria karsandra UN mae Common em Poritia hewitsoni hewitsoni

UP (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae BiSo oa Ancema ctesia ctesia NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae Comi aue Arhopala comica

(©Tshetsholo Naro NEN) Naro (©Tshetsholo mae odered aue Arhopala mae aiess Bushue Arhopala ganesa bazalus teetsa watsoni

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

UP mae Common inse Catapaecilma mae eodis aue major major

Arhopala singla NEN) Naro (©Tshetsholo

mae aiess ushue Flos areste

mae Common ash Rapala nissa mae Sierarea Bue Iraota timoleon nissoides timoleon (©Tshetsholo Naro NEN) Naro (©Tshetsholo © Kewekhrozo Thopi NEN (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae Cu Sierine Spindasis syama peguanus

mae Common aia Bue mae hai Sierine Spindasis rukmini Surendra quercetorum quercetorum (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae an meror Chitoria ulupi ulupi (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae ure meror Apatura ambica mae an meror Chitoria ulupi ambica ulupi

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi (©Tshetsholo Naro NEN) Naro (©Tshetsholo

(©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) mae oden meror Dilipia morgiana mae Serean meror Mimathyma chevana chevana mae Serean meror Mimathyma chevana chevana (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae Couresan Euripus nyctelius nyctelius mae Cire Hestinalis nama nama

mae asha Herona marathus marathus (©Tshetsholo Naro NEN)

(©Tshetsholo Naro NEN) (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae asern Courer Sephisa chandra mae Siren Hestina persimilis persimilis chandra

mae Ba rine Rohana parisatis parisatis (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae Common Casor Ariadne merione NEN) Naro (©Tshetsholo tapestrina mae rea Naa Charaxes eudamippus eudamippus mae Common Naa Charaxes athamas athamas

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

(©Tshetsholo Naro NEN)

(©Tshetsholo Naro NEN) mae ain ier Danaus chrysippus chrysippus mae Sried ier Danaus genutia

(©Tshetsholo Naro NEN) Naro (©Tshetsholo genutia mae Common a Cyrestis thyodamas (©Tshetsholo Naro NEN) thyodamas (©Tshetsholo Naro NEN)

(©Tshetsholo Naro NEN)

mae Sried Bue Cro femae Euploea mulciber mulciber

mae ass ier Parantica aglea melanoides mae Sried Bue Cro Euploea mulciber mulciber (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae Chooae ier Parantica melaneus plataniston

(©Tshetsholo Naro NEN) (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae ar Bue ier Tirumala septentrionis septentrionis (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae Parantica sita sita Chesnu ier

mae eo Coser Acraea issoria issoria NEN) Naro (©Tshetsholo

mae roia riar Argynnis hyperbius hyperbius mae are Siersri Argynnis childreni childreni

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

(©Tshetsholo Naro NEN) (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae ed aein femae female Cethosia biblis tisamena mae ed aein mae Cethosia biblis tisamena (©Tshetsholo Naro NEN) Naro (©Tshetsholo

male mae eoard aein Cethosia cyane

(©Tshetsholo Naro NEN) Naro (©Tshetsholo cyane

(©Tshetsholo Naro NEN) Naro (©Tshetsholo mae Common eoman Cirrochroa tyche mithila mae are eoman Cirrochroa aoris aoris male

UN (©Tshetsholo Naro NEN) Naro (©Tshetsholo

UP NEN) Naro (©Tshetsholo mae Common eoard Phalanta phalantha phalantha (©Tshetsholo Naro NEN) Naro (©Tshetsholo

female mae Cruiser Vindula erota erota (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae us Cupha erymanthis lotis

mae aran Vagrans egista sinha

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

(©Tshetsholo Naro NEN) (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae imaaan Serean Athyma mae Common Serean Athyma perius opalina orientalis perius mae Common Bea Libythea lepita lepita (©Tshetsholo Naro NEN)

mae rane Sa Serean femae (©Tshetsholo Naro NEN) Naro (©Tshetsholo Athyma cama cama (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae Sa Serean Athyma selenophora selenophora

mae rane Sa Serean mae Athyma cama cama (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae Bue ue Bassarona durga splendens (©Tshetsholo Naro NEN) mae rea Calinaga brahma brahma (©Tshetsholo Naro NEN) mae Bue ue Bassarona durga splendens

(©Tshetsholo Naro NEN) Naro (©Tshetsholo mae Bronze ue Euthalia nara mae renh ue Euthalia franciae nagaensis franciae (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae reen ue Euthalia sahadeva nadaka

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

(©Tshetsholo Naro NEN)

(©Tshetsholo Naro NEN) mae aud Baron Euthalia lubentina lubentina (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae hieeded Bue Baron Euthalia phemius phemius

(©Tshetsholo Naro NEN) Naro (©Tshetsholo mae Nama Saier Neptis f namba namba mae eo Saior Neptis ananta ochracea (©Tshetsholo Naro NEN) (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae Nama Saier Neptis f namba NEN) Naro (©Tshetsholo namba mae Broadanded Saier Neptis sankara amba mae Broadanded Saier Neptis sankara amba (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae aas Saier Neptis sappho mae Commander Moduza procris mae aas Saier Neptis sappho astola astola procris

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae ndian oroiseshe Aglais caschmirensis aesis (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae Cier Parthenos sylvia mae ono Araschnia prorsoides gambrisius dohertyi

(©Tshetsholo Naro NEN) Naro (©Tshetsholo mae uumn eaf Doleschallia mae ono Araschnia prorsoides bisaltide indica dohertyi mae rane aeaf Kallima inachus inachus (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae rane aeaf Kallima inachus inachus

mae rea fl Hypolimnas bolina jacintha (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae rea fl Hypolimnas bolina mae eao ans Junonia almana jacintha almana

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

(©Tshetsholo Naro NEN) (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae eo ans Junonia hierta mae Chooae ans Junonia iphita mae eoman ans Junonia lemonias hierta iphita lemonias (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae Bue ans Junonia orithya ocyale mae Bue dmira Kaniska canace canace (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) mae Sotted eser Symbrenthia hypselis cotanda (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae Sotted eser Symbrenthia hypselis cotanda mae Common eser Symbrenthia lilaea khasiana (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae Bueaied eser Symbrenthia niphanda niphanda

(©Tshetsholo Naro NEN) (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae Bueaied eser Symbrenthia mae ained ad Vanessa cardui niphanda niphanda cardui

mae ndian ed dmira Vanessa indica indica

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

mae Consae Dichorragia nesimachus nesimachus (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae are aun Faunis eumeus assama mae Common uer Discophora sondaica zal

(©Tshetsholo Naro NEN) (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae une or Thaumantis diores diores (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae Common amfl Elymnias hypermnestra undularis mae Sotted amfl Elymnias malelas malelas (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae Callerebia oria

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

Image 145. Spotted Palmfly Elymnias malelas malelas ©Tshetsholo Naro NEN) Naro ( ©Tshetsholo Image 147. Common Treebrown Image 146. Banded Treebrown Lethe rohria rohria Lethe confusa confusa ©Tshetsholo Naro NEN) Naro ( ©Tshetsholo ©Tshetsholo Naro NEN) Naro ( ©Tshetsholo

Image 148. Straight-banded Treebrown NEN) Naro ( ©Tshetsholo Lethe verma sintica Image 148. Straight-Banded Treebrown Lethe verma sintica

Image 149. Angled Red Forester Lethe chandica chandica ©Tshetsholo Naro NEN) Naro ( ©Tshetsholo ©Tshetsholo Naro NEN) Naro ( ©Tshetsholo

(©Tshetsholo Naro NEN) Image 150. Common Red Forester Image 150. Common Red Forester Lethe mekara mekara Lethe mekara mekara

Image 151. Tailed Red Forester Lethe sinorix sinorix

Image 153. Pale Forester Lethe latiaris Image 152. Bamboo Forester Lethe cf. kansa latiaris Image 153. Pale Forester Lethe latiaris latiaris

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 6627 Butterflies of Chizami Naro & Sondhi (©Tshetsholo Naro NEN) Naro (©Tshetsholo NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae iafor Lethe f sura mae Common enin Bron mae Common oodron Melanitis leda leda Lethe sidonis (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae Common enin Bron mae ar enin Bron Melanitis phedima bela Melanitis leda leda mae ar enin Bron Melanitis phedima bela

mae rea enin Bron Melanitis zitenius zitenius mae hieine Brushron Heteropsis malsara (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo

(©Tshetsholo Naro NEN) mae hieine Brushron Heterop- sis malsara mae iaine Bushron Mycalesis francisca sanatana mae eha Brushron Mycalesis lepcha kohimensis

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

mae eined arinh Neope pulaha pulaha mae hieeded Busron Mycalesis mestra sadona (© Sanjay Sondhi) mae eined arinh Neope pulaha pulaha

mae Ypthima s

mae imaaan ierin Ypthima

(©Tshetsholo Naro NEN) Naro (©Tshetsholo sakra sakra mae imaaan ierin Ypthima sakra sakra © Kewekhrozo Thopi NEN (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae reen raonai Lamproptera mae ass BueotteGraphium meges indistincta mae aied a Graphium agamemnon cloanthus cloanthus agamemnon

mae Common BueotteGraphium sarpedon sarpedon (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae ed een Papilio helenus mae eo een Papilio nephelus helenus chaon (©Tshetsholo Naro NEN) mae esser ime Papilio epycides epycides (©Tshetsholo Naro NEN) (©Tshetsholo Naro NEN) Naro (©Tshetsholo

mae Common ormon Papilio polytes mae edreas Papilio alcmenor romulus alcmenor (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae rea ormon Papilio memnon agenor

(©Tshetsholo Naro NEN) (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae ime Butterfl Papilio demoleus demoleus

mae Sane Papilio protenor mae Bue eao Papilo arcturus euprotenor arcturus

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi

mae aris eao Papilio paris paris NEN) Naro (©Tshetsholo mae Common Bain Atrophaneura varuna astorion mae Common eao Papilio bianor ganesa

(©Tshetsholo Naro NEN) NEN) Naro (©Tshetsholo mae hreeso rass eo mae Common indmi NEN) Naro (©Tshetsholo Eurema blanda silhetana Byasa polyeuctes polyeuctes mae Sma rass eo Eurema brigitta rubella (©Tshetsholo Naro NEN) Naro (©Tshetsholo NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae Soess rass eo mae Common rass eo mae Common aross Appias albina Eurema laeta sikkima Eurema hecabe hecabe darada

(©Tshetsholo Naro NEN) (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae rane aross Appias galba mae Chooae aross Appias lyncida eleonora (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae So un Appias lalage lalage

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae Common miran Catopsilia NEN) Naro (©Tshetsholo mae esser u Cepora nadina nadina pomona pomona mae otted miran Catopsilia pyranthe pyranthe (©Tshetsholo Naro NEN) Naro (©Tshetsholo

(©Tshetsholo Naro NEN) Naro (©Tshetsholo mae i ezee Delias belladonna mae edreas ezee Delias acalis lugens pyramus (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae edso ezee Delias descombesi descombesi (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae rea rane Hebomoia glaucippe glaucippe (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae eo rane Ixias pyrene mae edase ezee Delias pasithoe familiaris pasithoe (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae ae anderer Pareronia avatar mae are Caae hie Pieris mae ndian Caae hie brassicae nepalensis Pieris canidia indica

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 Butterflies of Chizami Naro & Sondhi (©Tshetsholo Naro NEN) Naro (©Tshetsholo NEN) Naro (©Tshetsholo

(©Tshetsholo Naro NEN) Naro (©Tshetsholo mae reeniened hie Pieris melete mae Sotted SaoohPrioneris ajaka thestylis thestylis mae Bah hie Pontia daplidice moorei (©Tshetsholo Naro NEN) Naro (©Tshetsholo (©Tshetsholo Naro NEN) Naro (©Tshetsholo mae Sried unh Dodona adonira NEN) Naro (©Tshetsholo mae ar ud Abisara fylla adonira mae unhineo Zemeros flegyas fleygas

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 6633 Butterflies of Chizami Naro & Sondhi

a b

endi rimari isied areas and heir haias sheshoo Naro NN a eu rai rue ier an ie of hesumi rai durin monsoon d ie of Norh as Neor NN omound

hreaened aa

Journal of Threatened Taxa | www.threatenedtaxa.org | 12 December 2014 | 6(13): 6593–6634 JOTT RECOGNIZES THIS CONSERVATION INITIATIVE

EMPOWERING YOUNGSTERS IN CONSERVING BIODIVERSITY - CHIZAMI EXPERIENCE

The Hoolock Gibbon Eco Club (HGEC) was formed in 2010. This was the first eco-club to be formed in Nagaland and has been a pioneering effort of an NGO called the North East Network (NEN). The club has 50 children in the age group of 10–14 years from the village of Chizami and the neighbouring village of Enhulumi. HGEC was an outcome of a wildlife education program to aid in mitigating wild animal hunting titled “Under the Canopy” that Dusty foot Productions (www.dustyfootindia.com) and Go Wild (payalmolur.blogspot.in) conducted in 2009. While filming a story on orphaned bear cubs, the Dusty Foot Production team came across rampant hunting and selling of wild meat in northeastern India. This led them to team up with a wildlife education group Go Wild Workshops to create an education program and curriculum to shift the focus from hunting to research, photography and enquiry. North East Network and its tryst with wildlife education and conservation began in 2009 when approached by the Rita Banerji (Dusty Foot Productions) and Payal Molur (Go Wild). North East Network (www.northeastnetwork. org) is a woman’s rights organization in northeast India, established in 1995. NEN’s role has been that of a facilitator to empower women of northeast A few of the publications of HGEC around issues of livelihood, health, conflict and governance through capacity building, awareness - First and second edition booklets with 171 and raising, networking, research and advocacy. North 217 species of butterflies of Chizami respectively East Network and its tryst with wildlife education (2012 and 2014) and conservation began in 2009 when approached - A booklet on the Birds of Chizami (2014) by the Rita Banerji (Dusty Foot Productions) and - Posters on wildlife of Chizami, Birds of Chizami Payal Molur (Go Wild) to be the local partner. Titli and Butterflies and Moths of Chizami trust (www.titlitrust.com), was also brought in to - Naro, T. (2012). Sighting of Aglais bring a new dimension to the scientific education cashmirensis aesis Fruhstorfer, 1912 (Nymphalidae) program. From there they have expanded the program from Nagaland, India. Journal of Threatened Taxa exponentially and now have trainers and clubs in 4(4): 2534–2535; http://dx.doi.org/10.11609/ three districts of Nagaland. JoTT.o3019.2534-5 The teachers and children trained in Nagaland, - Naro, T. & S. Sondhi (2013). Sightings of have not only understood the importance of Bath White Pontia daplidice moorei Röber, 1907 conserving their forests but also have taken it to the (Lepidoptera: Pieridae: Pierinae: Pierini) from next stage of maintaining biodiversity registers Arunachal Pradesh and Nagaland, India. Journal of and cataloguing the animals that are around them Threatened Taxa 5(7): 4122–4124; http://dx.doi. in a scientific manner. org/10.11609/JoTT.o3308.4122-4 Over the past four years the club has been churning out a series of wildlife publications in the Payal Molur form of posters, booklets, journal articles and popular [email protected] articles: Rita Banerji explaing the importance of lighting.

Chizami Village, Phek, Nagaland

Sanjay Sondhi describing the beauty of butterflies at the NEN campus. Understanding macro photography techniques

Photographing some animals is easier with a tripod

Diligently cataloging moths

Photo credits: Payal B Molur and Tshetsholo Naro

Payal Molur explaining how to map an area