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Molecular Phylogeny, Divergence Times and Biogeography of Spiders of the Subfamily Euophryinae (Araneae: Salticidae) ⇑ Jun-Xia Zhang A, , Wayne P
Molecular Phylogenetics and Evolution 68 (2013) 81–92 Contents lists available at SciVerse ScienceDirect Molec ular Phylo genetics and Evolution journal homepage: www.elsevier.com/locate/ympev Molecular phylogeny, divergence times and biogeography of spiders of the subfamily Euophryinae (Araneae: Salticidae) ⇑ Jun-Xia Zhang a, , Wayne P. Maddison a,b a Department of Zoology, University of British Columbia, Vancouver, BC, Canada V6T 1Z4 b Department of Botany and Beaty Biodiversity Museum, University of British Columbia, Vancouver, BC, Canada V6T 1Z4 article info abstract Article history: We investigate phylogenetic relationships of the jumping spider subfamily Euophryinae, diverse in spe- Received 10 August 2012 cies and genera in both the Old World and New World. DNA sequence data of four gene regions (nuclear: Revised 17 February 2013 28S, Actin 5C; mitochondrial: 16S-ND1, COI) were collected from 263 jumping spider species. The molec- Accepted 13 March 2013 ular phylogeny obtained by Bayesian, likelihood and parsimony methods strongly supports the mono- Available online 28 March 2013 phyly of a Euophryinae re-delimited to include 85 genera. Diolenius and its relatives are shown to be euophryines. Euophryines from different continental regions generally form separate clades on the phy- Keywords: logeny, with few cases of mixture. Known fossils of jumping spiders were used to calibrate a divergence Phylogeny time analysis, which suggests most divergences of euophryines were after the Eocene. Given the diver- Temporal divergence Biogeography gence times, several intercontinental dispersal event sare required to explain the distribution of euophry- Intercontinental dispersal ines. Early transitions of continental distribution between the Old and New World may have been Euophryinae facilitated by the Antarctic land bridge, which euophryines may have been uniquely able to exploit Diolenius because of their apparent cold tolerance. -
Further Study of Two Chinese Cave Spiders 77 Doi: 10.3897/Zookeys.870.35971 RESEARCH ARTICLE Launched to Accelerate Biodiversity Research
A peer-reviewed open-access journal ZooKeys 870: 77–100 (2019) Further study of two Chinese cave spiders 77 doi: 10.3897/zookeys.870.35971 RESEARCH ARTICLE http://zookeys.pensoft.net Launched to accelerate biodiversity research Further study of two Chinese cave spiders (Araneae, Mysmenidae), with description of a new genus Chengcheng Feng1, Jeremy A. Miller2, Yucheng Lin1, Yunfei Shu1 1 Key Laboratory of Bio-Resources and Eco-Environment of Ministry of Education, College of Life Sciences, Sichuan University, Chengdu 610065, Sichuan, China 2 Department of Biodiversity Discovery, Naturalis Biodiversity Center, Postbus 9517 2300 RA Leiden, The Netherlands Corresponding author: Yucheng Lin ([email protected]) Academic editor: Charles Haddad | Received 08 May 2019 | Accepted 09 July 2019 | Published 7 August 2019 http://zoobank.org/4167F0DE-2097-4F3D-A608-3C8365754F99 Citation: Feng C, Miller JA, Lin Y, Shu Y(2019) Further study of two Chinese cave spiders (Araneae, Mysmenidae), with description of a new genus. ZooKeys 870: 77–100. https://doi.org/10.3897/zookeys.870.35971 Abstract The current paper expands knowledge of two Chinese cave spider species originally described in the genus Maymena Gertsch, 1960: M. paquini Miller, Griswold & Yin, 2009 and M. kehen Miller, Griswold & Yin, 2009. With the exception of these two species, the genus Maymena is endemic to the western hemisphere, and new evidence presented here supports the creation of a new genus for the Chinese species, which we name Yamaneta gen. nov. The male of Y. kehen is described for the first time. Detailed illustrations of the habitus, male palps and epigyne are provided for these two species, as well as descriptions of their webs. -
Comparative Functional Morphology of Attachment Devices in Arachnida
Comparative functional morphology of attachment devices in Arachnida Vergleichende Funktionsmorphologie der Haftstrukturen bei Spinnentieren (Arthropoda: Arachnida) DISSERTATION zur Erlangung des akademischen Grades doctor rerum naturalium (Dr. rer. nat.) an der Mathematisch-Naturwissenschaftlichen Fakultät der Christian-Albrechts-Universität zu Kiel vorgelegt von Jonas Otto Wolff geboren am 20. September 1986 in Bergen auf Rügen Kiel, den 2. Juni 2015 Erster Gutachter: Prof. Stanislav N. Gorb _ Zweiter Gutachter: Dr. Dirk Brandis _ Tag der mündlichen Prüfung: 17. Juli 2015 _ Zum Druck genehmigt: 17. Juli 2015 _ gez. Prof. Dr. Wolfgang J. Duschl, Dekan Acknowledgements I owe Prof. Stanislav Gorb a great debt of gratitude. He taught me all skills to get a researcher and gave me all freedom to follow my ideas. I am very thankful for the opportunity to work in an active, fruitful and friendly research environment, with an interdisciplinary team and excellent laboratory equipment. I like to express my gratitude to Esther Appel, Joachim Oesert and Dr. Jan Michels for their kind and enthusiastic support on microscopy techniques. I thank Dr. Thomas Kleinteich and Dr. Jana Willkommen for their guidance on the µCt. For the fruitful discussions and numerous information on physical questions I like to thank Dr. Lars Heepe. I thank Dr. Clemens Schaber for his collaboration and great ideas on how to measure the adhesive forces of the tiny glue droplets of harvestmen. I thank Angela Veenendaal and Bettina Sattler for their kind help on administration issues. Especially I thank my students Ingo Grawe, Fabienne Frost, Marina Wirth and André Karstedt for their commitment and input of ideas. -
Opiliones, Palpatores, Caddoidea)
Shear, W. A. 1975 . The opilionid family Caddidae in North America, with notes on species from othe r regions (Opiliones, Palpatores, Caddoidea) . J. Arachnol . 2:65-88 . THE OPILIONID FAMILY CADDIDAE IN NORTH AMERICA, WITH NOTES ON SPECIES FROM OTHER REGION S (OPILIONES, PALPATORES, CADDOIDEA ) William A . Shear Biology Departmen t Hampden-Sydney, College Hampden-Sydney, Virginia 23943 ABSTRACT Species belonging to the opilionid genera Caddo, Acropsopilio, Austropsopilio and Cadella are herein considered to constitute the family Caddidae . The subfamily Caddinae contains the genu s Caddo ; the other genera are placed in the subfamily Acropsopilioninae. It is suggested that the palpatorid Opiliones be grouped in three superfamilies : Caddoidea (including the family Caddidae) , Phalangioidea (including the families Phalangiidae, Liobunidae, Neopilionidae and Sclerosomatidae ) and Troguloidea (including the families Trogulidae, Nemostomatidae, Ischyropsalidae an d Sabaconidae). North American members of the Caddidae are discussed in detail, and a new species , Caddo pepperella, is described . The North American caddids appear to be mostly parthenogenetic, an d C. pepperella is very likely a neotenic isolate of C. agilis. Illustrations and taxonomic notes ar e provided for the majority of the exotic species of the family . INTRODUCTION Considerable confusion has surrounded the taxonomy of the order Opiliones in North America, since the early work of the prolific Nathan Banks, who described many of ou r species in the last decade of the 1800's and the first few years of this century. For many species, no additional descriptive material has been published following the original de- scriptions, most of which were brief and concentrated on such characters as color and body proportions . -
Salticidae (Arachnida, Araneae) of Islands Off Australia
1999. The Journal of Arachnology 27:229±235 SALTICIDAE (ARACHNIDA, ARANEAE) OF ISLANDS OFF AUSTRALIA Barbara Patoleta and Marek ZÇ abka: Zaklad Zoologii WSRP, 08±110 Siedlce, Poland ABSTRACT. Thirty nine species of Salticidae from 33 Australian islands are analyzed with respect to their total distribution, dispersal possibilities and relations with the continental fauna. The possibility of the Torres Strait islands as a dispersal route for salticids is discussed. The studies of island faunas have been the ocean level ¯uctuations over the last 50,000 subject of zoogeographical and evolutionary years, at least some islands have been sub- research for over 150 years and have resulted merged or formed land bridges with the con- in hundreds of papers, with the syntheses by tinent (e.g., Torres Strait islands). All these Carlquist (1965, 1974) and MacArthur & Wil- circumstances and the human occupation son (1967) being the best known. make it rather unlikely for the majority of Modern zoogeographical analyses, based islands to have developed their own endemic on island spider faunas, began some 60 years salticid faunas. ago (Berland 1934) and have continued ever When one of us (MZ) began research on since by, e.g., Forster (1975), Lehtinen (1980, the Australian and New Guinean Salticidae 1996), Baert et al. (1989), ZÇ abka (1988, 1990, over ten years ago, close relationships be- 1991, 1993), Baert & Jocque (1993), Gillespie tween the faunas of these two regions were (1993), Gillespie et al. (1994), ProÂszynÂski expected. Consequently, it was hypothesized (1992, 1996) and Berry et al. (1996, 1997), that the Cape York Peninsula and Torres Strait but only a few papers were based on veri®ed islands were the natural passage for dispersal/ and suf®cient taxonomic data. -
The Short-Range Endemic Invertebrate Fauna of the Ravensthorpe Range
THE SHORT-RANGE ENDEMIC INVERTEBRATE FAUNA OF THE RAVENSTHORPE RANGE MARK S. HARVEY MEI CHEN LENG Department of Terrestrial Zoology Western Australian Museum June 2008 2 Executive Summary An intensive survey of short-range endemic invertebrates in the Ravensthorpe Range at 79 sites revealed a small but significant fauna of myriapods and arachnids. Four species of short-range endemic invertebrates were found: • The millipede Antichiropus sp. R • The millipede Atelomastix sp. C • The millipede Atelomastix sp. P • The pseudoscorpion Amblyolpium sp. “WA1” Atelomastix sp. C is the only species found to be endemic to the Ravensthorpe Range and was found at 14 sites. Antichiropus sp. R, Atelomastix sp. P and Amblyolpium sp. “WA1” are also found at nearby locations. Sites of high importance include: site 40 with 7 species; sites 7 and 48 each with 5 species; and sites 18 and 44 each with 4 species. WA Museum - Ravensthorpe Range Survey 3 Introduction Australia contains a multitude of terrestrial invertebrate fauna species, with many yet to be discovered and described. Arthropods alone were recently estimated to consist of approximately more than 250,000 species (Yeates et al. 2004). The majority of these belong to the arthropod classes Insecta and Arachnida, and although many have relatively wide distributions across the landscape, some are highly restricted in range with special ecological requirements. These taxa, termed short-range endemics (Harvey 2002b), are taxa categorised as having poor dispersal abilities and/or requiring very specific habitats, usually with naturally small distributional ranges of less than 10,000 km2 and the following ecological and life-history traits: • poor powers of dispersal; • confinement to discontinuous habitats; • usually highly seasonal, only active during cooler, wetter periods; and • low levels of fecundity. -
Higher-Level Phylogenetics of Linyphiid Spiders (Araneae, Linyphiidae) Based on Morphological and Molecular Evidence
Cladistics Cladistics 25 (2009) 231–262 10.1111/j.1096-0031.2009.00249.x Higher-level phylogenetics of linyphiid spiders (Araneae, Linyphiidae) based on morphological and molecular evidence Miquel A. Arnedoa,*, Gustavo Hormigab and Nikolaj Scharff c aDepartament Biologia Animal, Universitat de Barcelona, Av. Diagonal 645, E-8028 Barcelona, Spain; bDepartment of Biological Sciences, The George Washington University, Washington, DC 20052, USA; cDepartment of Entomology, Natural History Museum of Denmark, Zoological Museum, University of Copenhagen, Universitetsparken 15, DK-2100 Copenhagen, Denmark Accepted 19 November 2008 Abstract This study infers the higher-level cladistic relationships of linyphiid spiders from five genes (mitochondrial CO1, 16S; nuclear 28S, 18S, histone H3) and morphological data. In total, the character matrix includes 47 taxa: 35 linyphiids representing the currently used subfamilies of Linyphiidae (Stemonyphantinae, Mynogleninae, Erigoninae, and Linyphiinae (Micronetini plus Linyphiini)) and 12 outgroup species representing nine araneoid families (Pimoidae, Theridiidae, Nesticidae, Synotaxidae, Cyatholipidae, Mysmenidae, Theridiosomatidae, Tetragnathidae, and Araneidae). The morphological characters include those used in recent studies of linyphiid phylogenetics, covering both genitalic and somatic morphology. Different sequence alignments and analytical methods produce different cladistic hypotheses. Lack of congruence among different analyses is, in part, due to the shifting placement of Labulla, Pityohyphantes, -
First Records and Three New Species of the Family Symphytognathidae
ZooKeys 1012: 21–53 (2021) A peer-reviewed open-access journal doi: 10.3897/zookeys.1012.57047 RESEARCH ARTICLE https://zookeys.pensoft.net Launched to accelerate biodiversity research First records and three new species of the family Symphytognathidae (Arachnida, Araneae) from Thailand, and the circumscription of the genus Crassignatha Wunderlich, 1995 Francisco Andres Rivera-Quiroz1,2, Booppa Petcharad3, Jeremy A. Miller1 1 Department of Terrestrial Zoology, Understanding Evolution group, Naturalis Biodiversity Center, Darwin- weg 2, 2333CR Leiden, the Netherlands 2 Institute for Biology Leiden (IBL), Leiden University, Sylviusweg 72, 2333BE Leiden, the Netherlands 3 Faculty of Science and Technology, Thammasat University, Rangsit, Pathum Thani, 12121 Thailand Corresponding author: Francisco Andres Rivera-Quiroz ([email protected]) Academic editor: D. Dimitrov | Received 29 July 2020 | Accepted 30 September 2020 | Published 26 January 2021 http://zoobank.org/4B5ACAB0-5322-4893-BC53-B4A48F8DC20C Citation: Rivera-Quiroz FA, Petcharad B, Miller JA (2021) First records and three new species of the family Symphytognathidae (Arachnida, Araneae) from Thailand, and the circumscription of the genus Crassignatha Wunderlich, 1995. ZooKeys 1012: 21–53. https://doi.org/10.3897/zookeys.1012.57047 Abstract The family Symphytognathidae is reported from Thailand for the first time. Three new species: Anapistula choojaiae sp. nov., Crassignatha seeliam sp. nov., and Crassignatha seedam sp. nov. are described and illustrated. Distribution is expanded and additional morphological data are reported for Patu shiluensis Lin & Li, 2009. Specimens were collected in Thailand between July and August 2018. The newly described species were found in the north mountainous region of Chiang Mai, and Patu shiluensis was collected in the coastal region of Phuket. -
On the Australian Linyphiid Spider Alaxchelicera Ordinaria Butler, 1932 (Araneae)
Zootaxa 3750 (2): 193–196 ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Correspondence ZOOTAXA Copyright © 2013 Magnolia Press ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.3750.2.8 http://zoobank.org/urn:lsid:zoobank.org:pub:9515170F-60D0-43D3-A936-81E16EBEE6C3 On the Australian linyphiid spider Alaxchelicera ordinaria Butler, 1932 (Araneae) NIKOLAJ SCHARFF1 & GUSTAVO HORMIGA2 1Natural History Museum of Denmark, Zoological Museum and Center for Macroecology, Evolution and Climate, Universitetsparken 15, DK-2100 Copenhagen, Denmark. E-mail: [email protected] 2Department of Biological Sciences, The George Washington University, Washington, D.C. 20052, USA. E-mail: [email protected] Very few studies have addressed the linyphiid fauna of Australia. Most of the existing taxonomic work on Australian linyphiids consists of isolated species descriptions (e.g., Rainbow 1912) or at most are based on small number of species also described outside a revisionary context (e.g., Wunderlich 1976) (but see van Helsdingen 1972 for a revision of the Australian species of the genera Laperousea Dalmas, 1917 and Laetesia Simon, 1908). Microctenonyx subitaneus (O. P.-Cambridge, 1875) is a Holarctic erigonine (Linyphiidae) which has been introduced in many parts of the world, including Australia (Brennan 2004). In this paper we report a new junior synonym of Microctenonyx subitaneus described by Butler (1932) under the name Alaxchelicera ordinaria Butler, 1932. In 1932 L.S.G. Butler, a Melbourne-based arachnologist, published a paper in which he described six new spider genera from Victoria and New South Wales, all of them monotypic. Three of these new genera (Microlinypheus, Plectochetos and Alaxchelicera) he placed in the family Linyphiidae, the remaining three (Platycephala, Eterosonycha and Perissopmeros) were placed in Zodariidae. -
Spiders 27 November-5 December 2018 Submitted: August 2019 Robert Raven
Bush Blitz – Namadgi, ACT 27 Nov-5 Dec 2018 Namadgi, ACT Bush Blitz Spiders 27 November-5 December 2018 Submitted: August 2019 Robert Raven Nomenclature and taxonomy used in this report is consistent with: The Australian Faunal Directory (AFD) http://www.environment.gov.au/biodiversity/abrs/online-resources/fauna/afd/home Page 1 of 12 Bush Blitz – Namadgi, ACT 27 Nov-5 Dec 2018 Contents Contents .................................................................................................................................. 2 List of contributors ................................................................................................................... 2 Abstract ................................................................................................................................... 4 1. Introduction ...................................................................................................................... 4 2. Methods .......................................................................................................................... 4 2.1 Site selection ............................................................................................................. 4 2.2 Survey techniques ..................................................................................................... 4 2.2.1 Methods used at standard survey sites ................................................................... 5 2.3 Identifying the collections ......................................................................................... -
WO 2017/035099 Al 2 March 2017 (02.03.2017) P O P C T
(12) INTERNATIONAL APPLICATION PUBLISHED UNDER THE PATENT COOPERATION TREATY (PCT) (19) World Intellectual Property Organization International Bureau (10) International Publication Number (43) International Publication Date WO 2017/035099 Al 2 March 2017 (02.03.2017) P O P C T (51) International Patent Classification: BZ, CA, CH, CL, CN, CO, CR, CU, CZ, DE, DK, DM, C07C 39/00 (2006.01) C07D 303/32 (2006.01) DO, DZ, EC, EE, EG, ES, FI, GB, GD, GE, GH, GM, GT, C07C 49/242 (2006.01) HN, HR, HU, ID, IL, IN, IR, IS, JP, KE, KG, KN, KP, KR, KZ, LA, LC, LK, LR, LS, LU, LY, MA, MD, ME, MG, (21) International Application Number: MK, MN, MW, MX, MY, MZ, NA, NG, NI, NO, NZ, OM, PCT/US20 16/048092 PA, PE, PG, PH, PL, PT, QA, RO, RS, RU, RW, SA, SC, (22) International Filing Date: SD, SE, SG, SK, SL, SM, ST, SV, SY, TH, TJ, TM, TN, 22 August 2016 (22.08.2016) TR, TT, TZ, UA, UG, US, UZ, VC, VN, ZA, ZM, ZW. (25) Filing Language: English (84) Designated States (unless otherwise indicated, for every kind of regional protection available): ARIPO (BW, GH, (26) Publication Language: English GM, KE, LR, LS, MW, MZ, NA, RW, SD, SL, ST, SZ, (30) Priority Data: TZ, UG, ZM, ZW), Eurasian (AM, AZ, BY, KG, KZ, RU, 62/208,662 22 August 2015 (22.08.2015) US TJ, TM), European (AL, AT, BE, BG, CH, CY, CZ, DE, DK, EE, ES, FI, FR, GB, GR, HR, HU, IE, IS, IT, LT, LU, (71) Applicant: NEOZYME INTERNATIONAL, INC. -
Segmentation and Tagmosis in Chelicerata
Arthropod Structure & Development 46 (2017) 395e418 Contents lists available at ScienceDirect Arthropod Structure & Development journal homepage: www.elsevier.com/locate/asd Segmentation and tagmosis in Chelicerata * Jason A. Dunlop a, , James C. Lamsdell b a Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Invalidenstrasse 43, D-10115 Berlin, Germany b American Museum of Natural History, Division of Paleontology, Central Park West at 79th St, New York, NY 10024, USA article info abstract Article history: Patterns of segmentation and tagmosis are reviewed for Chelicerata. Depending on the outgroup, che- Received 4 April 2016 licerate origins are either among taxa with an anterior tagma of six somites, or taxa in which the ap- Accepted 18 May 2016 pendages of somite I became increasingly raptorial. All Chelicerata have appendage I as a chelate or Available online 21 June 2016 clasp-knife chelicera. The basic trend has obviously been to consolidate food-gathering and walking limbs as a prosoma and respiratory appendages on the opisthosoma. However, the boundary of the Keywords: prosoma is debatable in that some taxa have functionally incorporated somite VII and/or its appendages Arthropoda into the prosoma. Euchelicerata can be defined on having plate-like opisthosomal appendages, further Chelicerata fi Tagmosis modi ed within Arachnida. Total somite counts for Chelicerata range from a maximum of nineteen in Prosoma groups like Scorpiones and the extinct Eurypterida down to seven in modern Pycnogonida. Mites may Opisthosoma also show reduced somite counts, but reconstructing segmentation in these animals remains chal- lenging. Several innovations relating to tagmosis or the appendages borne on particular somites are summarised here as putative apomorphies of individual higher taxa.