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Chapter 1 - Introduction
EURASIAN MIDDLE AND LATE MIOCENE HOMINOID PALEOBIOGEOGRAPHY AND THE GEOGRAPHIC ORIGINS OF THE HOMININAE by Mariam C. Nargolwalla A thesis submitted in conformity with the requirements for the degree of Doctor of Philosophy Graduate Department of Anthropology University of Toronto © Copyright by M. Nargolwalla (2009) Eurasian Middle and Late Miocene Hominoid Paleobiogeography and the Geographic Origins of the Homininae Mariam C. Nargolwalla Doctor of Philosophy Department of Anthropology University of Toronto 2009 Abstract The origin and diversification of great apes and humans is among the most researched and debated series of events in the evolutionary history of the Primates. A fundamental part of understanding these events involves reconstructing paleoenvironmental and paleogeographic patterns in the Eurasian Miocene; a time period and geographic expanse rich in evidence of lineage origins and dispersals of numerous mammalian lineages, including apes. Traditionally, the geographic origin of the African ape and human lineage is considered to have occurred in Africa, however, an alternative hypothesis favouring a Eurasian origin has been proposed. This hypothesis suggests that that after an initial dispersal from Africa to Eurasia at ~17Ma and subsequent radiation from Spain to China, fossil apes disperse back to Africa at least once and found the African ape and human lineage in the late Miocene. The purpose of this study is to test the Eurasian origin hypothesis through the analysis of spatial and temporal patterns of distribution, in situ evolution, interprovincial and intercontinental dispersals of Eurasian terrestrial mammals in response to environmental factors. Using the NOW and Paleobiology databases, together with data collected through survey and excavation of middle and late Miocene vertebrate localities in Hungary and Romania, taphonomic bias and sampling completeness of Eurasian faunas are assessed. -
Stratigraphic and Paleoenvironmental Reconstruction of a Mid-Pliocene Fossil Site in the High Arctic (Ellesmere Island, Nunavut)
ARCTIC VOL. 69, NO. 2 (JUNE 2016) P. 185 – 204 http://dx.doi.org/10.14430/arctic4567 Stratigraphic and Paleoenvironmental Reconstruction of a Mid-Pliocene Fossil Site in the High Arctic (Ellesmere Island, Nunavut): Evidence of an Ancient Peatland with Beaver Activity William Travis Mitchell,1 Natalia Rybczynski,2,3 Claudia Schröder-Adams,1 Paul B. Hamilton,4 Robin Smith2 and Marianne Douglas5 (Received 12 June 2015; accepted in revised form 27 January 2016) ABSTRACT. Neogene terrestrial deposits of sand and gravel with preserved wood and peat accumulations occur in many areas of the High Arctic. The Pliocene-aged Beaver Pond fossil site (Ellesmere Island, NU) is one such site that differs from other sites in the great thickness of its peat layer and the presence of a rich vertebrate faunal assemblage, along with numerous beaver-cut sticks. Although the site has been the subject of intense paleontological investigations for over two decades, there has not been a reconstruction of its depositional history. In this study, measured sections within and surrounding the site established the stratigraphy and lateral continuity of the stratigraphic units. Grain size analysis, loss on ignition, and fossil diatom assemblages were examined to reconstruct paleoenvironmental changes in the sequence. The base of the section was interpreted as a floodplain system. Using modern peat accumulation rates, the maximum thickness (240 cm) of the overlying peat layer is estimated to represent 49 000 ± 12 000 years. From this evidence, we suggest that during the peat formation interval, beaver activity may have played a role in creating an open water environment. -
Cranial Morphology of the Oligocene Beaver Capacikala Gradatus from the John Day Basin and Comments on the Genus
Palaeontologia Electronica palaeo-electronica.org Cranial morphology of the Oligocene beaver Capacikala gradatus from the John Day Basin and comments on the genus Clara Stefen ABSTRACT The cranial morphology of the small Oligocene beaver Capacikala gradatus is described on the basis of a well preserved, nearly complete skull and partial mandibles from the John Day Formation, John Day Fossil Beds, Oregon, USA. The only nearly complete skull known so far from the same area as the type specimen is described here in detail. This is especially appropriate as the type specimen comes from an unknown locality within the John Day Formation and is only a fragmentary skull. The newly described specimen was found between dated marker beds, so that it can be no older than 28.7 Ma, nor younger than 27.89 Ma. Although Capacikala had been named 50 years ago (MacDonald, 1963), it is still not well known. Morphological comparisons are made to other mentioned or illustrated specimens of Capacikala, Palaeocastor and recent Castor; there are similarities and differences to both genera. The findings of the skull is discussed in comparison to the description of the genera Capacikala and Palaeocastor and some characters are revised. A phylogenetic analysis with few selected castorid species was performed, but resulted in poorly supported trees. How- ever, a complete revision of beaver phylogeny and of the characters used is beyond the scope of the paper. Clara Stefen. Senckenberg Naturhistorische Sammlungen Dresden, Museum of Zoology, Königsbrücker Landstrasse 159, 01109 Dresden, Germany, [email protected] KEY WORDS: Castoridae; Palaeocastorinae; skull; Tertiary INTRODUCTION Fremd et al., 1994; Hunt and Stepleton, 2004; Samuels and Zancanella, 2011). -
PROXY RECONSTRUCTIONS of PLIOCENE CLIMATE by Adam
DECIPHERING ARCTIC CLIMATE IN A PAST GREENHOUSE WORLD: MULTI- PROXY RECONSTRUCTIONS OF PLIOCENE CLIMATE by Adam Zoltan Csank _____________________ A Dissertation Submitted to the Faculty of the DEPARTMENT OF GEOSCIENCES In Partial Fulfillment of the Requirements For the Degree of DOCTOR OF PHILOSOPHY In the Graduate College THE UNIVERSITY OF ARIZONA 2011 2 THE UNIVERSITY OF ARIZONA GRADUATE COLLEGE As members of the Dissertation Committee, we certify that we have read the dissertation prepared by Adam Zoltan Csank entitled Deciphering Arctic Climate in a Past Greenhouse World: Multi-Proxy Reconstructions of Pliocene Climate and recommend that it be accepted as fulfilling the dissertation requirement for the Degree of Doctor of Philosophy _______________________________________________________________________ Date: Feb 4 2011 STEVEN LEAVITT _______________________________________________________________________ Date: Feb 4 2011 JONATHAN OVERPECK _______________________________________________________________________ Date: Feb 4 2011 JOELLEN RUSSELL _______________________________________________________________________ Date: Feb 4 2011 MALCOLM HUGHES _______________________________________________________________________ Date: Feb 4 2011 MICHAEL EVANS Final approval and acceptance of this dissertation is contingent upon the candidate’s submission of the final copies of the dissertation to the Graduate College. I hereby certify that I have read this dissertation prepared under my direction and recommend that it be accepted as fulfilling the dissertation requirement. ________________________________________________ Date: Mar 4, 2011 Dissertation Director: STEVEN LEAVITT 3 STATEMENT BY AUTHOR This document has been submitted in partial fulfillment of requirements for an advanced degree at the University of Arizona and is deposited in the University Library to be made available to borrowers under rules of the Library. Brief quotations from this document are allowable without special permission, provided that accurate acknowledgement of source is made. -
A Brief Overview of the Evolution of European Tertiary Beavers C
BALTIC FORESTRY A BRIEF OVERVIEW OF THE EVOLUTION OF EUROPEAN TERTIARY BEAVERS C. STEFEN REVIEW PAPER A Brief Overview of the Evolution of European Tertiary Beavers CLARA STEFEN Senckenberg Naturhistorische Sammlungen Dresden, Museum für Tierkunde, Königsbrücker Landstraße 159, 01109 Dresden, Germany, e-mail: [email protected] Baltic Forestry 17(1): 148153. Abstract Beavers are a well defined group of rodents that are known from the late Eocene to recent times. Today only one genus with two species, Castor fiber and Castor canadensis both with several described subspecies occur in Eurasia and North America respectively. During the Tertiary beavers were much more diverse in North America, Europe and Asia ranging in size from about a mice to a capybara. In Europe there is one major radiation starting in the Late Oligocene or Early Miocene with Propalaeocastor (= Astenofiber) or Steneofiber, with several subsequent species of Steneofiber, leading to Chalicomys, and Trogontherium. Some generic distinctions are still not quite clear here. Probable immigrants to Europe include Anchitheriomys and Dipoides. One difficulty in understanding castorid evolution is posed by the fact, that especially European beavers are often found in low numbers and the bulk of the fossil material consists of isolated teeth and mandibular or small maxillary fragments. Skulls and even substantial skull fragments are generally rare. As Recent Castor and larger samples of fossil beavers show great morphometric variability, it is strongly suggested to discuss beaver taxonomy on the basis of populations or representative statistical samples, not based on individual teeth or extremely fragmentary material. More names and descriptions will not help our understanding of this group of rodents. -
Late Neogene Beaver Dipoides from the Pipe Creek Sinkhole of Grant County, Indiana
Paludicola 9(4):167-175 April 2014 © by the Rochester Institute of Vertebrate Paleontology LATE NEOGENE BEAVER DIPOIDES FROM THE PIPE CREEK SINKHOLE OF GRANT COUNTY, INDIANA Robert A. Martin Department of Biological Sciences, Murray State University, Murray, KY 42071 ABSTRACT Two upper fourth premolars of the diminutive beaver Dipoides from the early Blancan Pipe Creek Sinkhole of Indiana were compared with new Dipoides wilsoni material from the Saw Rock Canyon 1 and Fallen Angel B assemblages of Seward County, Kansas and published accounts of other North American Dipoides. The Pipe Creek Sinkhole teeth are larger than those of most other late Neogene Dipoides. Although high-crowned and ever-growing, upper fourth premolars of the Pipe Creek Sinkhole beaver are less hypsodont than other samples with which they were compared. Despite this, both teeth are relatively advanced, with complete or nearly complete pseudostria 3 and the absence of dentine channels connecting lophs 2 and 3. The Pipe Creek Dipoides is probably not referable to D. wilsoni, but the small sample size does not currently allow allocation to other named species or the naming of a new species. A revised dental terminology is presented for the castoroidinin beavers. INTRODUCTION wilsoni Hibbard, 1949, D. rexroadensis Hibbard and Riggs, 1949, D. smithi Shotwell, 1955) D. vallicula The Pipe Creek Sinkhole in Grant County, Indiana, Shotwell, 1970, D. intermedius Zakrzewski, 1969 and D. uncovered accidentally during limestone quarrying tanneri Korth, 1998. D. intermedius has since been operations (Farlow et al., 2001; Martin et al., 2002; referred to Procastoroides (Ruez, Jr., 2009). Shotwell Farlow et al., 2010a), represents an important window (1955) observed that the holotype of D. -
Miocene Carnivorans from the Vallès-Penedès Basin (NE Iberian Peninsula)
Departament de Biologia Animal, de Biologia Vegetal i d’Ecologia Unitat d’Antropologia Biològica Miocene carnivorans from the Vallès-Penedès Basin (NE Iberian Peninsula) Josep Maria Robles Giménez Tesi Doctoral 2014 A mi padre y familia. INDEX Index .......................................................................................................................... 7 Preface and Acknowledgments [in Spanish] ....................................................... 13 I.–Introduction and Methodology ........................................................................ 19 Chapter 1. General introduction and aims of this dissertation .......................... 21 1.1. Aims and structure of this work .............................................................. 21 Motivation of this dissertation ................................................................ 21 Type of dissertation and general overview ............................................. 22 1.2. An introduction to the Carnivora ............................................................ 24 What is a carnivoran? ............................................................................. 24 Biology .................................................................................................... 25 Systematics and phylogeny ...................................................................... 28 Evolutionary history ................................................................................ 42 1.3. Carnivoran anatomy ............................................................................... -
Osteology of a Complete Skeleton of Dipoides Stirtoni (Rodentia, Castoridae) from the Late Miocene of Northern Oregon
Paludicola 10(4):189-206 June 2016 © by the Rochester Institute of Vertebrate Paleontology OSTEOLOGY OF A COMPLETE SKELETON OF DIPOIDES STIRTONI (RODENTIA, CASTORIDAE) FROM THE LATE MIOCENE OF NORTHERN OREGON James E. Martin1,2 and Shawna L. Johnson3 1University of Louisiana Geology Museum, School of Geosciences, University of Louisiana, Lafayette, Louisiana 70504 <[email protected]> 2J. E. Martin Geoscientific Consultation, 21051 Doral Court, Sturgis, South Dakota 57785 <[email protected]> 3Steven W. Corathers Associates, Sheridan, Wyoming 82801 <[email protected]> ABSTRACT The most complete skeleton of the small castorid, Dipoides stirtoni, was collected from late Miocene (early Hemphillian North American Land Mammal Age) deposits in northern Oregon. The specimen is spectacularly preserved, even including sesamoids; missing only the distal portion of the tail, which was eroded prior to discovery. The individual was found in a crouched position and appears to have been preserved in a burrow. The articulated skeleton exhibits incisors with rounded 4 enamel faces, parastriae/parastriids on P /P4, and elongate hind limbs, particularly the rear feet. The transverse processes of the caudal vertebrae are expanded laterally, but not so much as those of extant Castor canadensis. The proportions of the forelimbs are unlike those of C. canadensis, and could be interpreted as indicating that the Miocene beaver had a greater capability for burrowing. Overall, D. stirtoni appears to have been adapted for a swimming lifestyle, although perhaps not quite to the degree as that of C. canadensis. INTRODUCTION The castorid skeleton, University of Washington, complete skeleton, even the sesamoids were preserved. Burke Museum (UWBM) 59242, was collected in 1978 The dentition exhibits the typical occlusal S-pattern by the first author from south of the town of Arlington characteristic of Dipoides (Shotwell, 1955) and is in Gilliam County, northern Oregon (Figure 1). -
The Stratigraphic Sequence of North American Rodent Faunas
273 THE STRATIGRAPHIC SEQUENCE OF NORTH AMERICAN RODENT FAUNAS by Robert W. WILSON* CONTENT page Abstract, Zusammenfassung, Resume ................................ , . .. 274 Introduction ... .. 274 Wasatchian I Neustrian - Earliest Rhenanian I Earlier Eocene ... .. 275 Bridgerian - Uintan I Middle and Late Rhenanian - Early Headonian I Later Eocene . .. 276 Chadronian I Late Headonian - Early Suevian I Early Oligocene ..................... 277 Orellan - Whitneyan I Late Suevian - Arvernian I Later Oligocene . .. 278 Arikareean - Hemingfordian ! Agenian - Orleanian I Latest Oligocene - Earlier Miocene ................................................. 279 Barstovian I Astaracian I Middle Miocene .................................... 279 Clarendonian - Hemphillian I Vallesian - Turolian - Ruseinian I Late and Latest Miocene . .. 280 B1ancan I Villanyian I Pliocene . .. 280 Irvingtonian I Biharian I Earlier Pleistocene .................................. 281 Rancholabrean I Later Pleistocene . .. 281 Summary ......................................................... 282 Literature cited ..................................................... 282 * Associate, Mllseum of Natural History, The University of Kansas, Lawrence. Palaeovertebrata, Montpellier, Mem. Jubil. R. Lavocat : 273·283) (Acceptc le 7 Juin 1979, public le 31 Octobre 1980) 274 ABSTRACT Rodents first appear ill the latest Paleocene or earliest Eoccne as very fragmentary specimens (FamilY Para myidae) known largely from a single locality. After this sparse beginning. rodents are usually -
Giant Beaver (Castoroides) Palaeoecology Inferred from Stable Isotopes
Western University Scholarship@Western Electronic Thesis and Dissertation Repository 10-27-2016 12:00 AM Giant Beaver (Castoroides) Palaeoecology Inferred from Stable Isotopes Tessa Plint The University of Western Ontario Supervisor Fred J. Longstaffe The University of Western Ontario Graduate Program in Geology A thesis submitted in partial fulfillment of the equirr ements for the degree in Master of Science © Tessa Plint 2016 Follow this and additional works at: https://ir.lib.uwo.ca/etd Part of the Biogeochemistry Commons, Paleobiology Commons, and the Paleontology Commons Recommended Citation Plint, Tessa, "Giant Beaver (Castoroides) Palaeoecology Inferred from Stable Isotopes" (2016). Electronic Thesis and Dissertation Repository. 4236. https://ir.lib.uwo.ca/etd/4236 This Dissertation/Thesis is brought to you for free and open access by Scholarship@Western. It has been accepted for inclusion in Electronic Thesis and Dissertation Repository by an authorized administrator of Scholarship@Western. For more information, please contact [email protected]. Abstract Stable isotope analysis was used to explore unresolved questions about the palaeoecology of the extinct Pleistocene giant beaver (Castoroides). The 13C, δ15N, and 18O of bone collagen and structural carbonate from enamel served as proxies for palaeodiet and palaeoclimate. A new baseline for freshwater and terrestrial plant 13C and δ15N was utilized in a mixing model (SIAR) to assess rodent feeding niche. SIAR analysis indicated that Castoroides’ consumed a diet of predominantly macrophytes, making them reliant on wetland habitat for food. Based on isotopic data for potential modern analog species (Castor and Ondatra), SIAR also indicated that Castor and Castoroides occupied complementary dietary niches, allowing them to share habitat space throughout the Pleistocene. -
Qt4xr1s918.Pdf
UC Berkeley PaleoBios Title A Machairodont felid (Mammalia; Carnivora; Felidae) from the latest Hemphillian (Late Miocene/Early Pliocene) Bidahochi Formation, northeastern Arizona Permalink https://escholarship.org/uc/item/4xr1s918 Journal PaleoBios, 29(3) ISSN 0031-0298 Author Hodnett, John-Paul Publication Date 2010-06-01 DOI 10.5070/P9293021800 Peer reviewed eScholarship.org Powered by the California Digital Library University of California PaleoBios 29(3):76-91, June 1, 2010 © 2010 University of California Museum of Paleontology A Machairodont felid (Mammalia; Carnivora; Felidae) from the latest Hemphillian (Late Miocene/Early Pliocene) Bidahochi Formation, northeastern Arizona John-PaUl hodnett department of Biological Sciences, northern arizona University, Flagstaff, aZ 86011; [email protected] a lower jaw from the White Cone local fauna of the latest hemphillian Bidahochi Formation in northern arizona is the first description of a felid from this fauna and the first positively identified occurrence of the smilodontine mach- airodont Paramachairodus in north america. this lower jaw has characters identical to those seen in a similar sized machairodont felid from the Bone Valley Formation of Florida, suggesting that the same taxon is present in Florida. the diversity of the hemphillian machairodonts and the taxonomic status of Megantereon hesperus is reviewed. the characteristics of the ramus corpus and dentition places Paramachairodus firmly within the Smilodontini and adds further support that the more derived smilodontine machairodonts, Megantereon and Smilodon, had their origins in north america. IntRodUCtIon Ptychocheilus prelucius, and a new species of minnow, Evomus In the early 1950s the United States Geological Survey navaho (Uyeno and Miller 1965). Breed (1973) described conducted a geological survey on the navajo reservation aquatic bird tracks and fossils of Olor columbianus, the extant (Repenning et al. -
Evolution of Woodcutting Behaviour in Early Pliocene Beaver Driven by Consumption of Woody Plants Tessa Plint1,2*, Fred J
www.nature.com/scientificreports OPEN Evolution of woodcutting behaviour in Early Pliocene beaver driven by consumption of woody plants Tessa Plint1,2*, Fred J. Longstafe1, Ashley Ballantyne3, Alice Telka4,7 & Natalia Rybczynski5,6 Modern beavers (Castor) are prolifc ecosystem engineers and dramatically alter the landscape through tree harvesting and dam building. Little is known, however, about the evolutionary drivers of their woodcutting behaviour. Here we investigate if early woodcutting behaviour in Castoridae was driven by nutritional needs. We measured stable carbon and nitrogen isotopes (δ13C and δ15N) of coeval subfossil plants and beaver collagen (Dipoides sp.) from the Early Pliocene, High Arctic Beaver Pond fossil locality (Ellesmere Island), in order to reconstruct Dipoides sp. diet. Isotopic evidence indicates a diet of woody plants and freshwater macrophytes, supporting the hypothesis that this extinct semiaquatic beaver engaged in woodcutting behaviour for feeding purposes. In a phylogenetic context, the isotopic evidence implies that woodcutting and consumption of woody plants can be traced back to a small-bodied, semiaquatic Miocene castorid, suggesting that beavers have been consuming woody plants for over 20 million years. We propose that the behavioural complex (swimming, woodcutting, and consuming woody plants) preceded and facilitated the evolution of dam building. Dam building and food caching behaviours appear to be specializations for cold winter survival and may have evolved in response to late Neogene northern cooling. Beavers today are renowned for their woodcutting behaviour and construction abilities. Te extant genus Castor harvests trees and shrubs for sustenance (particularly during the winter1), but also for the purpose of lodge and dam building.