Surface Tension Dominates Insect Flight on Fluid Interfaces Haripriya Mukundarajan1, Thibaut C
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Nova Scotia Provincial Status Report Spotted Pondweed
Nova Scotia Provincial Status Report on Spotted Pondweed (Potamogeton pulcher Tuckerm.) prepared for Nova Scotia Species at Risk Working Group by David Mazerolle and Sean Blaney Atlantic Canada Conservation Data Centre P.O. Box 6416, Sackville, NB E4L 1C6 DRAFT Funding provided by Nova Scotia Department of Natural Resources Submitted December 2010 EXECUTIVE SUMMARY i TABLE OF CONTENTS EXECUTIVE SUMMARY ..................................................................................................i WILDLIFE SPECIES DESCRIPTION AND SIGNIFICANCE...........................................1 Name and Classification............................................................................................1 Morphological Description ........................................................................................2 Field identification......................................................................................................3 Designatable Units .....................................................................................................4 Special Significance...................................................................................................5 DISTRIBUTION ...............................................................................................................7 Global Range ..............................................................................................................7 Canadian Range .........................................................................................................8 -
The Evolutionary Significance of Body Size in Burying Beetles
Brigham Young University Masthead Logo BYU ScholarsArchive All Theses and Dissertations 2018-04-01 The volutE ionary Significance of Body Size in Burying Beetles Ashlee Nichole Momcilovich Brigham Young University Follow this and additional works at: https://scholarsarchive.byu.edu/etd BYU ScholarsArchive Citation Momcilovich, Ashlee Nichole, "The vE olutionary Significance of Body Size in Burying Beetles" (2018). All Theses and Dissertations. 7327. https://scholarsarchive.byu.edu/etd/7327 This Dissertation is brought to you for free and open access by BYU ScholarsArchive. It has been accepted for inclusion in All Theses and Dissertations by an authorized administrator of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. The Evolutionary Significance of Body Size in Burying Beetles Ashlee Nichole Momcilovich A dissertation submitted to the faculty of Brigham Young University in partial fulfillment of the requirements for the degree of Doctor of Philosophy Mark C. Belk, Chair Seth M. Bybee Jerald B. Johnson Steven L. Peck G. Bruce Schaalje Department of Biology Brigham Young University Copyright © 2018 Ashlee Nichole Momcilovich All Rights Reserved ABSTRACT The Evolutionary Significance of Body Size in Burying Beetles Ashlee Nichole Momcilovich Department of Biology, BYU Doctor of Philosophy Body size is one of the most commonly studied traits of an organism, which is largely due to its direct correlation with fitness, life history strategy, and physiology of the organism. Patterns of body size distribution are also often studied. The distribution of body size within species is looked at for suggestions of differential mating strategies or niche variation among ontogenetic development. Patterns are also examined among species to determine the effects of competition, environmental factors, and phylogenetic inertia. -
Flexural Stiffness Patterns of Butterfly Wings (Papilionoidea)
35:61–77,Journal of Research1996 (2000) on the Lepidoptera 35:61–77, 1996 (2000) 61 Flexural stiffness patterns of butterfly wings (Papilionoidea) Scott J. Steppan Committee on Evolutionary Biology, University of Chicago, Chicago, IL 60637, USA., E-mail: [email protected] Abstract. A flying insect generates aerodynamic forces through the ac- tive manipulation of the wing and the “passive” properties of deformability and wing shape. To investigate these “passive” properties, the flexural stiffness of dried forewings belonging to 10 butterfly species was compared to the butterflies’ gross morphological parameters to determine allom- etric relationships. The results show that flexural stiffness scales with wing 3.9 loading to nearly the fourth power (pw ) and is highly correlated with wing area cubed (S3.1). The generalized map of flexural stiffness along the wing span for Vanessa cardui has a reduction in stiffness near the distal tip and a large reduction near the base. The distal regions of the wings are stiffer against forces applied to the ventral side, while the basal region is much stiffer against forces applied dorsally. The null hypothesis of structural isom- etry as the explanation for flexural stiffness scaling is rejected. Instead, selection for a consistent dynamic wing geometry (angular deflection) in flight may be a major factor controlling general wing stiffness and deformability. Possible relationships to aerodynamic and flight habit fac- tors are discussed. This study proposes a new approach to addressing the mechanics of insect flight and these preliminary results need to be tested using fresh wings and more thorough sampling. KEY WORDS: biomechanics, butterfly wings, flight, allometry, flexural stiff- ness, aerodynamics INTRODUCTION A flying insect generates aerodynamic forces primarily through the ac- tive manipulation of wing movements and the “passive” morphological prop- erties of deformability and wing shape. -
Adhesion Performance in the Eggs of the Philippine Leaf Insect Phyllium Philippinicum (Phasmatodea: Phylliidae)
insects Article Adhesion Performance in the Eggs of the Philippine Leaf Insect Phyllium philippinicum (Phasmatodea: Phylliidae) Thies H. Büscher * , Elise Quigley and Stanislav N. Gorb Department of Functional Morphology and Biomechanics, Institute of Zoology, Kiel University, Am Botanischen Garten 9, 24118 Kiel, Germany; [email protected] (E.Q.); [email protected] (S.N.G.) * Correspondence: [email protected] Received: 12 June 2020; Accepted: 25 June 2020; Published: 28 June 2020 Abstract: Leaf insects (Phasmatodea: Phylliidae) exhibit perfect crypsis imitating leaves. Although the special appearance of the eggs of the species Phyllium philippinicum, which imitate plant seeds, has received attention in different taxonomic studies, the attachment capability of the eggs remains rather anecdotical. Weherein elucidate the specialized attachment mechanism of the eggs of this species and provide the first experimental approach to systematically characterize the functional properties of their adhesion by using different microscopy techniques and attachment force measurements on substrates with differing degrees of roughness and surface chemistry, as well as repetitive attachment/detachment cycles while under the influence of water contact. We found that a combination of folded exochorionic structures (pinnae) and a film of adhesive secretion contribute to attachment, which both respond to water. Adhesion is initiated by the glue, which becomes fluid through hydration, enabling adaption to the surface profile. Hierarchically structured pinnae support the spreading of the glue and reinforcement of the film. This combination aids the egg’s surface in adapting to the surface roughness, yet the attachment strength is additionally influenced by the egg’s surface chemistry, favoring hydrophilic substrates. -
UDC 595.786 E.V. Guskova, G.N. Kuftina NEW INFORMATION ON
Biological Bulletin of Bogdan Chmelnitskiy Melitopol State Pedagogical University 177 UDC 595.786 E.V. Guskova, G.N. Kuftina NEW INFORMATION ON THE TROPHIC SPECIALIZATION OF GALERUCELLA NYMPHAEAE (LINNAEUS, 1758) (COLEOPTERA, CHRYSOMELIDAE) OF ALTAI KRAI Altai State University, Lenin Str. 61, RUS-656049, Barnaul, Russia. Тел. (913) 2202577. E-mail: [email protected] Feeding of Galerucella nymphaeae (Linnaeus, 1758) (Coleoptera, Chrysomelidae) on leaves of water chestnut (Trapa natans L.) was registered in Siberia for the first time. Key words: Galerucella nymphaeae L., Chrysomelidae, Trapa natans L., trophic relations, Altai Krai. INTRODUCTION Galerucella nymphaeae (Linnaeus, 1758) (Coleoptera, Chrysomelidae) is a widespread holarctic species (Beenen, 2010). The literature reports that the basic food plants of this leaf beetle are white water lily (Nymphaea alba) and yellow water- lily (Nuphar lutea) (Nymphaeaceae) (Medvedev & Roginskaya, 1988; Guskova, 2001). However the representatives of this species can feed on other plants: Polygonum, Rumex (Polygonaceae), Potentilla (Rosaceae) and others (Brovdiy, 1973), and its nutrition on currants has been registered in Siberia (Dubeshko & Medvedev, 1989). RESULTS In Zmeinogorskiy district of Altai Krai on Lake Kolyvanskoye on the 25th of August, 2013, G. nymphaea beetles and instar larvae actively feeding on leaves of water chestnut Trapa natans L. (Trapaceae) were registered in large quantity (Figs 1−2). Fig. 1. Russia, Altai Krai, Zmeinogorskii district, lake Kolyvanskoe. ISSN 2225-5486 (Print), ISSN 2226-9010 (Online). Біологічний вісник МДПУ. 2016. №1 178 Бiологiчний вiсник Despite the availability of large amounts of yellow and white water lilies in the lake, the larvae and beetles fed only on floating leaves of water chestnut. -
The Flight of the Bumble
The Flight of the Bumble Bee Grade Span 4 Subject Area • Science • Math Materials • Fab@School Maker Studio • Digital fabricator or scissors • 65lb or 110lb cardstock • Pencils, pens, markers, or crayons • Stapler, tape, or glue Online Resources • Website: The Physics- There is a common myth that bumble bees defy the laws of Defying Flight of the physics as they apply to aerodynamics. Basically, they shouldn’t Bumblebee be able to fly. Using high-speed photography, Michael Dickinson, • Website: Bumblebees Can a professor of biology and insect flight expert at the University of Fly Into Thin Air Washington, published an article in 2005 all about the why and how the bumblebee takes flight. Through this Fab@School Maker Studio • Video: Bumble Bee in Slow activity, your students will examine the anatomy of a bumblebee or Motion other flying insect. • Article: Lasers Illuminate the Flight of the Bumblebee • PDF: Fab@School Quick Objective Start Guide Author • Using Fab@School Maker Studio, students will design a 2D Denine Jimmerson, or 3D prototype of a bumble bee (or another flying insect) Executive Director of that demonstrates how its structure serves it in its function Curriculum and Design, of flight. FableVision Learning © 2017 FableVisionLearning, LLC • The Flight of the Bumble Bee www.FableVisionLearning.com • [email protected] 1 Fab@School Maker Studio • www.FabMakerStudio.com Big Idea Challenge Functions help to determine form. Show the Bumble Bee in Slow Motion video to your students. Pose the question: How do bees fly? Driving Question How do bees fly? Explain to the students that they will be researching how bumble bees (or other insects/animals) fly. -
The Novel Aerodynamics of Insect Flight: Applications to Micro-Air Vehicles
The Journal of Experimental Biology 202, 3439–3448 (1999) 3439 Printed in Great Britain © The Company of Biologists Limited 1999 JEB2214 THE NOVEL AERODYNAMICS OF INSECT FLIGHT: APPLICATIONS TO MICRO-AIR VEHICLES C. P. ELLINGTON* Department of Zoology, University of Cambridge, Downing Street, Cambridge CB2 3EJ, UK *e-mail: [email protected] Accepted 15 September; published on WWW 16 November 1999 Summary The wing motion in free flight has been described for accompanies repeated clapping of the wings, but the spiral insects ranging from 1 to 100 mm in wingspan. To support LEV can be used to augment the lift production of the body weight, the wings typically produce 2–3 times propellers, rotors and micro-air vehicles (MAVs). Design more lift than can be accounted for by conventional characteristics of insect-based flying machines are aerodynamics. Some insects use the fling mechanism: the presented, along with estimates of the mass supported, the wings are clapped together and then flung open before the mechanical power requirement and maximum flight speeds start of the downstroke, creating a lift-enhancing vortex over a wide range of sizes and frequencies. To support a around each wing. Most insects, however, rely on a leading- given mass, larger machines need less power, but smaller edge vortex (LEV) created by dynamic stall during ones operating at higher frequencies will reach faster flapping; a strong spanwise flow is also generated by the speeds. pressure gradients on the flapping wing, causing the LEV to spiral out to the wingtip. Technical applications of the Key words: insect, flight, aerodynamics, flapping flight, micro-air fling are limited by the mechanical damage that vehicle. -
Galerucella Nymphaeae
PDF hosted at the Radboud Repository of the Radboud University Nijmegen The following full text is a publisher's version. For additional information about this publication click this link. http://hdl.handle.net/2066/19064 Please be advised that this information was generated on 2021-10-08 and may be subject to change. Evolution in action: Host race formation in Galerucella nymphaeae Stephanie Pappers 6WHOOLQJHQ behorende bij het proefschrift Evolution in action: host race formation in Galerucella nymphaeae Stephanie Pappers 1. Het waterleliehaantje, Galerucella nymphaeae, bestaat uit minstens twee gastheerrassen. 2. Differentiatie is mogelijk zonder geografische barrière, ook in de natuur. 3. Nothing in biology makes sense except in the light of evolution. 'RE]KDQVN\ $PHULFDQ %LRORJ\ 7HDFKHU 4. Als de vroege christenen beter naar Empedocles in plaats van Aristoteles hadden geluisterd had Darwin het niet zo moeilijk gehad. 5. De biologie heeft mij over de Schepper geleerd dat Hij in ieder geval een bijzondere voorkeur voor kevers heeft. 'H %ULWVH JHQHWLFXV -%6 +DOGDQH YROJHQV +XWFKLQVRQ $PHULFDQ 1DWXUDOLVW 6. Met de toenemende vergrijzing bij de universiteit zal ook het papierverbruik toenemen, al is het alleen al omdat een steeds groter lettertype nodig is. 7. Aangezien de meeste stadsduiven afstammen van verwilderde uitheemse rotsduiven zouden ze dus, alleen al uit oogpunt van natuurbescherming en het tegengaan van genetische vervuiling van de inheemse duiven- soorten, bestreden moeten worden. 8. Natuur is overal waar je mobieltje -
Study of Turning Takeoff Maneuver in Free-Flying Dragonflies
Study of turning takeoff maneuver in free-flying dragonflies: effect of dynamic coupling Samane Zeyghami1, Haibo Dong2 Mechanical and Aerospace Engineering, University of Virginia, VA, 22902 SUMMARY Turning takeoff flight of several dragonflies was recorded, using a high speed photogrammetry system. In this flight, dragonfly takes off while changing the flight direction at the same time. Center of mass was elevated about 1-2 body lengths. Five of these maneuvers were selected for 3D body surface reconstruction and the body orientation measurement. In all of these flights, 3D trajectories and transient rotations of the dragonfly’s body were observed. In oppose to conventional banked turn model, which ignores interactions between the rotational motions, in this study we investigated the strength of the dynamic coupling by dividing pitch, roll and yaw angular accelerations into two contributions; one from aerodynamic torque and one from dynamic coupling effect. The latter term is referred to as Dynamic Coupling Acceleration (DCA). The DCA term can be measured directly from instantaneous rotational velocities of the insect. We found a strong correlation between pitch and yaw velocities at the end of each wingbeat and the time integral of the corresponding DCA term. Generation of pitch, roll and yaw torques requires different aerodynamic mechanisms and is limited due to the other requirements of the flight. Our results suggest that employing DCA term gives the insect capability to perform a variety of maneuvers without fine adjustments in the aerodynamic torque. Key words: insect flight, aerial maneuver, dragonfly, pitch. INTRODUCTION Aerial maneuvers of different insect species are studied widely in the last decades and many aspects of maneuverability in insect flight are explored. -
Dragonfly Flight Quick Z
Dragonfly flight quick Z. Jane Wang Dragonflies have evolved for about 350 million years. What kinds of study aerodynamic tricks have they discovered? Jane Wang is an associate professor of theoretical and applied mechanics at Cornell University in Ithaca, New York. Summer in Ithaca, New York, is a good time to watch tional to the product of velocity squared, fluid density, and dragonflies. At a glance, you can see that a dragonfly has a wing area. That product can be interpreted as the rate of prominent head, an elongated body, and two pairs of slender momentum transfer from air particles that hit the wing and wings extending to each side. As it takes off, the wings appear bounce off. But if the particle picture were the whole story, as a blur. In air, the dragonfly dances in unpredictable steps, planes would not be able to carry much nor would they be hovering briefly then quickly moving to a new location. Just very efficient. Based on the particle picture, one would pre- when you think it might stay long enough in the viewfinder dict—as Isaac Newton did—a lift proportional to sin2α for a of your camera, poof! It is gone. In contrast, an airplane, noisy wing with angle of attack α. That’s much smaller than ob- and powerful, has a more straightforward way of going about served, at least for a small angle of attack. The same calcula- its business. Propelled by engines and lifted by wings, it wastes tion also predicts a maximum lift-to-drag ratio of 1 at an no time in going from one place to another. -
Forest Health Technology Enterprise Team Biological Control of Invasive
Forest Health Technology Enterprise Team TECHNOLOGY TRANSFER Biological Control Biological Control of Invasive Plants in the Eastern United States Roy Van Driesche Bernd Blossey Mark Hoddle Suzanne Lyon Richard Reardon Forest Health Technology Enterprise Team—Morgantown, West Virginia United States Forest FHTET-2002-04 Department of Service August 2002 Agriculture BIOLOGICAL CONTROL OF INVASIVE PLANTS IN THE EASTERN UNITED STATES BIOLOGICAL CONTROL OF INVASIVE PLANTS IN THE EASTERN UNITED STATES Technical Coordinators Roy Van Driesche and Suzanne Lyon Department of Entomology, University of Massachusets, Amherst, MA Bernd Blossey Department of Natural Resources, Cornell University, Ithaca, NY Mark Hoddle Department of Entomology, University of California, Riverside, CA Richard Reardon Forest Health Technology Enterprise Team, USDA, Forest Service, Morgantown, WV USDA Forest Service Publication FHTET-2002-04 ACKNOWLEDGMENTS We thank the authors of the individual chap- We would also like to thank the U.S. Depart- ters for their expertise in reviewing and summariz- ment of Agriculture–Forest Service, Forest Health ing the literature and providing current information Technology Enterprise Team, Morgantown, West on biological control of the major invasive plants in Virginia, for providing funding for the preparation the Eastern United States. and printing of this publication. G. Keith Douce, David Moorhead, and Charles Additional copies of this publication can be or- Bargeron of the Bugwood Network, University of dered from the Bulletin Distribution Center, Uni- Georgia (Tifton, Ga.), managed and digitized the pho- versity of Massachusetts, Amherst, MA 01003, (413) tographs and illustrations used in this publication and 545-2717; or Mark Hoddle, Department of Entomol- produced the CD-ROM accompanying this book. -
Flight of the Butterflies Educator Guide Table of Contents
Educator Guide 1 Flight of the Butterflies Educator Guide Table of Contents Welcome Monarch Butterflies – Background Information Educational Content in the Film Education Standards Planting A Butterfly Garden An Activity for All Grade Levels K-2 Classroom Activities Getting To Know Your Caterpillars What Is A Butterfly Habitat? Make a Wall Mural Monarch Migration Game 3-6 Classroom Activities Keying Out Kids How Far Can A Butterfly Glide? Insect Metamorphosis – A Bug’s Life The Very Hungry Caterpillar Warning Coloration You Don’t Taste the Way You Look: Understanding Mimicry 7-12 Classroom Activities Schoolyard Phenology Rearing Monarch Larvae Monarchs in the Balance Dilemma Cards How Many Grandchildren? Comparing Butterflies and Moths Vocabulary References Acknowledgements 2 Welcome to the Fascinating World of the Monarch Butterfly! Most school science curriculum includes the study of butterflies as well as concepts of migration, ecology, biodiversity and the process of scientific discovery. This Educator Guide summary provides science information for educators, a source of curriculum specific activities, vocabulary, and web and print resources for further investigation. More detailed educational activities and background information on the film, Flight of the Butterflies, are included on the enclosed disc of Educational Support Material and on the project web site www.flightofthebutterflies.com/learningcentre. Flight of the Butterflies is now available in Giant Screen/IMAX® 3D and 2D theaters worldwide and could be playing in your area. Some theaters are part of museums with live butterfly pavilions. We encourage you to schedule a field trip to see the film in conjunction with exploring activities in this Educator Guide and Support Materials.