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CURRICULUM VITAE Benjamin David Greenberg, MD Phd EDUCATION Undergraduate: Amherst College Medical School: University of Miami

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CURRICULUM VITAE Benjamin David Greenberg, MD Phd EDUCATION Undergraduate: Amherst College Medical School: University of Miami CURRICULUM VITAE Benjamin David Greenberg, MD PhD EDUCATION Undergraduate: Amherst College Medical School: University of Miami Graduate School: University of California, San Diego (UCSD), Neurosciences POSTGRADUATE TRAINING Residency: New York University VA Medical Center Internship in Internal Medicine Neurological Institute of New York, Columbia-Presbyterian Medical Resident in Neurology Johns Hopkins Hospital Residency in Psychiatry Fellowships: Harvard School of Public Health, Postdoctoral Fellowship, Interdisciplinary Programs in Health National Institute of Mental Health Intramural Program, Senior Staff Fellow (Commissioned Corps), Section on Clinical Neuropharmacology, Laboratory of Clinical Science POSTGRADUATE HONORS AND AWARDS Best Review Article of the Year, CNS Spectrums, 2003 Research Mentor Award, Brown University Department of Psychiatry and Human Behavior, 2001 Early Career Award, EEG and Clinical Neuroscience Society (ECNS), 10/2001 APA/SmithKline Beecham Young Faculty Award, Hon. Mention, 2000 Young Investigator Award, American Neuropsychiatric Assn, 1995 DISTA Fellowship, Society of Biological Psychiatry, 1991 PROFESSIONAL LICENCES AND BOARD CERTIFICATION Rhode Island Medical License Maryland Medical License Diplomate in Psychiatry, American Board of Psychiatry and Neurology ACADEMIC APPOINTMENTS Associate Professor, Department of Psychiatry and Human Behavior, Brown University 2/2000 - Present Greenberg 2 Chief, Adult Obsessive-Compulsive Disorder Research Unit, National Institute of Mental Health, Bethesda MD, 7/96-1/2000 Senior Staff Fellow, Laboratory of Clinical Science, Section on Clinical Neuropharmacology, National Institute of Mental Health, Bethesda MD, 7/92 – 1/2000 Instructor in Psychiatry, Johns Hopkins University, Baltimore MD, 1992-2000 OTHER APPOINTMENTS Board of Directors, Depression and Related Affective Disorders Association, 1997-2000 Executive Board, EEG and Clinical Neuroscience Society, 2004- ACADEMIC COMMITTEES Brown Medical Faculty Executive Committee, 2003 - Brown Collaboration for Translational Brain Research, 2002- Brown Psychiatry/Psychology Working Group, 2002- Brown Brain Science Program Executive Committee, 2000- Brown University MRI Facility Executive Committee, 2001- Brown University MRI Facility Scientific Advisory Committee, 2001- Interdepartmental Advisory Committee for Psychology, Brown University, 2001 Brown University Psychiatry Residency Curriculum Committee Brown University Brain Science Curriculum Committee, 2001-2 Brown University Psychiatry Residency UTP Clinic Committee, 2000 Director, Brown Psychiatry Residency PGY3 Rotation, Butler Hospital, 2000- Associate Chair, Butler Hospital Institutional Review Board, 2000 - NIMH Intramural Program Institutional Review Board, 1995-1999 NIMH Fellows Committee, 1994-1997 MEMBERSHIP IN SOCIETIES American College of Neuropsychopharmacology, Associate Member, 1999 Obsessive-Compulsive Foundation, 1998 Founding member, International Society for Transcranial Stimulation, 1997 American Neuropsychiatric Association, 1997 American Society of Clinical Psychopharmacology, 1996-1999 Society of Biological Psychiatry, 1992 American Psychiatric Association, Maryland Psychiatric Society, 1992 PUBLICATIONS (peer reviewed) 1. Vosbeck KD, Greenberg BD, and Awad WM: The proteolytic enzymes of the K-1 strain of Streptomyces griseus obtained from a commercial preparation (Pronase): Specificity and immobilization of aminopeptidase. Journal of Biological Chemistry 1975; 250(10): 3981-3987. Greenberg 3 2. Vosbeck KD, Greenberg BD, Ochoa MS, Whitney PL, and Award WM: Proteolytic enzymes of K-1 strain of Streptomyces griseus obtained from commercial preparation (Pronase): Effect of pH, metal ions, and amino acids on aminopeptidase activity. Journal of Biological Chemistry 1978; 253(1): 257-260. 3. Geyer MA, Segal DS, and Greenberg BD: Increased startle responding in rats treated with phencyclidine. Neurobehavioral Toxicology and Teratology 1984; 6: 161-164. 4. Russell JW, Greenberg BD, and Segal DS: Effects of phencyclidine on spontaneous aggressive behavior in the rat. Biological Psychiatry 1984; 19(2): 195-202. 5. Greenberg BD, Moore PM, Letz RE, and Baker EL: Computerized assessment of human neurotoxicity: Sensitivity to nitrous oxide exposure. Clinical Pharmacology and Therapeutics 1985; 38(6): 656-660. 6. Greenberg BD, and Segal DS: Acute and chronic behavioral interactions between phencyclidine and amphetamine: Evidence for involvement of dopaminergic mechanisms in some PCP-induced behaviors. Pharmacology, Biochemistry and Behavior 1985; 23(1): 99-105. 7. Greenberg BD, and Segal DS: Evidence for multiple opiate receptor involvement in different phencyclidine-induced unconditioned behaviors in rats. Psychopharmacology 1986; 88: 44- 53. 8. Benjamin J, Li L, Patterson C, Greenberg BD, Murphy, DL, Hamer DH: Population and familial association between the D4 dopamine receptor gene and measures of novelty seeking. Nature Genetics 1996; 12:81-84. 9. Michelson D, Altemus M, Galliven E, Hill D, Greenberg BD, Gold P: Naloxone-induced pituitary-adrenal activation does not differ in patients with depression, obsessive-compulsive disorder and healthy controls. Neuropsychopharmacology 1996; 15:207-12. 10. Keuler D, Altemus M, Galliven E, Hill D, Greenberg B, Murphy DL: Behavioral Effects of Naloxone Infusion in Obsessive-Compulsive Disorder. Biological Psychiatry 1996; 40: 154- 156. 11. Benjamin J, Greenberg BD, and Murphy DL: Daily administration of m-chlorophenylpiperazine to healthy human volunteers rapidly attenuates many of its behavioral, hormonal, cardiovascular and temperature effects. Psychopharmacology 1996; 127: 140-149. 12. Levine J, Witztum E, Greenberg BD, Barak Y: Inositol induced mania? (letter) American Journal of Psychiatry 1996;153: 839. 13. Lesch K-P, Bengel D, Heils A, Sabol SS, Greenberg BD, Petri S, Benjamin J, Muller CR, Hamer DH, Murphy DL: Association of anxiety-related traits with a polymorphism in the serotonin transporter gene regulatory region. Science, 1996; 274: 1527-1531. 14. Altemus M, Murphy DL, Greenberg B, Lesch P: Intact serotonin transporter gene in obsessive- compulsive disorder. American Journal of Medical Genetics (Neuropsychiatric Genetics) 1996; 409-411. Greenberg 4 15. Greenberg BD, George MS, Martin JD, Benjamin J, Schlaepfer TE, Altemus M, Wassermann EM, Post RM, and Murphy DL: Effect of prefrontal repetitive transcranial magnetic stimulation in obsessive-compulsive disorder: a preliminary study. American Journal of Psychiatry 1997; 154: 867-869. 16. Schmidt LA, Greenberg BD, Holzman GB, Schulkin J. Treatment of Depression by Obstetrician-Gynecologists: A Survey Study. Obstetrics and Gynecology 1997; 90: 296-300. 17. George MG, Wassermann EM, Kimbrell TK, Little JT, Williams WE, Danielson BA, Greenberg BD, Hallett M, Post RM: Mood improvement following daily left prefrontal rTMS mood in patients with depression: a placebo-controlled crossover trial. American Journal of Psychiatry 1997; 154: 1752-6. 18. Schlaepfer TE, Strain EC, Greenberg BD, Preston KL , Lancaster E , Bigelow GE, Barta PE, Pearlson GD: The Site of Opioid Action in the Human Brain: µ and κ Agonists Produce Different Cerebral Blood Flow Effects. American Journal of Psychiatry 1998; 155: 470-473. 19. Greenberg BD, Benjamin J, Martin JD, Keuler D, Huang S-J, Altemus M, and Murphy DL: Delayed obsessive-compulsive disorder symptom exacerbation after a single serotonin antagonist dose in fluoxetine-treated but not untreated patients. Psychopharmacology, 1998; 140(4): 434-44. 20. Murphy DL, Sims K, Eisenhofer G, Greenberg B, George T, Butler R, Berlin F, Zametkin A, Ernst M, Breakfield XO. Are MAO-A deficiency states in the general population and in putative high-risk populations highly uncommon? Journal of Neural Transmission (Suppl) 1998; 52: 29-38. 21. Enoch MA, Kaye WH, Rotondo A, Greenberg BD, Murphy DL, Goldman D. 5HT2A promoter Polymorphism -1438G/A, anorexia nervosa, and obsessive-compulsive disorder. Lancet 1998; 351: 1785-1786. 22. Greenberg BD, Tolliver TJ, Huang S-J, Li Q, Murphy DL. Genetic variation in the serotonin transporter promoter region affects serotonin uptake in human blood platelets. American Journal of Medical Genetics (Neuropsychiatric Genetics), 1999; 88: 83-87. 23. Greenberg BD, Ziemann U, Harmon A, Murphy DL, Wassermann EM. Reduced intracortical inhibition in obsessive-compulsive disorder on transcranial magnetic stimulation. Lancet 1998; 352: 881-882. 24. Schuback DE, Mulligan EL, Sims KB, Tivol EA, Greenberg BD, Chang S-F, Yang S-L, Mau Y- C, Shen C-Y, Ho M-S, Yang N-H, Butler MG, Fink S, Schwartz CE, Berlin F, Breakfield XO, Murphy DL, Hsu Y-PP. Screen for MAOA mutations in target human groups. American Journal of Medical Genetics (Neuropsychiatric Genetics), 1999; 88: 25-28. 25. Sabol SZ, Nelson ML, Fisher C, Gunzerath L, Brody CL, Hu S, Sirota LA, Marcus SE, Greenberg BD, Lucas FR 4th, Benjamin J, Murphy DL, Hamer DH. A genetic association for cigarette smoking behavior. Health Psychology 1999;18(1):7-13. 26. Ellingson T, Duddempudi S, Greenberg BD, Hooper D, Eisenhoffer G. Determination of differential activities of soluble and membrane-bound catechol-0-methyltransferase in tissues and erythrocytes. Journal of Chromatography, 1999; 729: 347-353. Greenberg 5 27. Sirota LA, Greenberg BD, Murphy DL, HamerDH. Non-linear association between the serotonin transporter promoter polymorphism and neuroticism: A caution against using extreme samples to identify quantitative trait loci. Psychiatric Genetics, 1999; 9 (1):35-8.
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