Chapter 27 the "Tetrapod"
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Tetrapod Biostratigraphy and Biochronology of the Triassic–Jurassic Transition on the Southern Colorado Plateau, USA
Palaeogeography, Palaeoclimatology, Palaeoecology 244 (2007) 242–256 www.elsevier.com/locate/palaeo Tetrapod biostratigraphy and biochronology of the Triassic–Jurassic transition on the southern Colorado Plateau, USA Spencer G. Lucas a,⁎, Lawrence H. Tanner b a New Mexico Museum of Natural History, 1801 Mountain Rd. N.W., Albuquerque, NM 87104-1375, USA b Department of Biology, Le Moyne College, 1419 Salt Springs Road, Syracuse, NY 13214, USA Received 15 March 2006; accepted 20 June 2006 Abstract Nonmarine fluvial, eolian and lacustrine strata of the Chinle and Glen Canyon groups on the southern Colorado Plateau preserve tetrapod body fossils and footprints that are one of the world's most extensive tetrapod fossil records across the Triassic– Jurassic boundary. We organize these tetrapod fossils into five, time-successive biostratigraphic assemblages (in ascending order, Owl Rock, Rock Point, Dinosaur Canyon, Whitmore Point and Kayenta) that we assign to the (ascending order) Revueltian, Apachean, Wassonian and Dawan land-vertebrate faunachrons (LVF). In doing so, we redefine the Wassonian and the Dawan LVFs. The Apachean–Wassonian boundary approximates the Triassic–Jurassic boundary. This tetrapod biostratigraphy and biochronology of the Triassic–Jurassic transition on the southern Colorado Plateau confirms that crurotarsan extinction closely corresponds to the end of the Triassic, and that a dramatic increase in dinosaur diversity, abundance and body size preceded the end of the Triassic. © 2006 Elsevier B.V. All rights reserved. Keywords: Triassic–Jurassic boundary; Colorado Plateau; Chinle Group; Glen Canyon Group; Tetrapod 1. Introduction 190 Ma. On the southern Colorado Plateau, the Triassic– Jurassic transition was a time of significant changes in the The Four Corners (common boundary of Utah, composition of the terrestrial vertebrate (tetrapod) fauna. -
Early Tetrapod Relationships Revisited
Biol. Rev. (2003), 78, pp. 251–345. f Cambridge Philosophical Society 251 DOI: 10.1017/S1464793102006103 Printed in the United Kingdom Early tetrapod relationships revisited MARCELLO RUTA1*, MICHAEL I. COATES1 and DONALD L. J. QUICKE2 1 The Department of Organismal Biology and Anatomy, The University of Chicago, 1027 East 57th Street, Chicago, IL 60637-1508, USA ([email protected]; [email protected]) 2 Department of Biology, Imperial College at Silwood Park, Ascot, Berkshire SL57PY, UK and Department of Entomology, The Natural History Museum, Cromwell Road, London SW75BD, UK ([email protected]) (Received 29 November 2001; revised 28 August 2002; accepted 2 September 2002) ABSTRACT In an attempt to investigate differences between the most widely discussed hypotheses of early tetrapod relation- ships, we assembled a new data matrix including 90 taxa coded for 319 cranial and postcranial characters. We have incorporated, where possible, original observations of numerous taxa spread throughout the major tetrapod clades. A stem-based (total-group) definition of Tetrapoda is preferred over apomorphy- and node-based (crown-group) definitions. This definition is operational, since it is based on a formal character analysis. A PAUP* search using a recently implemented version of the parsimony ratchet method yields 64 shortest trees. Differ- ences between these trees concern: (1) the internal relationships of aı¨stopods, the three selected species of which form a trichotomy; (2) the internal relationships of embolomeres, with Archeria -
Coastal and Ocean Engineering
May 18, 2020 Coastal and Ocean Engineering John Fenton Institute of Hydraulic Engineering and Water Resources Management Vienna University of Technology, Karlsplatz 13/222, 1040 Vienna, Austria URL: http://johndfenton.com/ URL: mailto:[email protected] Abstract This course introduces maritime engineering, encompassing coastal and ocean engineering. It con- centrates on providing an understanding of the many processes at work when the tides, storms and waves interact with the natural and human environments. The course will be a mixture of descrip- tion and theory – it is hoped that by understanding the theory that the practicewillbemadeallthe easier. There is nothing quite so practical as a good theory. Table of Contents References ....................... 2 1. Introduction ..................... 6 1.1 Physical properties of seawater ............. 6 2. Introduction to Oceanography ............... 7 2.1 Ocean currents .................. 7 2.2 El Niño, La Niña, and the Southern Oscillation ........10 2.3 Indian Ocean Dipole ................12 2.4 Continental shelf flow ................13 3. Tides .......................15 3.1 Introduction ...................15 3.2 Tide generating forces and equilibrium theory ........15 3.3 Dynamic model of tides ...............17 3.4 Harmonic analysis and prediction of tides ..........19 4. Surface gravity waves ..................21 4.1 The equations of fluid mechanics ............21 4.2 Boundary conditions ................28 4.3 The general problem of wave motion ...........29 4.4 Linear wave theory .................30 4.5 Shoaling, refraction and breaking ............44 4.6 Diffraction ...................50 4.7 Nonlinear wave theories ...............51 1 Coastal and Ocean Engineering John Fenton 5. The calculation of forces on ocean structures ...........54 5.1 Structural element much smaller than wavelength – drag and inertia forces .....................54 5.2 Structural element comparable with wavelength – diffraction forces ..56 6. -
Phylogeny of Basal Tetrapoda
Stuart S. Sumida Biology 342 Phylogeny of Basal Tetrapoda The group of bony fishes that gave rise to land-dwelling vertebrates and their descendants (Tetrapoda, or colloquially, “tetrapods”) was the lobe-finned fishes, or Sarcopterygii. Sarcoptrygii includes coelacanths (which retain one living form, Latimeria), lungfish, and crossopterygians. The transition from sarcopterygian fishes to stem tetrapods proceeded through a series of groups – not all of which are included here. There was no sharp and distinct transition, rather it was a continuum from very tetrapod-like fishes to very fish-like tetrapods. SARCOPTERYGII – THE LOBE-FINNED FISHES Includes •Actinista (including Coelacanths) •Dipnoi (lungfishes) •Crossopterygii Crossopterygians include “tetrapods” – 4- legged land-dwelling vertebrates. The Actinista date back to the Devonian. They have very well developed lobed-fins. There remains one livnig representative of the group, the coelacanth, Latimeria chalumnae. A lungfish The Crossopterygii include numerous representatives, the best known of which include Eusthenopteron (pictured here) and Panderichthyes. Panderichthyids were the most tetrapod-like of the sarcopterygian fishes. Panderichthyes – note the lack of dorsal fine, but retention of tail fin. Coelacanths Lungfish Rhizodontids Eusthenopteron Panderichthyes Tiktaalik Ventastega Acanthostega Ichthyostega Tulerpeton Whatcheeria Pederpes More advanced amphibians Tiktaalik roseae – a lobe-finned fish intermediate between typical sarcopterygians and basal tetrapods. Mid to Late Devonian; 375 million years old. The back end of Tiktaalik’s skull is intermediate between fishes and tetrapods. Tiktaalik is a fish with wrist bones, yet still retaining fin rays. The posture of Tiktaalik’s fin/limb is intermediate between that of fishes an tetrapods. Coelacanths Lungfish Rhizodontids Eusthenopteron Panderichthyes Tiktaalik Ventastega Acanthostega Ichthyostega Tulerpeton Whatcheeria Pederpes More advanced amphibians Reconstructions of the basal tetrapod Ventastega. -
1 Single-Layer Breakwater Armouring: Feedback on The
SINGLE-LAYER BREAKWATER ARMOURING: FEEDBACK ON THE ACCROPODE™ TECHNOLOGY FROM SITE EXPERIENCE GIRAUDEL Cyril1, GARCIA Nicolas2, LEDOUX Sébastien3 The single-layer technique appeared at the beginning of the 1980s, with the ACCROPODE™ unit, and is thus entering its third decade. At the time, this solution was a real innovation, reducing the amount of concrete and steepening armour facing slopes, hence reducing the volume of materials required. After three decades in use and more than 200 projects to date, it was important to summarize the lessons learned during this period and to inspect (above and below water) some of these structures in order to assess their behaviour and particularly to confirm the validity of the unit placing rules. In addition to the aspects related to armour stability, the focus has been given to the colonization by marine life of the structures, including the bedding layers, toe berms, underlayer, armour units. The purpose of this paper is to share the experience gained throughout the inspections undertaken since 2010 on structures built more than 10 years ago. A large panel of structures has been inspected, of different ages and at various locations worldwide. Keywords: rubble-mound breakwater; single-layer armouring; ACCROPODE™ units; biodiversity INTRODUCTION The ACCROPODE™ armour unit, well known today, is a plain concrete unit designed to protect the breakwaters, in aiming to reducing considerably the use of material while implementing steeper slopes and a single layer of concrete units (Figure 1). Invented in 1981 thanks to the bases and knowledge acquired with the Tetrapod invented by the same engineering company in 1953, the technology is still currently used and more than 200 applications have been built worldwide. -
Evolution of the Muscular System in Tetrapod Limbs Tatsuya Hirasawa1* and Shigeru Kuratani1,2
Hirasawa and Kuratani Zoological Letters (2018) 4:27 https://doi.org/10.1186/s40851-018-0110-2 REVIEW Open Access Evolution of the muscular system in tetrapod limbs Tatsuya Hirasawa1* and Shigeru Kuratani1,2 Abstract While skeletal evolution has been extensively studied, the evolution of limb muscles and brachial plexus has received less attention. In this review, we focus on the tempo and mode of evolution of forelimb muscles in the vertebrate history, and on the developmental mechanisms that have affected the evolution of their morphology. Tetrapod limb muscles develop from diffuse migrating cells derived from dermomyotomes, and the limb-innervating nerves lose their segmental patterns to form the brachial plexus distally. Despite such seemingly disorganized developmental processes, limb muscle homology has been highly conserved in tetrapod evolution, with the apparent exception of the mammalian diaphragm. The limb mesenchyme of lateral plate mesoderm likely plays a pivotal role in the subdivision of the myogenic cell population into individual muscles through the formation of interstitial muscle connective tissues. Interactions with tendons and motoneuron axons are involved in the early and late phases of limb muscle morphogenesis, respectively. The mechanism underlying the recurrent generation of limb muscle homology likely resides in these developmental processes, which should be studied from an evolutionary perspective in the future. Keywords: Development, Evolution, Homology, Fossils, Regeneration, Tetrapods Background other morphological characters that may change during The fossil record reveals that the evolutionary rate of growth. Skeletal muscles thus exhibit clear advantages vertebrate morphology has been variable, and morpho- for the integration of paleontology and evolutionary logical deviations and alterations have taken place unevenly developmental biology. -
Coastal Management Software to Support the Decision-Makers to Mitigate Coastal Erosion
Journal of Marine Science and Engineering Article Coastal Management Software to Support the Decision-Makers to Mitigate Coastal Erosion Carlos Coelho * , Pedro Narra, Bárbara Marinho and Márcia Lima RISCO & Civil Engineering Department, Aveiro University, Campus Universitário de Santiago, 3810-193 Aveiro, Portugal; [email protected] (P.N.); [email protected] (B.M.); [email protected] (M.L.) * Correspondence: [email protected] Received: 4 December 2019; Accepted: 8 January 2020; Published: 11 January 2020 Abstract: There are no sequential and integrated approaches that include the steps needed to perform an adequate management and planning of the coastal zones to mitigate coastal erosion problems and climate change effects. Important numerical model packs are available for users, but often looking deeply to the physical processes, demanding big computational efforts and focusing on specific problems. Thus, it is important to provide adequate tools to the decision-makers, which can be easily interpreted by populations, promoting discussions of optimal intervention scenarios in medium to long-term horizons. COMASO (coastal management software) intends to fill this gap, presenting a group of tools that can be applied in standalone mode, or in a sequential order. The first tool should map the coastal erosion vulnerability and risk, also including the climate change effects, defining a hierarchy of priorities where coastal defense interventions should be performed, or limiting/constraining some land uses or activities. In the locations identified as priorities, a more detailed analysis should consider the application of shoreline and cross-shore evolution models (second tool), allowing discussing intervention scenarios, in medium to long-term horizons. After the defined scenarios, the design of the intervention should be discussed, both in case of being a hard coastal structure or an artificial nourishment (third type of tools). -
Concrete Armour Units for Rubble Mound
ftt*attlsReserctt Mhlingrford CONCRETEARMOUR UNITS FOR RUBBLE MOUNDBREAKWATERS AND SEA WALLS: RECENTPROGRESS N I.f H Allsop BSc, C Eng, MICE Report SR 100 March 1988 Registered Office: Hydriulics Research Wallingford, Oxfordshire OXIO 8BA. Telephone: (X91 35381. Telex: 84.8552 This report describes work jointly funded by the Department of the Environment and the l,linistry of Agriculture Fisheries and Food. The Department of the Environment contract number was PECD7 16152 for which the nominated officer rras Dr R P Thorogood. The Ministry of Agriculture Fisheries and Food contribution was lrithin the Research Connission (Marine Flooding) CSA 557 for which the nominated officer was l{r A J ALlison. The work was carried out in the Maritime Engineering Department of Hydraulice Research, lfall-ingford under the rnanagementof Dr S I,l Huntington. The report is published on behalf of both DoE and I'|AFF, but any opinions expressed in this report are not necesearily those of the funding Departments. @ Crown copyright 1988 Publiehed by permission of the Controller of lter Majestyts Stationery Office. Goncrete,armour "unite.for' rubble mound breakwaters, sea val,ls and. revetuentsS recent :pfogf,eas, N lJ H Alleop Report SR 100, ilarch 1988 Eydraulics Reeearch, Wallingford Abetract ilany deep water breakrvatera constructed in,the laet 2O-50 years are of rubble sound construction, proteeted againat the effects of rave aetion by concrete armour nnite. Ttrese units are oftea of complex ehape; Ttrey are generally produced in unreinforced concrete ia eizee between around 2-50 tonnea depending apon the local water depth, the eeverity of the locel save eonditione, and on the efficiencl and: stsbiiity of the unit type selected. -
Tetrapod Phylogeny
© J989 Elsevier Science Publishers B. V. (Biomédical Division) The Hierarchy of Life B. Fernholm, K. Bremer and H. Jörnvall, editors 337 CHAPTER 25 Tetrapod phylogeny JACQUES GAUTHIER', DAVID CANNATELLA^, KEVIN DE QUEIROZ^, ARNOLD G. KLUGE* and TIMOTHY ROWE^ ' Deparlmenl qf Herpelology, California Academy of Sciences, San Francisco, CA 94118, U.S.A., ^Museum of Natural Sciences and Department of Biology, Louisiana State University, Baton Rouge, LA 70803, U.S.A., ^Department of ^oology and Museum of Vertebrate ^oology. University of California, Berkeley, CA 94720, U.S.A., 'Museum of ^oology and Department of Biology, University of Michigan, Ann Arbor, MI 48109, U.S.A. and ^Department of Geological Sciences, University of Texas, Austin, TX 78713, U.S.A. Introduction Early sarcopterygians were aquatic, but from the latter part of the Carboniferous on- ward that group has been dominated by terrestrial forms commonly known as the tet- rapods. Fig. 1 illustrates relationships among extant Tetrápoda [1-4J. As the clado- grams in Figs. 2•20 demonstrate, however, extant groups represent only a small part of the taxonomic and morphologic diversity of Tetrápoda. We hope to convey some appreciation for the broad outlines of tetrapod evolution during its 300+ million year history from late Mississippian to Recent times. In doing so, we summarize trees de- rived from the distribution of over 972 characters among 83 terminal taxa of Tetrápo- da. More than 90% of the terminal taxa we discuss are extinct, but all of the subter- minal taxa are represented in the extant biota. This enables us to emphasize the origins of living tetrapod groups while giving due consideration to the diversity and antiquity of the clades of which they are a part. -
Design & Construction of Coastal Structures
DESIGN & CONSTRUCTION OF COASTAL STRUCTURES A.V. SITARAMA SARMA, Research Officer Central Water & Power Research Station, Pune 1. INTRODUCTION Coastal structures are the marine structure located in coastal waters. Marine structures can be broadly classified into two types: a) Offshore Structures and b) Near-shore Structures The design of Near-shore structures and offshore structures varies greatly in terms of environmental factors, depth of water etc. Offshore structures are located in deepwater and are subjected to forces due to short crested multi-directional waves, which are predominant, apart from other forces due to wind, ocean currents etc. The different types of offshore structure are:- Gravity Type Structures Pile Supported Structures Floating Structures Submarine Pipelines The near-shore structures can be grouped into:- Rigid structure – Sheet piles, Walls Semi-rigid or Composite structures – Caissons or cells on rubble base Flexible Structures – Rubblemound breakwaters, Sea walls Fig 1: Types of Near-shore Structures Training Course on Coastal Engineering & Coastal Zone Management Thus these near shore structures are subjected to various marine environmental forces due to waves, winds and currents. The wave forces are the dominant forces and are decisive in the design of near shore coastal structures. Rubblemound structures are the most commonly applied type for breakwater and seawall. The stability of rubblemound coastal structures depends primarily upon the stability of individual armour units on its seawards slope. The other hydrodynamic aspects of the effect of waves on the rubblemound are wave run-up, rundown, overtopping, reflection and transmission. Design of flexible rubblemound structures is complex as it involves various aspects such as wave-structure interaction interlocking, characteristics of armour, friction between armour and secondary layer etc. -
Objectives Tetrapod Characteristics
1/22/2018 Becky Hardman University of Tennessee College of Veterinary Medicine [email protected] 416 360 300 250 200 146 65 Objectives Define Tetrapod/Amphibian Origin of Tetrapods Split of Amphibians Modern Amphibians Extant Families Simplification Tetrapod Characteristics Four Limbs Tetra= Four; Pod=Foot Some lost or vestigial “One bonetwo boneslittle blobsfingers/toes”- Neil Shubin Some lost or vestigial Includes all non-fish vertebrates 1 1/22/2018 Amphibian Characteristics “Tetrapod vertebrates that pass through a larval state and undergo metamorphosis into terrestrial adults.” • Anamniotes • Eggs need moist environment • Larval; metamorphosis • Permeable Skin • Cutaneous respiration • Two Gland Types • Mucous • Poison • Pedicellate Teeth • Amphibian papillae/Opercular bone • Can Hear Vibrations • Fat Bodies • Green Rods- fxn unknown • Singular Sacrum • Lost in caecilians Amphibian Characteristics As a Fossil… Articular surface of axis convex Exoccipital Bone articulates with dermal roofing Hand (Manus) 4 digits Foot (Pes) 5 digits Some Secondarily Lost Important to determine for fossil realtionships 2 1/22/2018 Phylogenetic Trees Phylogenetic Trees 3 1/22/2018 Darwin’s Tree Geologic Time Scale 416 360 300 250 200 146 65 Devonian: Age of Fishes Lobed-Finned Fishes Lungfishes; Coelacanths Tetrapodomorpha Panderichthyids Ichthyostega, Acanthostega Tetrapods 4 1/22/2018 416 360 300 250 200 146 65 Devonian: Fish to Tetrapod Panderichthyids 380 mya Predators in shallow water Eyes on top of head Lung and Gills -
Systems Biology Approach to the Origin of the Tetrapod Limb Koh
Systems biology approach to the origin of the tetrapod limb Koh Onimaru1*, Luciano Marcon2* 1. Laboratory for Phyloinformatics, RIKEN Center for Biosystems Dynamics Research (BDR), 2-1 Hirosawa, Wako city, Saitama prefecture, 351-0198, Japan. E-mail: [email protected] 2. Andalusian Centre for Developmental Biology (CABD), Universidad Pablo de Olavide, Carretera de Utrera km1, 41013, Sevilla, Spain. E-mail: [email protected] *Corresponding author July 2019 Abstract It is still not understood how similar genomic sequences have generated diverse and spectacular forms during evolution. The difficulty to bridge phenotypes and genotypes stems from the complexity of multicellular systems, where thousands of genes and cells interact with each other providing developmental non-linearity. To understand how diverse morphologies have evolved, it is essential to find ways to handle such complex systems. Here, we review the fin-to-limb transition as a case study for the evolution of multicellular systems. We first describe the historical perspective of comparative studies between fins and limbs. Second, we introduce our approach that combines mechanistic theory, computational modeling, and in vivo experiments to provide a mechanical explanation for the morphological difference between fish fins and tetrapod limbs. This approach helps resolve a long-standing debate about anatomical homology between the skeletal elements of fins and limbs. We will conclude by proposing that due to the counter- intuitive dynamics of gene interactions, integrative approaches that combine computer modeling, theory and experiments are essential to understand the evolution of multicellular organisms. Introduction Multi-cellular organisms have evolved a remarkable diversity of their forms. In the last decades, thanks to the advance of DNA sequencing technologies, the genome sequence data of various species have become available.