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CHAPTER 12 COLEOPTERA (Aquatic Beetles)

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CHAPTER 12 COLEOPTERA (Aquatic Beetles) Ù«·¼» ¬± ߯«¿¬·½ ײª»®¬»¾®¿¬»• ±º ¬¸» Ë°°»® Ó·¼©»•¬ | îððì CHAPTER 12 COLEOPTERA (Aquatic Beetles) Citation: Bouchard, R.W., Jr. 2004. Guide to aquatic macroinvertebrates of the Upper Midwest. Water Resources Center, University of Minnesota, St. Paul, MN. 208 pp. ݸ¿°¬»® ïî | ÝÑÔÛÑÐÌÛÎß 141 Ù«·¼» ¬± ߯«¿¬·½ ײª»®¬»¾®¿¬»• ±º ¬¸» Ë°°»® Ó·¼©»•¬ | îððì ORDER COLEOPTERA Aquatic Beetles 12 ̸» ±®¼»® ݱ´»±°¬»®¿ ·• ¿ ¸«¹» ±®¼»®ô ±º ©¸·½¸ ¬¸» ³¿¶±®·¬§ ±º ³»³¾»®• ¿®» ¬»®®»•¬®·¿´ò ر©»ª»®ô ¬¸»®» ¿®» •¬·´´ ¿ ¹®»¿¬ ²«³¾»® ±º ¾»»¬´»• ¿¼¿°¬»¼ ¬± ¿² ¿¯«¿¬·½ »¨·•¬»²½» »²½±³°¿••·²¹ ¿ ´¿®¹» ¼·ª»®•·¬§ ±º ¸¿¾·¬¿¬• ¿²¼ ´·º» ¸·•¬±®·»•ò ߯«¿¬·½ ¾»»¬´»• ½¿² ¾» º±«²¼ ·² ²»¿®´§ ¿²§ ¿¯«¿¬·½ ¸¿¾·¬¿¬ô ¾«¬ ¾»»¬´»• ®»¿½¸ ¬¸»·® ¹®»¿¬»•¬ ¼·ª»®•·¬§ ·² ´»²¬·½ ¸¿¾·¬¿¬• •«½¸ ¿• ©»¬´¿²¼• ¿²¼ °±²¼ ³¿®¹·²•ò ﮬ ±º ¬¸» ®»¿•±² º±® ¬¸»·® •«½½»•• ·² ¿¯«¿¬·½ ¸¿¾·¬• ·• ¬¸» ¿¾·´·¬§ ±º ¬¸» ¿¼«´¬• ¬± »²¬»® ±® ´»¿ª» ¬¸» ©¿¬»® ¬± •»¿®½¸ º±® ³¿¬»• ±® ·º ½±²¼·¬·±²• ¿®» ²±¬ ±°¬·³¿´ò ͱ³» ¾»»¬´»• ¿®» ¿¯«¿¬·½ ¿• ¾±¬¸ ´¿®ª¿» ¿²¼ ¿¼«´¬•ô ©¸·´» ±¬¸»®• ¿®» ¿¯«¿¬·½ ¿• ¿¼«´¬• ±® ¿• ´¿®ª¿»ò ر©»ª»®ô ¿´³±•¬ ¿´´ ¿¯«¿¬·½ ¿²¼ •»³·¿¯«¿¬·½ ݱ´»±°¬»®¿ °«°¿¬» ¬»®®»•¬®·¿´´§ ©·¬¸ ¬¸» »¨½»°¬·±² ±º ¿ º»© ¬¿¨¿ ¬¸¿¬ ¿®» ²±¬ ¼»¿´¬ ©·¬¸ ·² ¬¸·• ¹«·¼»ò ̸» ݱ´»±°¬»®¿ µ»§ ¼±»• ²±¬ ·²½´«¼» •»³·¿¯«¿¬·½ ¬¿¨¿ ©¸·½¸ ³¿§ ¾» ½±´´»½¬»¼ ·² ¿¯«¿¬·½ ·²ª»®¬»¾®¿¬» •¿³°´»•ô ¾«¬ ·¬ ©·´´ ¾» •«ºº·½·»²¬ º±® ¬¸» ³¿¶±® ¹®±«°•ò ß ³±®» ·²½´«•·ª» µ»§ ¬± ¿¯«¿¬·½ ¿²¼ •»³·¿¯«¿¬·½ ݱ´»±°¬»®¿ ½¿² ¾» º±«²¼ ·² Ó»®®·¬¬ ¿²¼ Ý«³³·²• øïççê÷ò Coleoptera Morphology Larvae: Ô¿®ª¿» ±º ¿¯«¿¬·½ ݱ´»±°¬»®¿ ½¿² ¾» ®»½±¹²·¦»¼ ¾§ ¬¸» °®»•»²½» ±º ¿ •½´»®±¬·¦»¼ ¸»¿¼ô ¬¸®»» °¿·®• ±º •»¹³»²¬»¼ ¬¸±®¿½·½ ´»¹•ô ¿²¼ ¬¸» ¿¾•»²½» ±º ©·²¹ °¿¼• øÚ·¹ò ïîòï÷ò ݸ¿®¿½¬»®• •«½¸ ¿• ¬¸» ²«³¾»® ±º ¬¿®•¿´ ½´¿©•ô ²«³¾»® ±º ´»¹ •»¹³»²¬•ô ¾±¼§ •¸¿°»ô ¿²¼ ¿²¬»²²¿´ ´»²¹¬¸ ¿®» ¼·¿¹²±•¬·½ ½¸¿®¿½¬»®• º±® ݱ´»±°¬»®¿ ´¿®ª¿»ò Ú·¹«®» ïîòïæ ܱ®•¿´ ª·»© ±º ½±´»±°¬»®¿² ´¿®ª¿ò ݸ¿°¬»® ïî | ÝÑÔÛÑÐÌÛÎß 143 Ù«·¼» ¬± ߯«¿¬·½ ײª»®¬»¾®¿¬»• ±º ¬¸» Ë°°»® Ó·¼©»•¬ | îððì Adults: ݱ´»±°¬»®¿ ¿¼«´¬• ½¿² ¾» ®»½±¹²·¦»¼ °®·³¿®·´§ ¾§ ¬¸» °®»•»²½» ±º ¸»¿ª·´§ •½´»®±¬·¦»¼ º±®» ©·²¹• ø»´§¬®¿÷ ©¸·½¸ ´¿½µ ª»·²• ¿²¼ ½±ª»® ¬¸» ³»³¾®¿²±«• ¸·²¼ ©·²¹• øÚ·¹ò ïîòî÷ò ײ ¿¼¼·¬·±²ô ¬¸» »²¬·®» ¾±¼§ ·• ¹»²»®¿´´§ ¸¿®¼»²»¼ ¿²¼ ¬¸®»» °¿·®• ±º •»¹³»²¬»¼ ´»¹• ¿®» °®»•»²¬ò ß¼«´¬ ݱ´»±°¬»®¿ º¿³·´·»• ½¿² ¾» •»°¿®¿¬»¼ ¾§ ½¸¿®¿½¬»®• •«½¸ ¿• ¬¸» •¸¿°» ±º ¬¸» »§»ô ¬¸» ¸·²¼ ½±¨¿»ô ¿²¼ ¬¸» ¿²¬»²²¿» øÚ·¹• ïîòîô ïîòí÷ò Ú·¹«®» ïîòîæ ܱ®•¿´ ª·»© ±º ½±´»±°¬»®¿² ¿¼«´¬ò Ú·¹«®» ïîòíæ Ê»²¬®¿´ ª·»© ±º ½±´»±°¬»®¿² ¿¼«´¬ò 144 ݸ¿°¬»® ïî | ÝÑÔÛÑÐÌÛÎß Ù«·¼» ¬± ߯«¿¬·½ ײª»®¬»¾®¿¬»• ±º ¬¸» Ë°°»® Ó·¼©»•¬ | îððì Key to Coleoptera Families (Larvae)* ïò Ì©± ¬¿®•¿´ ½´¿©• ¿¬ »²¼ ±º ´»¹ øÚ·¹ò ïîòì÷òòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòî Ú·¹«®» ïîòìæ Ô»¹ ±º θ¿²¬«• •°ò øθ¿²¬«•÷ ´¿®ª¿ô ܱ®•¿´ Ê·»©ò ïùò ß •·²¹´» ¬¿®•¿´ ½´¿© ¿¬ »²¼ ±º ´»¹ øÚ·¹ò ïîòë÷òòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòí Ú·¹«®» ïîòëæ Ô»¹ ±º ا¼®±½¸¿®¿ •°ò øا¼®±°¸·´·¼¿»÷ ´¿®ª¿ò îøï÷ò ß¾¼±³·²¿´ •»¹³»²¬ ïð ©·¬¸ î °¿·®• ±º ¸±±µ• øÚ·¹ò ïîòê÷å ïð °¿·®• ±º ´¿¬»®¿´ º·´¿³»²¬• ±² ¿¾¼±³»²òòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòGyrinidae °ò ïëì ¸±±µ• •»¹³»²¬ ïð Ú·¹«®» ïîòêæ Ì»®³·²¿´ ¿¾¼±³·²¿´ •»¹³»²¬• ±º Ù§®·²·¼¿» ´¿®ª¿ò îùò ر±µ• ¿¾•»²¬ ±² ¿¾¼±³·²¿´ •»¹³»²¬ ïð øÚ·¹ò ïîòé÷å ´¿¬»®¿´ º·´¿³»²¬• «•«¿´´§ ¿¾•»²¬ º®±³ ¿¾¼±³»² ¿´¬¸±«¹¸ ¬»®³·²¿´ º·´¿³»²¬• ¿®» ±º¬»² °®»•»²¬ øÚ·¹ò ïîòé÷òDytiscidae °ò ïëî •»¹³»²¬ ïð Ú·¹«®» ïîòéæ Ì»®³·²¿´ ¿¾¼±³·²¿´ •»¹³»²¬• ±º ܧ¬·•½·¼¿» ´¿®ª¿ò ݸ¿°¬»® ïî | ÝÑÔÛÑÐÌÛÎß 145 | 3(1'). Legs with 5 segments (not counting tarsal claw) (Fig. 12.8); abdomen terminating with 1-2 long filaments................................................................................ Haliplidae 3'. Legs with 4 segments (not counting tarsal claw) (Fig. 12.9); abdomen not terminating in 1-2 long filaments........................................................................................................4 4(3'). Mandibles large and easily seen from above (Fig. 12.10)........... Hydrophilidae 4'. Mandibles not easily seen from above...........................................................................5 5(4'). Antennae longer than head (Fig. 12.11)...............................................Scirtidae 5'. Antennae shorter than head .........................................................................................6 146 | Ù«·¼» ¬± ߯«¿¬·½ ײª»®¬»¾®¿¬»• ±º ¬¸» Ë°°»® Ó·¼©»•¬ | îððì êøëù÷ò Þ±¼§ º´¿¬¬»²»¼ ©·¬¸ ¬¸±®¿½·½ ¿²¼ ¿¾¼±³·²¿´ •»¹³»²¬• »¨°¿²¼»¼ •± ¬¸¿¬ ´»¹• ¿²¼ ¸»¿¼ ¿®» ±¾•½«®»¼ º®±³ ¿¾±ª» øÚ·¹ò ïîòïî÷ òòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòPsephenidae °ò ïëé Ú·¹«®» ïîòïîæ Û½¬±°®·¿ •°ò øЕ»°¸»²·¼¿»÷ ´¿®ª¿ô ܱ®•¿´ Ê·»©ò êùò Þ±¼§ ½§´·²¼®·½¿´ ±® •«¾ó½§´·²¼®·½¿´å ¸»¿¼ ¿²¼ ´»¹• ª·•·¾´» º®±³ ¿¾±ª» øÚ·¹ò ïîòïí÷òòòòòòòòòòòòòòò òòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòElmidae °ò ïëí Ú·¹«®» ïîòïíæ ͬ»²»´³·• •°ò øÛ´³·¼¿»÷ ´¿®ª¿ô ܱ®•¿´ Ê·»©ò ö ̸» º¿³·´§ Ü®§±°·¼¿» ·• ²±¬ ·²½´«¼»¼ ·² ¬¸» ݱ´»±°¬»®¿ ´¿®ª¿ µ»§ ¾»½¿«•» ¬¸»•» ´¿®ª¿» ´·ª» ·² ®·°¿®·¿² ¿®»¿• ¿²¼ ¿®» ²±¬ ¹»²»®¿´´§ ½±´´»½¬»¼ ·² ¿¯«¿¬·½ •¿³°´»•ò ݸ¿°¬»® ïî | ÝÑÔÛÑÐÌÛÎß 147 Ù«·¼» ¬± ߯«¿¬·½ ײª»®¬»¾®¿¬»• ±º ¬¸» Ë°°»® Ó·¼©»•¬ | îððì Key to Coleoptera Families (Adults)* ïò ݱ³°±«²¼ »§»• ¼·ª·¼»¼ ¿²¼ ¿°°»¿®·²¹ ¬± ¸¿ª» î °¿·®• ±º »§»• øÚ·¹ò ïîòïì÷òòòòòòòòòòòòòòòòòòòòòòòòòòòò òòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòGyrinidae °ò ïëì »§» Ú·¹«®» ïîòïìæ Ø»¿¼ ±º Ù§®·²«• •°ò øÙ§®·²·¼¿»÷ ¿¼«´¬ô Ô¿¬»®¿´ Ê·»©ò ïùò ݱ³°±«²¼ »§»• «²¼·ª·¼»¼ øÚ·¹ò ïîòïë÷ òòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòî »§» Ú·¹«®» ïîòïëæ Ø»¿¼ ±º Ô¿½½±°¸·´«• •°ò øܧ¬·•½·¼¿»÷ ¿¼«´¬ô Ô¿¬»®¿´ Ê·»©ò îøïù÷ò Ø·²¼ ½±¨¿» »¨°¿²¼»¼ ·²¬± °´¿¬»• ¬¸¿¬ ½±ª»® ¿¾¼±³·²¿´ •»¹³»²¬• ïóî ±® í ¿²¼ ¾¿•»• ±º ³»¬¿º»³±®¿ øÚ·¹ò ïîòïê÷òòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòHaliplidae °ò ïëë ¸·²¼ ½±¨¿ Ú·¹«®» ïîòïêæ Ó»¬¿¬¸±®¿¨ ¿²¼ ¿¾¼±³»² ±º л´¬±¼§¬»• •°ò øØ¿´·°´·¼¿»÷ ¿¼«´¬ô Ê»²¬®¿´ Ê·»©ò îùò Ø·²¼ ½±¨¿» ²±¬ »¨°¿²¼»¼ ·²¬± °´¿¬»• øÚ·¹•ò ïîòïéô ïîòïè÷òòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòò í ¸·²¼ ½±¨¿ ¸·²¼ ½±¨¿ Ú·¹«®» ïîòïéæ Ô¿½½±°¸·´«• •°ò Ú·¹«®» ïîòïèæ ا¼®±½¸¿®¿ •°ò øܧ¬·•½·¼¿»÷ ¿¼«´¬ô Ê»²¬®¿´ Ê·»©ò øا¼®±°¸·´·¼¿»÷ ¿¼«´¬ô Ê»²¬®¿´ Ê·»©ò 148 ݸ¿°¬»® ïî | ÝÑÔÛÑÐÌÛÎß Ù«·¼» ¬± ߯«¿¬·½ ײª»®¬»¾®¿¬»• ±º ¬¸» Ë°°»® Ó·¼©»•¬ | îððì íøîù÷ò Ø·²¼ ½±¨¿» »¨¬»²¼·²¹ °±•¬»®·±®´§ ¿²¼ ¼·ª·¼·²¹ ¿¾¼±³·²¿´ •»¹³»²¬ ï ·²¬± ¬©± •»½¬·±²• øÚ·¹ò ïîòïç÷òòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòDytiscidae °ò ïëî ¿¾¼±³·²¿´ •¬»®²·¬» ï Ú·¹«®» ïîòïçæ Ô¿½½±°¸·´«• •°ò øܧ¬·•½·¼¿»÷ ¿¼«´¬ô Ê»²¬®¿´ Ê·»©ò íùò Ø·²¼ ½±¨¿» ²±¬ »¨¬»²¼·²¹ °±•¬»®·±®´§ ¿²¼ ¼·ª·¼·²¹ ¿¾¼±³·²¿´ •»¹³»²¬ ï ·²¬± ¬©± •»½¬·±²• øÚ·¹ò ïîòîð÷ òòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòì ¿¾¼±³·²¿´ •¬»®²·¬» ï Ú·¹«®» ïîòîðæ ا¼®±½¸¿®¿ •°ò øا¼®±°¸·´·¼¿»÷ ¿¼«´¬ô Ê»²¬®¿´ Ê·»©ò ìøíù÷ò ß²¬»²²¿» ½´«¾¾»¼ ©·¬¸ ¿ ½«°ó´·µ» •»¹³»²¬ ¿¬ ¬¸» ¾¿•» ±º íó•»¹³»²¬»¼ ½´«¾ øÚ·¹ò ïîòîï÷òòòò òòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòHydrophilidae °ò ïëê Ú·¹«®» ïîòîïæ ß²¬»²²¿» ±º ا¼®±°¸·´·¼¿» ¿¼«´¬ò ìùò ß²¬»²²¿» ª¿®·¿¾´» ¾«¬ ²±¬ ©·¬¸ ¿ ½«°ó´·µ» •»¹³»²¬ ¿¬ ¬¸» ¾¿•» ±º íó•»¹³»²¬»¼ ½´«¾ øÚ·¹•ò ïîòîîô ïîòîí÷ òòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòë Ú·¹«®» ïîòîîæ ß²¬»²²¿ ±º Ú·¹«®» ïîòîíæ ß²¬»²²¿ ±º Ñ°¬·±•»®ª«• •°ò øÛ´³·¼¿»÷ ¿¼«´¬ò Ø»´·½¸«• •°ò øÜ®§±°·¼¿»÷ ¿¼«´¬ò ݸ¿°¬»® ïî | ÝÑÔÛÑÐÌÛÎß 149 Ù«·¼» ¬± ߯«¿¬·½ ײª»®¬»¾®¿¬»• ±º ¬¸» Ë°°»® Ó·¼©»•¬ | îððì ëøìù÷ò ß²¬»²²¿» •¸±®¬ ©·¬¸ ¿ °»½¬·²¿¬» ø½±³¾ó´·µ»÷ ½´«¾å ¾±¼§ •·¦» ëóêòë ³³ ´±²¹ øÚ·¹ò ïîòîì÷òòòò òòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòDryopidae °ò ïëï Ú·¹«®» ïîòîìæ ß²¬»²²¿ ±º Ø»´·½¸«• •°ò øÜ®§±°·¼¿»÷ ¿¼«´¬ò ëùò ß²¬»²²¿» ©·¬¸±«¬ °»½¬·²¿¬» ½´«¾å ¾±¼§ •·¦» ä ìòë ³³ ´±²¹ øÚ·¹ò ïîòîë÷òòòòòòòòòòòòòòòòòòòòòòòòòòòò òòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòòElmidae °ò ïëí Ú·¹«®» ïîòîëæ ß²¬»²²¿ ±º Ñ°¬·±•»®ª«• •°ò øÛ´³·¼¿»÷ ¿¼«´¬ò ö ̸» º¿³·´·»• Е»°¸»²·¼¿» ¿²¼ ͽ·®¬·¼¿» ¿®» ²±¬ ·²½´«¼»¼ ·² ¬¸» ¿¼«´¬ µ»§ ¿• ¬¸» ¿¼«´¬• ±º ¬¸»•» º¿³·´·»• ¿®» ²±¬ ¿¯«¿¬·½ ¿²¼ ¿®» ¬¸»®»º±®» ²±¬ ½±³³±²´§ ½±´´»½¬»¼ ·² ¿¯«¿¬·½ •¿³°´»•ò 150 ݸ¿°¬»® ïî | ÝÑÔÛÑÐÌÛÎß Ù«·¼» ¬± ߯«¿¬·½ ײª»®¬»¾®¿¬»• ±º ¬¸» Ë°°»® Ó·¼©»•¬ | îððì Coleoptera Family Descriptions Dryopidae ____________________________________________________________________ Common Name: Ô±²¹ó̱»¼ É¿¬»® Þ»»¬´»• Feeding Group: ͽ®¿°»®• Tolerance Value: ë øÓ±¼»®¿¬»÷ Habitat: Ü®§±°·¼ ¿¼«´¬• ±½½«® ·² ¬¸» •©·º¬ °±®¬·±²• ±º •¬®»¿³• ¿²¼ ¿®» ¹»²»®¿´´§ ½±´´»½¬»¼ «²¼»® ®±½µ• ¿²¼ ´±¹•ò Size: Adults: ͳ¿´´ øëóêòë ³³÷ Characteristics: Adults: Ø¿®¼ ¾±¼·»¼å ¿²¬»²²¿» •¸±®¬ ©·¬¸ ¿ °»½¬·²¿¬» ½´«¾å ¸·²¼ ½±¨¿» ²±¬ »¨¬»²¼·²¹ °±•¬»®·±®´§ ¿²¼ ¼·ª·¼·²¹ º·®•¬ ¿¾¼±³·²¿´ •»¹³»²¬ ·²¬± ¬©± •»½¬·±²•ò Notes: Ü®§±°·¼ ¿¼«´¬• ®»•»³¾´» »´³·¼•å ¸±©»ª»®ô ¼®§±°·¼• ¬»²¼ ¬± ¾» ´¿®¹»®ò ̸» ¿²¬»²²¿» ¿®» ¼·ºº·½«´¬ ¬± •»» ¾»½¿«•» ¬¸»§ ¿®» •¸±®¬»²»¼ °»½¬·²¿¬» ½´«¾• ¬¸¿¬ ¿®» •±³»¬·³»• ½±²½»¿´»¼ ©·¬¸ ¿ °±®¬·±² ±º ¬¸» ¸»¿¼ «²¼»® ¬¸» »²´¿®¹»¼ °®±²±¬«³ò ̸·• º¿³·´§ ±º ¾»»¬´»• ·• «²·¯«» ¾»½¿«•» ¬¸» ´¿®ª¿» ¿®» ¹»²»®¿´´§ ¬»®®»•¬®·¿´ Ú·¹«®» ïîòîêæ Ø»´·½¸«• •°ò øÜ®§±°·¼¿»÷ ¿¼«´¬ô ø•±³»¬·³»• •»³·¿¯«¿¬·½÷ ©¸»®»¿• ¬¸» ܱ®•¿´ Ê·»©ò ¿¼«´¬• ¿®» ¿¯«¿¬·½ò ݸ¿°¬»® ïî | ÝÑÔÛÑÐÌÛÎß 151 Ù«·¼» ¬± ߯«¿¬·½ ײª»®¬»¾®¿¬»• ±º ¬¸» Ë°°»® Ó·¼©»•¬ | îððì Dytiscidae ____________________________________________________________________ Common Name: Ю»¼¿½»±«• Ü·ª·²¹ Þ»»¬´»• Feeding
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    Bernhard J. van VONDEL1, Mogens HOLMEN2 & Pyotr N. PETROV3 1Natuurmuseum Rotterdam, The Netherlands 2Hillerød, Denmark 3Moscow South–West Gymnasium, Russia REVIEW OF THE PALAEARCTIC AND ORIENTAL SPECIES OF THE SUBGENUS HALIPLUS S.STR. (COLEOPTERA: HALIPLIDAE: HALIPLUS) WITH DESCRIPTIONS OF THREE NEW SPECIES Vondel, B.J. van, M. Holmen & P. N. Petrov, 2006. Review�������������������������������������� of the Palaearctic and Oriental species of the subgenus Haliplus s.str. (Coleoptera: Haliplidae: Haliplus) with descriptions of three new species. – Tijdschrift voor Entomologie 149: 227-273, figs. 1-231. [issn 0040-7496]. Published 1 December 2006. The species considered to belong to the subgenus Haliplus Latreille, 1802 and known to occur in the Palaearctic region are reviewed. Latreille’s selection of Dytiscus impressus Fabricius, 1787 as [misidentified] type species, is amended in the sense of the nominal species and not the misidentified identity, following iczn article 70.3.1. Dytiscus impressus is here formally selected as type species for Cnemidotus Illiger, 1802 and Hoplitus Clairville, 1806. Three new species are described: Haliplus (Haliplus) fuscicornis Holmen, Van Vondel & Petrov sp.n., H. kirgisiensis Holmen & Van Vondel sp.n. and H. turkmenicus Van Vondel sp.n.. Lectotypes are designated for H. ruficollis var. pedemontanus Fiori, H. lineolatus Mannerheim, H. brevis Wehncke, H. fluviatilis var. maculatus Seidlitz and H. minutus Takizawa. Three new synonymies are estab- lished: H. brevior Nakane, junior synonym of H. japonicus Sharp, H. fluviatilis var. maculatus Seidlitz, junior synonym of H. ruficollis (De Geer) and H. minutus Takizawa, junior synonym of H. simplex Clark. All species are described and figured. A key to the species is presented.
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  • Water Beetles
    Ireland Red List No. 1 Water beetles Ireland Red List No. 1: Water beetles G.N. Foster1, B.H. Nelson2 & Á. O Connor3 1 3 Eglinton Terrace, Ayr KA7 1JJ 2 Department of Natural Sciences, National Museums Northern Ireland 3 National Parks & Wildlife Service, Department of Environment, Heritage & Local Government Citation: Foster, G. N., Nelson, B. H. & O Connor, Á. (2009) Ireland Red List No. 1 – Water beetles. National Parks and Wildlife Service, Department of Environment, Heritage and Local Government, Dublin, Ireland. Cover images from top: Dryops similaris (© Roy Anderson); Gyrinus urinator, Hygrotus decoratus, Berosus signaticollis & Platambus maculatus (all © Jonty Denton) Ireland Red List Series Editors: N. Kingston & F. Marnell © National Parks and Wildlife Service 2009 ISSN 2009‐2016 Red list of Irish Water beetles 2009 ____________________________ CONTENTS ACKNOWLEDGEMENTS .................................................................................................................................... 1 EXECUTIVE SUMMARY...................................................................................................................................... 2 INTRODUCTION................................................................................................................................................ 3 NOMENCLATURE AND THE IRISH CHECKLIST................................................................................................ 3 COVERAGE .......................................................................................................................................................
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  • Bioindicators of Water Quality
    Ephemeroptera | Mayflies ACE-11 Coleoptera | Beetles Using this guide Coleoptera with the data sheets Bioindicators of Water Quality Beetles Quick–Reference Guide Coleoptera (Beetles) Authors: Julie Speelman and Natalie Carroll | Photographer (unless otherwise noted): Julie Speelman | Design and Layout: Purdue Agricultural Communication Family Tolerance Number Family Tolerance 4 3 7 Value Found Score 5 5 5 Dryopidae 5 0 0 Dryopidae (larvae) Baetidae Baetiscidae Dytiscidae Dytiscidae (adult) Caenidae Dytiscidae 5 2 10 This publication shows aquatic insects that can be used as Long-toed Water Beetle Predaceous Diving Beetle Predaceous Diving Beetle Small Minnow Mayfly Armored Mayfly Small Square-gill Mayfly Biotic Water Quality Degree of Organic Elmidae 5 0 0 bioindicators of water quality in Indiana waterways. Bioindicators 5 are biological systems that are sensitive to environmental changes Index Rating Pollution Gyrinidae 4 0 0 organic pollution Dryopidae and, therefore, can indicate when pollution is present in the water. 0.00–3.75 excellent Long-toed Water Beetle Haliplidae 7 0 0 unlikely A tolerance score is included for each insect in this publication. Hydrophilidae 5 3 15 slight organic The tolerance score, ranging from 0–10, represents the insect’s 3.76–4.25 very good Psephenidae 4 0 0 sensitivity to pollution and can be used to estimate the quality of pollution possible the water in which the insect was found. Insects with a score of some organic Order Total 5 25 4.26–5.00 good 0 are intolerant to pollution, meaning they cannot tolerate any pollution probable water pollution, while insects with a score of 10 are very tolerant of fairly substantial 5 5 4 1 polluted water.
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  • Hungerford's Crawling Water Beetle (Brychius Hungerfordi)
    COSEWIC Assessment and Status Report on the Hungerford’s Crawling Water Beetle Brychius hungerfordi in Canada ENDANGERED 2011 COSEWIC status reports are working documents used in assigning the status of wildlife species suspected of being at risk. This report may be cited as follows: COSEWIC. 2011. COSEWIC assessment and status report on the Hungerford’s Crawling Water Beetle Brychius hungerfordi in Canada. Committee on the Status of Endangered Wildlife in Canada. Ottawa. ix + 40 pp. (www.sararegistry.gc.ca/status/status_e.cfm). Production note: COSEWIC would like to acknowledge Colin Jones for writing the status report on Hungerford’s Crawling Water Beetle (Brychius hungerfordi) in Canada, prepared under contract with Environment Canada. This report was overseen and edited by Paul Catling, Co-chair of the COSEWIC Arthropods Specialist Subcommittee. For additional copies contact: COSEWIC Secretariat c/o Canadian Wildlife Service Environment Canada Ottawa, ON K1A 0H3 Tel.: 819-953-3215 Fax: 819-994-3684 E-mail: COSEWIC/[email protected] http://www.cosewic.gc.ca Également disponible en français sous le titre Ếvaluation et Rapport de situation du COSEPAC sur l’haliplide de Hungerford (Brychius hungerfordi) au Canada. Cover illustration/photo: Hungerford’s Crawling Water Beetle — Photo provided by S.A. Marshall, University of Guelph. ©Her Majesty the Queen in Right of Canada, 2011. Catalogue No. CW69-14/627-2011E-PDF ISBN 978-1-100-18679-5 Recycled paper COSEWIC Assessment Summary Assessment Summary – May 2011 Common name Hungerford’s Crawling Water Beetle Scientific name Brychius hungerfordi Status Endangered Reason for designation A probable early postglacial relict, this water beetle is endemic to the upper Great Lakes and is Endangered in the U.S.
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  • The Evolution and Genomic Basis of Beetle Diversity
    The evolution and genomic basis of beetle diversity Duane D. McKennaa,b,1,2, Seunggwan Shina,b,2, Dirk Ahrensc, Michael Balked, Cristian Beza-Bezaa,b, Dave J. Clarkea,b, Alexander Donathe, Hermes E. Escalonae,f,g, Frank Friedrichh, Harald Letschi, Shanlin Liuj, David Maddisonk, Christoph Mayere, Bernhard Misofe, Peyton J. Murina, Oliver Niehuisg, Ralph S. Petersc, Lars Podsiadlowskie, l m l,n o f l Hans Pohl , Erin D. Scully , Evgeny V. Yan , Xin Zhou , Adam Slipinski , and Rolf G. Beutel aDepartment of Biological Sciences, University of Memphis, Memphis, TN 38152; bCenter for Biodiversity Research, University of Memphis, Memphis, TN 38152; cCenter for Taxonomy and Evolutionary Research, Arthropoda Department, Zoologisches Forschungsmuseum Alexander Koenig, 53113 Bonn, Germany; dBavarian State Collection of Zoology, Bavarian Natural History Collections, 81247 Munich, Germany; eCenter for Molecular Biodiversity Research, Zoological Research Museum Alexander Koenig, 53113 Bonn, Germany; fAustralian National Insect Collection, Commonwealth Scientific and Industrial Research Organisation, Canberra, ACT 2601, Australia; gDepartment of Evolutionary Biology and Ecology, Institute for Biology I (Zoology), University of Freiburg, 79104 Freiburg, Germany; hInstitute of Zoology, University of Hamburg, D-20146 Hamburg, Germany; iDepartment of Botany and Biodiversity Research, University of Wien, Wien 1030, Austria; jChina National GeneBank, BGI-Shenzhen, 518083 Guangdong, People’s Republic of China; kDepartment of Integrative Biology, Oregon State
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  • Phylogeny of Psephenidae (Coleoptera: Byrrhoidea) Based on Larval, Pupal and Adult Characters
    Systematic Entomology (2007), 32, 502–538 DOI: 10.1111/j.1365-3113.2006.00374.x Phylogeny of Psephenidae (Coleoptera: Byrrhoidea) based on larval, pupal and adult characters CHI-FENG LEE1 , MASATAKA SATOˆ2 , WILLIAM D. SHEPARD3 and M A N F R E D A . J A¨CH4 1Institute of Biodiversity, National Cheng Kung University, Tainan 701, Taiwan, 2Dia Cuore 306, Kamegahora 3-1404, Midoriku, Nagoya, 458-0804, Japan, 3Essig Museum of Entomology, 201 Wellman Hall, University of California, Berkeley, CA 94720, U.S.A., and 4Natural History Museum, Burgring 7, A-1010 Wien, Austria Abstract. We conducted the first comprehensive phylogenetic analysis of Pse- phenidae, based on 143 morphological characters of adults, larvae and pupae and coded for 34 taxa, representing three outgroups and 31 psephenid genera, including four undescribed ones. A strict consensus tree calculated (439 steps, consistency index ¼ 0.45, retention index ¼ 0.75) from the two most-parsimonious cladograms indicated that the monophyly of the family and subfamilies is supported, with the exception of Eubriinae, which is paraphyletic when including Afroeubria. Here a new subfamily, Afroeubriinae (subfam.n.), is formally estab- lished for Afroeubria. The analysis also indicated that the ‘streamlined’ larva is a derived adaptive radiation. Here, suprageneric taxonomy and the evolution of some significant characters are discussed. Keys are provided to the subfamilies and genera of Psephenidae considering larvae, adults and pupae. Introduction type specimens and the association of adults and immature stages by rearing, five new genera were proposed for Ori- Psephenidae, the ‘water penny’ beetles, are characterized by ental taxa and a well-resolved phylogenetic tree was ob- the peculiar larval body shape (Figs 4–7).
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  • Revision of the Haliplidae of the Neotropical Region Including Mexico (HALIPLIDAE) 71
    ©Wiener Coleopterologenverein (WCV), download unter www.biologiezentrum.at 68 Koleopt. Rdsch. 78 (2008) Koleopterologische Rundschau 78 69–194 Wien, Juli 2008 VAZIRANI, T.G. 1969: Contribution to the study of aquatic beetles (Coleoptera) 2. A review of the subfamilies Noterinae, Laccophilinae, Dytiscinae and Hydroporinae (in part) from India. – Oriental Insects 2 (3–4): 221–341. Revision of the Haliplidae of the VAZIRANI, T.G. 1970: Contributions to the study of aquatic beetles (Coleoptera). 10. On a collection of Dytiscidae from Goa. – Oriental Insects 4: 441–446. Neotropical Region including Mexico VAZIRANI, T.G. 1972: Contributions to the study of aquatic beetles (Coleoptera). 13. A collection of (Coleoptera: Haliplidae) Dytiscidae from Nilgiri Hills, South India, with the description of a new species. – Proceedings of the Zoological Society of Calcutta 25: 117–122. B.J. van VONDEL & P.J. SPANGLER VAZIRANI, T.G. 1974: Two new species of Dytiscidae (Insecta: Coleoptera) from India. – Indian Museum Bulletin 7 (1) (1972): 16–20. VAZIRANI, T.G. 1975: Some new records of Dytiscidae (Coleoptera) from Tamil Nadu (India). – Newsletter of the Zoological Survey of India 1 (2): 20–21. Abstract VAZIRANI, T.G. 1977a: Catalogue of Oriental Dytiscidae. – Records of the Zoological Survey of India The species of the family Haliplidae (Coleoptera) occurring in the Neotropical Region (including Miscellaneous Publication Occasional Paper 6 (1976): 1–111. Mexico) are revised. Two genera, Haliplus LATREILLE and Peltodytes RÉGIMBART, and 53 species of Haliplidae are now recognized in the region of which the following 18 species are here described as VAZIRANI, T.G. 1977b: Some new records of Dytiscidae from South Gujarat.
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  • Scirtid Beetles (Insecta, Coleoptera, Scirtidae) from Sakhalin and the Kuril Lslands
    Title Scirtid Beetles (Insecta, Coleoptera, Scirtidae) from Sakhalin and the Kuril lslands Author(s) Yoshitomi, Hiroyuki; Nikitsky, Nikolai Citation 北海道大学総合博物館研究報告, 2, 45-47 Issue Date 2004-03 Doc URL http://hdl.handle.net/2115/47791 Type bulletin (article) Note Biodiversity and Biogeography of the Kuril Islands and Sakhalin vol.1 File Information v. 1-6.pdf Instructions for use Hokkaido University Collection of Scholarly and Academic Papers : HUSCAP Biodiversity and Biogeography of the KurU Islands and Sakhalin (2004) 1, 45-47. Scirtid Beetles (Insecta, Coleoptera, Scirtidae) from Sakhalin and the Kurillslands Hiroyuki Yoshitomi1 and Nikolai Nikitsky2 1 Bioindicator Co., Ltd., (Sapporo Branch), N 1, W 2-11, Chiio-ku, Sapporo, 060-0001 Japan. [email protected]; 2 Zoological Museum, Moscow Lomonosov State University, Herzen Street 6, Moscow K-9, 103009 Russia. [email protected] Abstract A total of fifteen species of the family Scirtidae (Insecta: Coleoptera) are recorded from Sakhalin and the Kuril Islands. Of these, four species, Elodes inornata, Elodes wilsoni, Sacodes minuta, and Cyphon kongsbergensis, are newly recorded from the Kuril Islands. Introduction Remarks. This species is newly recorded from the Kuril Islands. Klausnitzer (1982) recorded eleven species of the Distribution. Japan (Hokkaido, Honshu, Shikoku, family Scirtidae from Sakhalin and the Kuril Islands for Kyushu); Kuril Islands (Kunashir Isl.). the first time. Since then no other paper has been published until now. Elodes piceata Klausnitzer, 1982 In this short paper, we are going to show a faunal Helodes(!) piceata Klausnitzer, 1982,277, figs. 8-13 list of the family based on the previous papers and on [Type: Kunashir, not examined].
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  • World Catalogue of Haliplidae – Corrections and Additions, 2 (HALIPLIDAE) 25
    ©Wiener Coleopterologenverein (WCV), download unter www.biologiezentrum.at 22 Koleopt. Rdsch. 83 (2013) Koleopterologische Rundschau 83 23–34 Wien, September 2013 Laccophilus sordidus SHARP, 1882 First record from Iran. This is the most northern limit of the distribution of the species. It was World Catalogue of Haliplidae – previously known from Egypt, Saudi Arabia, and Yemen. corrections and additions, 2 Acknowledgements (Coleoptera: Haliplidae) We are grateful to Dr. H. Fery (Berlin) for his help with identification of some specimens and B.J. van VONDEL Dr. J. Hájek (Prague) for his help with literature. The deputy of research, Shahid Chamran University of Ahvaz is thanked for financial support of Abstract the project (# 101). A second series of corrections and additions to the World Catalogue of Haliplidae (Coleoptera) published as part of Volume 7 of the World Catalogue of insect series (VONDEL 2005) are presented. References All new taxa, new synonymies and new data on distribution are summarized. The number of species of the family Haliplidae is now 240, distributed in five genera. DARILMAZ, M.C., İNCEKARA, Ü. & VAFAEI, R. 2013: Contribution to the knowledge of Iranian Aquatic Adephaga (Coleoptera). – Spixiana 36 (1): 149–152. Key words: Coleoptera, Haliplidae, World Catalogue, additions, corrections. FERY, H. & HOSSEINIE, S.O. 1998: A taxonomic revision of Deronectes Sharp, 1882 (Insecta: Coleoptera: Dytiscidae) (part II). – Annalen des Naturhistorischen Museums Wien B 100: 219–290. Introduction FERY, H., PEŠIĆ, V. & DARVISHZADEH, I. 2012: Faunistic notes on some Hydradephaga from the Khuzestan, Hormozgan and Sistan & Baluchestan provinces in Iran, with descriptive notes on the The World Catalogue of the beetle family Haliplidae (VONDEL 2005) was published on June 24, female of Glareadessus franzi Wewalka & Biström 1998 (Coleoptera, Dytiscidae, Noteridae).
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  • A Bug's Life in the Columbia Slough
    A Bug’s Life in the Columbia Slough: Handbook of Invertebrates and Macroinvertebrate Monitoring in the Columbia Slough June 2005 Jeff Adams WWW.COLUMBIASLOUGH.ORG Contacts: The Xerces Society for Invertebrate The Columbia Slough Watershed Conservation Council Jeff Adams Ethan Chessin [email protected] [email protected] Director of Aquatic Programs Volunteer Coodinator 4828 SE Hawtorhne Blvd. 7040 NE 47th Avenue Portland, OR 97215-3252 Portland, OR 97218-1212 503-232-6639 503-281-1132 http://www.xerces.org http://www.columbiaslough.org Funding for this handbook and the education and monitoring activities associated with this project has been provided by: ! Metropolitan Greenspaces Program – a partnership between Metro and the U.S. Fish & Wildlife Service ! The Xerces Society for Invertebrate Conservation member contributions ! Northwest Service Academy ! Oregon Watershed Enhancement Board ! City of Portland Bureau of Environmental Services' Community Watershed Stewardship Program All image credits belong to Jeff Adams with the following exceptions: the Joseph D. Meyers map of Portland vicinity was downloaded from the Center for Columbia River History website; the image with line drawings of a water strider and a back swimmer is used with permission from the University of Illinois Department of Entomology; and the images of the creeping water bug, left-handed snail, and sponge are used with permission from Daniel Pickard of the California Department of Fish and Game. (Cover photo: restoration site on Columbia Slough near Interstate 205. The benches had recently been created, but had not yet been planted with native vegetation.) Handbook of Macroinvertebrate Monitoring in the Columbia Slough TABLE OF CONTENTS INTRODUCTION........................................................................................................................
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  • The Water Beetles of Miller Blue Hole, Sandusky County, Ohio (Insecta: Coleoptera)1-2
    THE WATER BEETLES OF MILLER BLUE HOLE, SANDUSKY COUNTY, OHIO (INSECTA: COLEOPTERA)1-2 BRIAN E. MELIN AND ROBERT C. GRAVES Department of Biology, Bowling Green State University, Bowling Green, Ohio 43408 ABSTRACT Thirty-one species of aquatic beetles, representing the families Dytiscidae, Hydro- philidae, Haliplidae, and Gyrinidae, were collected in the Miller Blue Hole, a large spring, between October 21, 1967, and January 27, 1969. The numbers of each species collected are tabulated by monthly intervals. Some of the species present in this unusual habitat are rare; Tropisternus columbianus is apparently recorded from Ohio for the first time. INTRODUCTION The Ohio "blue holes" are the outlets of an extensive underground drainage system occurring in limestone bedrock. They may best be thought of as gigantic springs reaching the surface as deep, conical pools or ponds which are drained by streams flowing into Lake Erie. There are several "blue holes" located in San- dusky and Erie counties, the best known being the Castalia Blue Hole, which has been "developed" as a commercial tourist attraction, with the result that its original natural environment has been destroyed. Closely adjacent to this one is the biggest of the "blue holes," the large spring-fed lake in the center of the town of Castalia. The Miller Blue Hole, six miles to the west of this town, is nearly two acres in size, and has been preserved by the state since 1932 in a relatively natural condi- tion. It is located in a 13-acre wooded tract surrounded by agricultural lands. Earlier works dealing with Miller Blue Hole have only briefly mentioned water beetles.
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  • The Insect Database in Dokdo, Korea: an Updated Version in 2020
    Biodiversity Data Journal 9: e62011 doi: 10.3897/BDJ.9.e62011 Data Paper The Insect database in Dokdo, Korea: An updated version in 2020 Jihun Ryu‡,§, Young-Kun Kim |, Sang Jae Suh|, Kwang Shik Choi‡,§,¶ ‡ School of Life Science, BK21 FOUR KNU Creative BioResearch Group, Kyungpook National University, Daegu, South Korea § Research Institute for Dok-do and Ulleung-do Island, Kyungpook National University, Daegu, South Korea | School of Applied Biosciences, Kyungpook National University, Daegu, South Korea ¶ Research Institute for Phylogenomics and Evolution, Kyungpook National University, Daegu, South Korea Corresponding author: Kwang Shik Choi ([email protected]) Academic editor: Paulo Borges Received: 14 Dec 2020 | Accepted: 20 Jan 2021 | Published: 26 Jan 2021 Citation: Ryu J, Kim Y-K, Suh SJ, Choi KS (2021) The Insect database in Dokdo, Korea: An updated version in 2020. Biodiversity Data Journal 9: e62011. https://doi.org/10.3897/BDJ.9.e62011 Abstract Background Dokdo, a group of islands near the East Coast of South Korea, comprises 89 small islands. These volcanic islands were created by an eruption that also led to the formation of the Ulleungdo Islands (located in the East Sea), which are approximately 87.525 km away from Dokdo. Dokdo is important for geopolitical reasons; however, because of certain barriers to investigation, such as weather and time constraints, knowledge of its insect fauna is limited compared to that of Ulleungdo. Until 2017, insect fauna on Dokdo included 10 orders, 74 families, 165 species and 23 undetermined species; subsequently, from 2018 to 2019, we discovered 23 previously unrecorded species and three undetermined species via an insect survey.
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