The LepidopterologicalSocietyLepidopterological Society of Japan
ue L maTmns.lopid. Soc. 1mpan 46 <2): 45-62, June 1995
On the LyeaenQpsis group of genera (Lepidoptera, Lycaenidae) of the Sulawesi region
Alan C. CAssiDy
18 Woodhurst Road, Maidenhead, Berkshire, SL6 8TF, England
Abstraet The status of the Sulawesi species ef the LycaenQPsis group of lycaenid genera (sensu Eliot & Kawazoe, 1983) is revised in the ]ight of new evidence from recent captures. A new species Sancten'ia drakei is described. Changes are given to the range and status of other taxa.
Key words Sulawesi, Lycaenidae, Polyommatini, lvcaenQPsis, Preject Wallace.
Introduetion
The butterflies of this group of genera have given problems to interested taxonomists for many years. A much-needed, authQritative and very detailed revision was undertaken `E by Eliot & Kawazoe and published in 1983 (hereinafter abbreviated to & K'). That revision is adopted as the baseline for further consideration of the representatives of the group from the Sulawesi region, including the islands of Banggai, Tukangbesi, Tana- jampea, Selayar, Sangihe, Talaud and the Sula archipelago as well as the mainland itself.
That the publication of further information on the group should be required so soon after a major revision is a result of discoveries made by the author in Northern Su!awesi whilst a member of Project Wallace, the commemorative expedition of the Royal Entomological Society of London.
Taxonomic and bionomic information is given in the form of an annotated checklist in accordance with the systematics of E & K. Where necessary, amendments are suggest- ed to update E & K's keys in the light of the new information given, Distributional inforrnation is based on material in the BMNH, RNH Leiden and on specimens collected during Project Wallace, together with data from original descriptions where applicable. The limited amount of an altitudinal and temporal data available is given as heights in metres and as months of the year (in Roman numerals).
Abbreyiations
AMEBMNHEIHUMMNHAllyn Museum of Entomo]ogy, Sarasota, Florida. British Museum (Natural History), London. Entomological Institute, Hokkaido University, Sapporo. Osaka Municipal Museum (Natural History), Osaka. MNHU Museum ftir Naturkunde der Humboldt-Universitat, Berlin. NSMRNHSMSMTZSBSZSINational Science Museum, Tokyo. Rijksmuseum von Natuurlijke Historie, Leiden,
Senckenberg Museum, Frankfurt. Staatliches Museum fttr Tierkunde, Dresden. Zoologische Sammulung des Bayerischen Staates, Munich. Zoological Survey of India, Calcutta,
NII-Electronic Library Service The LepidopterologicalSocietyLepidopterological Society of Japan
46･ Alan C, CAssmy
Genus IVleQpithecQps Distant
IVbcrpithecops sumbanus sumbanus Eliot & Kawazoe
IVleopithecaPs sumbanms sumbanus Eliot & Kawazoe, 1983: 47. Holotype male, Sumba (BMNH).
Locations. [Islands] : Tanajampea.
IVeqpithecQps utnbretta dorothea Eliot & Kawazoe
Nbopitltecops umbreha dorothea Eliot & Kawazoe, 1983 : 53. Holotype male, Sula Islands : Sula Besi, x. 1897 (W. Dohorty) (BMNH).
Although known from the Sula Islands, no representative of the species, or of the allied AL 2almom (Butler), appears yet to have been taken on mainland Sulawesi. This is perhaps surprising as the genus is widespread from Malaya to Borneo, and occurs in the Philippines and North Moluccas.
Locations. [Islands]: Sula Besi (x).
IVIiqpithecops umbretta tituria (Fruhstorfer)
PithecQPs 2almo,a titun'a Fruhstorfer, 1919: 84. Lectetype male, Tanajampea (BMNH).
Locations. [Islands] : Tanajampea (xii).
Genus Megisba Moore
Megisba mataya sikkima Moore
Magisba sthkima Moore, 1884: 21. Holotype male, India: Sikkim (?ZSI>. Ilathalia albidisca Moore, 1884: 21. Lectotype male, India (BMNH). Megisba malaya (Horsfield) : Nicevi]le, 1890 : 61, pl. 26, fig. 165 male
Fruhstorfer's original material was from Southern Sulawesi, but the species has now been recorded from several parts of the island. Males were regularly encountered drinking from wet sand beside lowland rivers in the Toraut district during the expedi- tion. Females could be found in nearby clearings or along paths through the forest.
Locations. [Mainland] : N Sulawesi : Dumoga-Bone Nat Pk (200 m) (i, ii, iii, iv), Palu (xi), Tawaya (ix). S Sulawesi:Bantimurung, Bonthain (viii, ix, x, xO. [Islands]: Tukangbesi (j), Sula Besi (x), Sangihe (iii).
Genus Sancterila Eliot & Kawazoe (=Celarchzts Eliot & Kawazoe, syn. llov. )
Subgenus Skencten'ki Eliot & Kawazoe. Type species: Cmpido deliciosa Pagenstecher, 1896. Subgenus Celarvhus E)iot & Kawazoe, stat. nov. Type species: Cyanin's tvanspectus archagathos Fruhstorfer, ]910. Subgenus Annentulus Eliot & Kawazoe, stat. nov. Type species : Cyanin's sheij2)ntii NicEville, 19e2.
E & K erected Cegarchzas on the basis of a deep inward fold in the vinculum, near the base
NII-Electronic LibraryMbrary Service The LepidopterologicalSocietyLepidopterological Society of Japan
Lo,caenopszs Group of Sulawesi 47
of the hump This unusual feature occurs also in Skenctenla but was overlooked by Eliot ipers comm ) during the preparation of his manuscript, and these two are considered as the same genus Additionally, E & K divided Celarchzts mto two subgenera, Celarchus and Armentztitts Smce Sanctenla and Cela7Thus were presented in the same work by E & K and I can not find any precedence for either of them excepting the priority due to their page order, I choose Stinctenla as the generic name for them as an act of first reviser It is therefore necessary to sink Celarchzts as a synenym of SZznctenla at the generic ]evel I propose the division of SZznctenla in thrs sense into three subgenera, Skenctenla, Celarchus and Armentulus in accordance with E & K's descriptions
Saneterita delictosa delictosa (Pagenstecher) (Figs 1, 2)
Cmpido (Cyanznsi delzciosa Pagenstecher, 1896 50 Holotype male, Sulawesi Mrnahassa {SM)
M " l ll ll IIt ttl l t tIt Iill aS III ltlly
w
IEFigsl-6
&2nctenlaspp 1,2 S delictosa delz"osa (Pagenstecher), di 3, 4 S drakez sp nov, holotype, [1 5,6 Dztto, paratype, ? (1,3,5 Upperside, 2,4,6 Under- side)
NII-Electronic Library Service The LepidopterologicalSocietyLepidopterological Society of Japan
48 Alan C. CAsslDy
Cmpidb delicr'osa Pagenstecher : Pagenstecher, 1897 : 416, pl. 20, fig. 8 male. Notarihn'ntts (Monodontidesi deliciosa (Pagenstecher) : Toxopeus, 1927 : 258.
The holotype remained the only known specimen until a long series was taken by Cassidy in 1985. E & K, p. 76, incorrectly included under this taxon three butterflies taken on Mt Tambusisi, Central Sulawesi in 1980. Unfortunately, the genitalia drawings on pp, 76- 77 of E & K were taken from this latter material. Following the rediscovery of the species in NDrth Sulawesi, the holotype was borrowed from Frankfurt and dissected. This showed the branched, spiked valvae shown in E & K, p. 81, fig. 48, as S. sohmai.
`Gunung During Project Wallace deliciosa was taken in two locations. One of these was 1440', an anomalously-named hilltop at 1,200 m elevation in the Toraut district of the Dumoga-Bone National Park. The other was the peak of Gunung Muajat, east of Kotamobagu, at 1,780m. All of the specimens collected were males, which were habitually involved in tailchases around the hillteps involving up to four or five individ- uals in a stream of blue. The form of the female remains unknown. Forewing length in the males varies frorn 14-16 mm.
Locations, [Mainland] : N Sulawesi : Dumoga-Bone Nat Pk (1,200 m) (iii), Gn Muajat (1,780 m) (iv). The species is unlikely to be found at elevations below 1,OOO m.
Sancterila deliciosa sohmai Eliot & Kawazo6, stat. nov.
Sancten7a sohmai Eliot & Kawazoe, 1983 : 80. Holotype male, Sulawesi : Toraja (MMNH).
This was described as a good species by E & K, but has proved to be conspecific with deliciosa by dissection of the male genitalia. The synonymy is not absolute, however, as the male of sohmai has enly a slight dusting of black subapical scales on the forewing upperside while deliciosa has a broad apical black triangle with superimposed blue scaling, As these taxa are geographically isolated in their respective montane regions, I suspect that intermediates will not be found. Therefore I retain sohmai as a valid subspecific name,
Locations. [Mainland] : C Sulawesi : Toraja (Pulu Pulu and Parado) (2,OOO-2,300 m) (i, vi, vii, x).
Sancterila drakei sp. noy. (Figs3-8)
Slancten'la deliciosa Pagenstecher: E & K, 1983: 76-77, Figs 42,43,470. Misidentification.
This new name is given to the two males and single female taken on Mt Tambusisi in 1980 by A. Bedford-Russell. The male is superficially similar te deliciosa with which it was associated by E & K. By way of description, I can do no better than to quote E & K and reproduce their genitalia drawings (Figs7 & 8). This I do with their kind permissien. It is evident from the study of this genus in Sulawesi that external charac- ters do not give satisfactory interspecific identification and that genitalic examination is necessary for each new population discovered.
In the males of drafeei, the fQrewing border is broader than in deliciosa, measuring 1.5 mrn at the tornus, the apical violet-blue patch is smaller, and on the hindwing the marginal lunules are conjoined and the submarginal lunules are more strongly developed. The androconia have an average of 14 ridges, In the male genitalia, the valva has a bulbous base with a single long, smooth apical process which is curved through almost 9e degrees.
In the female of dmkei the basal and discal areas are greyish blue overlaid basally with shining, dark blue scales. The forewing border is inwardly diffuse, measures 2.5 mm at
NII-Electronic Library Service The LepidopterologicalSocietyLepidopterological Society ofofJapan Japan
LycaenoPsisGroup of Sulawesi 49
e.
Kawazoe, 1983) Fig. 7. Male genitalia of Skencten'ladmkei sp. nov. (AfterEliot &
-i . g
dird 1,:pt1 L.qx} ,lt Xjlt;t?;i 'f 3'-, l.･,rif' ii A･!･t= ' "
Eliot & Kawazoe, 1983) Fig. 8.Female genitalia of Stlncten'ladmkeisp.nov.(After
NII-Electronic Library Service The LepidopterologicalSocietyLepidopterological Society of Japan
50 Alan C. CAssmy
vein Cu2 and increases slightly at the dorsum. There are four post-discal blackish spots in spaces lb to 4, that in space lb very faint. On the hindwing the marginal spots are contiguous, the submarginal lunules rather diffuse, the costal fuscous area fi11s spaces 8 and 7 and the basal half of space 6, and there are blackish post-discal spots in spaces 3 to 5. 0n the underside of both sexes the Celastn'na philipPina-like markings are com- plete and of moderate intensity.
At the suggestion of the collector, Bedford-Russell, the species is named after the famous English sailor/explorer Sir Francis Drake, to whom was dedicated Operation Drake, the expedition which provided the opportunity for much valuable collecting on Mt Tam- busisi.Typematerial.
Holotype. c7,East-CentralSulawesi,MtTambusisi,7,500ft,1.iv.1980, Operation Drake. (A. Bedford--Russel) (BMNH). Paratypes, 1 c{i, as holotype, in coll. Bedford-Russell. 1 \, Mt Tambusisi, 4,OOOft, 5-10. iii, 1980 (A. Bedford-Russell) (BMNH),
Sancterila russetli Eliot & Kawazoe
Skencten'la ntsselti Eliot & Kawazoe, 1983: 79, figs434"437. Holotype male, Sulawesi: East-central CBMNH),
This species is sympatric on Mt Tambusisi with drakei. No further information has become availab]e since the original description.
Location. [Mainland] : EC Sulawesi (Mt Tambu$isi, 760 m) (iii).
Genus Udara Toxopeus
Udara dilecta thoria (Fruhstorfer) (Figs9-12)
Clyanirts Planda thoha Fruhstorfer, 1910: 292. Lectotype male, Sulawesi (BMNH). I,ycaenopsis caniia thoria (Fruhstorfer): Fruhstorfer, 1917: 16 (`Sttdcelebes'); 1922 :' 865, pl. 152f female, 152g male. Cleldstrina (Uda7u) dilecla thoria (Fruhstorfer) : Toxopeus, 1928 : 230.
This is a fairly common species on hilltops above 1,OOO m. It is easily confused with CL rona Grose-Smith but can be distinguished from it by examination of the post-discal marks in spaces 6 and 7 on the underside of the hindwing. In tZ d. thoria these are of contrasting colours, that in space 7 darker, and usually oblique and well separated, In [L rona the spots are both brown, are aligned and close, almost forming a single line. As with the fellowing Udava spe ¢ ies, females are much harder to find than males unless a regular fiight path is discovered, often at a clear, treeless point where the butterflies cross a ridge. Females seem to use these paths, all travelling in the same direction, from about 11.30 am to around 14.30 pm, possibly during dispersal flights.
Locatiens. [Mainland] : N Sulawesi : Dumoga-Bone Nat Pk (1,200 m) (iii), Gn Muajat (1,IOO-1,780 m) (ii, iv). C Sulawesi : Pulu-Pulu (i, v). EC Sulawesj : Mt Tambusisi (ii), Lake Matano (iii). S Sulawesi : Bonthain (ii, x).
thlara rona rona Cb,anin's rona Grose-Smith, 1894 : 572. Holotype male, New Guinea: Irian Jaya (BMNH). L,),caenopsis rona (Grose-Smith) ; Chapman, 1909 : 456, fig. 97 rnale genitalia. Udam rona (Grose-Smith): D]Abrera, 1971: 383. Cekzstn'na (Uda?ul singalensis rona (Grose-Smith) : Toxopeus, 1928 : 230. NII-Electronic Library Service The LepldopterologloalLepidopterological SooletySociety of Japan L ／，caenopszs Group of Sulawesi 51 9 齏屡靂 塵巒層創糎鑾 瞥8 讐鬱 曲 蟹 暫 鍾謝麹屡 謡智崖● 營巒 轡童■ 釦讐 1 耳 壽 畧 逹津 11 羶 翹靂 匿蟇璽 騷璽■ 蘯■籔猶厳臨齏鷺 魘 ■ 璢膕 溷 靂醒 覆 露 瀝 ■ 量 轣 璽 瞎 匪翻溷雇置コ置●響鋼簒陰礬雪 腰醺［奮1■鉗礬護ぼ縢］遷誉n激 ヒ l l 15 匿暫腿 奮口窰讐鏨蟹冨蟹墜齏鬘擾聾 贈邇 龜彊覇 蘯奮看 猶塵■猶蠻暫潮 − Flgs 9 16 Udara spp g lO U dzlecta thona （Fruhstorfer），♂ 11，12 Ditto，♀ 13，14 rona Grose − 15 161 ）ttto 9 11 13 15 Upperslde 10 12 Urona （ Smlth） ♂ ， ，♀ （， ， ， ， ， 14，16 Underside） 一 NII-ElectronicN 工 工 Eleotronlo LibraryLlbrary Service The LepidopterologicalSocietyLepidopterological Society of Japan 52 Alan C CAsslDy Cyamms beretava Ribbe 1899 243, pl 4 fig 14 male Holotype male, Bisrnark Is New Britain (SMT) Cyanzns btcrgz Bethune-Baker 1908 117 pl 8 fig 11 male Helotype male, New Guinea Papua New Guinea This species was quite frequently encountered on Gn Mua]at m open areas at about 1,5eO m, but was not found on the summit at 1,780m Nor was it found at a well-collected 1,200m hilltop in the Teraut area of the Dumoga-Bone National Park It therefore appears more restricted in vertical distribution than the closely allied U d thona Locations [Mamland] N Sulawesi Gn Muajat (1,500m) (iv) C Sulawesi Pulu- Pulu (ix) EC Sulwesi Mt Tambusisi (1,200m) (ni) Udara placidula placidula (H H Druce) (Figs 17, 18) dyantnsPlaczdulaH H Druce,1895 572,pl 32,fig$6male 7female Holotypemale,Borneo(MNHU) ]bycaenQPsib tenella Plactdzala (H H Druce) Chapman, 1909 464, fig 109 male genitalta UZiara Plactdula Placzdula (H H Druce) Hayashi, 1976 152 This species can be separated from the two preceding by the great disparity in size between the post-discal spots in spaces 6 and 7 of the underside of the hmdwmg Indeed, that m space 6 may be almost obselete In the male the forewing upperside border is broader than in the other two species, being almost 1 mm wide During ProJect Wallace, Pla"dzala proved most elusive, only one specirnen bemg obtained This female was found at the forest edge, beside agricultural land at 1,200 m on the foothills of Gn MuaJat Locations [Mamland] N Sulawesi Gn MuaJat (1,200m) Illk }"mp ;f ., .kl ItE ,iil?" Il\fi\ ,k ii' vaIR-initrim b/: i-ewm s,es IX ii It ll ,, 19 w a,･iii t a ntfi/ew Figs17-20 Udamspp 17,18 UPlaczdulaPlacedula (H H Druce), 4 19,20 U cume nae eziphon Fruhstorfer, dny (17,19 Upperside,18,2e Underside) NII-Electronic Library Service The LepidopterologicalSocietyLepidopterological Society of Japan L,ycaenopsis Group of Sulawesi 53 Pulu (ix, xii), Palopo (iii), EC Sulawesi : Mt Tambusisi (1,200 m) (iiD. Udara camenae euphon (Fruhstorfer) (Figs 19, 20) dyanims' singalensis emphon Fruhstorfer, 1910: 291. Lectotype male, Sulawesi (BMNH). Lycaenopsis stmphis emphon (Fruhstorfer) : Fruhstorfer, 1917: 10, pl.1, fig.8 male genitalia. Clyanin's camenae valeria Fruhstorfer, 1910: 296. Lectotype female, Sulawesi Not previously recorded from the north of Sulawesi, this species was found to be locally "1440" quite common on hilltop (1,200 m), and on the summit on Gn Muajat. In these locations the forewing length varied from 15.0-15.5 mm, much the same as for examples from the Toraja highlands in Central Sulawesi. Specimens from the peak of Bonthain, in the south, average 17 mm. Locations. [Mainland] : N Sulawesi : Dumoga-Bone Nat Pk (1,200 m) (iii), Gn Muajat (1,780m) (iv). C Sulawesi : Pulu-Pulu (i, iii), Toraja. S Sulawesi : Bonthain (ii), Uiclara aristius aristius (Fruhstorfer) Clyaniri cesve an'stius Fruhstorfer, 1910: 294. Lectotype male, Sulawesi (BMNH). Lycaenepsis an'stius an'stitts Locations. [Mainland] : Sulawesi : Bonthain (1,OOO-2,OOO m) (ii, x). Udara aristius tetvari (Ribbe) byanin's lewan' Ribbe, 1926: 89. Holotype male, Sulawesi (SMT). This subspecies occurs in the Toraja highlands. The differences between it and aristius are well documented in E & K (p. 119). Lecations. [Mainland] : Sulawesi: Toraja highlands, Pulu-Pulu, Mt Osing (2,OOem) (vii), Udara etsuzoi Eliot & Kawazoe Udam etsuaoi Eliot & Kawazoe, 1983 : 119, figs 438, 439. Holotype male, Sulawesi : Central (NSM). No further information has come avasilable since the original description. Locations : [Mainland] : C Sulawesi : Toraja, Pulu-Pulu, Mt Osing (2,OOO m) (vii, x), Parado (2,eOO-2,300 m) (v, vi, vii, x). Genus Sidima EIiot & Kawazo6 Sidima sulawesiana Eliot & Kawazoe Sidima sulawesiana Eliot & Kawazoe, 1983 : 165, figs 376, 377, 392, 393. Holetype male, Sulawesi : Lake Dano (AME). This species was described from a pair of butterflies taken in Northern Sulawesi (prob- ably in the Tondano area) in 1940. A single male was taken on the summit of Gn Mogogonipa, 1,eOO m, during Project Wallace. Its distribution rnust be extremely local ; its apparent absence from other hilltops in the vicinity which were extensively collected at the same time cannot be explained. NII-Electronic LibraryMbrary Service The LepidopterologicalSocietyLepidopterological Society of Japan 54 Alan C CAssmy ll ' w tl lll{l 1W l' I ll tlI-Inv i" ll /wwilllIII:l l,#41 imaMibelfftinwwIMISffwwma23 Figs21-24 Aaytoloptsspp 21,22 A Puspakuehnz (Rober), c{n 23,24 A sanz`znga(Fmhs- torfer), \ {21,23 Upperside,22, 24 Underside) Locations [Mamland] N Sulawesi Tondano (vi, xii), Dumoga'Bone Nat Pk, Gn Mogogempa (1,OOO m) (iv) Genus Acytolepts Toxopeus Acytotepis puspa kuehni (Rober) (Figs 21, 22) Plebeius k"ehnz Rober, [1886] 60,pl 4 fig 29 Although Puspa is a widespread species, and kuehnt is known from South, East and North Sulawesi, mdividuals were not common during Pro)ect Wallace It is a lowland forest creature and did not appear to congregate anywhere in significant numbers A number of males were taken at the bathing place beside the Toraut river at the expedition Base Camp They appeared to be attracted by human presence although it was not clear whether this was to damp towellmg or to the soapy water among the riverside pebbles No female was discovered by the author during four months in the area Locations [Mainland] N Sulawesi Durnoga-Bone Nat Pk (20em) (ii, iv), Dongala, Palu (ix), Kalawara E Sulawesi Tombugu EC Sulawesi Lake Matano (m) S NII-Electronic Library Service The LepidopterologicalSocietyLepidopterological Society of Japan LycaenQPsis Group of Sulawesi S5 Sulawesi : Bantimurung (vii, viii, ix), Samanga (xi), [Islands] : Banggai. Acgtolepis pttspa deronda (Fruhstorfer) lvcaenQPsis pmspa, var.:Chapman, 1909:438. [o,caenopsis Puspa derondo Fruhstorfer, 1922: 871. Lectotype male, Sula Is (BMNH). In addition to the type series from the Sula Is, the BMNH has one specimen, ex Hewitson `Tondano' collection, from which is believed to be from the Minahassa peninsula. This location is considered doubtful as all other specimens seen from Minahassa are of ssp. leuehni. Locations. [Mainland]:N Sulawesi:Tondano (?). [Islands]:Sula Besi (x), Sula Mangole (x, xi), Acytolepis najTara (Fruhstorfer) Lfemale'). Clyanin's nop'artx Fruhstorfer, 1910: 287 (partim, male nec Holotype male, Sulawesi: Bua- Kraeng 5,OOO ft (BMNH). This species was not confirmed in the north of the island during Project Wallace, but two males were taken by Tennent at Lore-Lindu in August 1985. These vary in forewing length from 12 to 14 mm. Locations. [Mainland]:N Sulawesi:Tondano. C Sulawesi:Lore-Lindu (600- 1,2eO m) (viii), Tanah Toraja (viii), Bonthain (1,500 m). Aeutolepis samanga (Fruhstorfer) (Figs23,24) (lyaniris aktzsa (Hersfield>: Pagenstecher, 1897: 415. Misidentificatien. C5,anin's samanga Fruhsterfer, 1910:286. Lectotype female, Sulawesi (not located). Acytoler)is Puspa samanga (Fruhstorfer) : Toxopeus, 1928: 231 (partim, female nec male). Celastn'na samangtr (Fruhstorfer) : Eliot, 1956 : 35, fig. 2 male genitaJia. L,1,caenopsis Puspa haehni (ROber), dry season form female samdinga (Fruhstorfer) : Fruhstorfer, 1922 : 87e, pl. 152f female (entitled lyseas in error). `female' Oanin's nql'ava Fruhstorfer, 1910 : 287 (partim, recte maTe, nec male). `male' I,yczzenqPsis Puspa nql'ava (Fruhstorfer) : Fruhstorfer, 1922 : 870, pl. 152c recte female (partirn). LycaenQPsis akasa Parakczsa Fruhstorfer, 1922 : 862. Syntypes, Sulawesi : Minahassa Four females were taken during Project Wallace, from two locations. AII had the upperside hindwing rnargin much darker than that illustrated in E & K, fig. 267. Eliot ipers, comm.) states that some males also show similar characteristics. Certainly a male taken by Tennent in the Dumoga-Bone NP in July has a solid black border 4 mm wide on the hindwing. Alse, a male in the collection of Bedford-Russell, taken at Lake Matano in March 1980, has the upperside of the hindwing almest completely dark brown with only traces of blue scaling. It is likely that this represents nothing more significant than local variation, and is not accorded any taxonomic importance. Locations. [Mainland] : N Sulawesi : Dumoga-Bone Nat Pk (1,20e m) NII-Electronic Library Service The LepidopterologicalSocietyLepidopterological Society of Japan 56 Alan C, CAsslDy 'llli ,, .i,lliiil,il: iiil,, .l:ili/ i .,i .,llll1.,lllll,,::/, t･'zi:'li :w･l"･://///.l::!il::/11!:l･･:ill: ft-l' l' l'li･ii'l:･",i/i･llilil"ll::':/'..}I..ili ,Ill"'i"'l"'-/":::i! ,/#f,l:/::::t/lllA.･l:l, l"'ig･'\l,/,,iix"'i･ lil"ilil;. t... ･･25"" il tttl:ll ...:t, ,.,t/ ttttii. / ... ..t./ 'fS:'.'.',,,25 "::' ･--i:l:/ ,li･ ittv,...m.i.-whwhintwX Figs25,26. Celczstn'na PhilipPina gmdeniga (Fruhstorfer), cii. (25. Upperside ; 26. Under$ide.) Genus Cletastrina Tutt Clelastrina philippina phitippina (Semper) Clyanirz's PhilipPina Semper, 1889: 168, pl.2, figs16,18 male, 17 female (panim). Lectotype male, Philippines (SM). Cb,anin's PhilipPina PhilipPina Semper (partim) : Fruhstorfer, 1910 : 300 (Luzon>. Ll,caenopsis cardia Philipt,ina (Semper) : Fruhstorfer, 1917 : 14, I,ycaenopsis PhilipPina PhilipPina (Semper) : Toxopeus, 1926 : 369. Celastn'na PhilipPina PhilipPina (Semper) : Hayashi & Iwanaga, 1974: l8, figs 12, 13 male, lc-5c male genitalia. Locations. [Islands] : Talaud (ii, iii). Cetastrina philippina gradeniga (Fruhstorfer) (Figs 25, 26) ([lynan's gradeniga Fruhsterfer, 1910 : 3el. Lectotype male, Sulawesi (BMNH). LycaenQPsis neddo gnidbniga (Fruhstorfer): Fruhstorfer, 19I7: 35; 1922: 875, pl. 152e male. Lycaenopsis PhilipPina gvaclenigva [sic] (Fruhstorfer) : Toxopeus, 1926 : 369. lycaenopsis neddo Proba Fruhstorfer, 1922 : 875. Lectotype male, Sulawesi (ZSBS). A short series of males was taken by Tennent at Palu in August 1985, In all cases the upperside forewing border was 4 mm wide (compared with 2 mm quoted in E & K), whilst the hindwing border varied in width from 4 mm to 6 mm. Locations. [Mainland]: N Sulawesi: Palu (viii), E Sulawesi: Tombugu. S Sula- wesi: Bonthain (viii, ix). [Islands]: Sula Is (x). C:elastrina tavendularis lyce (Grose-Srnith) Cvanin's lyce Grose-Smith, 1895: 506. Lectotype male, Sulawesi (BMNH), Qyanin's Pellonia Frnhstorier, 1910 : 301. Lectotype male, Sulawesi {BMNH). Lo,caenopsis limbatus lyce, f. Pellonia (Fruhstorfer) : Fruhstorfer, 1917 : 32. The only specimen taken during Project Wallace was a male from the Toraut river bathing spot in March. It seemed attracted by human activity, as described for A. P. kuehni. Locations. [Mainland] : N Sulawesi : Dumoga-Bone Nat Pk (200 m) (iii).S Sulawesi : Bua-Kraeng (1,500 m) (ii, viii, ix, x). NII-Electronic LibraryMbrary Service The LepidopterologicalSocletyLepidopterological Society ofofJapan Japan L(ycaenopsts Group of Sularvesi 57 Il l, l''l l 'I ' ISI"W-fiWNfi1=ww-taXWtvell' ' ' I ' I l l ' l lll ; }l .l l l ' ll ll lltl l' tt tl l l ! ' lt i ,, ,/ 1!ii l ;I Yi Il l lt l' t I l' L t lI l' ' l t tl2, i l lll itan:gk"vaiiiuhvekKihtigki Figs27-32 Mbnodontzdes spp 27,28 M kolan 6q Dztto, \ 31, 32 M care (Niceville),di (27,29,31 Uppersicle Genus U)'anobothria Toxopeus U)'anobothria cetebica (Fruhstorfer) l4,caenQPsts aBothnndi celebtca Fruhstorfer, 1917 40, pl 1, fig 12 male genitalia, 1922877 pl152e male Holotype male, Sulawesi Bontham (ZSBS) Uranobothna cetebtca (Fruhstorfer) Toxopeus, 1927 261, text-fig 10 eye No further information has become available since the 1983 revision Loeations [Mamland] S Sulawesi Peak of Bontham (2,285m) (iv) NII-Electronic Library Service The LepidopterologicalSocietyLepidopterological Society of Japan 58 Alan C. CAsslDy Ch'anobothria tsukadai Eliot & Kawazoe Uvanobothn'a tsukadoi Eliot & Kawazoe, 1983 : 267, figs 454, 455, 460. Holotype male, Sulawesi : Toraja (NSM). This species was common on the summit of Gn Muajat (1,780 m), when visited in both March and July. Previously it had been known only from the Toraja district. Both genitalia and androconia of these northern examples agree with the original description of the species, although they exhibit considerable variability in size, with forewing lengths from 14 to 19 mm. Locations. [Mainland]: N Sulawesi: Gn Muajat (1,780m) (iii, vii), C Sulawesi: ･ Toraja, Pulu-Pulu&Parado (2,300 m) (v, vi, ix, x). Genus Monodontides Toxopeus Monodontides kolari (Ribbe) (Figs 27-30) Oanirz's (? itagizV feolan' Ribbe, 1926: 88. Holotype male, Sulawesi : Kalawara {SMT). "144e", Both sexes were taken on hilltop confirming the species presence in the north of the island. Locations. [Mainland] : N Sulawesi : Dumoga-Bone Nat Pk (1,200 m) (iii), Kalawara. C Sulawesi : Pulu-Pulu (i). Monodontides cara (Niceville) (Figs 31, 32) a,anin's cara Niceville, 1898 : 143, pl.Z, figs19,20. Holotype male, Sulawesi (ZSI). LycaenQPsis cara (Niceville): Fruhstorfer, 1917: 24, pl. 2, fig. 20; 1922: 868, pl. 152g male and female. A good series of both sexes was obtained from Gn Muajat in March and April 1985, thus extending the known range of the species, which was previously recorded from southern and central areas only. The males of this series show considerable variation in the valvae, as illustrated in Fig, 33, particularly at the apex and in the inner triangular process. However, I attach no taxonomic significance to this variation, and do not Fig.33. Left varvae ef Mbnodontidlas caTzz Nic6ville, showing individual variation in specimens from Gunung Muajat, 1,780 m, Kotamobagu, N Sulawesi. All specimens taken on 9. iv. 1985. NII-Electronic Library Service The LepidopterologicalSocietyLepidopterological Society of Japan Lo,caenQPsis Group of Sulawesi 59 consider that these specimens differ suMciently from typical cam to merit subspecific denomination. Locations. [Mainland] : N Sulawesi : Gn Muajat (1,780 m) (iii, iv). C Sulawesi : Pulu- Pulu (i, x, xi), S Sulawesi : Bonthain (ii, iv). Amendments to keys in Eliot & Kawazoe, 1983 Key to genera and subgenera based on male and female genita]ia 10 Valva without an apical process bearing subsidiary spines ...... Lestraniczts (p. 72) - Valva with an apical process bearing subsidiary spines ..,,.,,.Lycaenopsis (p. 36) 11 Vinculum deeply hollowed into in-turned, flap-like pouches. Valva with an apical process ,...... ,...,...... ,,...... ,...... ,..,...... 12 - Hollow of vinculum shallow, rounded, Valva without an apical process '''-''''''''''''''''''''''･--･-･････････-･････-･･-J･･･････････Plautelkz (p. 168) 12 Valva with apical process turned inwards towards the centreline of the armature .,..,...... ,...... 13 - Valva with apical process directed dorsally or distad. Sometimes also with a subapical or costal process...... Sancterila (StzncteriZa) (p. 74) 13 Valva ending in several scimitar-shaped spines. Lateral fenestrula incomplete ...... Skencterila (Armentulus) (p. 196) - Valva with apical process not as above. Lateral fenestrula complete ...... Sancterila rcelarchtt$) (p. 191) Key to species of Sbencten'la (SancteriILij 1 Stet - Stet 2 Valva with a long, forked, spinous costal process emerging just over halfway from base..,.,....,.,....,..,...,..,...,..,...,,...... ,.,,...... ,,...,..,..deliciosa - Valva not as above ,...... ,..,...... ,...... ･...... ･･-･･････3 3 Valva with a long apical process, bent through nearly 90 degrees ....dvaleei sp. n - Valva with a short, pointed subapical process,.,....,..,....,.,,....,.,,.rztsselli Key to subspecies of Sancten'la deliciosa 1 Male upperside with the forewing border expanding to a broad apical area bearing a vielet-blue patch. Minahassa .....,...... ,,...... degiciosa deliciosa - Male upperside with the forewing border enly 1 mm at the apex and with a detached blackish subapical fascia ...... ,..,...... ,...... ,.deliciosa sohmai Acknowledgements I wish to express thanks to the Staff of the BMNH for encouragement and facilities, to Dr Rienk de Jong of the RNH Leiden, for access to collections and to Mr A. Bedford- Russell, Mr T. W. Harman and Mr W. J. Tennent for making specimens available for examination, An especial debt is owed to Lt, Col. J. N. Eliot (Retd) for valuable comments during the preparation of this paper. Photographs were by the BMNH, and I would particularly like to thank Syntax Wordprocessing, Maidenhead, for production of the manuscript. NII-Electronic Library Service The LepidopterologicalSocietyLepidopterological Society of Japan 60 Alan C. CAsslDy References Bethune-Baker, G. T., 1908. Descriptiens of new species of butterflies of the division Rhopalocera from Africa and from New Guinea. Proc. zool. Soc. Lond. 1908: 110-126, 2pls. Chapman, T., 1909. A review of the species of the lepidopterous genus l4,caenQPsis Feld. {Cyanin's auct. nec Dalm.) on examination of the male ancilliary appendages. Proc. zool. Soc. Lod. 1909: 419- 476, 72 figs. D'Abrera, B., 1971. Butte,Y7ies of the Ablstralinn Ragion, edn 1. 415 pp., Melbourne. Druce, H. H., 1895. A monograph of the Bernean Lycaeniclae. PToc. zool. Soc. Lond. 1895 : 556-627, 4 pls. Eliot, J. N., 1956. New and Iittle-known Rhopalocera from the Oriental region. BulL Roj77es Mzas, 27 : 32-38. Eliot, J. N. and A. Kawazee, 1983. Blzae Butte7:i7ies of the Lycaenopsis Gromp. 309 pp., 560 figs. Lonclon. Fruhstorfer, H., 1910. Neue CbJaniris-Rassen und Ubersicht der bekannten Arten. Stettin. ent. Zig 71 : 282-305. Fruhsterfer,H., 1917. Revisionder Lycaeniclengattung Ll,caenopsis, A?Th. IVZ2turgesch. 82 (A) (1): 1-42, 2 pls. Frnhstorfer, H., 1918. Altes und Neues tiber Magisba maldya. A7vh. AJtzimrgesch. 82 (A) (12) : 4-8. Fruhstorfer, H., 1919. Revision der Artengruppe Pithecops und Grund der Morphologie der Klammeror- gane. Arch, IVLitzargesch. 83 Grese-Smith, H., 1895. Descriptions of new species of butterflies, captured by Mr. Doherty in the islands of the Eastern Archipelago, and new in the Museum of the Hon. Walter Rothschild at Tring. Nbvit. zool. 2: 505-514. Hayashi, H., 1976. New subspecies of ldmides and Udara frorn Palawan CLepidoptera : Lycaenidae). 7),O (]a 27: 151-155. Hayashi, H. and S. Iwanaga, 1974. A new species of Celastrina (Lepidoptera : Lycaenidae) from Borneo ancl Luzon. Konpt 7bkyo 42: 17･-23. Matsumura, S., 1919. 7]Vrousand insects Addit.3: 475-742, of.kipan. pls26-53. Moore, F., 1884. Descriptions ef some new Asiatic diurnal Lepidoptera; chiefiy from specimens contained in the Indian Museum, Calcutta. J asiat. Soc. Beng. 53 : 16-52. Niceville, L. de, 1890. Tlae BertteiY7ies of india, Bu,7nah and Cqylon 3. xi+505pp., 6pls. Calcutta. Niceville, L. de, 1898. 0n new ancl little-known butterfiies from the Indo-Malayan, Austro-Malayan and Australian regiens. J Bombay nat. Hist, Soc, 14 : 236-251, 1 pl. Pagenstecher, A., 1896. Neue malayische Lepidopteren. Ent. AJtichx BerL 22 : 49-54. Pagenstecher, A., 1897. in Ktikenthal, W., Ergebnisse einer zeologischen Forschungsreise in den Molukken und in Borneo. Lepidopteren. Abh. senckenb. natudersch. Ges. 23 : 353-467, 3pls. Ribbe, C., 1899, Beitrage zur Lepidopteren-Fauna des Bismarck- und Solomon-Archipels in der SUd- See. Dt. ent. Z Iris 12:219-260,1pl. Ribbe, C,, 1926. Neue Iycaenenformen hauptsaechlich von Celebes. Ent. Mitt. 15: 78-91. ROber, J., [1886]. Neue Tagschmetterlinge der indo-australischen Fauna. CorrespBl. ent. Vlar. Iri/g 1: 45-72, pls 2-5. Semper, G., 1886-1892. Die Schmettelinge der PhilipPinischen Inseln. 1. Tagfalter. 380+14 pp., 49+2 pls. Wiesbaden. Toxopeus, L. J., 1927. Eine Revision derjavanischen, zu L),caenopsis und verwanclten Genera gehUrigen Arten. Tijcischr. Ent. 70:232-302,1pl, Toxopeus, L. J., 1928. Eine Revision derjavanischen, zu lvcaenopsis und verwandten Genera geh6rigen Arten. TZidschr. Ent. 71: 179-265, 1pl. NII-Electronic Library Service The LepidopterologioalLepidopterological SooietySociety of Japan Lycaenopsis Group of Sulawesi 61 摘 要 ス ラ ウ ェ シ の LycaenopsdS 9e （鱗 翅 目， シ ジ ミ チ ョ ウ 科）（Alan C． Cassidy） ル リ シ ジ ミ類 を含 む Lycaenopsis 9eに つ い て は ， Eliot＆ Kawaze6 （1983）の 分類学 的再検 討 が あ り， こ の 研究 は 現在 で も重 要 な参 照 体 系 とな っ て い る ．筆者 は，1985 年イ ギ リ ス 王 立 昆 虫 学会 に よ っ て ‘ ’ こ 主 催 さ れ た 昆 虫 相 調 査 Project Wallace に 参 加 し ， 多 くの 材 料 を得 る とが で きた ．本 論 文 で は ， こ の に と つ い て ス ラ ェ シ に の に お の い れ ら 標本 も ウ な ら び そ 周辺 域 け る LycaenoPsisge 再 検討 を行 ， S 砌 碗 溺 σ drakei る と と に い つ か の っ た ． に そ の を 新種 を記 載 す も ， く 分 類学 的変 更 を行 以 下 概要 示 す ． Neopithecops umbretta dorothea Eliot＆ Kawazoe ス ラ 諸 島 に 分布 す る が ，近縁 の N ， z α lmora と 同 様 に ス ラ ウ ェ シ 本 島 で は 知 られ て い な い よ う で あ ． が マ ラ ヤ ボ ル オ フ ピ ン モ ル カ に る に か か ス ラ ウ ェ シ る 本属 ， ネ ， ィ リ ， 北 ッ 広 く分 布 す も わ らず に 分布 し な い の は 興 味 深 い ． − ＝ Sanctenla Eliot ＆ Kawazo6 （ Celarchms Eliot＆ Kawazo6 ） に こ と か た Eliot ＆ Kawazo6 （1983）は ， vinculum が 背 方 で 内側 深 く窪む ら Celarchvfs属 を創設 し が こ ら が に し た S αncteril α に も め ら れ る こ と が か っ た ， Eliotら は こ ， の 特徴 は彼 同時 創設 属 認 分 れ を 見落 と し て お り （Eliot私 信），前者 を後者 の シ ノ ニ ム と し た ． S．deliciosa deliciosa（Pagenstecher） い ス 本種 は ，著者 が 1985年 に 多 く の 個体 を採 集 す る ま で ， holotype1個体 の み が 知 られ て た ． ラ ウ ェ シ 北部 に 分布． S．deliciosasohmai Eliot＆ Kawazoe ， stat ．nov ． S，deliciosaの holotypeを解剖 し た 結 果 ，交 尾 器 の valva の 中 央 背 面 に 刺 状 突 起 が あ り， Eliot ら が 一 sohmai と し て 記 載 した種 は deliciosaと同 種 で あ る こ と が わ か っ た ． こ こ で は deliciosaの 亜 種 と した ． ス ラ ウ ェ シ 中部 に 分 布． S．drakei sp ． nov ． Eliot ら が deliciosaと し た も の （誤 同定） を新種 と し て 記載 し た ．本種 は deliciosaに 似 る が ， Eliot ら が 示 し た よ うに ♂ の 交 尾器 ，特 に valva の 形状 は 顕著 に 異 な っ て い る ．外 見 上 は 4π 惣 ∫の ♂ の 前翅 外縁黒 帯 は 幅 が よ り広 く，翅 頂部 の 紫青色 の 斑 紋 は よ り小 さ い こ と な ど で 区別 で き る ． Udara dilecta ’んo ガα （Fruhstorfer） 1 m ど で に ． rona と さ が 標 高 ，000 を越 え る 山頂 部 な 普 通 み られ る 次 ts 混 同 れ る ， 後翅 裏 面第 6，7 室 の 中室外 斑 で 区 別 で き る ．す な わ ち ，本種 で は 第 7 室 の 斑 紋 は 6 室 の も の よ り濃 く鮮 明 で 両者 が 分離 す る が ，次種 で は 両 者が 帯状 と な り連続 的 に 並 ぶ ． − U ．rona rona （Grose Smith） Muajat 山 の 標 高 1，500 m 付近 で 多 く見 ら れ た が ，1，780 m の 山 頂 で は 見 ら れ ず，前 種 と は 垂 直分 布 が 異 な る よ う で あ る ． U．Placidula PlaCttdula（H ． H ． Druce） 上 記 2 種 に 似 る が ，後翅 裏面第 6 室 の 中室外 斑 が ほ ぼ 消失 す る こ と， ♂ の 前翅 表面 外縁 黒 帯 が 広 い 一 NII-ElectronicN 工 工 Eleotronio Library Service The LepidopterologioalLepidopterological SooietySociety of Japan 62 Alan C． CAssIDY （約 lmrn ） こ と な ど で 区別 で き る ． Aaytolapis PuSPa kuehni （Rdber ） は で feuehni ス ラ ェ シ に ， か 1985 本種 広域 分布 種 ，亜 種 も ウ 広 く分 布 す る し し， 年 の調 査 で は あ ま り多 くな く 低 地 の 川 辺 特 に Toraut 川 キ ャ ン プ 地 の 入 浴 地 点 周辺 で 何 ら か の 人 工 物 に 誘 引 さ れ ， ， tt た と 思わ れ る多 くの ♂ が 見 られ た ． U ．tsuhadai Eliot＆ Kawazo6 こ れ ま で ト ラ ジ ャ 地 方 で し か 知 ら れ て い な か っ た が ，1985 年 の 調査 で は 北 ス ラ ウ ェ シ の Muajat 山 ・ （1，780 m ）の 頂上 で 3月 ，7 月 と も に 普通 で あ っ た ．今回得 られ た 個体 の 交尾器 発香鱗 の 形 状 は 原 一 − 記 載 も の と 致 した が ，個体 の 大 き さ に 顕 著 な変 異が あ っ た （前翅 長 14 19mm ）． Monodontides cara （Nic6ville） 一 ス ラ ウ ェ シ の 中 南部 か ら知 られ て い た が ，北 ス ラ ウ ェ シ の Muajat 山 で 3 月，4 月 に 採 集 さ れ た ． Fig．33 に 示 し た よ う に ， valva の 先 端 と 内方 の 突 起 の 形 状 に 個 体変 異 が認 め られ た． ［文責 ： 広渡俊哉 ］ （Accepted November 14，1994） 一 NII-ElectronicN 工 工 Eleotronio Library Service