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Nitrogen Fixation in Roots of Ceanothus
University of Richmond UR Scholarship Repository Biology Faculty Publications Biology Summer 2019 Nitrogen Fixation in Roots of Ceanothus W. John Hayden University of Richmond, [email protected] Follow this and additional works at: https://scholarship.richmond.edu/biology-faculty-publications Part of the Botany Commons, and the Plant Biology Commons Recommended Citation Hayden, John W. "Nitrogen Fixation in Roots of Ceanothus." Sempervirens: The Quarterly of the Virginia Native Plant Society (Summer 2019): 6-7. This Article is brought to you for free and open access by the Biology at UR Scholarship Repository. It has been accepted for inclusion in Biology Faculty Publications by an authorized administrator of UR Scholarship Repository. For more information, please contact [email protected]. 6 Sempervirens, Summer 2019 Nitrogen Fixation in Roots of Ceanothus Article by W. John Hayden, Botany Chair oots are usually out of sight and, nitrogen-containing compounds. Rtherefore, out of mind. But as But there is a paradox about any good gardener will tell you, it is nitrogen and life. You, me, the of utmost importance to understand chickens in my backyard, and the those unique plant organs, even if their Pawpaw tree that shades them are essential functions occur hidden from all constantly bathed in atmospheric cursory observation. The red roots of nitrogen gas, yet that form of nitrogen the 2019 VNPS Wildflower of the Year, is completely unavailable to our cells, Ceanothus americanus, are particularly or the cells of my chickens, or my important because they host symbiotic Pawpaw. Every breath I take brings a bacteria that perform the essential quantity of fresh air into my lungs, 78 function of nitrogen fixation. -
Central Appalachian Broadleaf Forest Coniferous Forest Meadow Province
Selecting Plants for Pollinators A Regional Guide for Farmers, Land Managers, and Gardeners In the Central Appalachian Broadleaf Forest Coniferous Forest Meadow Province Including the states of: Maryland, Pennsylvania, Virginia, West Virginia And parts of: Georgia, Kentucky, and North Carolina, NAPPC South Carolina, Tennessee Table of CONTENTS Why Support Pollinators? 4 Getting Started 5 Central Appalachian Broadleaf Forest 6 Meet the Pollinators 8 Plant Traits 10 Developing Plantings 12 Far ms 13 Public Lands 14 Home Landscapes 15 Bloom Periods 16 Plants That Attract Pollinators 18 Habitat Hints 20 This is one of several guides for Check list 22 different regions in the United States. We welcome your feedback to assist us in making the future Resources and Feedback 23 guides useful. Please contact us at [email protected] Cover: silver spotted skipper courtesy www.dangphoto.net 2 Selecting Plants for Pollinators Selecting Plants for Pollinators A Regional Guide for Farmers, Land Managers, and Gardeners In the Ecological Region of the Central Appalachian Broadleaf Forest Coniferous Forest Meadow Province Including the states of: Maryland, Pennsylvania, Virginia, West Virginia And parts of: Georgia, Kentucky, North Carolina, South Carolina, Tennessee a nappc and Pollinator Partnership™ Publication This guide was funded by the National Fish and Wildlife Foundation, the C.S. Fund, the Plant Conservation Alliance, the U.S. Forest Service, and the Bureau of Land Management with oversight by the Pollinator Partnership™ (www.pollinator.org), in support of the North American Pollinator Protection Campaign (NAPPC–www.nappc.org). Central Appalachian Broadleaf Forest – Coniferous Forest – Meadow Province 3 Why support pollinators? In theIr 1996 book, the Forgotten PollInators, Buchmann and Nabhan estimated that animal pollinators are needed for the reproduction “ Farming feeds of 90% of flowering plants and one third of human food crops. -
Diversity of Rhizobia Associated with Amorpha Fruticosa Isolated from Chinese Soils and Description of Mesorhizobium Amorphae Sp
International Journal of Systematic Bacteriology (1999), 49, 5 1-65 Printed in Great Britain Diversity of rhizobia associated with Amorpha fruticosa isolated from Chinese soils and description of Mesorhizobium amorphae sp. nov. E. T. Wang,lt3 P. van Berkum,2 X. H. SU~,~D. Beyene,2 W. X. Chen3 and E. Martinez-Romerol Author for correspondence : E. T. Wang. Tel : + 52 73 131697. Fax: + 52 73 175581. e-mail: [email protected] 1 Centro de lnvestigacidn Fifty-five Chinese isolates from nodules of Amorpha fruticosa were sobre Fijaci6n de characterized and compared with the type strains of the species and genera of Nitrdgeno, UNAM, Apdo Postal 565-A, Cuernavaca, bacteria which form nitrogen-f ixing symbioses with leguminous host plants. A Morelos, Mexico polyphasic approach, which included RFLP of PCR-amplified 165 rRNA genes, * Alfalfa and Soybean multilocus enzyme electrophoresis (MLEE), DNA-DNA hybridization, 165 rRNA Research Laboratory, gene sequencing, electrophoretic plasmid profiles, cross-nodulation and a Ag ricuI tu ra I Research phenotypic study, was used in the comparative analysis. The isolates Service, US Department of Agriculture, BeltsviI le, M D originated from several different sites in China and they varied in their 20705, USA phenotypic and genetic characteristics. The majority of the isolates had 3 Department of moderate to slow growth rates, produced acid on YMA and harboured a 930 kb Microbiology, College of symbiotic plasmid (pSym). Five different RFLP patterns were identified among Biology, China Agricultural the 16s rRNA genes of all the isolates. Isolates grouped by PCR-RFLP of the 165 University, Beijing 100094, People’s Republic of China rRNA genes were also separated into groups by variation in MLEE profiles and by DNA-DNA hybridization. -
Bee-Friendly Flowers: Aster
Bee-Friendly Flowers: Aster Like fireworks to celebrate the coming of fall, the vibrant pinks, purples, and whites of the star flowers burst into bloom just as summer flowers fade. They are ubiquitous, lighting up meadows, woodlands, river bottoms, salt marshes, sand dunes, roadsides, and waste places. There are many native species of asters in North America, but it’s hard to put a precise number on them. The problem is that asters used to be classified with their own genus, but recent strides in DNA analysis have made scientists rethink where to put them. Plants that used to be lumped into the genus Aster are now split into Symphiotrichum, Eurybia, Solidago, and Machaeranthera just to name a few. Not all taxonomists are onboard with the change, so many botanical New England aster sources list more than one name for the same plant. Despite the confusion about what to call them, the variety of asters is enormous. Hybridization between species frequently occurs in the wild and there are a plethora of human-created hybrids and cultivars. Some have clouds of tiny flowers and some have blossoms as large as daisies. What they all have in common is that each aster flower is a composite of numerous disc and ray florets, which collectively give the appearance of a single large flower. The center holds the disc florets, which are tubular, house the nectar, and are usually yellow, orange, or brownish in color. Those near the bullseye location have both stamens and pistils and can provide pollen to visiting insects. The outer discs are all females and only have pistils to receive pollen. -
"Pollinator Paradise" Garden at Chatham Mills
"Pollinator Paradise" Garden at Chatham Mills Created by Debbie Roos, North Carolina Cooperative Extension 206 species, 85% of them native to North Carolina More info at www.carolinapollinatorgarden.org Common Name Scientific Name Origin Perennial Flowers Yarrow Achillea millefolium 'Moonshine' NC Yarrow Achillea millefolium 'Paprika' NC Mexican giant hyssop Agastache mexicana 'Acapulco Orange' Mexico Anise hyssop Agastache x 'Blue Fortune' hybrid of U.S. native Mexican hyssop Agastache x 'Grape Nectar' Mexico Hummingbird mint Agastache x 'Red Happiness' southwest U.S. Licorice hyssop Agastache rupestris southwest U.S. Nodding onion Allium cernuum NC Dwarf indigo bush Amorpha herbacea NC Arkansas bluestar Amsonia hubrichtii Arkansas, Oklahoma Bluestar Amsonia tabernaemontana NC Tall anemone Anemone virginiana NC Eastern wild columbine Aquilegia canadensis NC Golden columbine Aquilegia chrysantha southwest U.S. Common leopardbane Arnica acaulis NC Swamp milkweed Asclepias incarnata NC Purple milkweed Asclepias purpurascens NC Common Name Scientific Name Origin Red milkweed Asclepias rubra NC Common milkweed Asclepias syriaca NC Butterfly weed Asclepias tuberosa NC Redring milkweed Asclepias variegata NC Whorled milkweed Asclepias verticillata NC Dwarf Tartarian aster Aster tataricus 'Jin Dai' exotic Wild indigo Baptisia x 'Carolina Moonlight' NC Wild indigo Baptisia x 'Purple Smoke' NC White wild indigo Baptisia alba NC Dwarf wild indigo Baptisia minor NC Downy wood mint Blephilia ciliata NC Decurrent false aster Boltonia decurrens central U.S. Bush's poppy mallow Callirhoe bushii central U.S. Fringed poppy mallow Callirhoe digitata central U.S. Prairie poppy mallow Callirhoe involucrata central U.S. Clustered poppy mallow Callirhoe triangulata NC Pink turtlehead Chelone lyonii NC Maryland golden aster Chrysopis mariana NC Field thistle Cirsium discolor NC Curlyheads Clematis ochroleuca NC Wild ageratum/mistflower Conoclinium coelestinum NC Palmleaf thoroughwort Conoclinium greggii southwest U.S. -
New Jersey Tea, Ceanothus Americanus, 2019 Virginia Wildlflower of the Year W
University of Richmond UR Scholarship Repository Biology Faculty Publications Biology 2019 New Jersey Tea, Ceanothus Americanus, 2019 Virginia Wildlflower of the Year W. John Hayden University of Richmond, [email protected] Follow this and additional works at: https://scholarship.richmond.edu/biology-faculty-publications Part of the Plant Sciences Commons Recommended Citation W. John Hayden. New Jersey Tea, Ceanothus americanus, 2019 Virginia Wildflower of the Year. Virginia Native Plant Society, 2019. This Brochure is brought to you for free and open access by the Biology at UR Scholarship Repository. It has been accepted for inclusion in Biology Faculty Publications by an authorized administrator of UR Scholarship Repository. For more information, please contact [email protected]. Ceanothus americanus ew Jersey Tea is a low shrub, generally less tips to the periphery of the flower. The five stamens Nthan 1 m tall and often profusely branched. are attached in radial alignment with the petals; fila- Stems are finely hairy, but may become smooth with ments are oriented vertically, positioning the anthers age. Vegetative stems are perennial, but flowering directly above the central portion of the flower. Ova- stems persist for just a single year. Leaves are mostly ries are three-lobed, superior, and positioned atop 5—10 cm long; leaf shape varies from narrowly to thick glandular disks; the short styles are topped with widely ovate, acuminate to acute at the apex, and cor- three-branched stigmas. Fruits possess a finely rugose date to rounded at the base; leaf margins are finely surface layer that is shed prior to ballistic dehiscence serrate; both leaf surfaces may be finely hairy, espe- of the inner layers; in this fashion the three seeds cially on the veins; vein pattern is pinnate with produced by each fruit are propelled a short distance In the Wild a pair of prominent secondary from the parent plant. -
Final Report
Final Report Final pre-release investigations of the gorse thrips (Sericothrips staphylinus) as a biocontrol agent for gorse (Ulex europaeus) in North America Date: August 31, 2012 Award Number: 10-CA-11420004-184 Report Period: June 1, 2010– May 31, 2012 Project Period: June 1, 2010– May 31, 2012 Recipient: Oregon State University Recipient Contact Person: Fritzi Grevstad Principal Investigator/ Project Director: Fritzi Grevstad Introduction Gorse (Ulex europaeus) is an environmental weed classified as noxious in the states of Washington, Oregon, California, and Hawaii. A classical biological control program has been applied in Hawaii with the introduction of 4 gorse-feeding arthropods, but only two of these (a mite and a seed weevil) have been introduced to the mainland U.S. The two insects that have not yet been introduced include the gorse thrips, Sericothips staphylinus (Thysanoptera: Thripidae), and the moth Agonopterix umbellana (Lepidoptera: Oecophoridae). With prior support from the U.S. Forest Service (joint venture agreement # 07-JV-281), we were able to complete host specificity testing of S. staphylinus on 44 North American plant species that were on the original test plant list. However, following review of the proposed Test Plant List, the Technical Advisory Group on Biocontrol of Weeds (TAG) recommended that we include an additional 18 plant species for testing. In this report, we present host specificity testing and related objectives necessary to bring the program to the implementation stage. Objectives (1) Acquire and grow the additional 18 species of plants recommended by the TAG. (2) Complete host specificity trials for the gorse thrips on the 18 plant species. -
The Vascular Plants of Massachusetts
The Vascular Plants of Massachusetts: The Vascular Plants of Massachusetts: A County Checklist • First Revision Melissa Dow Cullina, Bryan Connolly, Bruce Sorrie and Paul Somers Somers Bruce Sorrie and Paul Connolly, Bryan Cullina, Melissa Dow Revision • First A County Checklist Plants of Massachusetts: Vascular The A County Checklist First Revision Melissa Dow Cullina, Bryan Connolly, Bruce Sorrie and Paul Somers Massachusetts Natural Heritage & Endangered Species Program Massachusetts Division of Fisheries and Wildlife Natural Heritage & Endangered Species Program The Natural Heritage & Endangered Species Program (NHESP), part of the Massachusetts Division of Fisheries and Wildlife, is one of the programs forming the Natural Heritage network. NHESP is responsible for the conservation and protection of hundreds of species that are not hunted, fished, trapped, or commercially harvested in the state. The Program's highest priority is protecting the 176 species of vertebrate and invertebrate animals and 259 species of native plants that are officially listed as Endangered, Threatened or of Special Concern in Massachusetts. Endangered species conservation in Massachusetts depends on you! A major source of funding for the protection of rare and endangered species comes from voluntary donations on state income tax forms. Contributions go to the Natural Heritage & Endangered Species Fund, which provides a portion of the operating budget for the Natural Heritage & Endangered Species Program. NHESP protects rare species through biological inventory, -
Lepidoptera Learning Objective
QUARANTINE SIGNIFICANT LEPIDOPTERA OF CONCERN TO THE SOUTHERN UNITED STATES STEVEN PASSOA USDA/APHIS/PPQ 2007 1 LEPIDOPTERA GOALS . Learn techniques of specimen preparation and submission for CAPS Lepidoptera . Develop a list of Lepidoptera of regulatory concern to the southern USA . Learn to SCREEN samples for these species in the stage most likely to be seen by diagnostic labs using the MAJOR characters. Some species are only defined by a combination of features. In those cases, using the associated key and references listed is more accurate. Give examples from the major superfamilies . Distributions and hosts mentioned are the most likely pathways 2 DEVELOP A LIST . Criteria originally modified from biocontrol of weeds list in July 1991 memo, then modified by NEPSC committee . Now widely used in APHIS as mini-PRA . Survey methodology and taxonomic recognition added to economic criteria . Results are either threats (no pathway), CAPS targets (need to survey), or a dead survey (not practical to consider) 3 WHY LABS HATE TO IDENTIFY LEPIDOPTERA . Secret society of critical characters . Constant name changes . Characters hard to see, covered with scales, or both 4 EGGS . Two types . Do not kill important finds and sent urgent . Plan to rear them in a quarantine facility . Spodoptera and Lymantria (and others) cover the eggs with scales from the female’s body 5 LARVAE . Associate leaf miners with the mine and host . Mouthparts are the “genitalia” of the larval world . Fill vials so there is no air bubble when shipping . “Burp” rubber stoppers and parafilm screw top vials . Can kill and ship in vinegar . Put loose parts in small vials 6 PUPAE . -
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IN MEMORIAM: DONALD J. PINKAVA (29 AUGUST 1933–25 JULY 2017) Liz Makings Herbarium (ASU), School of Life Sciences Arizona State University, P.O. Box 874108 Tempe, Arizona 85287-4108, U.S.A. [email protected] My name is Liz Makings and I am the collections manager of the Arizona State University Herbarium. I was a graduate student at ASU in 2000 when I met Dr. Pinkava and he had just retired, so while I missed out on his talents as a teacher, I was lucky to get to know him as a mentor, colleague, and friend. Dr. Pinkava had a heart of gold, a mind like a trap, and a delightful collection of idiosyncrasies that was perfectly suited to his career path. He was hired at ASU in 1964 after completing his PhD. at Ohio State and was immediately responsible for teaching a 300 level botany class called “Flora of Arizona.” He undertook this responsibility with a meticulousness and attention to detail that can only be described as “Pinkavesque,” col- lecting the plants, learning the flora, and scouring the state for the best field trip sites. To his students he was simultaneously feared and adored. His exams turned men into boys and triggered anxiety attacks even among the best. He did not give grades, you earned them. There was no one more demanding, no one more thorough, yet no one more caring and helpful. Many former students have sung his praises and I’ll share this quote from one: “Dr. Pinkava was one of the kindest scientists I have ever interacted with, a trait that sometimes goes missing in our academic world. -
Chromosome Numbers in Compositae, XII: Heliantheae
SMITHSONIAN CONTRIBUTIONS TO BOTANY 0 NCTMBER 52 Chromosome Numbers in Compositae, XII: Heliantheae Harold Robinson, A. Michael Powell, Robert M. King, andJames F. Weedin SMITHSONIAN INSTITUTION PRESS City of Washington 1981 ABSTRACT Robinson, Harold, A. Michael Powell, Robert M. King, and James F. Weedin. Chromosome Numbers in Compositae, XII: Heliantheae. Smithsonian Contri- butions to Botany, number 52, 28 pages, 3 tables, 1981.-Chromosome reports are provided for 145 populations, including first reports for 33 species and three genera, Garcilassa, Riencourtia, and Helianthopsis. Chromosome numbers are arranged according to Robinson’s recently broadened concept of the Heliantheae, with citations for 212 of the ca. 265 genera and 32 of the 35 subtribes. Diverse elements, including the Ambrosieae, typical Heliantheae, most Helenieae, the Tegeteae, and genera such as Arnica from the Senecioneae, are seen to share a specialized cytological history involving polyploid ancestry. The authors disagree with one another regarding the point at which such polyploidy occurred and on whether subtribes lacking higher numbers, such as the Galinsoginae, share the polyploid ancestry. Numerous examples of aneuploid decrease, secondary polyploidy, and some secondary aneuploid decreases are cited. The Marshalliinae are considered remote from other subtribes and close to the Inuleae. Evidence from related tribes favors an ultimate base of X = 10 for the Heliantheae and at least the subfamily As teroideae. OFFICIALPUBLICATION DATE is handstamped in a limited number of initial copies and is recorded in the Institution’s annual report, Smithsonian Year. SERIESCOVER DESIGN: Leaf clearing from the katsura tree Cercidiphyllumjaponicum Siebold and Zuccarini. Library of Congress Cataloging in Publication Data Main entry under title: Chromosome numbers in Compositae, XII. -
Etude Sur L'origine Et L'évolution Des Variations Florales Chez Delphinium L. (Ranunculaceae) À Travers La Morphologie, L'anatomie Et La Tératologie
Etude sur l'origine et l'évolution des variations florales chez Delphinium L. (Ranunculaceae) à travers la morphologie, l'anatomie et la tératologie : 2019SACLS126 : NNT Thèse de doctorat de l'Université Paris-Saclay préparée à l'Université Paris-Sud ED n°567 : Sciences du végétal : du gène à l'écosystème (SDV) Spécialité de doctorat : Biologie Thèse présentée et soutenue à Paris, le 29/05/2019, par Felipe Espinosa Moreno Composition du Jury : Bernard Riera Chargé de Recherche, CNRS (MECADEV) Rapporteur Julien Bachelier Professeur, Freie Universität Berlin (DCPS) Rapporteur Catherine Damerval Directrice de Recherche, CNRS (Génétique Quantitative et Evolution Le Moulon) Présidente Dario De Franceschi Maître de Conférences, Muséum national d'Histoire naturelle (CR2P) Examinateur Sophie Nadot Professeure, Université Paris-Sud (ESE) Directrice de thèse Florian Jabbour Maître de conférences, Muséum national d'Histoire naturelle (ISYEB) Invité Etude sur l'origine et l'évolution des variations florales chez Delphinium L. (Ranunculaceae) à travers la morphologie, l'anatomie et la tératologie Remerciements Ce manuscrit présente le travail de doctorat que j'ai réalisé entre les années 2016 et 2019 au sein de l'Ecole doctorale Sciences du végétale: du gène à l'écosystème, à l'Université Paris-Saclay Paris-Sud et au Muséum national d'Histoire naturelle de Paris. Même si sa réalisation a impliqué un investissement personnel énorme, celui-ci a eu tout son sens uniquement et grâce à l'encadrement, le soutien et l'accompagnement de nombreuses personnes que je remercie de la façon la plus sincère. Je remercie très spécialement Florian Jabbour et Sophie Nadot, mes directeurs de thèse.