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A Molecular Phylogeny of Equatorial African Lacertidae, with the Description of a New Genus and Species from Eastern Democratic Republic of the Congo
Zoological Journal of the Linnean Society, 2011, 163, 913–942. With 7 figures A molecular phylogeny of Equatorial African Lacertidae, with the description of a new genus and species from eastern Democratic Republic of the Congo ELI GREENBAUM1*, CESAR O. VILLANUEVA1, CHIFUNDERA KUSAMBA2, MWENEBATU M. ARISTOTE3 and WILLIAM R. BRANCH4,5 1Department of Biological Sciences, University of Texas at El Paso, 500 West University Avenue, El Paso, TX 79968, USA 2Laboratoire d’Herpétologie, Département de Biologie, Centre de Recherche en Sciences Naturelles, Lwiro, République Démocratique du Congo 3Institut Superieur d’Ecologie pour la Conservation de la Nature, Katana Campus, Sud Kivu, République Démocratique du Congo 4Bayworld, P.O. Box 13147, Humewood 6013, South Africa 5Research Associate, Department of Zoology, Nelson Mandela Metropolitan University, Port Elizabeth, South Africa Received 25 July 2010; revised 21 November 2010; accepted for publication 18 January 2011 Currently, four species of the lacertid lizard genus Adolfus are known from Central and East Africa. We sequenced up to 2825 bp of two mitochondrial [16S and cytochrome b (cyt b)] and two nuclear [(c-mos (oocyte maturation factor) and RAG1 (recombination activating gene 1)] genes from 41 samples of Adolfus (representing every species), two species each of Gastropholis and Holaspis, and in separate analyses combined these data with GenBank sequences of all other Eremiadini genera and four Lacertini outgroups. Data from DNA sequences were analysed with maximum parsimony (PAUP), maximum-likelihood (RAxML) and Bayesian inference (MrBayes) criteria. Results demonstrated that Adolfus is not monophyletic: Adolfus africanus (type species), Adolfus alleni, and Adolfus jacksoni are sister taxa, whereas Adolfus vauereselli and a new species from the Itombwe Plateau of Democratic Republic of the Congo are in a separate lineage. -
Preliminary Analysis of Correlated Evolution of Morphology and Ecological Diversification in Lacertid Lizards
Butll. Soc. Cat. Herp., 19 (2011) Preliminary analysis of correlated evolution of morphology and ecological diversification in lacertid lizards Fèlix Amat Orriols Àrea d'Herpetologia, Museu de Granollers-Ciències Naturals. Francesc Macià 51. 08402 Granollers. Catalonia. Spain. [email protected] Resum S'ha investigat la diversitat morfològica en 129 espècies de lacèrtids i la seva relació amb l'ecologia, per mitjà de mètodes comparatius, utilitzant set variables morfomètriques. La mida corporal és la variable més important, determinant un gradient entre espècies de petita i gran mida independentment evolucionades al llarg de la filogènia dels lacèrtids. Aquesta variable està forta i positivament correlacionada amb les altres, emmascarant els patrons de diversitat morfològica. Anàlisis multivariants en les variables ajustades a la mida corporal mostren una covariació negativa entre les mides relatives de la cua i les extremitats. Remarcablement, les espècies arborícoles i semiarborícoles (Takydromus i el clade africà equatorial) han aparegut dues vegades independentment durant l'evolució dels lacèrtids i es caracteritzen per cues extremadament llargues i extremitats anteriors relativament llargues en comparació a les posteriors. El llangardaix arborícola i planador Holaspis, amb la seva cua curta, constitueix l’única excepció. Un altre cas de convergència ha estat trobat en algunes espècies que es mouen dins de vegetació densa o herba (Tropidosaura, Lacerta agilis, Takydromus amurensis o Zootoca) que presenten cues llargues i extremitats curtes. Al contrari, les especies que viuen en deserts, estepes o matollars amb escassa vegetació aïllada dins grans espais oberts han desenvolupat extremitats posteriors llargues i anteriors curtes per tal d'assolir elevades velocitats i maniobrabilitat. Aquest és el cas especialment de Acanthodactylus i Eremias Abstract Morphologic diversity was studied in 129 species of lacertid lizards and their relationship with ecology by means of comparative analysis on seven linear morphometric measurements. -
DUBAI NATURAL HISTORY GROUP Vol 35 No 4—April, 2020
DUBAI NATURAL HISTORY GROUP Vol 35 No 4—April, 2020 www.dnhg.org Inside this month: page Himalayan Butterfly Visitor 1 Announcements, 2 Errata/Update and Recorders Spotlight! 3 Vivid Red Starfish 4 Malaise Insect Trap Sorting 5 Portuguese Man of War 6 Paper Nautilus at the 6 East Coast Lectures and field trips 8 Contributors— Thanks to the following for their contributions this month: Roxanne Whelan, Lamjed El-Kefi, Angela Manthorpe, Ulrike Andorff, Alexis Biller, Binish Roobas and Gary Feulner. Send in your contributions by 25th May, for the attention of the: [email protected] Garden Watch! he periodic rains of the past fall and winter, which have continued into T spring, may prove to be the heaviest since the record-setting years of the mid-1990s. That may be an inconvenience to city-dwellers, but it presents a tremendous opportunity for naturalists because the rain promotes exceptional plant growth, which in turn is a boon to local fauna. Not only are plants and animals more abundant overall; there is also a greater opportunity to see rare species, many of which exist near their margins of tolerance in the UAE climate and thrive only in wetter years. In addition, a number of flying species, including butterflies, dragonflies and grasshoppers (and probably many others) are opportunistic migrants, visiting the UAE when times are good. This Hoopoe, one of a pair, was This latter phenomenon is believed to account for the UAE's latest "new" butterfly, seen on 28th April. the Indian Fritillary (Argynnis hyperbius), which was 'discovered' in Wadi Wurayah National Park (WWNP) in mid-February by DNHG Insect Recorder Binish Roobas, See inside for more garden during the course of an invited visit to update earlier surveys of WWNP's already observations that took place (Continued on page 4) during this month. -
Reconstruction of Past Distribution for the Mongolian Toad, Strauchbufo Raddei (Anura: Bufonidae) Using Environmental Modeling
Reconstruction of past distribution for the Mongolian toad, Strauchbufo raddei (Anura: Bufonidae) using environmental modeling Spartak N. Litvinchuk1,2, Natalya A. Schepina3 and Amaël Borzée4 1 Institute of Cytology of Russian Academy of Sciences, St. Petersburg, Russia 2 Biological Department, Dagestan State University, Makhachkala, Russia 3 Geological Institute, Siberian Branch of Russian Academy of Sciences, Ulan-Ude, Russia 4 Nanjing Forestry University, Nanjing, China ABSTRACT The use of ecological models enables determining the current distribution of species, but also their past distribution when matching climatic conditions are available. In specific cases, they can also be used to determine the likelihood of fossils to belong to the same species—under the hypothesis that all individuals of a species have the same ecological requirements. Here, using environmental modeling, we reconstructed the distribution of the Mongolian toad, Strauchbufo raddei, since the Last Glacial Maximum and thus covering the time period between the Late Pleistocene and the Holocene. We found the range of the species to have shifted over time, with the LGM population clustered around the current southern range of the species, before expanding east and north during the Pleistocene, and reaching the current range since the mid-Holocene. Finally, we determined that the ecological conditions during the life-time of the mid-Pleistocene fossils attributed to the species in Europe were too different from the one of the extant species or fossils occurring at the same period in Asia to belong to the same species. Subjects Submitted 17 February 2020 Biodiversity, Biogeography, Zoology Accepted 27 April 2020 Keywords Ecological modeling, North East Asia, Refugium, Distribution shift, Holocene, Published 5 June 2020 Pleistocene, Toads, Bufonidae Corresponding author Spartak N. -
Preliminary Analysis of Correlated Evolution of Morphology and Ecological Diversification in Lacertid Lizards
Butll. Soc. Cat. Herp., 19 (2011) Preliminary analysis of correlated evolution of morphology and ecological diversification in lacertid lizards Fèlix Amat Orriols Àrea d'Herpetologia, Museu de Granollers-Ciències Naturals. Francesc Macià 51. 08402 Granollers. Catalonia. Spain. [email protected] Resum S'ha investigat la diversitat morfològica en 129 espècies de lacèrtids i la seva relació amb l'ecologia, per mitjà de mètodes comparatius, utilitzant set variables morfomètriques. La mida corporal és la variable més important, determinant un gradient entre espècies de petita i gran mida independentment evolucionades al llarg de la filogènia dels lacèrtids. Aquesta variable està forta i positivament correlacionada amb les altres, emmascarant els patrons de diversitat morfològica. Anàlisis multivariants en les variables ajustades a la mida corporal mostren una covariació negativa entre les mides relatives de la cua i les extremitats. Remarcablement, les espècies arborícoles i semiarborícoles (Takydromus i el clade africà equatorial) han aparegut dues vegades independentment durant l'evolució dels lacèrtids i es caracteritzen per cues extremadament llargues i extremitats anteriors relativament llargues en comparació a les posteriors. El llangardaix arborícola i planador Holaspis, amb la seva cua curta, constitueix l’única excepció. Un altre cas de convergència ha estat trobat en algunes espècies que es mouen dins de vegetació densa o herba (Tropidosaura, Lacerta agilis, Takydromus amurensis o Zootoca) que presenten cues llargues i extremitats curtes. Al contrari, les especies que viuen en deserts, estepes o matollars amb escassa vegetació aïllada dins grans espais oberts han desenvolupat extremitats posteriors llargues i anteriors curtes per tal d'assolir elevades velocitats i maniobrabilitat. Aquest és el cas especialment de Acanthodactylus i Eremias Abstract Morphologic diversity was studied in 129 species of lacertid lizards and their relationship with ecology by means of comparative analysis on seven linear morphometric measurements. -
An Overview and Checklist of the Native and Alien Herpetofauna of the United Arab Emirates
Herpetological Conservation and Biology 5(3):529–536. Herpetological Conservation and Biology Symposium at the 6th World Congress of Herpetology. AN OVERVIEW AND CHECKLIST OF THE NATIVE AND ALIEN HERPETOFAUNA OF THE UNITED ARAB EMIRATES 1 1 2 PRITPAL S. SOORAE , MYYAS AL QUARQAZ , AND ANDREW S. GARDNER 1Environment Agency-ABU DHABI, P.O. Box 45553, Abu Dhabi, United Arab Emirates, e-mail: [email protected] 2Natural Science and Public Health, College of Arts and Sciences, Zayed University, P.O. Box 4783, Abu Dhabi, United Arab Emirates Abstract.—This paper provides an updated checklist of the United Arab Emirates (UAE) native and alien herpetofauna. The UAE, while largely a desert country with a hyper-arid climate, also has a range of more mesic habitats such as islands, mountains, and wadis. As such it has a diverse native herpetofauna of at least 72 species as follows: two amphibian species (Bufonidae), five marine turtle species (Cheloniidae [four] and Dermochelyidae [one]), 42 lizard species (Agamidae [six], Gekkonidae [19], Lacertidae [10], Scincidae [six], and Varanidae [one]), a single amphisbaenian, and 22 snake species (Leptotyphlopidae [one], Boidae [one], Colubridae [seven], Hydrophiidae [nine], and Viperidae [four]). Additionally, we recorded at least eight alien species, although only the Brahminy Blind Snake (Ramphotyplops braminus) appears to have become naturalized. We also list legislation and international conventions pertinent to the herpetofauna. Key Words.— amphibians; checklist; invasive; reptiles; United Arab Emirates INTRODUCTION (Arnold 1984, 1986; Balletto et al. 1985; Gasperetti 1988; Leviton et al. 1992; Gasperetti et al. 1993; Egan The United Arab Emirates (UAE) is a federation of 2007). -
Scientific Report Water Research and Learning Programme Wadi Wurayah National Park Field Season 2016–2017 EWS-WWF
Scientific Report | Water Research and Learning Programme | Wadi Wurayah National Park | Field Season 2016–2017 Wurayah | Wadi and Learning Programme Scientific Research Report | Water Scientific Report Water Research and Learning Programme Wadi Wurayah National Park Field Season 2016–2017 EWS-WWF 1 Marine Turtle Conservation Project Final Scientifc Report 1 Marine Turtle Conservation Project Final Scientifc Report Marine Turtle Conservation Project Final Scientifc Report 1 PROJECT PARTNERS HSBC Bank Middle East Ltd. One of the largest international banks in the Middle East and a key financial partner and supporter of Wadi Wurayah National Park since 2006, HSBC Bank Middle East Ltd. established the Water Research and Learning Programme as part of its Global Water Programme. Fujairah Municipality Strategic partner and driver of Wadi Wurayah National Park development. The mission of Fujairah Municipality is to provide advanced infrastructure, a sustainable environment, and excellence in services to the people of Fujairah. Emirates Wildlife Society-WWF Emirates Wildlife Society-WWF is a UAE environmental non-governmental organisation established under the patronage of H. H. Sheikh Hamdan bin Zayed Al Nahyan, ruler’s representative in the western region and chairman of Environmental Agency Abu Dhabi. Since its establishment, Emirates Wildlife Society has been working in association with WWF, one of the largest and Prepared by most respected independent global conservation organisations, to initiate and Patricia Cabrera, EWS-WWF implement environmental conservation and education projects in the region. Altaf Habib, EWS-WWF EWS-WWF has been active in the UAE since 2001, and its mission is to work with Anne V. Bourbon, EWS-WWF people and institutions within the UAE and the region to conserve biodiversity and tackle climate change through education, awareness, policy, and science-based Reviewed by conservation initiatives. -
The Terrestrial Mammals, Reptiles and Amphibians of the Uae – Species List and Status Report
THE TERRESTRIAL MAMMALS, REPTILES AND AMPHIBIANS OF THE UAE – SPECIES LIST AND STATUS REPORT January 2005 TERRESTRIAL ENVIRONMENT RESEARCH CENTRE ENVIRONMENTAL RESEARCH & WILDLIFE DEVELOPMENT AGENCY P.O. Box 45553 Abu Dhabi DOCUMENT ISSUE SHEET Project Number: 03-31-0001 Project Title: Abu Dhabi Baseline Survey Name Signature Date Drew, C.R. Al Dhaheri, S.S. Prepared by: Barcelo, I. Tourenq, C. Submitted by: Drew, C.R. Approved by: Newby, J. Authorized for Issue by: Issue Status: Final Recommended Circulation: Internal and external File Reference Number: 03-31-0001/WSM/TP007 Drew, C.R.// Al Dhaheri, S.S.// Barcelo, I.// Tourenq, C.//Al Team Members Hemeri, A.A. DOCUMENT REVISION SHEET Revision No. Date Affected Date of By pages Change V2.1 30/11/03 All 29/11/03 CRD020 V2.2 18/9/04 6 18/9/04 CRD020 V2.3 24/10/04 4 & 5 24/10/04 CRD020 V2.4 24/11/04 4, 7, 14 27/11/04 CRD020 V2.5 08/01/05 1,4,11,15,16 08/01/05 CJT207 Table of Contents Table of Contents ________________________________________________________________________________ 3 Part 1 The Mammals of The UAE____________________________________________________________________ 4 1. Carnivores (Order Carnivora) ______________________________________________________________ 5 a. Cats (Family Felidae)___________________________________________________________________ 5 b. Dogs (Family Canidae) __________________________________________________________________ 5 c. Hyaenas (Family Hyaenidae) _____________________________________________________________ 5 d. Weasels (Family Mustelidae) _____________________________________________________________ -
Felis Margarita, Sand Cat
The IUCN Red List of Threatened Species™ ISSN 2307-8235 (online) IUCN 2008: T8541A50651884 Felis margarita, Sand Cat Assessment by: Sliwa, A., Ghadirian, T., Appel, A., Banfield, L., Sher Shah, M. & Wacher, T. View on www.iucnredlist.org Citation: Sliwa, A., Ghadirian, T., Appel, A., Banfield, L., Sher Shah, M. & Wacher, T. 2016. Felis margarita. The IUCN Red List of Threatened Species 2016: e.T8541A50651884. http://dx.doi.org/10.2305/IUCN.UK.2016-2.RLTS.T8541A50651884.en Copyright: © 2016 International Union for Conservation of Nature and Natural Resources Reproduction of this publication for educational or other non-commercial purposes is authorized without prior written permission from the copyright holder provided the source is fully acknowledged. Reproduction of this publication for resale, reposting or other commercial purposes is prohibited without prior written permission from the copyright holder. For further details see Terms of Use. The IUCN Red List of Threatened Species™ is produced and managed by the IUCN Global Species Programme, the IUCN Species Survival Commission (SSC) and The IUCN Red List Partnership. The IUCN Red List Partners are: BirdLife International; Botanic Gardens Conservation International; Conservation International; Microsoft; NatureServe; Royal Botanic Gardens, Kew; Sapienza University of Rome; Texas A&M University; Wildscreen; and Zoological Society of London. If you see any errors or have any questions or suggestions on what is shown in this document, please provide us with feedback so that we can correct or extend the information provided. THE IUCN RED LIST OF THREATENED SPECIES™ Taxonomy Kingdom Phylum Class Order Family Animalia Chordata Mammalia Carnivora Felidae Taxon Name: Felis margarita Loche, 1858 Regional Assessments: • Mediterranean Common Name(s): • English: Sand Cat, Sand Dune Cat • French: Chat des sables • Spanish: Gato de las Arenas, Gato del Sahara Taxonomic Notes: Taxonomy is currently under review by the IUCN SSC Cat Specialist Group (2014). -
Contents: FRITZ, U.: Herpetology and Herpetological Type Specimens At
Contents: FRITZ, U.: Herpetology and herpetological type specimens at the Museum für Tierkunde Dresden with a bibliography of herpetological contributions by FRITZ JÜRGEN OBST (Amphibia, Reptilia) GÜNTHER, R.: Westernmost records of the Papuan frog genus Copiula with descriptions of two new species (Amphibia: Anura: Microhylidae) FRITZ, U., ZIEGLER, T., HERRMANN, H.-W. & E. LEHR: Intergradation between subspecies of Cuora galbinifrons BOURRET, 1939 and Pyxidea mouhotii (GRAY, 1862) in southern North Vietnam (Reptilia: Testudines: Geoemydidae) GUICKING, D., FRITZ, U., WINK, M. & E. LEHR: New data on the diversity of the Southeast Asian leaf turtle genus Cyclemys BELL, 1834. Molecular results (Reptilia: Testudines: Geoemydidae) PRASCHAG, P. & R. GEMEL: Identity of the black soft-shell turtle Aspideretes nigricans (ANDERSON, 1875), with remarks on related species (Reptilia: Testudines: Trionychidae) BÖHME, W. & W. WEITSCHAT: New finds of lizards in Baltic amber (Reptilia: Squamata: Sauria: Lacertidae) MORAVEC, J.: A new subspecies of the Sinai Agama Pseudotrapelus sinaitus from southern Syria and northern Jordan (Reptilia: Squamata: Sauria: Agamidae) SCHMIDTLER, J. F., HECKES, U., BISCHOFF, W. & M. FRANZEN: Altitude dependent character variation in rock lizards of the Darevskia clarkorum (DAREVSKY & VEDMEDERJA, 1977)/D. dryada (DAREVSKY & TUNIYEV, 1997) complex: a case of climate parallel variation of pholidosis? (Reptilia: Squamata: Sauria: Lacertidae) SEGOLI, M., COHEN, T. & Y. L. WERNER: A new lizard of the genus Mesalina from Mt. Sinai, Egypt (Reptilia: Squamata: Sauria: Lacertidae) ORLOV, N., ANANJEVA, N., BARABANOV, A., RYABOV, S. & R. KHALIKOV: Diversity of vipers (Azemiopinae, Crotalinae) in East, Southeast, and South Asia: Annotated checklist and natural history data (Reptilia: Squamata: Serpentes: Viperidae) ZIEGLER, T., UNGER, J., FEILER, A. -
A Phylogeny and Revised Classification of Squamata, Including 4161 Species of Lizards and Snakes
BMC Evolutionary Biology This Provisional PDF corresponds to the article as it appeared upon acceptance. Fully formatted PDF and full text (HTML) versions will be made available soon. A phylogeny and revised classification of Squamata, including 4161 species of lizards and snakes BMC Evolutionary Biology 2013, 13:93 doi:10.1186/1471-2148-13-93 Robert Alexander Pyron ([email protected]) Frank T Burbrink ([email protected]) John J Wiens ([email protected]) ISSN 1471-2148 Article type Research article Submission date 30 January 2013 Acceptance date 19 March 2013 Publication date 29 April 2013 Article URL http://www.biomedcentral.com/1471-2148/13/93 Like all articles in BMC journals, this peer-reviewed article can be downloaded, printed and distributed freely for any purposes (see copyright notice below). Articles in BMC journals are listed in PubMed and archived at PubMed Central. For information about publishing your research in BMC journals or any BioMed Central journal, go to http://www.biomedcentral.com/info/authors/ © 2013 Pyron et al. This is an open access article distributed under the terms of the Creative Commons Attribution License (http://creativecommons.org/licenses/by/2.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly cited. A phylogeny and revised classification of Squamata, including 4161 species of lizards and snakes Robert Alexander Pyron 1* * Corresponding author Email: [email protected] Frank T Burbrink 2,3 Email: [email protected] John J Wiens 4 Email: [email protected] 1 Department of Biological Sciences, The George Washington University, 2023 G St. -
Karyologische Untersuchungen an Der Omaneidechse Lacerta Jayakari Boulenger, 1887
© Biodiversity Heritage Library, http://www.biodiversitylibrary.org/; www.zoologicalbulletin.de; www.biologiezentrum.at Bonn. zool. Beitr. Bd. 42 H. 1 S. 67—73 Bonn, März 1991 Karyologische Untersuchungen an der Omaneidechse Lacerta jayakari Boulenger, 1887 Barbara Fritz, Wolfgang Bischoff & Johannes-Peter Fritz Abstract. The diploid chromosome complement of Lacerta jayakari consists of 38 chromosomes with 36 acrocentric macrochromosomes and 2 microchromosomes. The presence of female heterogamety (ZZ/ZW system), where W is a microchromosome, is reported. One of the small pairs (probably the 15th) are the NOR bearing macrochromoso- mes. The DNA content/cell is 3.03 pg in the female and 2.88 pg in the male. L. jayakari is not closely related to the genus Gallotia but to the modern majority of the Lacertidae. Key words. Sauria, Lacertidae, Lacerta jayakari, karyotype, NOR. Einleitung Die Omaneidechse Lacerta jayakari Boulenger, 1887 gehört zu den vielen Endemiten der Arabischen Halbinsel. Bis vor kurzem war über dieses Tier, von den morphologi- schen Angaben Boulengers (1887, 1920) abgesehen, kaum etwas bekannt (Bischoff & Schmidtler 1981). Dies hat sich in den letzten Jahren grundlegend geändert. Auf der Basis neuen Sammlungsmaterials wurden die morphologischen und anatomi- schen Merkmale genauer beschrieben (Arnold 1972, 1973, 1989). Auch wurden erste ökologische Daten (Arnold 1972, Arnold & Gallagher 1977) und Beschreibungen des Verhaltens und der Fortpflanzungsbiologie veröffentlicht (Bischoff 1981, 1987). Die systematische Stellung von L. Jayakari war lange Zeit unklar und wurde dem- zufolge unterschiedlich bewertet (vgl. den Überblick bei Lutz et al. 1986). Analysen des Paarungsverhaltens zeigten Parallelen zu den Kanareneidechsen der Gattung Gallotia (Böhme & Bischoff 1976, Bischoff 1981) und ließen eine sehr lange Isola- tionszeit vermuten.