Timing the Extant Avian Radiation: the Rise of Modern Birds, and The
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Comparative Anatomy of Mu50phagidae (Turacos)
Comparative Anatomy of MU50phagidae (Turacos) by Georgann B. Johnston Sacramento, CA Introduction proVides the red colored feathers in he 20 species which comprise species of the first two genera. Both the avian order Musophagidae' pigments contain copper and spectral (commonly called turacos) data demonstrates that the former is have a number of physical and likely an oxidized version of the latter. anatomical characteristics that set them swered and many generalizations sus (Dyck, 1992) In fact, the two pigments apaI1 from many other birds. While pect in light of new information about are intermingled within individual uniformity among the 20 species is not these species' ecology including feathers in the breast patches and crests complete, ceI1ain generalizations can behavior and diet. of some species and turacoverdin be made. One ofthese is that the sexes occurs only in the presence of turacin. are visually indistinguishable in all of Feathers Other species outside the the species save C. Jeucogaster, in Probably the most distinguishing fea Musophagidae order have turacoverdin which the males have a black beak ture of these birds are two unique pig pigment, including Ithaginis (pheasant) and the females a green beak. ments deposited in their feather keratin. and Rollolus (paI1ridge), both members Unfortunately, most of the literature One, turacoverdin, is a green pigment of the Galliformes. An additional inter regarding the anatomy of these birds found in the rami in all species of esting note is that both pigments are was developed more than 40 years Tauraco and Musophaga, and in soluble in a weak base - which may ago, leaving many questions unan- Corythaeola cristata. -
PTAČÍ FYLOGENEZE Dovnitř Patří Šplhavci
E Afroaves – řada linií této skupiny je velmi rozrůzněna v Africe a pro Afriku stepokur kropenatý typická (vlhy, zoborožci, mandelíci), ale fosilie africký původ nepodporují. Mnoho (Pterocles burchelli) linií bylo řazeno do řádu srostloprstých, ale ten není monofyletický, poněvadž PTAČÍ FYLOGENEZE dovnitř patří šplhavci. Patří sem i myšáci (Coliiformes) a trogoni. F Australaves – obdobně jako u Afroaves dnes sice řada linií žije jen FYLOGENEZE NA HRUBÉ ÚROVNI A PTAČÍ ŘÁDY v australasijské oblasti, ale fosilní zástupci byli nalezeni všude možně, takže o původu skupiny těžko něco soudit. Zobrazený fylogenetický stromeček zahrnuje jen velké skupiny, přičemž co je „velká“ skupina, je docela arbitrární. Aby vůbec bylo možné dělat O Passerimorphae (nebo Psittacopasserae) – papoušci korektní fylogenetické analýzy, nesmí být těch skupin moc a zároveň je potřeba, aby to byly skupiny, o nichž se předpokládá, že jsou a pěvci sdílejí řadu znaků, například schopnost učit se zpěvu, která byla monofyletické. Reálně se totiž sekvenují samozřejmě jednotliví zástupci (tedy jeden druh, respektive přímo jeden jedinec z dané skupiny), považovaná za klíčovou pro jejich sloučení (pokládala se za znak vzniklý jen a potřebujeme tedy, aby skutečně skupinu reprezentovali. Proto by nestačilo brát zástupce tradičních ptačích řádů, které známe z učebnic, jednou u jejich společného předka, tedy synapomorfii). Ovšem vzhledem k tomu, jelikož se ukázalo, že řada z nich monofyletických není. Tradiční „dravci“ byli například polyfyletičtí, poněvadž zahrnovali monofyletickou že u pěvců je s jistotou známa jen ve skupině Oscines (zpěvní) a navíc se vyskytuje skupinu složenou z orlů, jestřábů, káňat apod. (včetně kondorů), ale taky sokolovité (poštolky, sokoly apod.), kteří jsou ve skutečnosti blízce také u zcela nezávislých kolibříků, musela vzniknout víckrát konvergentně, a to příbuzní pěvcům a papouškům. -
The Tarsometatarsus of the Middle Eocene Loon Colymbiculus Udovichenkoi
– 17 – Paleornithological Research 2013 Proceed. 8th Inter nat. Meeting Society of Avian Paleontology and Evolution Ursula B. Göhlich & Andreas Kroh (Eds) The tarsometatarsus of the Middle Eocene loon Colymbiculus udovichenkoi GERALD MAYR1, LEONID GOROBETS2 & EVGENIJ ZVONOK3 1 Senckenberg Research Institute and Natural History Museum Frankfurt, Frankfurt am Main, Germany; E-mail: [email protected] 2 Taras Shevchenko National University of Kiev, Dept. of Ecology and Environmental Protection, Kiev, Ukraine 3 Institute of Geological Sciences of NAS of Ukraine, Branch of Paleontology and Stratigraphy, Kiev, Ukraine Abstract — We describe the previously unknown tarsometatarsus of the earliest unambiguously identified loon, Colymbiculus udovichenkoi, from the Middle Eocene of the Ukraine. Except for being more elongate and apart from details of the hypotarsus morphology, the bone resembles the tarsometatarsus of the Early Miocene Colym- boides minutus. We consider the hypotarsus morphology of Colymbiculus to be plesiomorphic for Gaviiformes. Colymboides and crown group Gaviiformes are each characterized by an autapomorphic hypotarsus morphology, which precludes the former from being directly ancestral to the latter. The similarities shared by C. udovichenkoi and C. minutus, including their small size, are likely to be plesiomorphic for Gaviiformes. Although the disap- pearance of small stem group Gaviiformes may be related to the retreat of loons to cold Northern latitudes, more data are needed to firmly establish this hypothesis. We finally note that early Paleogene stem group Gaviiformes markedly differ from putative Late Cretaceous loons, whose identification needs to be verified by further fossil specimens. Key words: Colymbiculus, Colymboides, fossil birds, Gaviiformes, Eocene, Lutetian, Ukraine Introduction 1982 from the Early Miocene of the Czech Republic (ŠVEC 1982), the earliest stem line- Loons (Gaviiformes) have a fairly comprehen- age representative, being distinctly smaller than sive Neogene fossil record (OLSON 1985; MAYR its modern congeners. -
Lowest Paleogene Strata on Vega Island, Antarctica
Palaeogeography, Palaeoclimatology, Palaeoecology 402 (2014) 55–72 Contents lists available at ScienceDirect Palaeogeography, Palaeoclimatology, Palaeoecology journal homepage: www.elsevier.com/locate/palaeo Stratigraphy and vertebrate paleoecology of Upper Cretaceous–?lowest Paleogene strata on Vega Island, Antarctica Eric M. Roberts a,⁎, Matthew C. Lamanna b, Julia A. Clarke c, Jin Meng d, Eric Gorscak e, Joseph J.W. Sertich f, Patrick M. O'Connor e, Kerin M. Claeson g,RossD.E.MacPheeh a School of Earth and Environmental Sciences, James Cook University, Townsville, QLD 4811, Australia b Section of Vertebrate Paleontology, Carnegie Museum of Natural History, 4400 Forbes Ave., Pittsburgh, PA 15213, USA c Department of Geological Sciences, Jackson School of Geosciences, University of Texas at Austin, 1 University Station C1100, Austin, TX 78712, USA d Department of Paleontology, American Museum of Natural History, Central Park West at 79th St., New York, NY 10024, USA e Department of Biomedical Sciences, Heritage College of Osteopathic Medicine, Ohio University, Athens, OH 45701, USA f Department of Earth Sciences, Denver Museum of Nature and Science, 2001 Colorado Blvd., Denver, CO 80205, USA g Department of Biomedical Sciences, Philadelphia College of Osteopathic Medicine, Philadelphia, PA 19131, USA h Department of Mammalogy, American Museum of Natural History, Central Park West at 79th St., New York, NY 10024, USA article info abstract Article history: The Upper Cretaceous (Maastrichtian) Sandwich Bluff Member of the López de Bertodano Formation is well Received 24 October 2013 exposed on Vega Island in the James Ross Basin off the northeastern coast of the Antarctic Peninsula. Although Received in revised form 18 February 2014 this unit is one of the richest sources of end-Cretaceous vertebrate fossils in Antarctica, it is also one of the Accepted 3 March 2014 least sedimentologically and stratigraphically characterized units in the basin. -
Dieter Thomas Tietze Editor How They Arise, Modify and Vanish
Fascinating Life Sciences Dieter Thomas Tietze Editor Bird Species How They Arise, Modify and Vanish Fascinating Life Sciences This interdisciplinary series brings together the most essential and captivating topics in the life sciences. They range from the plant sciences to zoology, from the microbiome to macrobiome, and from basic biology to biotechnology. The series not only highlights fascinating research; it also discusses major challenges associated with the life sciences and related disciplines and outlines future research directions. Individual volumes provide in-depth information, are richly illustrated with photographs, illustrations, and maps, and feature suggestions for further reading or glossaries where appropriate. Interested researchers in all areas of the life sciences, as well as biology enthusiasts, will find the series’ interdisciplinary focus and highly readable volumes especially appealing. More information about this series at http://www.springer.com/series/15408 Dieter Thomas Tietze Editor Bird Species How They Arise, Modify and Vanish Editor Dieter Thomas Tietze Natural History Museum Basel Basel, Switzerland ISSN 2509-6745 ISSN 2509-6753 (electronic) Fascinating Life Sciences ISBN 978-3-319-91688-0 ISBN 978-3-319-91689-7 (eBook) https://doi.org/10.1007/978-3-319-91689-7 Library of Congress Control Number: 2018948152 © The Editor(s) (if applicable) and The Author(s) 2018. This book is an open access publication. Open Access This book is licensed under the terms of the Creative Commons Attribution 4.0 International License (http://creativecommons.org/licenses/by/4.0/), which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license and indicate if changes were made. -
Onetouch 4.0 Scanned Documents
/ Chapter 2 THE FOSSIL RECORD OF BIRDS Storrs L. Olson Department of Vertebrate Zoology National Museum of Natural History Smithsonian Institution Washington, DC. I. Introduction 80 II. Archaeopteryx 85 III. Early Cretaceous Birds 87 IV. Hesperornithiformes 89 V. Ichthyornithiformes 91 VI. Other Mesozojc Birds 92 VII. Paleognathous Birds 96 A. The Problem of the Origins of Paleognathous Birds 96 B. The Fossil Record of Paleognathous Birds 104 VIII. The "Basal" Land Bird Assemblage 107 A. Opisthocomidae 109 B. Musophagidae 109 C. Cuculidae HO D. Falconidae HI E. Sagittariidae 112 F. Accipitridae 112 G. Pandionidae 114 H. Galliformes 114 1. Family Incertae Sedis Turnicidae 119 J. Columbiformes 119 K. Psittaciforines 120 L. Family Incertae Sedis Zygodactylidae 121 IX. The "Higher" Land Bird Assemblage 122 A. Coliiformes 124 B. Coraciiformes (Including Trogonidae and Galbulae) 124 C. Strigiformes 129 D. Caprimulgiformes 132 E. Apodiformes 134 F. Family Incertae Sedis Trochilidae 135 G. Order Incertae Sedis Bucerotiformes (Including Upupae) 136 H. Piciformes 138 I. Passeriformes 139 X. The Water Bird Assemblage 141 A. Gruiformes 142 B. Family Incertae Sedis Ardeidae 165 79 Avian Biology, Vol. Vlll ISBN 0-12-249408-3 80 STORES L. OLSON C. Family Incertae Sedis Podicipedidae 168 D. Charadriiformes 169 E. Anseriformes 186 F. Ciconiiformes 188 G. Pelecaniformes 192 H. Procellariiformes 208 I. Gaviiformes 212 J. Sphenisciformes 217 XI. Conclusion 217 References 218 I. Introduction Avian paleontology has long been a poor stepsister to its mammalian counterpart, a fact that may be attributed in some measure to an insufRcien- cy of qualified workers and to the absence in birds of heterodont teeth, on which the greater proportion of the fossil record of mammals is founded. -
An Avian Femur from the Late Cretaceous of Vega Island, Antarctic Peninsula: Removing the Record of Cursorial Landbirds from the Mesozoic of Antarctica
An avian femur from the Late Cretaceous of Vega Island, Antarctic Peninsula: removing the record of cursorial landbirds from the Mesozoic of Antarctica Abagael R. West1,2,3, Christopher R. Torres4, Judd A. Case5, Julia A. Clarke4,6, Patrick M. O’Connor7,8 and Matthew C. Lamanna1 1 Section of Vertebrate Paleontology, Carnegie Museum of Natural History, Pittsburgh, PA, USA 2 Section of Mammals, Carnegie Museum of Natural History, Pittsburgh, PA, USA 3 Department of Biological Sciences, University of Pittsburgh, Pittsburgh, PA, USA 4 Department of Integrative Biology, University of Texas at Austin, Austin, TX, USA 5 Department of Biology, Eastern Washington University, Cheney, WA, USA 6 Jackson School of Geosciences, University of Texas at Austin, Austin, TX, USA 7 Department of Biomedical Sciences, Ohio University Heritage College of Osteopathic Medicine, Athens, OH, USA 8 Ohio Center for Ecology and Evolutionary Studies, Ohio University, Athens, OH, USA ABSTRACT In 2006, a partial avian femur (South Dakota School of Mines and Technology (SDSM) 78247) from the Upper Cretaceous (Maastrichtian) Sandwich Bluff Member of the López de Bertodano Formation of Sandwich Bluff on Vega Island of the northern Antarctic Peninsula was briefly reported as that of a cariamiform—a clade that includes extant and volant South American species and many extinct flightless and cursorial species. Although other authors have since rejected this taxonomic assignment, SDSM 78247 had never been the subject of a detailed description, hindering a definitive assessment -
WORKING with TURACOS Musophagidae Spp
125 AL WABRA WILDLIFE PRESERVATION, QATAR: WORKING WITH TURACOS Musophagidae spp. by Simon Bruslund Jensen The turacos are a primitive, almost primeval-looking family of medium to large-sized birds, most closely related to the cuckoos. The 23 species are distributed exclusively in tropical African forests and savannahs. Most species are beautifully coloured in shades of green, blue and red, and are well known for their unique feather pigments, the so-called turacin and turacoverin, found only in this family. Sven Hammer/AWWP A Prince Ruspoli’s Turaco Tauraco ruspolii approaching a waterhole close to Arero, Ethiopia. Due to their appealing appearance, many of the species have for many years been popular in zoos and private collections. However, just a few years back, captive breeding would never have been considered as an option for the conservation management of turacos, as they had long been considered as delicate and rather difficult to breed in captivity. In recent years the methods used for keeping and breeding turacos have improved a great deal, due largely to the systematic approach by dedicated individuals and public collections such as the zoos in Cologne, Germany and Houston, Texas, USA. Now for the first time, it seems possible to establish self-sustainable captive populations, in particular within the 126 JENSEN - TURACOS framework of regional and international cooperation, with studbooks being maintained to ensure the necessary exchange of bloodlines. This is in line with the identification of more and more threats to several turaco species in the wild, especially to some of the unique subspecies with limited distribution. -
Modern Birds Classification System Tinamiformes
6.1.2011 Classification system • Subclass: Neornites (modern birds) – Superorder: Paleognathae, Neognathae Modern Birds • Paleognathae – two orders, 49 species • Struthioniformes—ostriches, emus, kiwis, and allies • Tinamiformes—tinamous Ing. Jakub Hlava Department of Zoology and Fisheries CULS Tinamiformes • flightless • Dwarf Tinamou • consists of about 47 species in 9 genera • Dwarf Tinamou ‐ 43 g (1.5 oz) and 20 cm (7.9 in) • Gray Tinamou ‐ 2.3 kg (5.1 lb) 53 cm (21 in) • small fruits and seeds, leaves, larvae, worms, and mollusks • Gray Tinamou 1 6.1.2011 Struthioniformes Struthioniformes • large, flightless birds • Ostrich • most of them now extinct • Cassowary • chicks • Emu • adults more omnivorous or insectivorous • • adults are primarily vegetarian (digestive tracts) Kiwi • Emus have a more omnivorous diet, including insects and other small animals • kiwis eat earthworms, insects, and other similar creatures Neognathae Galloanserae • comprises 27 orders • Anseriformes ‐ waterfowl (150) • 10,000 species • Galliformes ‐ wildfowl/landfowl (250+) • Superorder Galloanserae (fowl) • Superorder Neoaves (higher neognaths) 2 6.1.2011 Anseriformes (screamers) Anatidae (dablling ducks) • includes ducks, geese and swans • South America • cosmopolitan distribution • Small group • domestication • Large, bulky • hunted animals‐ food and recreation • Small head, large feet • biggest genus (40‐50sp.) ‐ Anas Anas shoveler • mallards (wild ducks) • pintails • shlhovelers • wigeons • teals northern pintail wigeon male (Eurasian) 3 6.1.2011 Tadorninae‐ -
Norntates PUBLISHED by the AMERICAN MUSEUM of NATURAL HISTORY CENTRAL PARK WEST at 79TH STREET, NEW YORK, NY 10024 Number 3265, 36 Pp., 15 Figures May 4, 1999
AMERICANt MUSEUM Norntates PUBLISHED BY THE AMERICAN MUSEUM OF NATURAL HISTORY CENTRAL PARK WEST AT 79TH STREET, NEW YORK, NY 10024 Number 3265, 36 pp., 15 figures May 4, 1999 An Oviraptorid Skeleton from the Late Cretaceous of Ukhaa Tolgod, Mongolia, Preserved in an Avianlike Brooding Position Over an Oviraptorid Nest JAMES M. CLARK,I MARK A. NORELL,2 AND LUIS M. CHIAPPE3 ABSTRACT The articulated postcranial skeleton of an ovi- presence of a single, ossified ventral segment in raptorid dinosaur (Theropoda, Coelurosauria) each rib as well as ossified uncinate processes from the Late Cretaceous Djadokhta Formation associated with the thoracic ribs. Remnants of of Ukhaa Tolgod, Mongolia, is preserved over- keratinous sheaths are preserved with four of the lying a nest. The eggs are similar in size, shape, manal claws, and the bony and keratinous claws and ornamentation to another egg from this lo- were as strongly curved as the manal claws of cality in which an oviraptorid embryo is pre- Archaeopteryx and the pedal claws of modern served, suggesting that the nest is of the same climbing birds. The skeleton is positioned over species as the adult skeleton overlying it and was the center of the nest, with its limbs arranged parented by the adult. The lack of a skull pre- symmetrically on either side and its arms spread cludes specific identification, but in several fea- out around the nest perimeter. This is one of four tures the specimen is more similar to Oviraptor known oviraptorid skeletons preserved on nests than to other oviraptorids. The ventral part of the of this type of egg, comprising 23.5% of the 17 thorax is exceptionally well preserved and pro- oviraptorid skeletons collected from the Dja- vides evidence for other avian features that were dokhta Formation before 1996. -
AOU Classification Committee – North and Middle America
AOU Classification Committee – North and Middle America Proposal Set 2016-C No. Page Title 01 02 Change the English name of Alauda arvensis to Eurasian Skylark 02 06 Recognize Lilian’s Meadowlark Sturnella lilianae as a separate species from S. magna 03 20 Change the English name of Euplectes franciscanus to Northern Red Bishop 04 25 Transfer Sandhill Crane Grus canadensis to Antigone 05 29 Add Rufous-necked Wood-Rail Aramides axillaris to the U.S. list 06 31 Revise our higher-level linear sequence as follows: (a) Move Strigiformes to precede Trogoniformes; (b) Move Accipitriformes to precede Strigiformes; (c) Move Gaviiformes to precede Procellariiformes; (d) Move Eurypygiformes and Phaethontiformes to precede Gaviiformes; (e) Reverse the linear sequence of Podicipediformes and Phoenicopteriformes; (f) Move Pterocliformes and Columbiformes to follow Podicipediformes; (g) Move Cuculiformes, Caprimulgiformes, and Apodiformes to follow Columbiformes; and (h) Move Charadriiformes and Gruiformes to precede Eurypygiformes 07 45 Transfer Neocrex to Mustelirallus 08 48 (a) Split Ardenna from Puffinus, and (b) Revise the linear sequence of species of Ardenna 09 51 Separate Cathartiformes from Accipitriformes 10 58 Recognize Colibri cyanotus as a separate species from C. thalassinus 11 61 Change the English name “Brush-Finch” to “Brushfinch” 12 62 Change the English name of Ramphastos ambiguus 13 63 Split Plain Wren Cantorchilus modestus into three species 14 71 Recognize the genus Cercomacroides (Thamnophilidae) 15 74 Split Oceanodroma cheimomnestes and O. socorroensis from Leach’s Storm- Petrel O. leucorhoa 2016-C-1 N&MA Classification Committee p. 453 Change the English name of Alauda arvensis to Eurasian Skylark There are a dizzying number of larks (Alaudidae) worldwide and a first-time visitor to Africa or Mongolia might confront 10 or more species across several genera. -
Leptosomiformes ~ Trogoniformes ~ Bucerotiformes ~ Piciformes
Birds of the World part 6 Afroaves The core landbirds originating in Africa TELLURAVES: AFROAVES – core landbirds originating in Africa (8 orders) • ORDER ACCIPITRIFORMES – hawks and allies (4 families, 265 species) – Family Cathartidae – New World vultures (7 species) – Family Sagittariidae – secretarybird (1 species) – Family Pandionidae – ospreys (2 species) – Family Accipitridae – kites, hawks, and eagles (255 species) • ORDER STRIGIFORMES – owls (2 families, 241 species) – Family Tytonidae – barn owls (19 species) – Family Strigidae – owls (222 species) • ORDER COLIIFORMES (1 family, 6 species) – Family Coliidae – mousebirds (6 species) • ORDER LEPTOSOMIFORMES (1 family, 1 species) – Family Leptosomidae – cuckoo-roller (1 species) • ORDER TROGONIFORMES (1 family, 43 species) – Family Trogonidae – trogons (43 species) • ORDER BUCEROTIFORMES – hornbills and hoopoes (4 families, 74 species) – Family Upupidae – hoopoes (4 species) – Family Phoeniculidae – wood hoopoes (9 species) – Family Bucorvidae – ground hornbills (2 species) – Family Bucerotidae – hornbills (59 species) • ORDER PICIFORMES – woodpeckers and allies (9 families, 443 species) – Family Galbulidae – jacamars (18 species) – Family Bucconidae – puffbirds (37 species) – Family Capitonidae – New World barbets (15 species) – Family Semnornithidae – toucan barbets (2 species) – Family Ramphastidae – toucans (46 species) – Family Megalaimidae – Asian barbets (32 species) – Family Lybiidae – African barbets (42 species) – Family Indicatoridae – honeyguides (17 species) – Family