Western North American Naturalist

Volume 62 Number 4 Article 1

10-28-2002

Stoneflies () of National Park,

B. C. Kondratieff Colorado State University, Fort Collins

Richard A. Lechleitner Mount Ranier National Park, Ashford, Washington

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Recommended Citation Kondratieff, B. C. and Lechleitner, Richard A. (2002) "Stoneflies (Plecoptera) of Mount Rainier National Park, Washington," Western North American Naturalist: Vol. 62 : No. 4 , Article 1. Available at: https://scholarsarchive.byu.edu/wnan/vol62/iss4/1

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STONEFLIES (PLECOPTERA) OF MOUNT RAINIER NATIONAL PARK, WASHINGTON

B.C. Kondratieff1 and Richard A. Lechleitner2

ABSTRACT.—Mount Rainier National Park, with an area of 95,356 ha, is approximately one-third as large as the state of Rhode Island. The lowest point is 490 m in elevation in the southeastern corner near where the Ohanapecosh crosses the southern boundary. Columbia Crest is the highest point at 4392 m. The entire park is a rugged landscape marked by the major topographical feature, Mount Rainier, comprising over 25,899 ha, almost one-third of the park. The park lies entirely west of the crest line of the . Most streams in the park originate on Mount Rainier; however, several large through the park near its boundaries. One of the first attempts to summarize the stoneflies of Washington, including Mount Rainier National Park, was Hoppe’s 1938 work that reported ca 8 species. Jewett (1959) reviewed the stoneflies of the Pacific Northwest and listed 7 species that had type localities in the park: kincaidi Frison, Doddsia occidentalis (Banks), Soliperla fenderi ( Jewett), Frisonia picticeps (Hanson), Isoperla rainiera Jewett, Megarcys irregularis (Banks), and M. subtruncata (Hanson). Subsequently, Kathroperla takhoma Stark and Surdick (1987) was described from the park. Samples of adult stoneflies from 1994 to 2001 indicate the presence of at least 82 species, with 64% of these typical Pacific Northwest species, and 30 species, or 36%, widespread western North American species. Seventeen new Washington state records are listed, including a substantial range extension for Lednia tumana (Ricker). One undescribed species in the Sweltsa borealis complex was also discovered. We also present illustrations of male terminalia for augusta (Banks) and infuscata (Claassen) to aid in the identification of these species.

Key words: stoneflies, Plecoptera, Mount Rainier National Park, Washington.

Mount Rainier National Park is in Lewis collapse, and erosion. It has been estimated and Pierce counties on the western slope of that the cone was once about 600 m higher than the Cascade Range of Washington State. Mount it is now. Even before Mount Rainier reached Rainier is located approximately 65–110 km its greatest height, rivers and glaciers were from the Seattle-Tacoma metropolitan area cutting deep valleys and huge, bowl-shaped known as (Fig. 1). The park area cirques into its sides. During the last major is 95,356 ha, and elevations range from 490 m glaciation, which ended about 10,000 years above sea level to 4392 m at the summit. ago, the valley glaciers grew to as much as 60 Mount Rainier, one of the highest and topo- km long, and smaller glaciers nestled in cirques graphically most impressive of the world’s above 1400 m. The peak has essentially re- volcanoes, owes its scenic beauty to many fea- mained the same in appearance since the last tures. The broad cone rises about 2300 m major glaciations. The last major eruption of above its 2100-m foundation and stands in the volcano occurred about 2000 years ago, solitary splendor as the highest peak in the although the last eruption was approximately Cascade Range. Its rocky ice/-mantled 150 years ago (Mullineaux 1974). slopes above timberline contrast with the dense Mount Rainier is situated within a temper- green forests, giving Mount Rainier the appear- ate, maritime climate. Several climatic zones ance of an arctic island in a temperate sea. exist elevationally and geographically around There are 26 major glaciers on Mount Rainier the park. However, the east-southeast side of and numerous unnamed snow or ice patches. the park is generally the driest, and the north- Mount Rainier was born during the Pleisto- west side is the wettest sector (especially dur- cene, and gradually the high main cone was ing spring and summer months). Annual precip- built up. As eruptions diminished, Mount itation is heavy, ranging from about 1.5 m at Rainier began to deteriorate by explosion, lowest elevations to over 2.5 m in the subalpine

1Department of Bioagricultural Sciences and Pest Management, Colorado State University, Fort Collins, CO 80523. 2Mount Rainier National Park, Tahoma Woods, Star Route, Ashford, WA 98304.

385 386 WESTERN NORTH AMERICAN NATURALIST [Volume 62

Fig. 1. Map of western Washington, indicating Mount Rainier National Park, Washington.

regions. Because southwesterly winds bear rivers are the exception, flowing into the Cow- much of the moisture, a rain shadow occurs on litz River, outside the park, and then draining the northeast side of the park. Over 90% of the into the and on to the Pacific precipitation occurs between November and Ocean. Each major river occupies a deep can- April. Much of the winter precipitation is snow yon, its floor 300–1000 m below the adjacent that accumulates into depths of 6 to 8 m at divides. Valley floor gradients are steep and in- higher elevations. At Paradise the average crease markedly upstream, especially in Tahoma annual snowfall is about 17 m, with over 26 m Creek, North and South Puyallup and Mowich of snow during the winter of 1998–99. Winter rivers. The park includes 470 mapped rivers temperatures are relatively warm (mean January and streams, some 400 mapped lakes and ponds, temperatures of about –4° to –1°C). Summers and over 1200 ha of other wetland types, in- tend to be cool (mean July temperatures of 10° cluding numerous thermal and mineral springs. to 20°C), and extended periods of cloudiness About 800 species of plants are known from are not uncommon. July and August are usu- the park (Franklin et al. 1988), reflecting the ally comparatively dry. Fog and high winds may varied climatic and environmental conditions be expected any day of the year. encountered across the 3800-m elevation gra- The topography of the park is rugged and dient. Approximately 58% of the park is covered precipitous, consisting mainly of peaks and by forests, mostly old-growth stands ranging valleys. Nine major rivers and their tributaries from 200 to 1000 years old. The subalpine drain the flanks of the mountain, with all but 2 parkland covers approximately 23% of the flowing into Puget Sound near Tacoma, Wash- park; vegetation in this zone is a mosaic of tree ington. The Muddy Fork and Ohanapecosh clumps and herbaceous meadows extending 2002] MOUNT RAINIER NATIONAL PARK STONEFLIES 387 from 1500 m to about 2200 m elevation. The Jewett (1959) in his review of the stoneflies alpine zone extends from treeline to the of the Pacific Northwest listed 7 species that mountain’s summit and covers approximately had type localities in the park: Megaleuctra 19% of the park. Permanent snow and ice cover kincaidi Frison, D. occidentalis, Soliperla fend- half of the alpine zone, with fragile alpine vege- eri ( Jewett), Frisonia picticeps, (Hanson), I. tation growing on rock and soil outcrops. rainiera Jewett, M. irregularis, and M. sub- According to historic records, coho (silver) truncata (Hanson). Szczytko and Stewart (1979) salmon, chinook, and steelhead were once pres- included a paratype female for I. bifurcata ent in many of the park’s major rivers before from Longmire; and Stark and Surdick (1987) the construction of dams outside the park. described Kathroperla takhoma from Falls Chinook, steelhead, and coho are now trans- Creek. Table 1 summarizes the primary type ported around some dams in an effort to re- localities in the park. Paradise Valley was rela- store these anadromous fish runs, thereby tively easy to access in the early history of the allowing access to the headwaters in the park. park, and a resort was established there. These fish species are thought to occur in the Our study had 2 main objectives: to docu- White and Carbon rivers inside the park. Bull ment the general distribution of stoneflies of trout are present throughout the park. In addi- Mount Rainier National Park, and to provide information on endemic, rare, or potentially tion to native fish species, nonnative and threatened species. Stoneflies are known to be hatchery fish have been introduced to lakes major indicators of water quality (Baumann and streams in the park. Nonnative and hatch- 1979) and are often dominant food-web com- ery fish stocks have probably altered natural ponents of most temperate lotic ecosystems lake and stream ecosystems. (Stewart and Stark 1988). Knowledge of the One of the first attempts to summarize the occurrence and distribution of these stoneflies for the state of Washington, including will help serve as a basis for future biomoni- Mount Rainier National Park, was the work of toring programs as dramatic land-use changes Hoppe (1938). She reported at least 8 species occur in adjacent areas. from the park: brevis (Banks) (as Pel- toperia [sic] brevis), Megarcys irregularis (Banks) METHODS (as Perlodes irregularis), Claassenia sabulosa (Banks) (as Perla languida Needham and Claas- We sampled over 110 sites between 1994 sen), Isoperla sordida Banks, I. fulva Claassen and 2001 (Fig. 2). These sites were primarily (as I. cascadensis Hoppe), Ostrocerca dimicki (Claassen) (as Nemoura dimicki), Zapada cinc- tipes (Banks) (as Nemoura cinctipes), and TABLE 1. Primary types originally described from Mount Doddsia occidentalis (Banks) (as Taeniopteryx Rainier National Park, Washington. occidentalis). Stark and Nelson (1994) did not report Y. brevis from Washington, and Hoppe’s Megaleuctra kincaidi Frison 1942. Type locality: Frying- records probably refer to Y. nigrisoma (Banks) pan Creek TAENIOPTERYGIDAE or Y. mariana (Ricker). Hoppe’s record may be Doddsia occidentalis (Banks) 1900. Type locality: Mount the one originally cited by Needham and Rainier National Park Claassen (1925). Ostrocerca dimicki was not collected from the park during the present Soliperla fenderi ( Jewett) 1955. Type locality: St. study, and the reported specimens were not Andrews Creek located. Her record may refer to O. foersteri Kathroperla takhoma Stark and Surdick 1987. Type (Ricker); however, both species are known locality: Falls Creek from Washington. These small nemourids are PERLODIDAE often difficult to specifically identify (Young et Frisonia picticeps (Hanson) 1942. Type locality: Paradise al. 1989). Additionally, we have been unable to River Isoperla rainiera Jewett 1954. Type locality: Mount confirm the presence of the widespread and Rainier National Park common I. fulva from the park. Hoppe (1938) Megarcys irregularis (Banks) 1900. Type locality: Mount listed it as I. cascadensis from the Nisqually Rainier National Park River, a stream draining the park. Szczytko and Megarcys subtruncata (Hanson) 1942. Type locality: Paradise Valley Stewart (1979) examined the female specimen. 388 WESTERN NORTH AMERICAN NATURALIST [Volume 62

Fig. 2. Map of Mount Rainier National Park, Washington, indicating major streams and roads. Major collecting sites of stoneflies during this study are indicated by ▲.

chosen to cover all relatively accessible major typical Pacific Northwest species (Ricker 1943, drainages. Large areas of the park (Fig. 2) are Jewett 1959, Ricker and Scudder 1975), and roadless or difficult to access due to extreme 30 species, or 36%, are widespread western topography. All adult and larval specimens North American species (Table 3; Baumann were preserved in 80% ethanol. We employed et al. 1977). One undescribed species in the standard collection techniques, including beat- Sweltsa borealis complex was also discovered ing sheets, aerial nets, and collection by hand. and will be described later. Specimens collected during the study were Seventeen new stonefly state records for deposited in the C.P. Gillette Museum of Washington were collected during the study Diversity, Colorado State Univer- (Table 2; see Nelson and Baumann 1989, Stark sity. We also present illustrations of the male 1998). Sixteen of these species are known from terminalia for 2 common leuctrid species, Des- adjacent states or British Columbia (Ricker and paxia augusta (Banks) (Figs. 3, 4) and Moselia Scudder 1975, Stark 1998) and were expected infuscata (Claassen) (Figs. 5, 6) to aid in their records. However, Lednia tumana (Ricker) rep- identification. resents a major range extension. The remarkable Nearctic nemourid genus Lednia is currently RESULTS AND DISCUSSION known only in the literature from Glacier National Park, Montana (Stewart and Stark We collected over 3600 adult specimens 1988). R.W. Baumann (personal communica- representing at least 82 species of stoneflies in tion) indicates that specimens of this genus are the park during the study; they are listed in also known from North Cascades National Table 2. Sixty-four percent of these taxa are Park, Washington, and a site in the California 2002] MOUNT RAINIER NATIONAL PARK STONEFLIES 389

Figs. 3–4. (Banks), male terminalia: 4, lateral; 5, dorsal.

Sierra Nevada. There may be additional relict outside localities is M. yosemite (Needham and populations of Lednia along the Coast and Claassen). This species was originally described Cascade mountains of the Pacific Northwest. from Mt. Lyell, Yosemite National Park, Cali- Jewett (1955) described S. fenderi from a fornia, and was collected during this study from single male collected from St. Andrews Creek Fryingpan Creek (Van Wieren et al. 2001). in the park. Stark (1983) reported this species Additionally, M. irregularis, known only from only from Mount Rainier, mostly from the St. Washington and British Columbia, occurs Andrews Creek area (B.P. Stark, personal com- abundantly in the park. Ricker (1943) once munication, has specimens from Mount Adams, described a flight of adults at Fryingpan Creek Shamama Co.). This species can be collected as “a small canyon filled with flying adults to a from spring seeps and rheocrenes throughout depth of possibly 100 feet, resembling the the park (Appendix). Isoperla rainiera was originally described swarms of large termites. . . .” by Jewett (1954) from the park from a single The nymphs of Setvena tibialis (Banks) are male. The female was described later from Mt. the dominant stonefly predators of most rheo- Hood, Oregon ( Jewett 1962). Szczytko and crenes and small streams of the park. Stewart Stewart (1979) did not report any additional and Stanger (1985) previously reported this specimens from the park. Recently, I. rainiera species from the park. Jewett (1959) considered was collected in Montana (S.W. Szczytko per- S. tibialis a rare species of the Pacific Northwest, sonal communication). Another rare species of and collections from the park represent sub- stonefly known from the park and only a few stantial new material for study. 390 WESTERN NORTH AMERICAN NATURALIST [Volume 62

Figs. 5–6. (Claassen), male terminalia: 6, lateral; 7, dorsal.

Despaxia augusta was found to be an abun- in most small streams of the park. Adult emer- dant autumn- and early winter-emerging spe- gence is generally from April to July. Illustra- cies in the park, occurring in small streams tions of the distinctive male terminalia are and rheocrenes. This species ranges from presented in Figures 5 and 6. Alaska to Montana, southwest to northern Cal- No major distributional patterns for the 82 ifornia. Ricker (1954) clarified the of stonefly species were discernible for the park. this species. Baumann et al. (1977) indicated Of the species with an adequate number of adults of D. augusta are uncommonly collected, records, a few species such as Mesocapnia probably because of their late emergence. Fig- oenone, Malenka californica, Doroneuria bau- ures 3 and 4 present illustrations of the male manni, and Kogotus nonus appear restricted to terminalia. drainages on the west side of the park (Appen- The other abundant leuctrid in the park is dix). Most other species are generally distrib- Moselia infuscata. However, very little biologi- uted throughout the park. cal information is available for this species Despite 7 years of relatively intensive col- (Stewart and Stark 1988). Nymphs are common lecting, at least 11 species are represented only 2002] MOUNT RAINIER NATIONAL PARK STONEFLIES 391

TABLE 2. Stonefly species recorded from Mount Rainier National Park, Washington. An asterisk (*) indicates a new state record for Washington (see Stark [1998] and Nelson and Baumann [1989]). Zapada cinctipes (Banks) is not indi- cated from Washington in Stark (1998); however, the type locality of this species is Olympia, Washington. EUHOLOGNATHA SYSTELLOGNATHA CHLOROPERLIDAE Bolshecapnia sasquatchi (Ricker)* Chloroperlinae Capnia confusa Claassen Alloperla fraterna Frison C. elongata Claassen A. serrata Needham & Claassen C. excavata Claassen Plumiperla diversa (Frison) C. gracilaria Claassen Suwallia dubia (Frison) C. licina Jewett S. forcipata (Neave) C. melia Frison S. pallidula (Banks) C. nana Claassen Sweltsa borealis (Banks) C. sextuberculata Jewett S. exquisita (Frison) Eucapnopsis brevicauda Claassen* S. occidens (Frison) Isocapnia agassizi Ricker S. revelstoka ( Jewett) I. grandis (Banks)* S. n. sp. I. spenceri Ricker* Paraperlinae I. vedderensis (Ricker)* Kathroperla perdita (Banks) Mesocapnia oenone (Neave) K. takhoma Stark & Surdick Paracapnia oswegaptera ( Jewett) Paraperla frontalis (Banks) P. wilsoni Ricker LEUCTRIDAE Leuctrinae PELTOPERLIDAE Despaxia augusta (Banks) Soliperla fenderi ( Jewett) Moselia infuscata (Claassen) Yoraperla mariana (Ricker) forcipata (Frison)* Y. nigrisoma (Banks) P. occidentalis (Banks)* Y. siletz Stark & Nelson P. projecta (Frison) PERLIDAE P. vershina Gaufin & Ricker Doroneuria baumanni Stark & Gaufin Perlomyia collaris Banks* Hesperoperla pacifica (Banks) P. utahensis Needham & Claassen* Claassenia sabulosa (Banks) Megaleuctrinae Megaleuctra kincaidi Frison PERLODIDAE Isoperlinae Isoperla bifurcata Szczytko & Stewart Amphinemurinae I. fusca Needham & Claassen Malenka californica (Claassen)* I. gravitans (Needham & Claassen) M. cornuta (Claassen) I. petersoni Needham & Christenson* I. rainiera Jewett Lednia tumana (Ricker)* I. sobria (Hagen) Ostrocerca foersteri (Ricker) I. sordida Banks Podmosta decepta (Frison)* I. tilasqua Szczytko & Stewart* P. delicatula (Claassen)* Perlodinae Prostoia besametsa (Ricker)* Arcynopterygini Soyedina interrupta (Claassen) Frisonia picticeps (Hanson) S. producta (Claassen) Megarcys irregularis (Banks) Visoka cataractae (Neave) M. subtruncata Hanson Zapada cinctipes (Banks) M. yosemite (Needham & Claassen) Z. columbiana (Claassen) Setvena tibialis (Banks) Z. cordillera (Baumann & Gaufin) Skwala americana (Klapalek) Z. frigida (Claassen) Diploperlini Z. haysi (Ricker) Kogotus nonus (Needham & Claassen) Z. oregonensis (Claassen)* PTERONARCYIDAE TAENIOPTERYGIDAE Pteronarcys princeps Banks Brachypteryinae Doddsia occidentalis (Banks) Taenionema kincaidi (Hoppe) T. pallidum (Banks) 392 WESTERN NORTH AMERICAN NATURALIST [Volume 62

Fig. 7. Seasonal distribution of adult stoneflies occurring at Mount Rainier National Park, Washington. 2002] MOUNT RAINIER NATIONAL PARK STONEFLIES 393

TABLE 3. Stonefly species occurring at Mount Rainier National Park that are considered widespread western North American species. Capnia confusa Claassen Z. columbiana (Claassen) C. gracilaria Claassen Z. frigida (Claassen) C. nana Claassen Z. haysi (Ricker) Eucapnopsis brevicauda Claassen Z. oregonensis (Claassen) Isocapnia grandis (Banks) Doddsia occidentalis (Banks) Paraleuctra occidentalis (Banks) Sweltsa borealis (Banks) P. projecta (Frison) S. revelstoka ( Jewett) P. vershina Gaufin & Ricker Suwallia pallidula (Banks) Perlomyia utahensis Needham & Claassen Kathroperla perdita Banks Malenka californica (Claassen) Paraperla frontalis (Banks) Podmosta decepta (Frison) Isoperla petersoni Needham & Christenson P. delicatula (Claassen) I. sobria (Hagen) Prostoia besametsa (Ricker) Skwala americana (Klapalek) Taenionema pallidum (Banks) Hesperoperla pacifica (Banks) Zapada cinctipes (Banks Claassenia sabulosa (Banks)

by single collections (Appendix). Included are Dr. Edward A. Lisowski, Yakima, Washington, Capnia excavata, Paracapnia oswegaptera, Per- and Robert Dopiriak, Mount Rainier National lomyia utahensis, L. tumana, Hesperoperla paci- Park, for making valuable collections of stone- fica, Claassenia sabulosa, I. bifurcata, I. fusca, flies from Mount Rainier available for study. I. petersoni, I. sobria, and Frisonia picticeps. Other populations of these species surely occur LITERATURE CITED in the park, yet to be discovered. The emergence phenology of comparable BAUMANN, R.W. 1979. Nearctic stonefly genera as indica- tors of ecological parameters (Plecoptera: Insecta). species of stoneflies of Mount Rainier National Great Basin Naturalist 39:241–244. Park is very similar to what Jewett (1959), BAUMANN, R.W., A.R. GAUFIN, AND R.F. SURDICK. 1977. Ricker (1943), and Kerst and Anderson (1974) The stoneflies (Plecoptera) of the Rocky Mountains. reported from streams of southwestern British Memoirs of the American Entomological Society 31. 208 pp. Columbia and Oregon (Fig. 7). For many FRANKLIN, J.F., W.H. MOIR, M.A. HEMSTROM, S.E. GREENE, species, adults can be collected for long peri- AND B.G. SMITH. 1988. The forest communities of ods. Generally, there are winter-emerging Mount Rainier National Park. Scientific Monograph species, including the capniids and 2 species Series 19. U.S. Department of Interior, Washington, of Zapada, Z. cinctipes and Z. columbiana DC. 194 pp. HOPPE, G.N. 1938. Plecoptera of Washington. University (Fig. 7). Adults of numerous species can be of Washington Publication in Biology 4:139–174. collected beginning in March and April, with JEWETT, S.G. 1954. New stoneflies (Plecoptera) from west- some of these species, for example Taenionema ern North America. Journal of the Fisheries Research Board of Canada 11:543–549. kincaidi, still present in August. The largest ______. 1955. Notes and descriptions concerning western group of species begins to emerge throughout North American stoneflies (Plecoptera). Wasmann May and June (Fig. 7). There is also a rather Journal of Biology 13:145–153. distinct late summer- and fall-emerging group ______. 1959. The stoneflies (Plecoptera) of the Pacific of species, typified by M. oenone, D. augusta, Northwest. Oregon State Monographs, Studies in Entomology 3:1–95. Alloperla serrata, and K. nonus. ______. 1962. New stoneflies and records from the Pacific coast of the . Pan-Pacific Entomologist ACKNOWLEDGMENTS 38:15–20. KERST, C.D., AND N.H. ANDERSON. 1974. Emergence pat- terns of Plecoptera in a stream in Oregon, USA. We thank Drs. Richard W. Baumann, Brig- Freshwater Biology 4:205–212. ham Young University, Stanley W. Szczytko, MULLINEAUX, D.R. 1974. Pumice and other pyroclastic University of Wisconsin, and Bill P. Stark, deposits in Mount Rainier National Park, Washing- Mississippi College, for verifying species ton. U.S. Geological Survey Bulletin 1326. 83 pp. identifications and providing records. Drs. NEEDHAM, J.G., AND P. W. C LAASSEN. 1925. A monograph of the Plecoptera or stoneflies of America north of Baumann and Stark also provided prepublica- Mexico. Thomas Say Foundation, Entomological tion reviews of the manuscript. We also thank Society of America 2. 397 pp. 394 WESTERN NORTH AMERICAN NATURALIST [Volume 62

NELSON, C.R., AND R.W. BAUMANN. 1989. Systematics and APPENDIX. Localities for collections records of distribution of the winter stonefly genus Capnia (Ple- Plecoptera of Mount Rainier National Park, Wash- coptera: Capniidae) in North America. Great Basin ington. Naturalist 49:289–363. RICKER, W.E. 1943. Stoneflies of southwestern British Order Plecoptera Columbia. Indiana University Publications in Science, Suborder Series 12. 145 pp. Group Euholognatha ______. 1954. Nomenclatorial notes on Plecoptera. Pro- Family Capniidae ceedings of the Entomological Society of British Genus Capnia Columbia 51:37–39. RICKER, W.E., AND G.G.E. SCUDDER. 1975. An annotated Bolshecapnia sasquatchi checklist of the Plecoptera (Insecta) of British Colum- bia. Syesis 8:333–348. , Hwy 143, 17-March-70, R.A. Hite & D.S. Potter, 2 males (BYU); Ohanapecosh River, Hwy 143, STARK, B.P. 1983. A review of the genus Soliperla (Ple- coptera: Peltoperlidae). Great Basin Naturalist 43: 16-March-73, D.S. Potter & L.M. Preble, 1 male (BYU). 30–44. ______. 1998. North American stonefly list. www.mc.edu/ Capnia confusa ~stark/stonefly.html 0.6 mile S of Ipsut Campground, 04-Apr-97, 1 female; STARK, B.P., AND C.R. NELSON. 1994. Systematics, phy- White River Highway 410, 29-May-97, 2 males, 1 female; logeny and zoogeography of genus Yoraperla (Ple- Tahoma Creek, 30-May-97, 1 male; Tahoma Creek at coptera: Peltoperlidae). Entomologica Scandinavica Nisqually to Paradise Road, 30-May-97, 2 males; stream 25:241–273. by South Camp, 04-May-98, 2 males, 1 STARK, B.P., AND R.F. SURDICK. 1987. A new Kathroperla female; entrance, 15-Aug-99, 1 female; Fry- species from western North America (Plecoptera: ingpan Creek at Sunrise Road Bridge, 16-Aug-99, 3 Chloroperlidae). Proceedings of the Entomological females; Tahoma Creek at Fish Creek, 08-Jul-00, 1 female; Society of Washington 89:527–531. Cataract Creek at , 09-Jul-00, 1 male, 9 STEWART, K.W., AND J.A. STANGER. 1985. The nymphs, and females; St. Andrews Creek at Westside Road, 14-Jul-00, a new species, of North American Setvena Illies 1 male, 2 females; Fryingpan Creek at Sunrise Road, 16- (Plecoptera: Perlodidae). Pan-Pacific Entomologist Jul-00, 1 female; Tahoma Creek, 13-May-01, 1 female; 61:237–244. seeps into White River on Highway 410, 14-May-01, 3 STEWART, K.W., AND B.P. STARK. 1988. Nymphs of North males, 4 females; Longmire, 15-May-01, 1 female; American stonefly genera (Plecoptera). Thomas Say Ohanapecosh Campground, 16-May-01, 1 female; White Foundation, Entomological Society of America River Highway 410, 0.5 mile S of park boundary, 16-May- 12:1–460. 01, 3 males, 4 females. SZCZYTKO, S.W., AND K.W. STEWART. 1979. The genus Isoperla (Plecoptera) of western North America: Capnia elongata holomorphology and systematics, and a new stonefly 0.7 mile SE of Ipsut Campground, 25-Mar-95, 1 male; genus Cascadoperla. Memoirs of the American Paradise River in Paradise Valley, 02-Aug-00, 2 females; Entomological Society 32. 120 pp. Ohanapecosh River at the Grove of the Patriarchs, 16-May- VANWIEREN, B.J., B.C. KONDRATIEFF, AND B.P. STARK. 2001. 01, 1 male; White River Highway 410, 0.5 mile S of park A review of the North American species of Megarcys boundary, 16-May-01, 2 males. (Plecoptera: Perlodidae). Proceedings of the Ento- mological Society of Washington 103:409–427. Capnia excavata YOUNG, D.C., B.C. KONDRATIEFF, AND R.F. KIRCHNER. 1989. Description of male Ostrocerca Ricker (Plecoptera: Boggy stream tributary to Tahoma Creek, 07-May-01, 1 Nemouridae) using the scanning electron micro- male, 1 female. scope. Proceedings of the Entomological Society of Washington 91:257–268. Capnia gracilaria Paradise River at Fourth Crossing, 21-Apr-96, 1 male; Received 1 May 2001 south of Chenuis Falls parking lot, 23-Feb-97, 1 male; Accepted 15 September 2001 Westside Road at Paradise Road, 29-Mar-97, 1 male, 1 female; Kautz Creek at Nisqually to Paradise Road, 04- Apr-01, 18 males, 6 females.

Capnia licina at creek from Tato Falls, 14-Apr-94, 2 males, 1 female; 2.2 miles SE Ipsut Campground, 25-Mar-95, 2 males; Nahunta Falls, 16-Apr-95, 3 males; Glacier View Bridge, 16-Apr-95, 2 males, 1 female; 0.1 mile NE Glacier View Bridge, 14-Apr-96, 5 males; Snow Lake inflow, 15- Jun-97, 7 males, 2 females; Shaw Creek at White River Road, 07-Jul-99, 2 males; Paradise River in Paradise Val- ley, 10-Jul-00, 4 females; Paradise River in Paradise Valley, 02-Jul-01, 4 males, 3 females. 2002] MOUNT RAINIER NATIONAL PARK STONEFLIES 395

Capnia melia Canyon Road, 05-Jun-99, 3 females; Falls Creek at Stevens Canyon Road, 11-Jun-99, 2 males, 1 female; trib- Ipsut Falls, 23-Feb-95, 54 males, 19 females; 2.2 miles SE utary to Tahoma Creek at Westside Road, 08-Jul-99, 1 of Ipsut Campground, 25-Mar-95, 1 male; 0.7 mile SE of female; Paradise River in Paradise Valley, 16-Aug-99, 1 Ipsut Campground, 25-Mar-95, 1 male, 4 females; Nahunta male, 1 female; Paradise River above , 16- Falls, 16-Apr-95, 3 males; Nisqually River at creek from Aug-99, 1 female; Falls Creek at Stevens Canyon Road, Tato Falls, 18-Feb-96, 2 males; Pinnacle Peak Trail stream, 25-Jun-00, 4 females; Ohanapecosh River at the Grove of 14-Apr-96, 5 males, 3 females; Nisqually River at creek the Patriarchs, 16-May-01, 2 females; White River High- from Tato Falls, 14-Apr-96, 2 males; south of Chenuis Falls way 410, 0.5 mile S of park boundary, 16-May-01, 4 males, parking lot, 23-Feb-97, 4 males, 1 female; 0.7 mile SE of 3 females; Falls Creek at Stevens Canyon Road, 07-Jun- Ipsut Campground, 23-Feb-97, 4 males, 1 female; 0.6 mile 01, 1 male, 2 females; Panther Creek at Highway 123, 09- S of Ipsut Campground, 23-Feb-97, 6 males; Westside Jun-01, 1 male, 1 female; Deer Creek at Eastside Trail, Road 1.3 miles N of Paradise Road, 29-Mar-97, 2 males, 6 09-Jun-01, 2 males, 1 female; Ohanapecosh River 0.5 mile females; Tahoma Creek at Nisqually to Paradise Road, 29- S of Shriner Peak Trailhead, 09-Jun-01, 1 male, 2 females; Mar-97, 1 male; 0.6 mile S of Ipsut Campground, 04-Apr- Paradise River above Narada Falls, 02-Jul-01, 1 male, 1 97, 2 males, 2 females; 0.7 mile SE of Ipsut Campground, female. 04-Apr-97, 5 males, 9 females; Ipsut Falls on Ipsut Creek, 04-May-97, 9 males, 10 females; Nickel Creek at Stevens Genus Isocapnia Canyon Road, 28-May-97, 1 male; Crystal Creek at High- way 410, 29-May-97, 6 males, 5 females; unnamed stream Isocapnia abbreviata N of Crystal Creek, Highway 410, 29-May-97, 2 males, 11 Seeps into White River on Highway 410, 14-May-01, 1 females; seeps into White River on Highway 410, 29- male, 3 females; White River Highway 410, 0.5 mile S of May-97, 2 males; Glacier View Bridge, 30-May-97, 4 park boundary, 16-May-01, 2 males, 6 females. females; Eagle Peak Creek at Nisqually River, 30-May-97, 3 males, 3 females; Eagle Peak Creek 4360 ft, 14-Jun-97, Isocapnia agassizi 3 males; Snow Lake inflow, 15-Jun-97, 1 male; Olallie Creek, 28-Jun-97, 2 males; Fryingpan Creek at Sunrise White River on Highway 410, 29-May-97, 1 male; Tahoma Road Bridge, 01-Apr-98, 1 male, 1 female; White River Creek at Nisqually to Paradise Road, 30-May-97, 1 Campground, 01-Apr-98, 1 male, 1 female; St. Andrews female; seeps into White River on Highway 410, 14-May- Creek at Westside Road, 29-Apr-98, 2 males, 6 females; 01, 2 males, 1 female; White River Highway 410, 0.5 mile Mountain Meadow Stream at Paul Peak Trail, 07-Jul-98, 1 S of park boundary, 16-May-01, 1 male, 3 females. male; Falls Creek at Stevens Canyon Road, 05-Jun-99, 4 males, 1 female; Falls Creek at Stevens Canyon Road, 11- Isocapnia grandis Jun-99, 3 males; Fryingpan Creek at Sunrise Road Bridge, White River on Highway 410, 28-May-97, 1 female; Kautz 06-Jul-99, 4 males; South at Westside Creek at Nisqually to Paradise Road, 04-Apr-01, 1 male; Road, 24-Sep-00, 1 female; Eagle Peak Creek at Nisqually seeps into White River on Highway 410, 14-May-01, 6 River, 13-May-01, 5 males, 9 females; Ipsut Falls at Ipsut males, 2 females; White River Highway 410, 0.5 mile S of Creek, 14-May-01, 1 male; Falls Creek at Stevens Canyon park boundary, 16-May-01, 2 males, 1 female. Road, 07-Jun-01, 1 female; White River at Sunrise Road, 15-Jun-01, 1 male; seeps into Fryingpan Creek 0.5 mile Isocapnia spenceri upstream from Sunrise Road, 15-Jun-01, 1 male, 2 females. Ohanapecosh Campground, 28-May-97, 1 female; seeps into White River on Highway 410, 14-May-01, 1 male, 1 Capnia nana female. Tahoma Creek at Nisqually to Paradise Road, 29-Mar-97, 1 female; Glacier View Bridge, 30-May-97, 3 females; Genus Mesocapnia Longmire, 15-May-01, 1 female. Mesocapnia oenone Capnia sextuberculata Carbon River Entrance, 20-Sep-97, 8 males, 3 females; Carbon River Entrance, 15-Aug-99, 1 male; west fork of Deer Creek at Eastside Trail, 09-Jun-01, 1 female; Par- White River, 13-Sep-99, 1 female. adise River above Narada Falls, 02-Jul-01, 1 female. Genus Paracapnia Genus Eucapnopsis Paracapnia oswegaptera Eucapnopsis brevicauda Seeps into Eagle Peak Creek, 28-May-97, 1 male. Laughingwater Creek, 27-Jun-95, 1 female; Nisqually River at Longmire, 28-May-97, 2 males; Ohanapecosh Family Leuctridae Hot Springs, 28-May-97, 1 male; Ohanapecosh Camp- Genus Despaxia ground, 28-May-97, 6 males, 5 females; Nickel Creek at Stevens Canyon Road, 28-May-97, 19 males, 28 females; Despaxia augusta Falls Creek at Stevens Canyon Road, 28-May-97, 11 Westside Road, 0.2 mile NE of Paradise Road, 18-Dec-94, males, 10 females; Ohanapecosh River at the Grove of the 1 male; Pinnacle Peak trail near road (Reflection Lakes), Patriarchs, 28-May-97, 15 males, 4 females; Ohanapecosh 28-Oct-95, 1 female; Paradise Valley Road 0.7 mile S of River at Sheep Creek, 29-May-97, 13 males, 7 females; Paradise, 28-Oct-95, 3 males, 6 females; 0.1 mile S of Panther Creek at Highway 123, 29-May-97, 22 males, 17 , 24-Nov-95, 1 female; Fish Creek near females; Olallie Creek, 28-Jun-97, 2 females; Huckleberry Tahoma Creek, 19-Dec-95, 1 female; 0.1 mile N of Frying- Creek Bridge, 28-Apr-98, 1 male; Falls Creek at Stevens pan Creek Bridge, 03-Nov-96, 1 male, 3 females; 0.2 mile 396 WESTERN NORTH AMERICAN NATURALIST [Volume 62

NE of Fryingpan Creek bridge, 03-Nov-96, 1 female; 0.1 females; Seep into Fryingpan Creek, 08-Jul-99, 3 males, mile E of Ranger Creek Bridge, 17-Nov-96, 1 female; Kautz 10 females; small tributary on Westside Road 2 miles N of Creek Trail 0.7 mile N of Nisqually Road, 08-Dec-96, 2 Paradise Road, 08-Jul-99, 1 female; tributary to Tahoma females; Ipsut Creek at Carbon River Road Bridge, 08- Creek at Paradise Road, 08-Jul-99, 1 female; high-gradi- Sep-97, 2 males; Carbon River Entrance, 20-Sep-97, 8 ent tributary at Nisqually River at Longmire, 09-Jul-99, 1 males, 3 females; small tributary to Shaw Creek, 07-Jul- male, 1 female; Paradise River in Paradise Valley, 16-Aug- 99, 6 males, 6 females; tributary to Tahoma Creek at Par- 99, 1 female; Paradise River at Fourth Crossing, 16-Aug- adise Road, 08-Jul-99, 1 female; Carbon River Entrance, 99, 1 female; creek behind Mystic Lake Ranger Station, 15-Aug-99, 1 female; Carbon River at Alice Falls, 15-Aug- 04-Sep-99, 1 male, 2 females; Falls Creek at Stevens 99, 1 male; stream by Lake James, 13-Sep-99, 1 male, 1 Canyon Road, 25-Jun-00, 5 males, 6 females; stream on female; west fork of White River, 13-Sep-99, 1 male; Van the Westside Road 1.5 miles from the Paradise Road, 26- Horn Creek at Waterfall, 13-Sep-99, 2 males; Fryingpan Jun-00, 1 male, 3 females; Paradise River in Paradise Val- Creek at Wonderland Trail Bridge, 29-Jun-00, 1 male, 1 ley, 10-Jul-00, 2 males, 4 females; St. Andrews Creek at female; seeps 0.9 mile S of St. Andrews Creek, 16-Sep-00, Westside Road, 14-Jul-00, 6 males, 3 females; Paradise 6 males, 2 females; seeps 0.8 mile S of St. Andrews Creek, River at Fourth Crossing, 23-Jul-00, 1 male; Paradise River 16-Sep-00, 2 males, 2 females; stream 0.4 mile S of St. in Paradise Valley, 02-Aug-00, 2 males; seeps into Eagle Andrews Creek, 16-Sep-00, 15 males, 10 females; stream Peak Creek, 07-May-01, 10 males, 2 females; boggy 1.75 miles S of St. Andrews Creek, 16-Sep-00, 9 males; St. stream tributary to Tahoma Creek 07-May-01, 17 males, Andrews Creek at Westside Road, 16-Sep-00, 18 males, 2 15 females; seeps by Nisqually River Bridge in Longmire, females; stream 1.5 miles S of St. Andrews Creek, 16-Sep- 13-May-01, 2 males, 2 females; Eagle Peak Creek at 00, 2 males, 3 females; stream 1.15 miles S of St. Andrews Nisqually River, 13-May-01, 13 males, 3 females; Ipsut Creek, 16-Sep-00, 4 males, 1 female; Dick Creek at Won- Falls at Ipsut Creek, 14-May-01, 3 males, 4 females; derland Trail, 23-Sep-00, 1 female; St. Andrews Creek at Huckleberry Creek Bridge, 14-May-01, 7 males; Eagle Westside Road, 17-Oct-00, 2 females; boggy stream tribu- Peak Creek at Nisqually River, 15-May-01, 1 male; small tary to Tahoma Creek, 07-May-01, 1 male. stream near Ohanapecosh Visitor Center, 16-May-01, 1 male, 3 females; stream into White River at Highway 410 Genus Moselia 0.5 mile S of park boundary, 16-May-01, 2 males, 2 Moselia infuscata females; Panther Creek at Highway 123, 09-Jun-01, 2 females; Deer Creek at Eastside Trail, 09-Jun-01, 2 males, Carbon River Entrance, Rainforest Loop Trail, 08-May- 1 female; St. Andrews Creek at Westside Road, 16-Jun-01, 95, 2 females; Reflection Lake, stream near Pinnacle Peak 2 males, 1 female; boggy stream tributary to Tahoma Trail, 14-Apr-96, 1 male; Tahoma Creek at Nisqually to Creek, 17-Jun-01, 7 males, 6 females; small stream near Paradise Road, 29-Mar-97, 1 male; Carbon River Entrance, Narada Falls, 23-Jul-01, 1 male. Rainforest Loop Trail, 27-May-97, 3 males, 2 females; Carbon River Entrance, 27-May-97, 4 males, 5 females; Genus Paraleuctra Ohanapecosh River at the Grove of the Patriarchs, 28- May-97, 1 female; Ohanapecosh Campground, 28-May- Paraleuctra forcipata 97, 1 female; Falls Creek at Stevens Canyon Road, 28- Westside Road 2.7 miles NNE of Nisqually Entrance, 29- May-97, 8 males, 5 females; seeps into Eagle Creek, 28- Mar-97, 1 male; Tahoma Creek, 1.8 miles NE of Nisqually May-97, 3 males, 6 females; Ohanapecosh River at Sheep Entrance, 29-Mar-97, 1 male, 2 females; Carbon River at Creek, 29-May-97, 1 male, 1 female; unnamed stream N Alice Falls, 06-Jun-97, 5 males; Huckleberry Creek of Crystal Creek, Highway 410, 29-May-97, 14 males, 11 Bridge, 28-Apr-98, 2 males; Mountain Meadow Stream at females; unnamed stream 3.4 miles S of park boundary Paul Peak Trail, 07-May-98, 2 males; Falls Creek at Highway 410, 29-May-97, 9 males, 5 females; seeps into Stevens Canyon Road, 05-Jun-99, 1 male, 1 female; Falls White River on Highway 410, 29-May-97, 12 males, 7 Creek at Stevens Canyon Road, 11-Jun-99, 2 males; Fry- females; Panther Creek at Highway 123, 29-May-97, 3 ingpan Creek at Sunrise Road Bridge, 06-Jul-99, 2 males; females; Nisqually River at Glacier View Bridge, 07-Jul- stream into White River at Highway 410 N of White 99, 7 males, 6 females; Eagle Peak Creek at Nisqually River Entrance, 07-Jul-99, 3 males; small tributary to River, 30-May-97, 10 males, 6 females; Ipsut Creek at Shaw Creek, 07-Jul-99, 3 males; small tributary on West- Carbon River Road Bridge, 06-Jun-97, 2 males; Eagle side Road 2 miles N of Paradise Road, 08-Jul-99, 1 male; Peak Creek 4360 ft, 14-Jun-97, 5 males, 1 female; St. Falls Creek at Stevens Canyon Road, 25-Jun-00, 2 males, Andrews Creek at Westside Road, 19-Jun-97, 6 males, 15 4 females; Tahoma Creek at Fish Creek, 08-Jul-00, 1 male; females; Olallie Creek, 28-Jun-97, 1 female; White River St. Andrews Creek at Westside Road, 14-Jul-00, 2 males, Campground, 23-Jul-97, 4 males, 7 females; White River 1 female; Fryingpan Creek at Sunrise Road Bridge, 16- at Sunrise Road Bridge, 02-Jul-98, 1 male, 2 females; Jul-00, 1 male. Mountain Meadow Stream at Paul Peak Trail, 07-Jul-98, 2 males; Falls Creek at Stevens Canyon Road, 05-Jun-99, 4 Paraleuctra occidentalis males; Falls Creek at Stevens Canyon Road, 11-Jun-99, 3 White River Road 0.5 mile N of ranger station, 21-Apr-96, males, 4 females; Falls Creek at Stevens Canyon Road, 06- 1 male; Westside Road 1.3 miles N of Paradise Road, 29- Jul-99, 4 males, 8 females; Fryingpan Creek at Sunrise Mar-97, 3 males; Tahoma Creek 1.8 miles NE of Paradise Road Bridge, 06-Jul-99, 36 males, 38 females; tributary to Road, 29-Mar-97, 3 males, 4 females; Ohanapecosh Ohanapecosh River, 07-Jul-99, 1 male, 4 females; stream Campground, 28-May-97, 1 male; Eagle Peak Creek at into White River at Highway 410 N of White River Nisqually River, 30-May-97, 4 females; Huckleberry Creek Entrance, 07-Jul-99, 1 male; Nisqually River at Glacier Bridge, 21-Apr-98, 1 male; Huckleberry Creek Bridge, View Bridge, 30-May-97, 38 males, 11 females; tributary 28-Apr-98, 2 males; St. Andrews Creek at Westside Road, to Tahoma Creek at Westside Road, 08-Jul-99, 7 males, 10 29-Apr-98, 1 male, 2 females; stream by South Mowich 2002] MOUNT RAINIER NATIONAL PARK STONEFLIES 397

River Camp, 07-May-98, 1 male; Fryingpan Creek at Sun- female; Huckleberry Creek Bridge, 14-May-01, 2 males, 1 rise Road Bridge, 06-Jul-99, 1 female; tributary to female. Ohanapecosh River, 07-Jul-99, 4 males; tributary to Tahoma Creek at Westside Road, 08-Jul-99, 1 male, 1 Perlomyia utahensis female; Falls Creek at Stevens Canyon Road, 08-Jul-99, 1 Eagle Peak Creek at Bridge, 30-May-97, 1 male. male; Paradise River in Paradise Valley, 16-Aug-99, 1 male, 1 female; Falls Creek at Stevens Canyon Road, 25- Genus Megaleuctra Jun-00, 2 males, 4 females; Paradise River in Paradise Val- Megaleuctra kincaidi ley, 10-Jul-00, 3 females; Falls Creek at Stevens Canyon Road, 16-Jul-00, 1 female; Fryingpan Creek 0.25 mile Ohanapecosh Campground, 09-May-96, 1 female; Frying- upstream of Sunrise Road, 16-Jul-00 1 male; Ipsut Falls at pan Creek at Sunrise Road Bridge, 06-Jul-99, 1 male; Ipsut Creek, 14-May-01, 1 male, 2 females; Ohanapecosh seep into Fryingpan Creek, 08-Jul-99, 1 male, 2 females; River at the Grove of the Patriarchs, 16-May-01, 1 male, 1 boggy stream tributary to Tahoma Creek, 07-May-01, 1 female; White River Highway 410, 0.5 mile S of park male. boundary, 16-May-01, 2 males, 2 females; Paradise River in Paradise Valley, 02-Jul-01, 1 male. Family Nemouridae Genus Malenka Paraleuctra projecta Malenka californica Boggy stream tributary to Tahoma Creek, 07-May-01, 3 Carbon River Entrance, 27-May-97, 1 male; Carbon River males; seeps by Nisqually River Bridge in Longmire, 07- Entrance, 21-Sep-97, 1 male; Carbon River Entrance, 15- May-01, 8 males; Eagle Peak Creek at Nisqually River, Aug-99, 2 females; Carbon River Entrance, 17-Aug-99, 6 13-May-01, 35 males, 18 females; seeps by Nisqually males. River Bridge in Longmire, 13-May-01, 1 male; Ipsut Falls at Ipsut Creek, 14-May-01, 1 male; Eagle Peak Creek at Malenka cornuta Nisqually River, 15-May-01, 2 males, 7 females; Nisqually River, 2 miles W of Longmire, 15-May-01, 1 male. Seeps into Eagle Creek, 28-May-97, 3 females; unnamed stream 3.4 miles S of park boundary on Highway 410, 29- Paraleuctra vershina May-97, 1 male; White River on Highway 410, 29-May-97, 1 male, 3 females; seeps into White River on Highway 410, Rainforest Loop Trail at Carbon River Entrance, 08-May- 29-May-97, 1 female; Ipsut Creek at Campground, 08- 95, 7 males, 1 female; Carbon River Entrance, 24-May-95, Sep-97, 1 female; Falls Creek at Stevens Canyon Road, 1 male; Carbon River Entrance, Rainforest Loop Trail, 27- 06-Jul-99, 1 male; Nisqually River at Glacier View Bridge, May-97, 5 males, 3 females; Nisqually River at Longmire, 07-Jul-99, 1 female; small tributary on Westside Road 2 28-May-97, 1 male, 2 females; Ohanapecosh Campground, miles N of Paradise Road, 08-Jul-99, 1 male; tributary to 28-May-97, 1 male; Ohanapecosh River at the Grove of Tahoma Creek at Paradise Road, 08-Jul-99, 10 males, 6 the Patriarchs, 28-May-97, 1 male, 1 female; Ohanapecosh females; Carbon River Entrance, 15-Aug-99, 2 females; River at Sheep Creek, 29-May-97, 1 male; seeps into White small tributary on Westside Road 2 miles N of Paradise River on Highway 410, 29-May-97, 4 males, 3 females; Road, 17-Aug-99, 3 females; West fork of White River, 13- Panther Creek at Highway 123, 29-May-97, 2 males; stream Sep-99, 1 female; Falls Creek at Stevens Canyon Road, into White River at Highway 410 N of White River 25-Jun-00, 1 male, 1 female; stream on Westside Road 1.5 entrance, 07-Jul-99, 2 males; Fryingpan Creek at Sunrise miles from Paradise Road, 26-Jun-00, 1 male; Fish Creek Road Bridge, 29-Jun-00, 1 female; Fish Creek at Westside at Westside Road, 14-Jul-00, 1 female; Falls Creek at Road, 14-Jul-00, 1 female; Fish Creek at Dry Creek, 07- Stevens Canyon Road, 16-Jul-00, 1 male; seep 0.8 mile S May-01, 1 male, 1 female; seeps into White River on of St. Andrews Creek, 16-Sep-00, 1 female; White River Highway 410, 14-May-01, 3 males, 2 females; White River at Sunrise Road, 15-Jun-01, 1 male; boggy stream tributary Highway 410, 0.5 mile S of park boundary, 16-May-01, 1 to Tahoma Creek, 17-Jun-01, 4 males, 7 females; boggy male, 2 females; Ohanapecosh Campground, 16-May-01, stream tributary to Tahoma Creek, 09-Jul-01, 1 male. 2 males, 1 female; White River at Sunrise Road, 15-Jun- 01, 7 males, 4 females; Paradise River in Paradise Valley, Genus Ledina 02-Jul-01, 1 male; Fryingpan Creek at Sunrise Road Bridge, 02-Jul-01, 2 males, 3 females; seeps into Fryingpan Creek Ledina tumana 0.5 mile upstream from Sunrise Road, 15-Jun-01, 6 males, Fryingpan Creek at Sunrise Road Bridge, 16-Jul-00, 1 male. 1 female. Genus Ostrocerca Genus Perlomyia Ostrocerca foersteri Perlomyia collaris Ipsut Creek at campground, 18-May-95, 3 males; Falls Huckleberry Creek Bridge, 28-Apr-98, 1 male; stream by Creek on Carbon River Road, 27-May-97, 1 male; South Mowich River Camp, 04-May-98, 1 male; North Nisqually River at Longmire, 28-May-97, 3 males; Falls Mowich River at Wonderland Trail, 07-May-98, 1 male; Creek at Stevens Canyon Road, 06-Jul-99, 1 male; small Mountain Meadow Stream at Paul Peak Trail, 07-May-98, stream near Longmire, 09-Jul-99, 1 male. 1 male; seeps by Nisqually River Bridge in Longmire, 07- May-01, 10 males, 1 female; seeps by Nisqually River Genus Podmosta Bridge in Longmire, 13-May-01, 2 males, 1 female; Eagle Podmosta decepta Peak Creek at Nisqually River, 13-May-01, 17 males, 4 females; seeps into White River on Highway 410, 14- Small stream into Ohanapecosh River near Highway 123, May-01, 1 male; Ipsut Falls at Ipsut Creek, 14-May-01, 1 07-Jul-99, 1 female; Paradise River in Paradise Valley, 16- 398 WESTERN NORTH AMERICAN NATURALIST [Volume 62

Aug-99, 3 males, 5 females; Paradise River above Narada Bridge, 06-Jul-99, 8 males, 4 females; Paradise River in Falls, 16-Aug-99, 1 female; Paradise River in Paradise Val- Paradise Valley, 16-Aug-99, 13 males, 3 females; Paradise ley, 07-Sep-99, 2 females. River above Narada Falls, 16-Aug-99, 3 males, 4 females; Paradise River in Paradise Valley, 02-Aug-00, 1 male; Podmosta delicatula boggy stream tributary to Tahoma Creek, 07-May-01, 3 Paradise River above Narada Falls, 16-Aug-99, 1 male; males, 2 females; seeps into White River on Highway 410, Paradise River in Paradise Valley, 16-Aug-99, 9 males, 7 14-May-01, 2 males; stream into White River at Highway females; Paradise River in Paradise Valley, 02-Aug-00, 1 410, 0.5 mile S of park boundary, 16-May-01, 5 males, 4 male; Eagle Peak Creek at Nisqually River, 13-May-01, 1 females; St. Andrews Creek at Westside Road, 16-Jun-01, female. 1 male; boggy stream tributary to Tahoma Creek, 17-Jun- 01, 5 males; Paradise River above Narada Falls, 02-Jul-01, Genus Prostoia 5 males, 4 females. Prostoia besametsa Genus Visoka Ohanapecosh River at the Grove of the Patriarchs, 28- Visoka cataractae May-97, 1 male, 1 female; Ohanapecosh Campground, 28- May-97, 4 males, 5 females; Panther Creek at Highway 0.6 mile S of Ipsut Campground, 04-Apr-97, 1 male; Ipsut 123, 29-May-97, 2 males, 5 females; Fryingpan Creek at Falls on Ipsut Creek, 04-May-97, 17 males, 1 female. Sunrise Road Bridge, 06-Jul-99, 1 male; Fryingpan Creek at Sunrise Road Bridge, 16-Jul-00, 1 male, 1 female; seeps Genus Zapada into White River on Highway 410, 14-May-01, 7 males, 6 Zapada cinctipes females; Longmire, 15-May-01, 1 male, 1 female; Ohana- pecosh Campground, 16-May-01, 2 males, 5 females; 0.7 mile SE of Ipsut Campground, 25-Mar-95, 4 males, 2 White River on Highway 410, 0.5 mile S of park bound- females; Tahoma Creek at Westside Road, 02-Mar-96, 1 ary, 16-May-01, 17 males, 8 females; Ohanapecosh River male, 3 females; Tahoma Creek 0.1 mile N of Dry Creek, at the Grove of the Patriarchs, 16-May-01, 2 males, 1 02-Mar-96, 1 male, 3 females; Reflection Lake, stream female; Deer Creek at Eastside Trail, 09-Jun-01, 1 female. near Pinnacle Peak Trail, 14-Apr-96, 2 males, 2 females; White River Road 0.5 mile N of ranger station, 21-Apr-96, Genus Soyedina 1 male, 4 females; Ohanapecosh Campground, 12-May- 96, 1 female; Ipsut Falls at Ipsut Creek, 23-Feb-97, 1 Soyedina interrupta female; south of Chenuis Falls parking lot, 23-Feb-97, 2 Summerland, 23-Jul-95, 1 male; 0.5 mile W of White males, 13 females; 0.7 mile SE of Ipsut Campground, 23- River Ranger Station, 21-Apr-96, 1 male; seeps into Eagle Mar-97, 1 female; Westside Road 1.3 miles N of Paradise Creek, 28-May-97, 2 males, 1 female; unnamed stream 3.4 Road, 29-Mar-97, 4 females; Tahoma Creek, 1.8 miles NE miles S of park boundary on Highway 410, 29-May-97, 1 of Nisqually Entrance, 29-Mar-97, 4 males, 8 females; male; seeps into White River on Highway 410, 29-May- Westside Road, 2.6 miles NNE of Nisqually Entrance, 29- 97, 4 males, 1 female; Nisqually River at Glacier View Mar-97, 5 males, 6 females; 0.6 mile S of Ipsut Camp- Bridge, 30-May-97, 11 males, 3 females; St. Andrews ground, 04-Apr-97, 1 male, 1 female; Shaw Creek at Creek at Westside Road, 19-Jun-97, 1 male, 2 females; White River Road, 21-Apr-97, 1 male; White River Camp- White River across from the White River Campground, ground, 01-Apr-98, 1 female; tributary to Tahoma Creek at 02-Jul-98, 1 male; Falls Creek at Stevens Canyon Road, Westside Road, 08-Jul-99, 1 female; Paradise River in Par- 05-Jun-99, 1 male, 2 females; Falls Creek at Stevens adise Valley, 16-Aug-99, 9 males, 8 females; Paradise River Canyon Road, 11-Jun-99, 1 male, 5 females; Falls Creek at above Narada Falls, 07-Sep-99, 1 male, 2 females; Par- Stevens Canyon Road, 06-Jul-99, 4 males; Fryingpan adise River at Fourth Crossing, 16-Jul-00, 2 males, 1 Creek at Sunrise Road Bridge, 06-Jul-99, 4 males, 5 female; Paradise River at Fourth Crossing, 23-Jul-00, 1 females; Nisqually River at Glacier View Bridge, 07-Jul- male; Paradise River in Paradise Valley, 02-Aug-00, 2 99, 3 males, 5 females; seep into Fryingpan Creek, 08-Jul- males; boggy stream tributary to Tahoma Creek, 07-May- 99, 3 males, 1 female; stream on Westside Road 1.5 miles 01, 3 females; Longmire, 15-May-01, 1 male. from Paradise Road, 26-Jun-00, 1 male, 1 female; Frying- pan Creek at Wonderland Trail, 29-Jun-00, 1 male; South Zapada columbiana Puyallup River at Westside Road, 14-Jul-00, 1 male, 1 2.2 miles SE of Ipsut Campground, 25-Mar-95, 2 males, 2 female; Longmire, 13-May-01, 1 male; Eagle Peak Creek females; Nahunta Falls, 16-Apr-95, 12 males; Nisqually at Nisqually River, 13-May-01, 1 male; Ipsut Falls at Ipsut River at Longmire, 16-Apr-95, 1 male; 0.15 mile NE of Creek, 14-May-01, 1 male, 1 female; Eagle Peak Creek at Glacier View Bridge, 16-Apr-95, 2 males; Squaw Lake, 07- Nisqually River, 15-May-01, 1 male; White River High- May-95, 1 male; Nisqually River at creek from Tato Falls, way 410, 0.5 mile S of park boundary, 16-May-01, 1 male. 14-Apr-96, 5 males, 5 females; Reflection Lake, stream near Pinnacle Peak Trail, 14-Apr-96, 11 males, 5 females; Soyedina producta 0.1 mile NE of Glacier View Bridge, 14-Apr-96, 1 male; Unnamed stream 3.4 miles S of park boundary on High- Paradise River at Fourth Crossing, 21-Apr-96, 22 males, 9 way 410, 29-May-97, 3 males; seeps into White River on females; Edith Creek at Fourth Crossing, 05-May-96, 2 Highway 410, 29-May-97, 1 male; Nisqually River at males, 1 female; Edith Creek at Fourth Crossing, 16-May- Glacier View Bridge, 30-May-97, 24 males, 3 females; 96, 15 males, 6 females; Ipsut Falls, 23-Feb-97, 10 males, Carbon River at Alice Falls, 06-Jun-97, 3 males, 4 females; 3 females; Ipsut Creek, 0.6 mile S of Ipsut Campground, Ipsut Creek at Carbon River Road Bridge, 06-Jun-97, 1 23-Feb-97, 1 male; Tahoma Creek, 02-Mar-97, 1 male, 1 male, 1 female; North Mowich River at Wonderland Trail, female; Westside Road, 2.6 miles NNE of Nisqually 07-May-98, 2 males; Fryingpan Creek at Sunrise Road Entrance, 29-Mar-97, 4 males, 3 females; Westside Road 2002] MOUNT RAINIER NATIONAL PARK STONEFLIES 399

1.3 miles N of Paradise Road, 29-Mar-97, 2 males; 0.6 females; tributary to Tahoma Creek at Westside Road, 08- mile S of Ipsut Campground, 04-Apr-97, 1 male; 0.7 mile Jul-99, 3 males, 1 female; Paradise River in Paradise Val- SE of Ipsut Campground, 04-Apr-97, 10 males, 9 females; ley, 07-Sep-99, 1 male; Ohanapecosh River at the Grove Ipsut Falls on Ipsut Creek, 04-May-97, 1 female; Crystal of the Patriarchs, 16-May-01, 1 male; White River High- Creek at Highway 410, 29-May-97, 1 male, 2 females; Eagle way 410, 0.5 mile S of park boundary, 16-May-01, 2 males; Peak Creek at 4360 ft, 14-Jun-97, 2 males, 2 females; boggy stream tributary to Tahoma Creek, 17-Jun-01, 4 White River Campground, 01-Apr-98, 15 males, 12 females; males, 1 female. Fryingpan Creek at Sunrise Road Bridge, 01-Apr-98, 5 males, 4 females; Paradise River in Paradise Valley, 10-Jul- Family Taeniopterygidae 00, 3 males, 11 females; Paradise River in Paradise Valley, Genus Doddsia 16-Jul-00, 2 males, 3 females; Eagle Peak Creek at Nis- Doddsia occidentalis qually River, 13-May-01, 1 male, 1 female; Deer Creek at Eastside Trail, 09-Jun-01, 1 male; Paradise River in Par- Nisqually River at Longmire, 16-Apr-95, 2 males; Paradise adise Valley, 02-Jul-01, 1 male. Ice Caves, 20-Aug-95, 2 females; Westside Road 2.6 miles NNE of Nisqually Entrance, 29-Mar-97, 6 males, 5 females; Zapada cordillera Nisqually River at Longmire, 28-May-97, 5 females; seeps into White River on Highway 410, 29-May-97, 7 females; Westside Road 1.3 miles N of Paradise Road, 29-Mar-97, 1 Carbon River at Alice Falls, 06-Jun-97, 2 males; Glacier male, 1 female; Tahoma Creek, 1.8 miles NE of Nisqually Basin, 20-Jul-97, 3 females; Glacier Basin, 21-Jul-97, 3 Entrance, 29-Mar-97, 1 female; boggy stream tributary to females; Glacier Basin, 27-Jul-97, 16 females; Huckle- Tahoma Creek, 07-May-01, 1 female; seeps into Eagle berry Creek Bridge, 17-Apr-98, 2 males, 2 females; Long- Peak Creek, 07-May-01, 1 male. mire, 20-Apr-98, 1 male; Huckleberry Creek Bridge, 21- Apr-98, 1 male, 5 females; South Puyallup River at Westside Zapada frigida Road, 21-Apr-98, 1 female; St. Andrews Creek at Westside 0.6 mile WSW of Shaw Creek Bridge, 03-Nov-96, 1 female; Road, 29-Apr-98, 3 males, 1 female; North Mowich River Carbon River Entrance, Rainforest Loop Trail, 08-May- at Wonderland Trail, 07-May-98, 1 male, 1 female; Glacier 95, 1 male, 1 female; Carbon River Entrance, Rainforest Basin, 06-Jun-98, 24 females; Falls Creek at Stevens Can- Loop Trail, 27-May-97, 1 male; Ohanapecosh Campground, yon Road, 11-Jun-99, 2 females; Falls Creek at Stevens 28-May-97, 2 males; Falls Creek at Stevens Canyon Road, Canyon Road, 06-Jul-99, 2 females; small stream into 28-May-97, 1 female; Carbon River at Alice Falls, 06-Jun- Ohanapecosh River near Highway 123, 07-Jul-99, 1 female; 97, 4 males, 2 females; Ipsut Creek at Carbon River Road stream into White River at Highway 410 N of White Bridge, 06-Jun-97, 1 male; St. Andrews Creek at Westside River Entrance, 07-Jul-99, 1 female; Pebble Creek at Road, 19-Jun-97, 5 males; White River across from the 7200 feet, 20-Jul-99, 6 females; Muir Snowfield at 8000 ft, White River Campground, 02-Jul-98, 3 males, 1 female; 23-Jul-99, 1 female; Fryingpan Creek at Wonderland Trail, Fryingpan Creek at Sunrise Road Bridge, 06-Jul-99, 11 29-Jun-00, 1 male, 1 female; Paradise River in Paradise males, 5 females; Falls Creek at Stevens Canyon Road, 06- Valley, 10-Jul-00, 3 females; Fryingpan Creek at Sunrise Jul-99, 4 males; seep into Fryingpan Creek, 08-Jul-99, 3 Road Bridge, 10-Jul-00, 2 females; Fryingpan Creek at males; tributary to Tahoma Creek at Westside Road, 08- Sunrise Road Bridge, 16-Jul-00, 19 females; Kautz Creek Jul-99, 2 males, 1 female; Falls Creek at Stevens Canyon at Nisqually to Paradise Road, 04-Apr-01, 1 male, 1 female; Road, 08-Jul-99, 2 males; high-gradient tributary at Paradise Ice Caves, 12-May-01, 2 females; Kautz Creek at Nisqually River Longmire, 09-Jul-99, 2 males; Paradise Nisqually to Paradise Road, 13-May-01, 1 female; Tahoma River in Paradise Valley, 16-Aug-99, 5 males, 4 females; Creek at Nisqually to Paradise Road, 13-May-01, 1 male, Falls Creek at Stevens Canyon Road, 25-Jun-00, 1 male, 2 1 female; Ipsut Falls at Ipsut Creek, 14-May-01, 3 females; females; Paradise River in Paradise Valley, 10-Jul-00, 2 seeps into White River on Highway 410, 14-May-01, 3 females; St. Andrews Creek at Westside Road, 14-Jul-00, males, 7 females; Eagle Peak Creek at Nisqually River, 15- 1 male; Paradise River in Paradise Valley, 02-Aug-00, 1 male, May-01, 1 male; Nisqually River, 2 miles W of Longmire, 1 female; seeps into Fryingpan Creek 0.5 mile upstream 15-May-01, 8 males, 6 females; Longmire, 15-May-01, 13 from Sunrise Road, 15-Jun-01, 4 males, 1 female. males, 4 females; Paradise River at Stevens Canyon Road, 15-May-01, 1 male, 1 female; Ohanapecosh River at the Zapada haysi Grove of the Patriarchs, 15-May-01, 1 male, 1 female; White River Highway 410, 0.5 mile S of park boundary, White River across from the White River Campground, 16-May-01, 1 male, 2 females; Ohanapecosh Campground, 02-Jul-98, 1 male; White River Highway 410, 0.5 mile S 16-May-01, 4 females. of park boundary, 16-May-01, 2 males; Ohanapecosh River at the Grove of the Patriarchs, 16-May-01, 1 male; Genus Taenionema boggy stream tributary to Tahoma Creek, 04-Jul-01, 1 male. Taenionema kincaidi Kautz Creek Trail 0.7 mile N of Nisqually Road, 08-May- Zapada oregonensis 95, 1 male, 2 females; South Mowich River Camp, 31-May- Carbon River Entrance, Rainforest Loop Trail, 27-May- 95, 2 males, 1 female; Tahoma Woods, 29-Apr-96, 1 male; 97, 4 males, 1 female; unnamed stream N of Crystal Creek, Ohanapecosh Campground, 28-May-97, 6 males, 5 females; Highway 410, 29-May-97, 3 males; seeps into White River Ohanapecosh River at the Grove of the Patriarchs, 28- on Highway 410, 29-May-97, 1 male; White River High- May-97, 3 males, 3 females; Nisqually River at Longmire, way 410, 29-May-97, 1 male; Mountain Meadow Stream 28-May-97, 4 males, 1 female; Panther Creek at Highway at Paul Peak Trail, 07-May-98, 2 males; White River across 123, 29-May-97, 5 males, 2 females; seeps into White River from the White River Campground, 02-Jul-98, 2 males, 2 on Highway 410, 29-May-97, 17 males, 7 females; Deer 400 WESTERN NORTH AMERICAN NATURALIST [Volume 62

Creek on Highway 123, 29-May-97, 5 males, 3 females; Alloperla serrata Ohanapecosh River at Sheep Creek, 29-May-97, 5 males, Edith Creek at Fourth Crossing, 13-Sep-97, 1 male; Para- 4 females; Tahoma Creek at Nisqually to Paradise Road, dise River in Paradise Valley, 16-Aug-99, 3 males; Paradise 30-May-97, 6 females; Carbon River at Alice Falls, 06- River above Narada Falls, 16-Aug-99, 9 males; Paradise Jun-97, 6 males, 6 females; Ipsut Creek at Carbon River River at Fourth Crossing, 16-Aug-99, 2 males; Paradise Road Bridge, 06-Jun-97, 5 males, 4 females; Mountain River in Paradise Valley, 07-Sep-99, 1 male, 2 females; Meadow Stream at Paul Peak Trail, 07-May-98, 7 males, 8 Paradise River in Paradise Valley, 02-Aug-00, 2 males, 2 females; North Mowich River at Wonderland Trail, 07- females. May-98, 14 males, 4 females; Fryingpan Creek at Sunrise Road Bridge, 03-Jul-98, 1 female; Fryingpan Creek at Sun- Genus Plumiperla rise Road Bridge, 06-Jul-99, 18 males, 7 females; Nisqually Plumiperla diversa River at Glacier View Bridge, 07-Jul-99, 1 male, 2 females; stream into White River at Highway 410 N of White Carbon River Entrance, 27-May-97, 15 males, 16 females; River Entrance, 07-Jul-99, 1 female; Shaw Creek at White Nisqually River at Longmire, 28-May-97, 1 male; seeps River Road, 07-Jul-99, 2 males; tributary to Tahoma Creek into White River on Highway 410, 29-May-97, 2 males, 1 at Westside Road, 08-Jul-99, 2 females; Falls Creek at female; Tahoma Creek at Nisqually to Paradise Road, 30- Stevens Canyon Road, 08-Jul-99, 1 male; small stream May-97, 3 males, 2 females; Longmire, 27-Jul-98, 1 male; near Longmire, 09-Jul-99, 1 female; Paradise River at Falls Creek at Stevens Canyon Road, 06-Jul-99, 5 males, 5 Fourth Crossing, 16-Aug-99, 1 male; Paradise River above females; small stream near Longmire, 07-Jul-99, 2 males; Narada Falls, 16-Aug-99, 2 males, 27 females; Fryingpan Nisqually River at Glacier View Bridge, 07-Jul-99, 14 Creek at Sunrise Road Bridge, 16-Aug-99, 5 females; Falls males, 11 females; tributary to Ohanapecosh River, 07- Creek at Stevens Canyon Road, 25-Jun-00, 1 male; Fry- Jul-99, 1 male, 6 females; Tahoma Creek, 08-Jul-99, 1 ingpan Creek at Wonderland Trail, 29-Jun-00, 2 females; male, 3 females; tributary to Tahoma Creek at Westside Cataract Creek at Wonderland Trail, 09-Jul-00, 2 males, 3 Road, 08-Jul-99, 1 female; Paradise River above Narada females; Paradise River in Paradise Valley, 10-Jul-00, 1 Falls, 16-Aug-99, 2 males; Fryingpan Creek at Sunrise male, 2 females; Fryingpan Creek at Sunrise Road, 10-Jul- Road Bridge, 16-Aug-99, 2 males, 2 females; Falls Creek 00, 1 female; Fish Creek at Westside Road, 14-Jul-00, 5 at Stevens Canyon Road, 25-Jun-00, 2 females; Fryingpan females; Fryingpan Creek at Sunrise Road, 16-Jul-00, 4 Creek at Sunrise Road Bridge, 29-Jun-00, 1 male, 2 females; Fryingpan Creek 0.25 mile upstream from Sun- females; Tahoma Creek at Fish Creek, 08-Jul-00, 5 males, rise Road, 16-Jul-00, 1 male, 2 females; Falls Creek at 2 females; Cataract Creek at Wonderland Trail, 09-Jul-00, Stevens Canyon Road, 16-Jul-00, 1 female; Paradise River 1 male, 1 female; South Puyallup River at Westside Road, at Fourth Crossing, 16-Jul-00, 4 males, 1 female; Fish Creek 14-Jul-00, 20 males, 7 females; Fryingpan Creek at Sun- at Westside Road, 05-Aug-00, 1 female; seeps by Nisqually rise Road Bridge, 16-Jul-00, 2 males, 5 females; Fryingpan River Bridge in Longmire, 07-May-01, 2 males, 1 female; Creek 0.25 mile upstream of Sunrise Road, 16-Jul-00, 3 Fish Creek at Dry Creek, 07-May-01, 8 males; Kautz males, 4 females; Tahoma Creek at Nisqually to Paradise Creek at Nisqually to Paradise Road, 13-May-01, 8 males; Road, 13-May-01, 1 male; White River at Sunrise Road Eagle Peak Creek at Nisqually River, 13-May-01, 1 male; Bridge, 15-Jun-01, 3 males, 2 females; boggy stream trib- Tahoma Creek at Nisqually to Paradise Road, 13-May-01, utary to Tahoma Creek, 17-Jun-01, 2 males, 1 female; 6 males; Ipsut Falls at Ipsut Creek, 14-May-01, 2 females; small stream near Narada Falls, 23-Jul-01, 3 males. seeps into White River on Highway 410, 14-May-01, 2 Genus Suwallia males; Huckleberry Creek Bridge, 14-May-01, 1 male; Paradise River at Stevens Canyon Road, 15-May-01, 1 Suwallia dubia male; Nisqually River, 2 miles W of Longmire, 15-May- Ipsut Creek at Carbon River Road Bridge, 08-Sep-97, 1 01, 4 males, 1 female; White River Highway 410, 0.5 mile female; Carbon River Entrance, 15-Aug-99, 1 male, 1 S of park boundary, 16-May-01, 4 males; Ohanapecosh female; Carbon River Entrance, 16-Aug-99, 1 male; Falls Campground, 16-May-01, 2 males; Panther Creek at High- Creek at Stevens Canyon Road, 16-Aug-99, 1 male; small way 123, 09-Jun-01, 1 female; Ohanapecosh River 0.5 tributary on Westside Road 2 miles N of Paradise Road, mile S of Shriner Peak Trailhead, 09-Jun-01, 4 males, 3 17-Aug-99, 1 male; Van Horn Creek at Waterfall, 13-Sep- females; Deer Creek at Eastside Trail, 09-Jun-01, 1 male; 99, 3 males, 2 females; West fork of White River, 13-Sep- White River at Sunrise Road, 15-Jun-01, 1 male, 1 female; 99, 1 female; South Puyallup River at Westside Road, 29- Paradise River above Narada Falls, 02-Jul-01, 2 females. Sep-00, 16 males, 6 females.

Taenionema pallidum Suwallia forcipata North Mowich River at Wonderland Trail, 07-May-98, 2 Nisqually River at Longmire, 04-Apr-96, 1 male, 8 females; males, 1 female; Tahoma Creek at Nisqually to Paradise Carbon River Entrance, 20-Sep-97, 10 males, 13 females; Road, 13-May-01, 1 male; seeps into White River on Huckleberry Creek Bridge, 27-Sep-97, 8 females. Highway 410, 14-May-01, 1 male. Suwallia pallidula Group Systellognatha South Mowich River Camp, 31-May-95, 4 males, 5 Family Chloroperlidae females. Genus Alloperla Alloperla fraterna Genus Sweltsa Falls Creek at Stevens Canyon Road, 16-Aug-99, 6 males; Sweltsa borealis small stream near Ohanapecosh Visitor Center, 16-May- Carbon River Entrance, Rainforest Loop Trail, 08-May- 01, 1 male. 95, 3 males, 3 females; Wonderland Trail 0.5 mile E of Paul 2002] MOUNT RAINIER NATIONAL PARK STONEFLIES 401

Peak Trail junction, 01-Jun-95, 1 male, 1 female; Carbon River in Paradise Valley, 16-Aug-99, 8 males, 4 females; River Entrance, 27-May-97, 1 male, 2 females; Carbon Paradise River in Paradise Valley, 07-Sep-99, 2 males, 2 River Entrance, Rainforest Loop Trail, 27-May-97, 2 males, females; Falls Creek at Stevens Canyon Road, 25-Jun-00, 2 females; Nisqually River at Longmire, 28-May-97, 1 2 males; stream on Westside Road 1.5 miles from Paradise female; Falls Creek at Stevens Canyon Road, 28-May-97, Road, 26-Jun-00, 2 males, 4 females; Fryingpan Creek at 5 males, 3 females; White River Highway 410, 29-May-97, Sunrise Road Bridge, 29-Jun-00, 1 male; Cataract Creek 1 male, 1 female; unnamed stream N of Crystal Creek, at Wonderland Trail, 09-Jul-00, 10 males, 3 females; Fry- Highway 410, 29-May-97, 10 males, 4 females; seeps into ingpan Creek at Sunrise Road Bridge, 10-Jul-00, 2 males, White River on Highway 410, 29-May-97, 3 females; Pan- 2 females; Fish Creek at Westside Road, 14-Jul-00, 2 ther Creek at Highway 123, 29-May-97, 1 female; Eagle males, 11 females; St. Andrews Creek at Westside Road, Peak Creek at Nisqually River, 30-May-97, 1 female; Ipsut 14-Jul-00, 10 males, 5 females; Fryingpan Creek at Sun- Creek at Carbon River Road Bridge, 06-Jun-97, 1 male, 1 rise Road Bridge, 16-Jul-00, 1 male, 2 females; Fryingpan female; St. Andrews Creek at Westside Road, 14-Jun-97, 1 Creek 0.25 mile upstream from Sunrise Road, 16-Jul-00, 4 male; Olallie Creek, 28-Jun-97, 2 males; Falls Creek at males; Paradise River at Fourth Crossing, 23-Jul-00, 3 Stevens Canyon Road, 06-Jul-99, 4 males, 2 females; stream males; Paradise River in Paradise Valley, 02-Aug-00, 1 into White River at Highway 410 N of White River en- male, 6 females; Fish Creek at Westside Road, 05-Aug-00, trance, 07-Jul-99, 1 female; tributary to Ohanapecosh 1 male, 2 females; boggy stream tributary to Tahoma Creek, River, 07-Jul-99, 7 females; Nisqually River at Glacier 01-Jun-01, 8 males, 2 females; Falls Creek at Stevens Can- View Bridge, 07-Jul-99, 3 females; small stream into yon Road, 07-Jun-01, 1 male, 1 female; Panther Creek at Ohanapecosh River near Highway 123, 07-Jul-99, 1 male; Highway 123, 09-Jun-01, 1 male; boggy stream tributary tributary to Tahoma Creek at Westside Road, 08-Jul-99, 3 to Tahoma Creek, 17-Jun-01, 6 males; Paradise River males, 4 females; tributary to Tahoma Creek at Paradise above Narada Falls, 02-Jul-01, 1 male; boggy stream tribu- Road, 08-Jul-99, 3 females; small tributary on Westside tary to Tahoma Creek, 09-Jul-01, 2 males, 1 female; small Road 2 miles N of Paradise Road, 08-Jul-99, 1 female; stream near Narada Falls, 23-Jul-01, 2 males. high-gradient tributary at Nisqually River Longmire, 09- Jul-99, 1 male; Paradise River above Narada Falls, 16-Aug- Sweltsa occidens 99, 1 male, 3 females; Paradise River in Paradise Valley, Laughingwater Creek, 27-Jun-95, 5 males; St. Andrews 16-Aug-99, 5 males, 7 females; Paradise River in Paradise Creek at Westside Road, 24-Jul-97, 4 males, 32 females; Valley, 07-Sep-99, 1 male, 1 female; Van Horn Creek at high-gradient tributary at Nisqually River Longmire, 09- Waterfall, 13-Sep-99, 1 female; Falls Creek at Stevens Jul-99, 3 males; Paradise River in Paradise Valley, 16-Aug- Canyon Road, 25-Jun-00, 4 males, 2 females; Cataract 99, 4 males, 7 females; Paradise River above Narada Falls, Creek at Wonderland Trail, 09-Jul-00, 1 female; Paradise 16-Aug-99, 52 males, 15 females; Paradise River at Fourth River in Paradise Valley, 10-Jul-00, 3 males; Fish Creek at Crossing, 16-Aug-99, 1 male; Falls Creek at Stevens Canyon Westside Road, 14-Jul-00, 1 male; South Puyallup River Road, 25-Jun-00, 1 male; Cataract Creek at Wonderland at Westside Road, 14-Jul-00, 1 female; Paradise River in Trail, 09-Jul-00, 2 males; St. Andrews Creek at Westside Paradise Valley, 23-Jul-00, 1 female; Paradise River in Par- Road, 14-Jul-00, 4 females; Fryingpan Creek at Sunrise adise Valley, 02-Aug-00, 1 male, 4 females; Olallie Creek Road Bridge, 16-Jul-00, 1 male, 2 females; Fryingpan Creek at Olallie Creek Camp, 03-Aug-00, 1 female; boggy stream 0.25 mile upstream Sunrise Road, 29-Jun-00, 1 male; Par- tributary to Tahoma Creek, 07-May-01, 2 males; Ipsut adise River at Fourth Crossing, 23-Jul-00, 2 males, 1 female; Falls at Ipsut Creek, 14-May-01, 1 male; Ohanapecosh Tamanos Creek at trail, 03-Aug-00, 10 males, 5 females; River at the Grove of the Patriarchs, 16-May-01, 1 male; Panther Creek at Highway 123, 09-Jun-01, 1 male; boggy boggy stream tributary to Tahoma Creek, 17-Jun-01, 3 stream tributary to Tahoma Creek, 17-Jun-01, 1 male; Fry- males, 4 females; small stream near Narada Falls, 23-Jul- ingpan Creek at Sunrise Road Bridge, 02-Jul-01, 1 male; 01, 2 males, 2 females. small stream near Narada Falls, 23-Jul-01, 1 male.

Sweltsa exquisita Sweltsa revelstoka Carbon River Entrance, Rainforest Loop Trail, 27-May- White River Campground, 23-Jul-97, 7 males, 6 females; 97, 1 male; Carbon River Entrance, 27-May-97, 1 male, 1 inflow upper Palisade Lake, 08-Aug-97, 17 males, 8 females; female; Carbon River at Chenuis Falls Trailhead, 06-Jun- White River across from White River Campground, 02- 97, 8 males; St. Andrews Creek at Westside Road, 24-Jul- Jul-98, 4 males, 1 female; seep into Fryingpan Creek, 08- 97, 3 males, 1 female; Edith Creek at Fourth Crossing, 13- Jul-99, 1 male; Fryingpan Creek at Sunrise Road Bridge, Sep-97, 2 males; Falls Creek at Stevens Canyon Road, 06- 16-Aug-99, 1 female; Paradise River above Narada Falls, Jul-99, 7 males, 13 females; Fryingpan Creek at Sunrise 16-Aug-99, 3 males, 5 females; Paradise River in Paradise Road Bridge, 06-Jul-99, 1 male; tributary to Ohanapecosh Valley, 16-Aug-99, 5 males, 3 females; Paradise River at River, 07-Jul-99, 2 males; Nisqually River at Glacier View Fourth Crossing, 16-Aug-99, 5 males, 7 females; creek be- Bridge, 07-Jul-99, 2 males; small stream into Ohana- hind Mystic Lake Ranger Station, 04-Sep-99, 1 female; pecosh River near Highway 123, 07-Jul-99, 9 males; tribu- Cataract Creek at Wonderland Trail, 09-Jul-00, 1 female; tary to Tahoma Creek at Westside Road, 08-Jul-99, 2 Fryingpan Creek at Sunrise Road, 10-Jul-00, 6 males, 2 males, 3 females; Falls Creek at Stevens Canyon Road, 08- females; Fryingpan Creek at Sunrise Road, 16-Jul-00, 1 Jul-99, 4 males; small stream near Longmire, 09-Jul-99, 2 female; Paradise River at Fourth Crossing, 23-Jul-00, 7 males, 3 females; High gradient tributary at Nisqually males, 1 female; Paradise River in Paradise Valley, 02-Aug- River at Longmire, 09-Jul-99, 1 male; Paradise River at 00, 4 males, 4 females; Tamanos Creek at Trail, 03-Aug-00, Fourth Crossing, 16-Aug-99, 1 male; Paradise River above 2 males; Paradise River in Paradise Valley, 02-Jul-01, 1 male; Narada Falls, 16-Aug-99, 7 males, 4 females; Falls Creek at Fryingpan Creek at Sunrise Road, 02-Jul-01, 1 female; Stevens Canyon Road, 16-Aug-99, 1 female; Paradise Dick Creek at Wonderland Trail, 27-Jul-01, 1 female. 402 WESTERN NORTH AMERICAN NATURALIST [Volume 62

Sweltsa new species White River across from the White River Campground, Tributary to Tahoma Creek at Westside Road, 08-Jul-99, 1 02-Jul-98, 1 female; boggy stream tributary to Tahoma male. Creek, 07-May-01, 1 male, 2 females; White River High- way 410, 0.5 mile S of park boundary, 16-May-01, 1 Genus Kathroperla female; stream into White River at Highway 410, 0.5 mile S of park boundary, 16-May-01, 1 male; Ohanapecosh Kathroperla perdita River 0.5 mile S of Shriner Peak Trailhead, 09-Jun-01, 5 White River Highway 410, 29-May-97, 2 males; unnamed females. stream 3.4 miles S of park boundary on Highway 410, 29- May-97, 1 female; Crystal Creek at Highway 410, 29- Yoraperla nigrisoma May-97, 1 male, 1 female; unnamed stream N of Crystal Mowich Road at Grindstone Trail Crossing, 07-Jul-95, 1 Creek, Highway 410, 29-May-97, 2 females; Panther female; Ipsut Creek at Carbon River Road Bridge, 06- Creek at Highway 123, 29-May-97, 1 female; Tahoma Jun-97, 1 female; Falls Creek at Stevens Canyon Road, 11- Creek at Nisqually to Paradise Road, 30-May-97, 2 males; Jun-99, 2 males; Fryingpan Creek at Sunrise Road Bridge, Mountain Meadow Stream at Paul Peak Trail, 07-May-98, 06-Jul-99, 1 male, 1 female; small tributary on Westside 1 male; seeps into White River on Highway 410, 14-May- Road 2 miles N of Paradise Road, 08-Jul-99, 1 female; 01, 1 female; Ipsut Falls at Ipsut Creek, 14-May-01, 1 tributary to Tahoma Creek at Westside Road, 08-Jul-99, male, 1 female; White River Highway 410, 0.5 mile S of 25 males, 12 females; tributary to Tahoma Creek at Par- park boundary, 16-May-01, 1 male, 2 females; Deer Creek adise Road, 08-Jul-99, 2 males, 1 female; Paradise River in at Eastside Trail, 09-Jun-01, 1 male; White River at Sun- Paradise Valley, 16-Aug-99, 1 male, 1 female; Paradise rise Road, 15-Jun-01, 1 female. River above Narada Falls, 16-Aug-99, 2 females; Falls Kathroperla takhoma Creek at Stevens Canyon Road, 16-Aug-99, 2 females; small tributary on Westside Road 2 miles N of Paradise Falls Creek at Stevens Canyon Road, 06-Jul-99, 1 female; Road, 17-Aug-99, 1 male; creek behind Mystic Lake tributary to Ohanapecosh River, 07-Jul-99, 1 female; Falls Ranger Station, 04-Sep-99, 2 males, 3 females; Falls Creek Creek at Stevens Canyon Road, 25-Jun-00, 1 female; at Stevens Canyon Road, 25-Jun-00, 9 males, 4 females; small stream near Ohanapecosh Visitor Center, 16-May- Paradise River in Paradise Valley, 10-Jul-00, 6 males; St. 01, 1 male. Andrews Creek at Westside Road, 14-Jul-00, 3 males, 2 females; Falls Creek at Stevens Canyon Road, 16-Jul-00, 1 Genus Paraperla male, 3 females; Paradise River in Paradise Valley, 02-Aug- Paraperla frontalis 00, 5 females; Olallie Creek at Olallie Creek Camp, 03- Ohanapecosh Ranger Station, 22-Jun-96, 1 female; Car- Aug-00, 1 female; boggy stream tributary to Tahoma bon River Entrance, 27-May-97, 2 females; Ohanapecosh Creek, 07-May-01, 1 male, 2 females; small stream near River at the Grove of the Patriarchs, 28-May-97, 1 male; Ohanapecosh Visitor Center, 16-May-01, 1 male; Ohana- White River Highway 410, 29-May-97, 1 male. pecosh River 0.5 mile S of Shriner Peak Trailhead, 09- Jun-01, 1 female; boggy stream tributary to Tahoma Creek, Paraperla wilsoni 17-Jun-01, 8 males, 2 females; boggy stream tributary to Tahoma Creek, 04-Jul-01, 1 male; boggy stream tributary 0.15 mile NE of Glacier View Bridge, 16-Apr-95, 1 female; to Tahoma Creek, 09-Jul-01, 2 females. 0.6 mile S of Ipsut Campground, 04-Apr-97, 1 female; Nisqually River at Longmire, 28-May-97, 3 females, 1 Yoraperla siletz nymph; Panther Creek at Highway 123, 09-Jun-01, 1 male. Falls Creek at Stevens Canyon Road, 06-Jul-99, 9 males, Family Peltoperlidae 11 females; Falls Creek at Stevens Canyon Road, 08-Jul- Genus Soliperla 99, 5 males, 5 females; small tributary on Westside Road 2 miles N of Paradise Road, 08-Jul-99, 1 male, 1 female; Soliperla fenderi Falls Creek at Stevens Canyon Road, 25-Jun-00, 2 males, Tributary to Tahoma Creek, 08-Jul-99, 4 males, 1 female; 2 females. small tributary on Westside Road 2 miles N of Paradise Road, 08-Jul-99, 1 female; Falls Creek at Stevens Canyon Family Perlidae Road, 08-Jul-99, 1 male, 1 female; small tributary on Genus Doroneuria Westside Road 2 miles N of Paradise Road, 17-Aug-99, 4 males, 1 female; boggy stream tributary to Tahoma Creek, Doroneuria baumanni 17-Aug-99, 1 female; stream on Westside Road 1.5 miles Carbon River Entrance at June Creek, 13-Jul-95, 2 nymphs; from Paradise Road, 26-Jun-00, 1 male; Cataract Creek at tributary to Tahoma Creek at Paradise Road, 08-Jul-99, 1 Wonderland Trail, 09-Jul-00, 1 male; Dick Creek at Won- female; Carbon River Entrance, 15-Aug-99, 5 males, 1 derland Trail, 27-Jul-01, 4 males. female.

Genus Yoraperla Genus Hesperoperla Yoraperla mariana Hesperoperla pacifica Ohanapecosh Campground, 28-May-97, 1 male, 4 females; Ohanapecosh Ranger Station, 17-Jul-97, 1 female. Panther Creek at Highway 123, 29-May-97, 2 males, 1 female; Ohanapecosh River at Sheep Creek, 29-May-97, 1 Genus Claassenia male, 1 female; unnamed stream N of Crystal Creek, Claassenia sabulosa Highway 410, 29-May-97, 9 males, 7 females; Mountain Meadow Stream at Paul Peak Trail, 07-May-98, 1 male; Ohanapecosh Ranger Station, 09-Sep-97, 1 female. 2002] MOUNT RAINIER NATIONAL PARK STONEFLIES 403

Family Perlodidae Bridge, 06-Jul-99, 1 male, 2 females; Fryingpan Creek at Genus Isoperla Sunrise Road Bridge, 16-Aug-99, 1 female; Cataract Creek Isoperla bifurcata at Wonderland Trail, 09-Jul-00, 1 female; Fryingpan Creek at Sunrise Road Bridge, 10-Jul-00, 2 females; Nisqually Carbon River Entrance, 15-Aug-99, 1 female. River at Longmire, 13-May-01, 1 male, 1 female; seeps into White River on Highway 410, 14-May-01, 9 males, 2 Isoperla fusca females; Paradise River above Narada Falls, 02-Jul-01, 1 fe- Seeps into White River on Highway 410, 29-May-97, 1 male; Cataract Creek at Wonderland Trail, 20-Jul-01, 1 fe- nymph. male; Puyallup River at Wonderland Trail, 28-Jul-01, 1 male.

Isoperla gravitans Megarcys subtruncata Carbon River Entrance, Rainforest Loop Trail, 08-May- Deer Creek on Highway 123, 27-May-97, 2 males; Ohana- 95, 1 male; Falls Creek at Stevens Canyon Road, 25-Jun- pecosh River at the Grove of the Patriarchs, 28-May-97, 1 00, 1 male. male, 1 female; Goat Creek on Highway 410, 29-May-97, 4 males, 1 female; Panther Creek at Highway 123, 29- Isoperla petersoni May-97, 1 male; Carbon River at Chenuis Falls Trailhead, 06-Jun-97, 2 females; Ipsut Creek at Carbon River Road Fryingpan Creek at Sunrise Road Bridge, 16-Jul-00, 3 Bridge, 06-Jul-97, 2 females; Glacier Basin, 20-Jul-97, 4 males, 1 female. females; Glacier Basin, 21-Jul-97, 4 females; Glacier Basin, 27-Jul-97, 2 females; Fryingpan Creek at Sunrise Road Isoperla rainiera Bridge, 06-Jul-99,1 male, 1 female; Paradise River above White River Campground, 23-Jul-97, 1 female; Nisqually Narada Falls, 16-Aug-99, 1 female; Paradise River in Para- River at Glacier Vista Bridge, 07-Jul-99, 1 female; Paradise dise Valley, 16-Aug-99, 3 females; Fryingpan Creek at River above Narada Falls, 16-Aug-99, 2 males, 1 female; Sunrise Road Bridge, 16-Aug-99, 1 male, 2 females; Falls Paradise River in Paradise Valley, 02-Aug-00, 16 males, 2 Creek at Stevens Canyon Road, 25-Jun-00, 2 males; Frying- females; Fryingpan Creek at Wonderland Trail, 31-Oct-00, pan Creek at Sunrise Road Bridge, 29-Jun-00, 1 female; 1 female. Huckleberry Creek Bridge, 14-May-01, 2 males; Long- mire, 15-May-01, 1 male; Ohanapecosh Campground, 16- Isoperla sobria May-01, 1 male; Panther Creek at Highway 123, 09-Jun- 01, 1 male, 2 females; boggy stream tributary to Tahoma Fryingpan Creek at Sunrise Road Bridge, 16-Jul-00, 1 Creek, 17-Jun-01, 1 male. male, 1 female; White River Highway 410, 0.5 mile S of park boundary, 16-May-01, 1 male. Megarcys yosemite Isoperla sordida Fryingpan Creek at Sunrise Road Bridge, 16-Aug-99, 1 male; Fryingpan Creek at Sunrise Road Bridge, 16-Jul-00, Fryingpan Creek at Wonderland Trail, 03-Jul-98, 1 female; 4 females. Longmire, 08-Sep-98, 1 female; stream into White River at Highway 410 N of White River entrance, 07-Jul-99, 3 Genus Setvena males; small tributary on Westside Road 2 miles N of Par- adise Road, 08-Jul-99, 1 male; boggy stream tributary to Setvena tibialis Tahoma Creek, 17-Aug-99, 1 female; Fryingpan Creek at Mowich Road at Grindstone Trail Crossing, 07-Jul-95, 1 Sunrise Road Bridge, 05-Sep-99, 1 female; Paradise River female; Paradise Ice Caves, 20-Aug-95, 1 male, 1 female; in Paradise Valley, 07-Sep-99, 2 males; Fryingpan Creek at St. Andrews Creek at Westside Road, 24-Jul-97, 2 females; Sunrise Road Bridge, 13-Sep-99, 1 male; North Puyallup Paradise Ice Caves, 13-Sep-97, 1 female; Sunrise Visitor River at Wonderland Trail, 24-Sep-00, 2 males, 32 females. Center, 13-Aug-98, 1 male; Falls Creek at Stevens Canyon Road, 06-Jul-99, 3 males, 1 female; Paradise River in Par- Isoperla tilasqua adise Valley, 16-Aug-99, 2 males; Paradise River at Fourth Stream into White River at Highway 410 N of White Crossing, 16-Aug-99, 1 male; Falls Creek at Stevens Can- River Entrance, 07-Jul-99, 3 males; tributary to Tahoma yon Road, 16-Aug-99, 1 female; small tributary on Westside Creek at Paradise Road, 08-Jul-99, 2 males, 3 females; Road 2 miles N of Paradise Road, 17-Aug-99, 1 female; boggy stream tributary to Tahoma Creek, 17-Aug-99, 1 Paradise River in Paradise Valley, 10-Jul-00, 1 male; Par- female. adise River at Fourth Crossing, 23-Jul-00, 1 male.

Genus Frisonia Genus Skwala Frisonia picticeps Skwala americana Muir Snowfield at 8000 ft, 28-Jul-95, 1 female. Ohanapecosh Ranger Station, 05-May-96, 1 female; Ohana- pecosh Campground, 28-May-97, 2 females; Ohanapecosh Genus Megarcys Hot Springs, 28-May-97, 1 female; Ohanapecosh Ranger Station, 28-May-97, 2 males. Megarcys irregularis Carbon River Entrance, 27-May-97, 8 males, 3 females; Genus Kogotus Nisqually River at Longmire, 28-May-97, 2 males, 1 Kogotus nonus female; White River Highway 410, 29-May-97, 8 males, 1 female; Tahoma Creek at Nisqually to Paradise Road, 30- Ipsut Creek at Carbon River Road Bridge, 08-Sep-97, 1 May-97, 2 females; Fryingpan Creek at Sunrise Road male; Carbon River Entrance, 15-Aug-99, 1 male, 2 females; 404 WESTERN NORTH AMERICAN NATURALIST [Volume 62

Fryingpan Creek at Sunrise Road Bridge, 16-Jul-00, 1 female.

Family Pteronarcyidae Genus Pteronarcys Pteronarcys princeps Stevens Canyon Entrance, 15-May-95, 1 female; Panther Creek at Highway 123, 29-May-97, 1 male; small stream near Ohanapecosh Visitor Center, 16-May-01, 3 nymphs.