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Acta Biologica Benrodis Band 13/2006 Band 13 2006 Acta Biologica Benrodis und Mitteilungen aus dem Museum für Naturkunde in der Stiftung Schloss und Park Benrath ISSN 0177-9214 Acta Biologica Benrodis Acta Biologica Benrodis und Mitteilungen aus dem Museum für Naturkunde in der Stiftung Schloss und Park Benrath Band 13 – 2006 Gedruckt mit Unterstützung der Stiftung Schloss und Park Benrath sowie dem Freundeskreis Museum für Naturkunde Benrath e.V. Acta Biologica Benrodis 13 (2006) Acta Biologica Benrodis + Mitteilungen aus dem Museum für Naturkunde in der Stiftung Schloss und Park Benrath Herausgeber Prof Dr. H. Greven Institut für Zoomorphologie und Zellbiologie der Heinrich-Heine-Universität Düsseldorf Universitätsstr.1 D-40225 Düsseldorf Dr. H. Heuwinkel Prof. Dr. R. Wittig Museum für Naturkunde Botanisches Institut der Stiftung Schloss und Park Benrath Johann Wolfgang Goethe-Universität Benrather Schloßallee 102 Siesmayerstr. 70 D-40597 Düsseldorf D-60323 Frankfurt am Main Schriftleitung Prof. Dr. H. Greven Institut für Zoomorphologie und Zellbiologie der Heinrich-Heine-Universität Düsseldorf Universitätsstr. 1 D-40252 Düsseldorf [email protected] Produktion und Satz: Verlag Natur & Wissenschaft, Postfach 170209, D-42624 Solingen ISBN 978-3-936616-51-4 Acta Biologica Benrodis erscheint einmal im Jahr mit ca. 120 Seiten Umfang pro Heft und kann vom Museum für Naturkunde in der Stiftung Schloss und Park Benrath, Benrather Schlossallee 102, Schloss Benrath, D-40597 Düsseldorf, oder vom Verlag zum Abonnementspreis von 15,00 Euro (10,00 Euro für Studenten) pro Jahrgang (zuzüglich Versandkosten) oder im wissenschaftlichen Zeitschriftenaustausch bezogen werden. Titelfoto: Warnender Dreifarbglanzstar in einer Akazie. Aus dem Beitrag von H. Heuwinkel und U. Beigel-Heuwinkel (S. 249-252). Cover photo: Alarm calling Superb Starling in an Acacia. From the article of H. Heuwinkel und U. Beigel-Heuwinkel (S. 249-252). Inhalt Band 13 – 2006 STEVENS, M., WALDMANN, G.: Endemics of the Lesser Antillean Island of Montserrat, British West Indies/Endemiten der Insel Montserrat, Kleine Antillen (Britisch Westindien) ...............................................................................1 FUCHS, R., KUTZELNIGG, H., KEIL, P.: 20 Jahre Entwicklung der Flora und Vegetation im NSG „Steinbruch Hofermühle-Süd“ (Gemeinde Heiligenhaus, Kreis Mettmann)/20 years development of flora and vegetation in the nature conservation area „Limestone-Quarry Hofermühle- Süd“ (community Heiligenhaus, Mettmann district) ....................................................23 DIETRICH, W.: Pflanzliche Neubürger (Neophyten) im Raum Düsseldorf/Neuss (Deutschland, Nordrhein-Westfalen)/New plant species (neophytes) in the Düsseldorf/Neuss area (Germany, North Rhine-Westphalia) ...............................51 FUCHS, R., KUTZELNIGG, H., FEIGE, G. B.: Seminatural ancient forest in urban agglomeration area „Ruhrgebiet“/Naturnahe Altwälder im Ballungsraum Ruhrgebiet ................................................................................................91 WELTJE, L., OEHLMANN, J.: Effects of endocrine disrupting compounds and temperature on the moulting frequency of the freshwater isopod Asellus aquaticus L. (Isopoda: Asellota)/Effekte von hormonähnlich wirksamen Substanzen und Temperatur auf die Häutungsfrequenz der Süßwasserassel Asellus aquaticus L. (Isopoda: Asellota) ........................................105 DIAS, A.N., HASSALL, M.: The abundance, life history and reproduction traits of peracarid macro-decomposers in a Ria Formosa lagoon salt marsh, Southern Portugal/Abundanz, “life history” und Charakteristika der Fortpflanzung von Makrozersetzern (Peracarida) in einer Salzwiese (Ria Formosa Lagune, Südportugal) ............................................................................117 ZIMMER, M.: The role of animal-microbe interactions in isopod ecology and evolution/Die Bedeutung tierlich-mikrobieller Wechselwirkungen für die Ökologie und Evolution von Asseln ...............................................................127 BEYL, F.E.: Morphometrische Untersuchungen am Gemüsespargel, Asparagus officinalis L./Morphometric studies in branches of Common Asparagus, Asparagus officinalis L. ...............................................................169 KREUELS, M.: Die Webspinnen (Arachnida: Araneae) aus Beifängen des NSG Teverener Heide (NRW, Kreis Heinsberg)/The spiders (Arachnidaa: Araneae) from bycatches of the nature reserve Teverener Heath (North Rhine-Westphalia, district Heinsberg) ..........................................................................185 Acta Biologica Benrodis 13 (2006) GREVEN, H., JÖMANN, N., CLEMEN, G.: Dentigerous bones, dentition and dental laminae in the hynobiid salamander Ranodon sibiricus Kessler, 1866 (Amphibia, Urodela)/Zahntragende Knochen, Bezahnung und Zahnleisten bei dem Hynobiiden Ranodon sibiricus Kessler, 1866 (Amphibia, Urodela) ..............................................................................195 DASHDAMIROW, S.: A new genus and species of false scorpion from Vietnam showing remarkable chelal modifications (Archnida: Chelonetida) / Eine neue Gattung und Art von Pseudoskorpionen aus Vietnam mit bemerkenswerten Modifikationen der Chelae (Arachnida: Chelonethida) .............................................................................................219 TIEMANN, I., REHKÄMPER, G.: Rasseunterschiede bei Haushühnern (Gallus gallus f. d.) bei klassischer und operanter Konditionierung in der Skinner-Box/Breedspecific differences among domesticated chickens (Gallus gallus f. d.) revealed by classical and operant conditioning in the Skinner-Box ....................................................................................................................231 Kurze Mitteilungen/Short communications SCHÜTT, H.: Wie kam die Nördliche Grundwasserschnecke Bythiospeum acicula moenanum (Hydrobiidae) an den Niederhein?/How came the Northern Groundwater Snail Bythiospeum acicula moenanum (Hydrobiidae) to the Lower Rhine? ..............................................................................241 KRECHEL, R., BRAUN, TH., STEVENS, M.: Erstnachweis des Springfrosches (Rana dalmatina Bonaparte, 1840) im Regierungsbezirk Düsseldorf/ First record of the Agile Frog (Rana dalmatina Bonaparte, 1840) in the administrative district of Düsseldorf ...........................................................................245 HEUWINKEL, H., BEIGEL-HEUWINKEL, U.: Alarmrufe von Dreifarbglanzstaren (Lamprotornis superbus) bei Entdeckung eines Bodenfeindes/Alarm calls of Superb Starlings (Lamprotornis superbus) detecting a ground predator ...................249 Buchbesprechungen ............................................22, 50, 90, 116, 126, 194, 218, 230, 244, 253 Mitteilungen aus dem Museum für Naturkunde in der Stiftung Schloss und Park Benrath ...........................................................................................M1 Endemics of the Lesser Antillean Island of Montserrat, British West Indies 1 Acta Biologica Benrodis 13 (2006): 1-21 Endemics of the Lesser Antillean Island of Montserrat, British West Indies Endemiten der Insel Montserrat, Kleine Antillen (Britisch Westindien) MICHAEL STEVENS1 & GEORG WALDMANN2 1 Institut für Zoomorphologie und Zellbiologie, Heinrich-Heine-Universität Düsseldorf, Universitätsstraße 1, D-40225 Düsseldorf, Germany; [email protected] 2 Am Trietenbroich 24, D-41352 Korschenbroich, Germany; [email protected] Summary: Montserrat (British West Indies, Lesser Antilles), is part of the Caribbean biodiversity hotspot region with numerous endemic species including the following families: Lauraceae, Pipe- raceae, Theaceae, Marcgraviaceae, Clusiaceae, Elaeocarpaceae, Begoniaceae, Styracaceae, Melasto- mataceae, Rubiaceae, Cyclanthaceae, Bromeliacae, Agavaceae, Hesperiidae, Leptodactylidae, Gek- konidae, Polychrotidae, Teiidae, Psittacidae (fossil only), Trochilidae, Mimidae, Icteridae, Criceti- dae (fossil only) and Phyllostomidae. Natural hazards like hurricanes and recent volcanism probab- ly affect the survival of endemic species on Montserrat. Lesser Antilles, Montserrat, single-island endemics, regional island endemics Zusammenfassung: Die Karibikinsel Montserrat (Kleine Antillen; Britisch Westindien) liegt inmitten eines Zentrums globaler Biodiversität und beherbergt zahlreiche lokale und regionale Endemiten aus folgenden Familien: Lauraceae, Piperaceae, Theaceae, Marcgraviaceae, Clusiaceae, Elaeocarpaceae, Begoniaceae, Styracaceae, Melastomataceae, Rubiaceae, Cyclanthaceae, Bromeli- acae, Agavaceae, Hesperiidae, Leptodactylidae, Gekkonidae, Polychrotidae, Teiidae, Psittacidae (nur fossil), Trochilidae, Mimidae, Icteridae, Cricetidae (nur fossil) und Phyllostomidae. Naturer- eignisse wie tropische Wirbelstürme oder rezenter Vulkanismus beeinflussen möglicherweise das Überleben von Endemiten auf Montserrat. Kleine Antillen, Montserrat, Endemiten, regionale Endemiten 1. Introduction land endemics. Regional endemics currently known, with a distribution on the Lesser The Caribbean Region is among the eight Antillean islands, in some cases reaching the leading world biodiversity hotspots. More Virgin Islands and Puerto Rico or Trinidad than 12,000 vascular plants and about 1,500 and Tobago are included. vertebrates (excluding teleosts) have been described from the Caribbean (DAVIS et al. 2. The Lesser Antilles and the island of 1997; MYERS et al. 2000). Comprising only Montserrat 0.018 % of the Earth’s land surface, some 2.3 % of all higher plants and 2.9
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