Gekkotan Lizard Taxonomy
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3% 5% 2% 4% 3% 5% H 2% 4% A M A D R Y 3% 5% A GEKKOTAN LIZARD TAXONOMY 2% 4% D ARNOLD G. KLUGE V O 3% 5% L 2% 4% 26 NO.1 3% 5% 2% 4% 3% 5% 2% 4% J A 3% 5% N 2% 4% U A R Y 3% 5% 2 2% 4% 0 0 1 VOL. 26 NO. 1 JANUARY, 2001 3% 5% 2% 4% INSTRUCTIONS TO CONTRIBUTORS Hamadryad publishes original papers dealing with, but not necessarily restricted to, the herpetology of Asia. Re- views of books and major papers are also published. Manuscripts should be only in English and submitted in triplicate (one original and two copies, along with three cop- ies of all tables and figures), printed or typewritten on one side of the paper. Manuscripts can also be submitted as email file attachments. Papers previously published or submitted for publication elsewhere should not be submitted. Final submissions of accepted papers on disks (IBM-compatible only) are desirable. For general style, contributors are requested to examine the current issue of Hamadryad. Authors with access to publication funds are requested to pay US$ 5 or equivalent per printed page of their papers to help defray production costs. Reprints cost Rs. 2.00 or 10 US cents per page inclusive of postage charges, and should be ordered at the time the paper is accepted. Major papers exceeding four pages (double spaced typescript) should contain the following headings: Title, name and address of author (but not titles and affiliations), Abstract, Key Words (five to 10 words), Introduction, Material and Methods, Results, Discussion, Acknowledgements, Literature Cited (only the references cited in the paper). Ap- pendices follow the main paper. Descriptions of new taxa will be considered as major papers regardless of size. Ab- stracts (up to 150 words) should summarize the important findings of the paper and should avoid references. In case of descriptions of new taxa, diagnoses should be provided in the abstract. Special attention should be paid to accents and diacritical marks: if fonts are not available in the software or typewriter, these may be put directly on the hard copy of the manuscript by hand. References should be in the following format: Papers: GAULKE, M. 1994. Notes on the herpetofauna of Panaon and Sámar, East Visayans, Philippines. Hamadryad 19:1-10. Books: RUSSELL, P. 1796. An account of Indian serpents collected on the coast of Coromandel; containing descriptions and drawings of each species; together with experiments and remarks on their several poisons. George Nicol, Lon- don. viii + 90 pp + Pl. I-XLVI. Chapters in books: KIEW, B. H. 1984. Terrestrial vertebrate fauna of Lambir Hills National Park. In: An ecological survey of Lambir Hills National Park, Sarawak. pp: 55-63. E. Soepadmo & K. I. Sudderuddin (Eds). University of Malaya, Kuala Lumpur. Unpublished reports: WILKINSON, J. W., J. W. ARNTZEN & R. S. THORPE. 1995. Amphibian populations on organic farms: their role in pest control. M.A.F.F. Report. iv + 48 pp. Publications in languages other than English: SZCZERBAK, N. N. & M. L. GOLUBEV. 1986. [The gekkonid fauna of the USSR and adjacent countries. ] Sci. Acad. Ukranian SSR Zool. Inst. 1986: 1-232. [In Russian.] Tables should be comprehensive without reference to the text. Tables should follow the main body of the paper, and their desired location within the text indicated on the manuscript. Footnotes are generally not allowed. Line art and half tones are preferred as electronic files saved on compact disks or attached to email files. Black and white photo- graphic prints and slide transparencies are accepted if they are sharp and show good contrast. Black and white photos should be printed on glossy paper at least 152 x 204 mm after being trimmed at right angles and should carry the name of the author and of the manuscript at the back. Illustrations should be preferably done with Indian ink on good quality tracing paper. All lettering should be done using high quality computer printouts or transfers. Illustrations should be planned to fit either one column (6.5 mm) or one page (14 mm) width, after suitable reduction. All manuscripts received for Hamadryad are sent to two or more referees. Authors of papers describing range exten- sions and descriptions of new taxa should deposit their material in established systematic institutions and cite mu- seum registration numbers. Range extensions not based on specimens, such as endangered or other protected species, may be based on photographs that are deposited in a recognized systematic institution. Manuscripts should be sent by registered post to: Indraneil Das, Institute of Biodiversity and Environmental Conser- vation, Universiti Malaysia Sarawak, 94300 Kota Samarahan, Sarawak, Malaysia. Email: [email protected] HAMADRYAD Journal of the Centre for Herpetology, Madras Crocodile Bank Trust EDITOR Indraneil Das AESSOCIATE DITORS Harry V. Andrews & Romulus Whitaker EBDITORIAL OARD Kraig Adler Roger A. Avery Aaron M. Bauer Cornell University Bristol University Villanova University U.S.A. United Kingdom U.S.A. J. C. Daniel Anslem De Silva Alain Dubois Bombay Natural History Society University of Peradeniya Musé e National d’Histoire India Sri Lanka Naturelle, France Sushil K. Dutta John G. Frazier M. S. Khan Utkal University Cinvestav Lauderville, Florida India México U.S.A. Kelvin K. P. Lim Edward O. Moll Peter C. H. Pritchard National University of Singapore Eastern Illinois University Florida Audubon Society Singapore U.S.A. U.S.A. James Perran Ross Lala A. K. Singh Garth Underwood Florida Museum of Natural History Simlipal Tiger Reserve Natural History Museum U.S.A. India United Kingdom LFAYOUT AND ORMAT Luc Gastmans Front cover: File-eared tree frog (Polypedates otilophus ). Adult female that was found at the base of Gunung Buri, Sarawak, East Malaysia (Borneo), next to a shallow rainwater pool. This is a large species, females attaining a snout- vent length of 97 mm, while males reach 80 mm. It occurs on Borneo and Sumatra, the saw-edged bony ridge over the tympanum being diagnostic of the species. © Indraneil Das. (Fujichrome Velvia50 ASA). Back cover: Spiny hill turtle (Heosemys spinosa ). A widespread terrestrial turtle from southern Myanmar, Thailand, the Malay Peninsula, Sumatra, Borneo, the Natunas to the Sulu Archipelago and Mindanao. This partially-burnt individual was found in a patch of kerangas forest, Kilometre 13, Labi Road, Belait District, Brunei Darussalam (Borneo). A length of 225 mm is reached by the species, which has strikingly serrated shell margins as juveniles. © Joseph K. Charles. (Fujichrome Velvia50 ASA). C ENTER FOR H ERPETOLOGY M ADRAS C ROCODILE B ANK T RUST TRUSTEES Harry V. Andrews G. Dattatri Raj K. Gupta David Lobo J. Krishnadev Rao EX-OFFICIO TRUSTEES REGIONAL DEPUTY DIRECTOR JOINT DIRECTOR WILDLIFE WARDEN Wildlife Preservation Tamil Nadu Tamil Nadu Forest Southern Region Tourism Department Department Romulus Whitaker FOUNDER DIRECTOR: Romulus Whitaker JOINT DIRECTOR: Harry V. Andrews SCIENTIFIC OFFICER: Indraneil Das ASSISTANT CURATOR: Nikhil Whitaker SCIENTIFIC RESEARCH COMMITTEE Harry V.Andrews, Bharat Bhushan, Indraneil Das, G. Dattatri, A. K. Kumaraguru and Romulus Whitaker RESEARCH ASSOCIATES Manish Chandi,Ravi Kailas, Madhuri Ramesh and Brian Stacy RESEARCH COLLABORATORS M. Farid Ahsan, Rauf Ali, Christopher Austin, Aaron M. Bauer, Joseph K. Charles, Binod C. Choudhury, Anslem De Silva, John G. Frazier, Walter Gastmans, Maren Gaulke, Fritz Huchzermeyer, M. S. Khan, A. K. Kumaraguru, Harvey B. Lillywhite, Kelum Manamendra-Arachchi, Edward O. Moll, Rohan Pethiyagoda, Lala A. K. Singh and Bruce A. Young ASSOCIATIONS AND OFFICIAL MEMBERSHIPS Amphibia and Reptile Research Organization of IUCN/SSC Captive Breeding Specialist Group Sri Lanka IUCN/SSC Specialist Group on Sustainable Use Andaman Science Association of Wild Species Asiatic Herpetological Research IUCN/SSC Declining Amphibian Populations Bombay Natural History Society Central Zoo Authority of India Task Force Flora and Fauna International International Species Information System Food and Agricultural Organization of the Indian National Trust for Art and Cultural United Nations Heritage Indian Herpetological Society Palni Hills Conservation Council International Trade Centre/UNCTAD Society for Southeast Asian Herpetology IUCN/SSC Crocodile Specialist Group Society for the Study of Amphibians and IUCN/SSC Tortoise and Freshwater Turtle Specialist Group Reptiles IUCN/SSC Marine Turtle Specialist Group Wildlife Heritage Trust of Sri Lanka IUCN/SSC South Asian Reptile and Amphibian World Congress of Herpetology Specialist Group World Wide Fund for Nature HAMADRYAD Journal of the Centre for Herpetology Subscription form for 2001/2002 Name: Address: City/State/Country: Fax/Phone/Email: Annual Subscription (inclusive of air mail postage for foreign members) Indian* and other SAARC (individuals): Indian Rs. 150.00 (Bangladesh, Bhutan, India, Maldives, Nepal, Pakistan and Sri Lanka) Indian* and other SAARC (institutions): Indian Rs. 250.00 (Bangladesh, Bhutan, India, Maldives, Nepal, Pakistan and Sri Lanka) ASEAN**: US$ 20.00 (Brunei, Indonesia, Malaysia, Myanmar, Philippines, Singapore, Thailand and Vietnam) All other countries (individuals)**: US$ 35.00 All other countries (institutions)**: US$ 50.00 Donations: The Centre for Herpetology invites donations to help defray production costs of Hamadryad and to support other herpetological activities * For outstation cheques within India, please add Rs. 20.00. ** Foreign subscribers can pay in their currency via a personal cheque. Non-US subscribers can pay equivalent in their currency. Space for communication: Cheques or Money Orders made out in favour of “Madras Crocodile Bank Trust” should be sent to: Centre for Herpetology Madras Crocodile Bank Trust Post Bag 4 Mamallapuram 603 104 Tamil Nadu India HAMADRYAD Vol. 25. No. 2. December, 2000 Date of issue: 1 December, 2000 ISSN 0972-205X CONTENTS S. K. DUTTA,M.F.AHMED & I. DAS. Kalophrynus, a new genus for India, with the description of a new species, K. orangensis, from Assam State .......................67–74 M. VENCES,S.WANKE,G.ODIERNA,J.KOSUCH & M.