Coniacian andSantonian belemnite faunasfrom Bornholm, De k
Walter Kegel Christensenand Max-Gotthard Se ulz
Santonianto Maastrichtian ammonit s from Scania, south.em weden
William James Kennedy andWalter Kegel Christensen
SCANDINAVIAN UNI VERSITYPRESS
r-I�II Oslo � Copenhagen � Stockholm � Boston FOSSILS AND STRATA
An international monograph series of palaeontology and stratigraphy
Owner Lethaia Foundation, Oslo. Administrative Council: Hans Jørgen Hansen, Copenhagen, David Bruton, Oslo, Christina Franzen, Stockholm, Stefan Bengtson, Uppsala.
Editor Stefan Bengtson, Dept. Palaeozoology, Swedish Museum of Natural History, Box 5000?, S-104 05 Stockholm, Sweden; tel. +46-8 666 4220; fax +46-8 666 4184; e-mail [email protected].
Publisher Scandinavian University Press, P.O. Box 2959, Tøyen, N-0608 Oslo 6, Norway.
Programme Fossils and Strata is an international series of monographs and memoirs in palaeontology and stratigraphy, published in cooperation between the Scandinavian countries. It is issued in Numbers with individual pagination. Fossils and Strata forms part of the same structured publishingprogramme as the journals Lethaia and Boreas. These two journals are fully international and accept papers within their respective sectors of sciencewithout nationallimitations or preferences. Fossils and Strata, however, is an outlet for more comprehensive systematic and regional monographs emanating primarily from the five countries of Norden. Contributions from other countries may also be included if this series is deemed appropriate with regard to distribution and availability. Articles can normally only be accepted if they are heavily subsidized by the national Research Council in their country of origin or by other funds. All income is re-invested in forthcoming numbers of the series. Although articles in German and French may be accepted, the use of English is stronglypreferred. An English abstract should always be provided, and non-English articles should have English versions of the figure captions. Abstracts or summaries in one or more additional languages may be added. Many regional or systematic descriptions and revisions contain a nucleus of results which are of immediate and general interest in international palaeontology and stratigraphy. It is expected that authors of such papers will to some extent duplicate their publication in the form of an article for a journal, in the first place Lethaia or Boreas.
Sales Individual numbers and standing subscriptions may be ordered from Scandinavian University Press (postal address as above; e-mail address [email protected]). Prices (subject to revision) are listed on the back side of each issue. IPA members generally have a 50% discount on older issues (ask for information fromScandinavian University Press). Allprices exclude postage and handling.
LET HAlA
An international journal of palaeontology and stratigraphy An international journal of Quaternary research
Scandinavian University Press, p.o. Box 2959, Tøyen, N-0608 Oslo 6, Norway Santonian to Maastrichtian ammonites from Scania, southern Sweden
WILLIAM JAMES KENNEDY AND WALTER KEGEL CHRISTENSEN
Kennedy, W.). & Christensen, W.K. 1997 08 22: Santonian to Maastrichtian ammonites from Scania, southern Sweden. Fossils and Strata, No. 44, pp. 75-128. Oslo. ISSN 0300-9491. ISBN 82-00-37695-8.
Eighteen genera of ammonites are recorded from the Santonian to Maastrichtian rocks of Scania in southern Sweden. Ammonites are fairly com mon in the Vomb Trough, rare in the Kristianstad Basin and not recorded from the Båstad Basin. The chronostratigraphical assign ment of the outcrops on ammonite and be!emnite evidence is in agreement. DAmmonites, Upper Cretaceous, Santonian, Campanian, Maastrichtian, Scania, southern Sweden.
W. James Kennedy, UniversityMuseum Oxford, Parks Road, Oxfo rd OX1 3PW, UK; Walter K. Christensen [[email protected], Geological Museum, University of Copenhagen, 0ster Voldgade 5-7, DK-1350 Copenhagen, Denmark; 6th September, 1995; revised 28th August, 1996.
Contents Introduction ...... 77 Malmo area ...... 83 Geological setting ...... 77 Limhamn pit ...... 83 Stratigraphy ...... 78 Ulrice!und near Nasbyholm ...... 83 Framework ...... 78 Deep boreholes ...... 83 The conventional' belemnite zonation ...... 78 Systematic palaeontology...... 83 The subdivision of the Upper Cretaceous of Sweden ...... 78 Location of specimens ...... 83 Locality details ...... 80 Order Ammonoidea Zitte!, 1884; Suborder Lytoceratina Vomb Trough ...... 80 Hyatt, 1889; Superfamily Tetragonitaceae Hyatt, Eriksdal ...... 81 1900; Family Gaudryceratidae Spath, 1927 ...... 83 Kåseberga ...... 81 Genus Gaudryceras de Grossouvre, 1894 ...... 83 Kopinge district ...... 81 Gaudrycerassp ...... 83 Kullemolla-Lyckås ...... 81 Family Tetragonitidae Hyatt, 1900 ...... 84 Rodmolla-Tosterup ...... 81 Genus Saghalinites Wright & Matsumoto, 1954 ...... 84 Kopingsberg- l borehole ...... 81 Saghalinites sp ...... 84 Kristianstad Basin ...... 82 Genus Pseudophyllites Kossmat, 1895 ...... 85 Åhus ...... 82 Pseudophyllites indra (Forbes, 1846) ...... 85 Balsberg ...... 82 Suborder Ammonitina Hyatt, 1889; Superfamily Bjarnum ...... 82 Desmocerataceae Zitle!, 1895; Family Desmoceratidae Ignaberga new quarry ...... 82 Zittel, 1895; Subfamily Puzosiinae Spath, 1922 ...... 85 Ivo Klack (=Blaksudden, Ivobrottet) ...... 83 Genus and subgenus Hauericeras de Grossouvre, 1894 ...... 85 Hauericeras (Hauericeras) pseudogardeni (Schluter, 1872) .....85 Glyptoxoceras crispatum (Moberg, 1885) ...... 107 Family Pachydiscidae Spath, 1922 ...... 88 Family Baculitidae Gill, 1871 ...... 108 Genus and subgenus Pachydiscus Zittel, 1884 ...... 88 Genus Baculites Lamarck, 1799 ...... 108 Pachydiscus (Pachydiscus) colligatus (Binkhorst, 1861) ...... 88 Baculites suecicus Moberg, 1885 ...... 108 Pachydiscus (Pachydiscus) haldemsis (Schluter, 1867) ...... 92 Baculites incurvatus Moberg, 1885 ...... 108 Pachydiscus (Pachydiscus) cf. subrobustus Baculites brevicosta? Moberg, 1885 ...... 109 Seunes, 1892 ...... 94 Baculites sp. Moberg, 1885 ...... 109 Genus Patagiosites Spath, 1953 ...... 95 Baculites cf. aquilaensis Reeside, 1927 ...... 110
Patagiosites stobaei (Nilsson, 1827) ...... •...•...•.••...... 95 Baculites vertebralis Moberg, 1885 ...... 110 Genus Nowakites Spath, 1922 ...... 102 Baculites angustus Moberg, 1885 ...... 110 Nowakites hernensis (Schluter, 1867) ...... 102 Baculites schlueteri Moberg, 1885 ...... 111 Family Muniericeratidae Wright, 1952 ...... 104 Baculites knorrianus (Desmarest, 1817) ...... III
Genus Tragodesmoceras Spath, 1922 ...... 104 Baculites spp ...... 112 Tragodesmoceras clypeale (Schltiter, 1867) ...... 104 Aptychi of Baculites ...... 112 Superfamily Hoplitaceae H. DouvilIe, 1890; Family Superfamily Scaphitaceae GiB, 1871: Family Scaphitidae Placenticeratidae Meek, 1876 ...... 104 Gill, 1871; Subfamily Scaphitinae Gill, 1871 ...... 115 Genus and subgenus Hoplitoplacenticeras Paulcke, 1907 ...... 104 Genus and subgenus Scaphites Parkinson, 1811 ...... 115 Hoplitoplacenticeras (Hoplitoplacenticeras) cf. Scaphites (Scaphites) hippocrepis (DeKay, 1828) form III coesfeldiense (Schltiter, 1867) ...... 104 of Cobban, 1969 ...... 115 Suborder Ancyloceratina Wiedmann, 1966; Superfamily Scaphites (Scaphites) binodosus Roemer, 1841 ...... 115
Turrilitaceae Gill, 1871; Family Nostoceratidae Scaphites (Scaphites) spp ...... 116 Hyatt, 1894 ...... 104 Genus Trachyscaphites Cobban & Scott, 1964 ...... 116 Genus BostrychocerasHyatt, 1900 ...... 104 Trachyscaphites spiniger spiniger (Schluter, 1872) ...... 116 Bostrychoceraspoly plocum (Roemer, 1841) ...... 104 Genus Hoploscaphites Nowak, 1911 ...... 121 Genus Nostoceras Hyatt, 1894 ...... 105 Hoploscaphites ikorfatensis Birkelund, 1965 ...... 121 Nostoceras junior (Moberg, 1885) ...... 105 Hoploscaphites constrictus (J. Sowerby, 1817) ...... 123 Family Diplomoceratidae Spath, 1926; Subfamily Hoploscaphites tenuistriatus (Kner, 1848) ...... 124 Diplomoceratinae Spath, 1926 ...... 106 Genus Acanthoscaphites Nowak, 1911 ...... 124
Genus LewyitesMatsumoto & Miyauchi, 1984 ...... 106 Acanthoscaphites sp ...... 124
Lewyites elegans (Moberg, 1885) ...... 106 Acknowledgements ...... 124
Genus Glyptoxoceras Spath, 1925 ...... 107 References ...... 125 Introduction
More than a century has passed since Moberg (1885) Geological setting monographed the Santonian to Maastrichtian ammonites of southern Sweden. He described 21 species and two The block-faulted Fennoscandian Border Zone acted as a types of aptychus, the majority of which came from the transition zone between the stable Fennoscandian Shield Vomb Trough. and the subsiding Danish Subbasin during the Permian As early as the 1650s, the Danish polyhistor Ole Worm and Mesozoic (Fig. 1). (1655) figured an ammonite fromthe Upper Cretaceous Upper Lower and Upper Cretaceous deposits occur in ofScania. Stobaeus (1732, Pl. 1:7-9) figuredan ammonite Sweden in the Båstad and Kristianstad Basins north east of as Cornu Ammonis (=Ammonites stobaei Nilsson, the border zone, in the Vomb Trough within the border 1827=Patagiosites stobaei herein). In the early part of the zone, and in the Malmo area southwest of the border zone 19th century, ammonites were dealt with by Wahlenberg (Fig. 1). The latter area belongs to the Danish Subbasin (1818) and Nilsson (1826, 1827), and subsequently by Schliiter (1870, 1871-1876), Lundgren (1881), Stolley (1896, 1897, 1930), Hagg (1930, 1935, 1943, 1947, 1954), Lundegren (1933, 1935), Brotzen (1945, 1958), 0dum (1953), Birkelund (1973) and Birkelund & Bromley l2iSI Upper Cretaceous (1979). Recently, Santonian ammonites from the ® To wn
Kopingsberg-l borehole in the Vomb Trough were o Locality Borehole described by Kennedy & Christensen (1993) and those • � fromthe uppermost Maastrichtian of the Limhamn pit by Birkelund (1993). FENNOSCANDIAN SHIELD Ammonites are fairly common in the Vomb Trough, rare in the Kristianstad Basin and not recorded from the Båstad Basin. A few ammonites are recorded from the Malmo area. A single specimen referred to Lewesiceras woodi Wright, 1979 (p. 312) (=Pseudopuzosia sp. of Bir kelund, 1973, Pl. 12: 1), an U pp er Turonian species, is known fromSa rdal on the Swedish west coast. Christensen (1973, 1975, 1986, 1993) described the belemnites from the temporary outcrop at Sardal, the Kristianstad and Båstad Basins, in addition to the Vomb Trough, and placed the localities in the international stratigraphical framework on belemnite evidence. Chris tensen (1985) reviewed the Albian to Maastrichtian of southern Sweden and Bornholm. The aim of the present paper is to revise the Santonian to Maastrichtian ammonite faun as of Scania in southern Fig. 1. Map of southern Sweden, showing the distribution of the Upper Sweden. Most of the specimens described here were col Cretaceous sedimentary rocks in the Båstad Basin, Kristianstad Basin, Vomb Trough and Malmiiarea, boreholes and the approximate location lected in the last part of the previous century and the of the Fennoscandian Border Zone. Modified fr om Kennedy & Chris beginning of this century. tensen (1993). 78 W,J. Kennedy & w.K. Christensen FOSSILS AND STRATA 44 (1997) from a tectonic and sedimentological point of view. in the B. mucronata group owing to its size, surface mark Upper Cretaceous deposits are also recorded fromSa rdal ings, and internal characters. Moreover, Christensen rec on the Swedish west coast north of Båstad, Province of ognized three chronological subspecies of B. minor, which Halland, and from two boreholes in Hano Bay. form an evolutionary line age. B. minor I and B. minor Il Biocalcarenites and glauconitic sands tones predomi Christensen, 1995, predominate in the upper Upper nate in the basins northeast of the border zone, and the Campanian of Norfolk, whereas B. minor III Christensen, thickness of the strata is less than 250 m. The sedimentary 1995, occurs rarely in the lower Lower Maastrichtian. deposits of the Vomb Trough are characterized by detrital The concept of B. langei Jeletzky is open to question, clastic sediments - glauconitic, calcareous and clayey silt because it is understood only in the sense of the holotype sto nes - and the thickness is a little less than 800 m. Lime and paratype. Thus, the concept differs from one author stones, including chalk and marly chaik, prevail in the to another. For instance, B. langei sensu Birkelund (1957) Malmo area, where the thickness is about 1650 m. and B. langei sensu Schulz (1978) are not conspecificwith B. langei o[Jeletzky, 1948. Christensen (1995) offered a Belemnitella zonation for Stratigraphy the Upper Campanian and lower Lower Maaastrichtian of Norfolk, including five informal zones, in as cen ding Framework. - The Santonian to Maastrichtian of Sweden order, the B. mucronata and B. woodi zones in the lower is of markedly boreal aspect and can be calibrated against part of the Upper Campanian, the B. minor I and B. minor the standard successions of northwest Germany, which Il zones in the upper part of the Upper Campanian, and have been carefully studied during the last fo rty years by the B. minor III zone in the lower Lower Maastrichtian. numerous workers (see reviews by Ernst et al. 1979; The zones were regarded as informal, because the bases of Schulz et al. 1984; and Schulz 1996). The Coniacian to most of them cannot be precisely defined,owing to strati Maastrichtian white chaik of Lagerdorf, Kronsmoor and graphical gaps between the exposures. It was noted that Hemmor, northwest Germany, has been subdivided in to further work is necessary to see if these local zones can be 33 faunal zones on the basis of inoceramid bivalves, extended to other are as in Europe. Keutgen (1996) has ammonites, belemnites, echinoids, and crinoids (Schulz shown subsequently that the zonation recognized in Nor et al. 1984; Schulz 1996). The faunal zones of the Santo folk is also applicable in northeast Belgium. nian to lower Lower Maastrichtian provide the strati Schulz (1979) subdivided the Lower Maastrichtian of graphical framework for the present paper (Fig. 2). northwest Germany into six Belemnella zones on the basis of species of two evolutionary lineages of Belemnella: the The conventional belemnite zonation. - The conventional slender subgenus B. (Belemnella) Nowak and the sto ut belemnite zonation of the Santonian to Lower Maastrich subgenus B. (Pachybelemnella) Schulz. tian of northwest Europe is based upon species of the genus Gonioteuthis Bayle fo r the Santonian-Lower Cam The subdivision of the Upp er Cretaceous of Sweden. - panian, species of the genus Belemnitella d'Orbigny fo r When discussing the subdivision of the U pper Cretaceous the uppermost Lower and Upper Campanian, and species ofSweden it should be borne in mind that up to 1930, the of the genus Belemnella Nowak fo r the Lower Maastrich zonation was concerned only with the Santonian to Maas tian (Fig. 2). trichtian. Cenomanian deposits were first recorded from The Santonian and Lower Campanian of northwest the Båstad Basin by Lundegren (1932) and Stolley (1932), Germany is subdivided in to 10 belemnite zones (Ernst and the Kristanstad Basin by Christensen (1970). Brotzen 1964). The conventional Upper Campanian Belemnitella (1945) reported Cenomanian, Turonian and Coniacian zonation includes three interval zones, and these are, in strata from the Hollviken borehoies. ascending order, the B. mucronata Zone in the lower part In the Swedish Santonian to Maastrichtian, the belem of the Upper Campanian and the B. minor and B. langei nites, above all other macrofossils, have been shown to be Zones in the upper part of the Upper Campanian. This of fundamental importance in biostratigraphy and corre zonation was introduced by Jeletzky (1951) and has been lation. Christensen (1975, 1986) reviewed the belemnite us ed subsequently by numerous authors. It was discussed zonation in great detail, and consequently only a brief by Christensen (1995), who argued that the conventional outline is given here. B. minor and B. langei Zones should not be maintained for The first reliable subdivision was made by Schluter the following reasons. The concepts of B. minor Jeletzky (1870), who recognized two belemnite zones, the Belem based on the diagnosis and the holotype are rather differ nitella mucronata Zone above and the Belemnitella sub
ent. In contrast to earlier authors, who interpreted B. ventricosa Zone ( = Belemnellocamax mammillatus Zone) minor on the basis of the diagnosis, Christensen (1995) below. More belemnite zones were added to Schliiter's interpreted this species with respect to its holotype. In this stratigraphical scherne by subsequent authors, including respect, B. minor is a very large species, which was placed Moberg (1885). Stolley (1897, 1930) subdivided the Seno- FOSSILS AND STRATA 44 (1997) Santonian to Maastrichtian ammonites 79
Fig. 2. Stratigraphical scheme of the Santonian to lower Lower ,ei 1) 2) 3) LOCALITIES 4) o C!) BELEMNITE lONAL FAUNAL ZONES, Maastrichtian, showing faunal tjF lONES, BELEMNITES, nian of Germany and Sweden on the basis of belemnites The German Quadraten-Kreide is broadly equivalent to into, from the bottom to top, the Westfalicus-Kreide= the Lower Campanian. Stolley (1897, 1930) subdivided Emscher, Granulaten-Kreide, Quadraten-Kreide = Mam the Mucronaten-Kreide into three zones, mainly on the millaten-Kreide, and Mucronaten-Kreide. This subdivi basis of ammonites. The lower and middle Mucronaten sion was us ed subsequently in Sweden, but translated into Kreide equate with the Upper Campanian, and the upper westfalicuskrita, granulatuskrita, mammillatuskrita, and Mucronaten-Kreide with the Maastrichtian. Stolley corre mucronatakrita. lated the German Quadraten-Kreide with the Swedish 80 W,J. Kennedy & w.K. Christensen FOSSILS AND STRATA 44 (1997) Fig. 3. Map of the Vomb Trough, showing outerops and borehoies. AfterChristensen (1986). N D Danian 1 � Upper Cretaceous • Locality e Town o Borehole • RODMOLLA I--�--,-�--,-��-r--�-r��,-���--+-r-�TOSTERUP ING ELSTORP e VALLE�BERGA-,--'-.,--'-,..-'-I KAs 20 km I Mammillaten-Kreide. Christensen (1975) discussed at Locality details length the age of the zone of Belemnellocamax mammilla tus and concluded that this zone equates with the German The localities mentioned in the text are listed below. The gracilis/mucronata Zone, the uppermost zone of the outcrops of the Kristanstad Basin were described in detail Lower Campanian (Fig. 2). Christensen (1986) noted, by Christensen (1975) and those of the Vomb Trough by however, that specimens of Belemnellocamax balsvikensis, Christensen (1986), who also reviewed earlier work. The a species firstrecognized by Brotzen (1960) and described stratigraphical age of the localities based on their belem later by Christensen (1975), were placed in Belemnelloca nite and ammonite faunas is discussed below. Most out max mammillatus by earlier authors. B. balsvikensis occurs crops only expose a few metres of sediments, which can be in the lower Upper Campanian in beds that can be corre referred to a specificbelemnite zone. lated with the German conica/mucronata Zone and the lower part of the overlying basiplana/spiniger Zone (Christensen & Schulz 1976). The Swedish mammilla tuskrita, therefore, spans the uppermost Lower and lower Vo mb Trough Upper Campanian, and the upper part of the mammilla Fig. 3 tuskrita (zone of B. balsvikensis) can be correlated with the lower part of the German Mucronaten-Kreide. Severai informal lithostratigraphical units, including the Christensen (1975, 1986, 1993) did not establish fo rmal Eriksdal marl, Kullemolla marl, Lyckås marl, Kåseberga belemnite zones for the Upper Cretaceous of Sweden, marl, Kopinge sandstone, Valleberga sandstone, and because extensive, continuous sections are not available Tosterup conglomerate, have been established for various in the Kristianstad Basin, Båstad Basin and Vomb sedimentary rocks in the trough. Some of these units have Trough. He correlated the belemnite faunas of southern been used subsequently as fo rmal formations. They are, Sweden with those of northwest Europe, in particular however, poorly de fined or not defined at all and are northwest Germany (Fig. 2) and the Russian Platform. therefore used informally here. FOSSILS AND STRA TA 44 (1997) Santonian to Maastrichtian ammonites 81 Eriksdal. - The abandoned pit at Eriksdal is now covered but this may be due to lack of exposures (Christensen by scree and overgrown. The Eriksdal marl has yielded a 1986, p. 18). belemnite assemblage consisting of Gonioteuthis westfali The following ammonites are recorded: Gaudryceras cagranulata, Belemnitella propinqua, and Actinocamax sp., Pachydiscus haldemsis, Patagiosites stobaei, Nostoceras verus, which indicate the upper Middle Santonian G. junior, Lewyites elegans, Baculites cf. aquilaensis, B. angus westfalicagranulata Zone (Christensen 1986) (Fig. 2). It tus, B. schlueteri, B. spp., Trachyscaphites spiniger spiniger should be noted, however, that Christensen also reported and Hoploscaphites ikorfatensis. This is a lower Upper a few specimens of Gonioteuthis, which are closely compa Campanian ammonite association, a dating compatible rable to G. westfalica, possibly indicating that lower mid with the higher belemnite assemblage. dIe Santonian strata are also present at Eriksdal. Moberg (1885) recorded the Maastrichtian ammonites The following ammonites are here recorded from Scaphites constrictus and Anagaudryceras cf. lueneburgense Eriksdal: Hauericeras pseudogardeni, Nowakites hernensis, from the Kopinge sandstone. These determinations are in Glyptoxoceras crispatum, and Baculites ex gr. capensis (= B. error. The record of S. constrictus may be based on frag brevicosta? Moberg and B. incurvatus Moberg); belemnite ments or microconchs of Hoploscaphites ikorfatensis, and and ammonite datings are in agreement. that of A. luenebergense on Gaudryceras sp. (see below). Kåseberga. - Loose blocks oflight, porous, yellow to grey Ku llemolla-Lyckås. - Severai closely spaced outcrops are ish-yellow Upper Cretaceous marlstone are known from recorded at Kullemolla and Lyckås (see review by Chris immediately west of the small village of Kåseberga at the tensen 1986). These are now very poorly exposed or not sea-shore on the south coast. Christensen (1986) exposed at all. The Kullemolla and Lyckås marls have recorded Belemnitella ex gr. alpha/praecursor and Gonio yielded a basal Lower Campanian belemnite assemblage teuthis sp., possibly G. granulataquadrata or G. quadrata, of Gonioteuthis granulataquadrata, Belemnitella alpha, from the loose blocks, which are thus basal Lower Cam Actinocamax verus and Belemnellocamax ex gr. grossouv panian on belemnite evidence. It is noteworthy, however, rei, indicating the G. granulataquadrata Zone (Chris that a single specimen of the Cenomanian Actinocamax ex tensen 1986) (Fig. 2). We record Hauericeras pseudogar gr. primus/p lenus has been collected recently from a loose deni from Lyckås, an occurrence compatible with the block at this locality (Christensen, unpublished). Some of belemnite dating. the blocks are therefore Cenomanian. The ammonites recorded from the blocks are: Scaphites Rodmolla-To sterup. - Five outcrops are recorded from hippocrepis Ill, Baculites suecicus Moberg, B. ex gr. capen Rodmolla-Tosterup (see review by Christensen 1986). sis (=B. incurvatus Moberg). Belemnite and ammonite datings are in agreement. The Campanian part of the sequence consists of alter nating calcareous sandstones (Kopinge sandstone) and Kopinge distriet. - The Kopinge sandstone, a yellow, conglomerates (Tosterup conglomerate). Christensen highly calcareous, glauconitic sandstone, has been (1986) recognized two belemnite assemblages from the recorded from many small outcrops in the Kopinge dis outcrops. Localities 2 and 4 have yielded an uppermost trict, including Kopingemolla, Svenstorp, Valleberga, Lower Campanian belemnite assemblage of Belemnelloca lngelstorp, Herrestad and Fredriksberg. The majority of max mammillatus, Belemnitella mucronata and Gonio these outcrops are now inaccessible or very poorly teuthis quadrata scaniensis? Loose blocks of the Tosterup exposed. The Kopinge sandstone was placed in the conglomerate at Ioc. 3 yielded the lower Upper Campa Mucronaten-Kreide or equivalent zones by authors in the nian Belemnellocamax balsvikensis (Fig. 2). last part of the 19th century and the beginning of the 20th The following ammonites are recorded from Rod century (Schliiter 1870; Moberg 1885; Stolley 1897; Hagg molla-Tosterup: Pachydiscus cf. subrobustus, Patagiosites 1930; and others). Lundegren (1933), however, recorded stobaei, Lewyites elegans, Trachyscaphites spiniger spini Belemnellocamax mammillatus from the Kopinge sand ger, Hoploscaphites ikorfatensis and Baculites cf. aqui stone, and he therefore assigned the oldest part of the laensis. This is a lower Upper Campanian ammonite sandstone to the mammillatuskrita. association, a dating compatible with the higher belem Christensen (1986) recorded two belemnite assem nite assemblage. blages from the Kopinge sandstone: (l) an assemblage consisting of Belemnellocamax mammillatus and Belem Kopingsberg-l borehole. - The Santonian ammonites of nitella mueronata, which is from the uppermost Lower this borehole were described by Kennedy & Christensen Campanian, and (2) an assemblage of Belemnitella mucro (1993), who recorded Hauericeras cf. pseudogardeni, Sca nata and B. aff.B. langei, which is from the middle Upper larites sp., Baculites sp. group of capensis, B. sp. l, Boeh Campanian (Fig. 2). The zone of Belemnellocamax bals moceras krekeleri, B. arculus and Scaphites kieslingswalden vikensis and Belemnitella mucronata was not recognized, sis fiseheri. 82 W,f. Kennedy & W.K. Christensen FOSSILS AND STRATA 44 (1997) Fig. 4. Map of the Kristianstad Basin, showing the location of the sites dealt • BJARNUM with he rein. Modifiedfr om Siverson (1992). HASSLEHOLM • KRISTIANSTAD N 1 AHUS BA LTIC SEA • Town • Locality � Upper Cretaceous o 10 , , km Kristianstad Basin of calcisiltite with detrital crystalline material and phos Fig. 4 phatic nodules. The uppermost Lower Campanian Belemnellocamax Ahus. - Loose blocks and boulders of the so-called Ahus mammillatus occurs in the conglomerate. The calcisiltite sandstone are recorded from Revhaken, immediately yielded a basal Lower Maastrichtian belemnite assem south of Ahus. The age of the Ahus sandstone has been the blage of Belemnella lanceolata and Belemnitella minor subject of much discussion; it is probably fromthe earliest sensu Christensen (1975), indicating the lower part of Bel Maastrichtian but may also be late st Campanian (Chris emnella lanceolata Zone sensu Schulz (1979). We record tensen 1975). We record Baculites vertebralis Moberg non Baculites knorrianus, a Lower Maastrichtian species, from Lamarek from the Ahus sandstone. The type material is this locality. Belemnite and ammonite datings are thus in inadequate, and this taxon is best treated as a nomen agreement. dubium (see below). Ignaberga new quarry. - In the early 1970s, this pit showed Balsberg. - This locality is a naturai cave, which shows about 10 m of calcarenite with a conglomerate in the mid biocalcarenites, resting partly on the crystalline basement dIe part of the section (Christensen 1975, Fig. 7). The calc and partly upon quartz sand. Christensen (1975) arenites have yielded an uppermost Lower Campanian recorded the uppermost Lower Campanian Belemnelloca belemnite assemblage of Belemnellocamax mammillatus, max mammillatus and Belemnitella mucronata from this Belemnellocamax ex gr. grossouvrei, Belemnitella mucro outerop. It should be noted, however, that Christensen nata and Gonioteuthis quadrata scaniensis (Christensen (1975, p. 46) also recorded speeimens of Belemnellocamax 1975). with a rather deep pseudoalveolus and many closely In the spring of 1977, downward exploitation revealed spaced conellae. These speeimens may be transitional a hardground at a horizon not exposed previously (Birke fo rms between B. mammillatus and B. balsvikensis or lund & Bromley 1979). A speeimen of Hauericeras cf. belong to the lower Upper Campanian B. balsvikensis. We pseudogardeni (=H.pseu dogardeni herein) was recorded record 'Baculites vertebralis' Moberg non Lamarek from from 5 cm below the hardground by Birkelund & Brom this outcrop. ley (1979), implying that the calcarenites below the hardground are Upper Santonian - basal Lower Campa Bjiirnum. - This abandoned pit showed a O.s m thick con nian. There is thus a considerable hiatus in the sequence glomerate at the base, which rested on weathered crystal at the hardground, spanning the G. q. quadrata and G. line basement. The conglomerate is overlain by about 6 m quadrata gracilis Zones (Fig. 2). Ignaberga new quarry, as FOSSILS AND STRA T A 44 (1997) Santonian to Maastrichtian ammonites 83 well as abandoned pits and mines at Ignaberga, were dealt Ulricelund near Niisbyholm. - The abandoned pit at with in detail by Erlstrom & Gabrielson (1992). Ulricelund in the southeastern part of the Malmo area was dug in a glacially transported mass of Maastrichtian Ivo Klack (=Blaksudden, Ivobrottet). - The coarse-grained chalk (Hagg 1954). This chalk yielded the brachiopod calcarenites of this pit have yielded an uppermost Lower Trigonosemus pulehellus, which is the index fossil of the Campanian belemnite assemblage of Belemnellocamax pulchellus-pulchellus Zone of the brachiopod zonation of mammillatus, Belemnellocamax ex gr. grossouvrei, Belem Surlyk (1984). This zone is placed in the basal part of the nitelIa mucronata and Gonioteuthis quadrata scaniensis upper Lower Maastrichtian. We record Scaphites constric (Christensen 1975) (Fig. 2). tus from this locality, an occurrence compatible with the We record the following ammonites from Ivo Klack: brachiopod dating. Pseudophyllites indra, Pachydiscus colligatus, Scaphites bin odosus, and Baculites spp. Belemnite and ammonite dat ings are in agreement. Deep borehoies. - Ammonites have been studied from the following deep borehoies: the Hollviken, Svedala and Trelleborg borehoies. Malmo area We record Bostrychoceras polyplocum (Upper Campa Fig. l nian) from the Trelleborg borehole and Nowakites hern Limhamn pit. - This abandoned pit lies in the southern ensis (Santonian) from the Svedala borehole. The follow outskirts of the town of Malmo. Birkelund (1993) ing Upper Campanian to Maastrichtian ammonites are recorded Scaphites constrictus from the chalk, which was recorded from the Hollviken borehoies: Saghalinites sp., placed in the stevensis-chitonifo rmis Zone of the brachio Tragodesmoceras clypeale, Hoplitoplacenticeras coesfeldi pod zonation ofSurlyk (1984); this is the uppermost bra ense, Hop loscaphites ikorfatensis, H. tenuistriatus and chiopod zone of the Maastrichtian. Acanthoscaphites sp. Systematie palaeontology Location of speeimens. - This is indicated by the following Gaudryceras Sp. abbreviations: BMNH, Naturai History Museum, Lon Fig. SA-B don; IPB, Palaontologisches Institut der Universitat, Bonn; LO, Geological Institute, University of Lund; Description. - The specimen is a well-preserved fragment MGUH, Geological Museum, University of Copenhagen; of body chamber from Kopinge, with calcite-replaced MNB, Museum fur Naturkunde, Berlin; PI, Palaeonto shell preserved. The maximum preserved whorl height is 46 mm; the whorl breadth to height ratio is 0.76, the logical Institute, Uppsala; RM, Naturhistoriska riksmu seet, Stockholm (Swedish Museum of Naturai History, whorl section ovoid, with feebly convex inner flanks, flat Section of Paleozoology); SGU, Sveriges Geologiska ten ed, convergent outer flanks, broadly rounded ventro Undersokning, Uppsala (Geological Survey of Sweden). lateral shoulders and a feebly convex venter. Ornament is of very finelirae, convex across the inner flank, concave on the outer flank, stronglypr ojected on the outer ventrolateral shoulder and broadly convex Order Ammonoidea Zittel, 1884 across the venter. There are periodic scale-like ridges and shallow grooves. Suborder Lytoceratina Hyatt, 1889 Discussion. - This fragment is specifically indeterminate Superfamily Tetragonitaceae Hyatt, 1900 and might even be referred to Anagaudryceras Shimizu, 1934. Moberg (1885) recorded Anagaudryceras cf. luene Family Gaudryceratidae Spath, 1927 burgense (Schhiter, 1872) from the Kopinge sandstone. This is a Maastrichtian species; the present specimen may explain the record as being based on one of the long-rang Genus Gaudryceras de Grossouvre, 1894 ing, finelyribb ed Gaudryceras species. Type speeies. -Ammonites mitis Hauer, 1866, by subse Occurrence. - Kopinge sandstone, Kopinge, probably quent designation of Boule et al. (1906, p. 183 [Il]). middle Upper Campanian. 84 W,f. Kennedy & w.K. Christensen FOSSILS AND STRATA 44 (1997) Fig. 5. DA-B. Gaudryceras sp. SGU unregistered, probably middle Upper Campanian, Kopinge sandstone, Kopinge. DC Saghalinites sp. SGU unregis tered, Lower Maastrichtian, Hollviken-1 borehole, 540.65-540.75 m. DO. Pseudophyllites indra (Forbes, 1846). SGU unregistered, uppermost Lower Campanian of lvo Klack. DE. Hauericeras (Hauericeras) pseudogardeni (Schliiter, 1872). SGU Type 3875, the original of Ammonites n.sp.? of Moberg (1885, Pl. 2:7), from the upper Middle Santonian ofEriksdal. All xl. Family Tetragonitidae Hyatt, 1900 Saghalinites sp. Fig. 5C Genus Saghalinites Wright & Description. - The speeimen is a crushed composite Matsumoto, 1954 mould of 120° of phragmocone with a maximum pre served whorl height of 15.5 mm. The shell surface is Ty pe speeies. - Ammonites cala Forbes, 1846, p. 104, Pl. smooth, but fo r widely spaced prorsiradiate constrictions, 87:4, by original designation of Wright & Matsumoto convex on the inner flank, concave on the outer flankand (1954, p. llO). fe ebly convex across the venter. FOSSILS AND STRATA 44 (1997) Santonian to Maastrichtian ammonites 85 Discussion. - This fragment is specifically indeterminate, Brazil, western Australia, New Zealand, South India, but may belong in Saghalinites wrighti Birkelund, 1965 Madagascar, Zululand (South Africa). (see revisions in Birkelund 1993, p. 45, PIs. 1:6-7; 2:1, 3- 4, 6, and Ward & Kennedy 1993, p. 21, Figs. 19.3-19.4, 19.8, 19.12, 20.1-20.3), a species that ranges from upper Suborder Ammonitina Hyatt, 1889 Lower to lower Upper Maastrichtian. Occurrence. - Lower Maastrichtian, Hollviken-1 bore Superfamily Desmocerataceae hole, 540.65-540.75 m. Zittel, 1895 Family Desmoceratidae Zittel, 1895 Genus Pseudophyllites Kossmat, 1895 Subfamily Puzosiinae Spath, 1922 Ty pe species. -Ammonites indra Forbes, 1846, p. 105, Pl. 11:7, by original designation of Kossmat (1895, p. Genus and subgenus Hauericeras de 137[41]). Grossouvre, 1894 (=Schlueteria Rollier, 1922, p. 359, non Fritsch in Fritsch & Kafka, 1887, p. 33; Pseudogardenia Tomlin, 1930, p. 23) Pseudophyllites indra (Forbes, 1846) Discussion. - Matsumoto in Matsumoto, Toshimitsu & Fig. SD Kawashita (1990) distinguished two subgenera in Hau ericeras, the nominotypical subgenus, with feeble ventro Synonymy. - 01846 Ammonites Indra - Forbes, p. 105, Pl. lateral riblets/nodes, and Gardeniceras Matsumoto & 11:7. 01992 Pseudophyllites indra (Forbes) - Kennedy & Obata, 1955 (p. 134) (type species Ammonites gardeni Henderson, p. 398, PIs. 3:7-9, 13-27; 4:1-3 (with fullsyn Baily, 1855, p. 450, Pl. 11:3), which lacks riblets/nodes. onymy). 01993 Pseudophyllites indra (Forbes) - Ward & He also concluded that Hauericeratinae Matsumoto, Kennedy, p. 22, Figs. 17.8, 18.9-10, 19.7, 19.9, 19.13, 1938, should be incorporated into Puzosiinae, a view 19.21, 21.2, 22.1, 27.6. 01993 Pseudophyllites indra accepted here. (Forbes) - Hancock & Kennedy, p. 153, Pl. 1:3-4. 01993 Pseudophyllites indra (Forbes) - Kennedy & Hancock, p. 577, Pl. 1:4, 7. Hauericeras (Hauericeras) pseudogardeni Ty pes. - Lectotype, designated by Kennedy & Klinger (Schhiter, 1872) (1977, p. 182), is BMNH C51068, the original of Forbes Figs. SE, 6 (1846, Pl. 11:7). The paralectotypes are C51 069-73, all from the Upper Maastrichtian Valudavur Group ofPon Synonymy. - 01872 Ammonites pseudo-Gardeni - Schlii dicherry, south India. ter, p. 54, Pl. 16:3-6. 01875 Haploceras pseudogardeni Schliiter - Neumayr, p. 915. 01885 Ammonites n.sp.? - Description. - The specimen, from Ivo Klack, is a worn Moberg, p. 25, Pl. 2:7. 01894 Hauericeras pseudo-Gardeni internal mould of a single chamber with a maximum pre Schliiter- De Grossouvre, p. 220, Text-fig. 81. O non 1898 served whorl height of 120 mm. The whorl section is com Hauericeras pseudo-gardeni Schliiter - Mariani, p. 8 (1), pressed oval, with a massive septal lobe. Fig. 6. 01899 Ammonites pseudogardeni Schliiter var. Discussion. - The massive septal lobe alone serves to nodatus - Schliiter,p. 411. 01899 Ammonites pseudo-gar identify this fragment as Pseudophyllites indra, rather deni Schliit. - Schliiter, p. 411. 01905 Hauericeras pseu dogardeni Schliiter- Wegner, p. 207. 01905 Hauericeras than Pseudophyllites loryi (Kilian & Reboul, 1909, p. 18, Pl. 1:4-5) and its synonyms P. latus (Marshall, 1926, p. buszii - Wegner, p. 208, Pl. 8:1a-b. 01905 Hauericeras 149, PIs. 20:6, 6d; 32:1-2), and P. peregrinus Spath, 1953, buszii var. nodata - Wegner, p. 209, Pl. 8:1a. 01905 Hau p. 7, Pl. 1:6-9), as discussed by Henderson & McNamara ericeras buszii var. costata - Wegner, p. 208, Pl. 8:1b. (1985, p. 50). 01906 Hauericeras pseudo-Garden i Schliiter - Miiller & Wollemann, p. 14, PIs. 4:1-4; 8:3. 01920 Hauericeras Occurrence. - Pseudophyllites may first appear in the pseudogardeni Schliiter - Koplitz, p. 64, Pl. 8:24. 01920 Upper Santonian and ranges with certainty from Lower Hauericeras buszii Wegner - Koplitz, p. 64. 01925 Hau Campanian to up perm ost Maastrichtian. P. indra is ericeras Pseudo-Gardeni Schliiter - Diener, p. 95. 01930 known from Sweden, Northern Ireland, southwestern Hauericeras cf. pseudo-Gardeni Schliiter - Hagg, p. 60. France, Spain, Poland, Austria, New Jersey, The U.S. Gulf 01931 Hauericeras pseudogardeni Schliit- Riedel, p. 694, Coast, British Columbia, Alaska, Saghalin, Japan, Chile, Pl. 78:7. 01931 Hauericeras buszii Wegner - Riedel, p. 86 W.]. Kennedy & w.K. Christensen FOSSILS AND STRATA 44 (1997) Fig. 6. Hauericeras (Hauericeras) pseudogardeni (Schliiter, 1872). Lectotype, IPB 48, the original of Schliiter (1872, Pl. 16:5-6), fromDiilmen, West phalia. xO.75. 695. 01935 Hauericeras pseudo-Gardeni (Schliiter) - 23, Pl. 4:11. 01957 Hauericeras pseudogardeni (Schliiter) Hiigg, p. 59. 01938 Hauericeras pseudogardeni Schliit. - - Wright, p. L37l, Fig. 485:1a-1d. 01964 Hauericeras Roman, p. 406, Pl. 41:393. O?non 1951 Hauericeras cf. pseudogardeni - Arnold, p. 12. 01979 Hauericeras cf. pseudogardeni Schliiter - Mikhailov, p. 81, Pl. 12:50. pseudogardeni (Schliiter) - Birklund & Bromley, p. 173, 0?1953 Hauericeras pseudo-gardeni Schliit. - 0dum, p. Fig. 3. 01987 Hauericeras pseudogardeni (Schliiter) - FOSSILS AND STRATA 44 (1997) Santonian to Maastrichtian ammonites 87 Kennedy & Summesberger, p. 28. 01990 Hauericeras ventrai ribs. One specimen shows delicate biconcave con pseudogardeni (Schliiter) - Matsumoto in Matsumoto, strictions, four per half whorl, at a diameter of 57 mm, Toshimitsu & Kawashita, p. 439, Fig. 1. 01993 Hauer together with indications of delicate outer flank ribs. iceras cf. pseudogardeni (Schliiter) - Kennedy & Chris Beyond 60 mm diameter, ornament is much more tensen, p. 152, Fig. 2a. marked, with delicate biconcave prorsiradiate ribs that increase by branching and intercalation, strengthening Ty pe. - Lectotype, by the subsequent designation of Mat markedly on the ventolateral shoulder, where they project sumoto in Matsumoto et al. (1990, p. 440), is IPB 48, the strongly forwards, and may strengthen into crescentic original of Schliiter (1872, Pl. 16:5-6), from Diilmen, outer lateral-ventrolateral nodes on some but not all ribs. Westphalia, Germany (Fig. 6). MNHP R51 769 shows a combination of constrictions, Description and discussion. - Moberg (1885, p. 25, Pl. 2:7) outer flank ribs and tubercles extending to a phragmo figured as Ammonites n.sp. a juvenile of this species 46 cone diameter of 130 mm; MNHP R51770 is, in contrast, mm in diameter from Eriksdal (SGU Type 3875; Fig. 5E virtually smooth at the same diameter. What may be a herein). A second specimen described by him as Ammo small adult with ribb ed, nodate and constricted phragmo nites sp. (1885, p. 25, Pl. 2:6) from Eriksdal was referred to cone has half a whorl of body chamber, with three nar Ammonites clypealis of Schliiter by Hagg (1930), but row, biconvex constrictions with associated low broad seems rather to be a Nowakites hernensis (Schliiter, 1867), collar ribs, of which the adapical rib bears a small bullate as discussed below. Birkelund & Bromley (1979, p. 173, to feebly crescentic outer lateral node. There are traces of Fig. 3) described as H. cf. pseudogardeni a phragmocone delicate riblets and striae on the interspaces between, but of a juvenile from Ignaberga that also belongs here (PI Sk no tubercles. A second small adult is 155 mm in diameter; 70), as follows: 'The fully septate nucleus is 85 mm in it has half a whorl ofbody chamber preserved. The phrag diameter. It is rather involute, the umbilical ratio being mocone is again ribbed and constricted, with numerous about 22%. The venter is sharp and high and the sides are delicate outer lateral nodes. On the body chamber, there flat. The umbilical shoulder is not preserved. The speci are fivebiconca ve prorsiradiate constrictions, again asso men carries six or seven constrictions on the outermost ciated with collar-ribs, the adapical strengthened into a whorl preserved. They are deep and slightly sigmoid. The crescentic outer lateral tubercle. The interspaces are sutures are fairly well preserved and extremely incised. nearly smooth. MNHP 51772 shows comparable body In spite of the poor preservation, the specimen can be chamber characteristics at a much smaller size. Although referred to the genus Hauericeras on the basis of the cross the shape is distorted into an ellipse, the major diameter section, constrictions and suture lines. Its closest relative is only 110 mm. If the last three specimens are correctly seems to be the type of the genus, Hauericeras pseudogar interpreted as adult H. (H.) pseudogardeni, they may be deni (Schliiter, 1872). Thus, a comparison with material microconchs of the species. from Diilmen, Westphalen, kept in Miinster Museum, The lectotype (Fig. 6) is an internal mould, still septate and figured material from that region (Schliiter 1872; at 237 mm. Coiling is involute, with 62% of the previous Miiller & Wollemann 1906) shows good agreement with whorl covered, the umbilicus small (24% of the diame the specimen. The only difference lies in the constrictions, ter), shallow, with a flattened-outward-inclined umbilical which may be slightly more curved in our specimen. The wall and narrowly rounded, sharp umbilical shoulder. closest comparable specimen is figuredby Miiller& Wol The whorl section is compress ed, lanceolate, with the lemann (1906, Pl. 4:4).' greatest breadth around mid-flank.A pronounced facet is We also examined six specimens from Braunschweig in associated with a break in the even profile of the whorl the Institut fur Palaontologie, Bonn (unregistered), and section on the outer flank; the venter is acute. The outer four in the collection of the Museum National d'Histoire flank region between facet and venter bear scarcely dis Naturelle, Paris (MNHP R5 1 769-51772). cernible traces of delicate, markedly prorsiradiate ribs on The early growth stages are shown by the Eriksdal spec the adapical 90° sector of the outer whorl. There are traces imen and the specimens from Braunschweig, all of which of three constrictions on the adapertural 120° of the outer are preserved as crushed composite moulds. Co iling is whorl, 60° apart. They are narrow, shallow, straight and involute, the umbilicus small (17-20% of diameter), shal prorsiradiate on the innermost flank,then flexed and fe e low, with a low, flattenedwall and sharp umbilical shoul bly concave on the inner flank, fe ebly convex across the der. The whorl section is very compressed, lanceolate, middle of the flank, more markedly concave on the outer, with the greatest breadth below mid-flank; the degree of and projected strongly forward on the outermost flank.A compression has been greatly accentuated by post-mortem further large, wholly septate specimen from Diilmen in compaction. The moulds show very clearly that the the Bonn collections is 290 mm in diameter and retains phragmocones had a solid, sharp keel, demarcated from calcite-replaced shell over much of its surface. There are the venter by a groove. Specimens may be smooth to a indications of a further 240° of umbilical seam of a now diameter of 60 mm or may have delicate umbilical and missing outer whorl to an umbilical diameter of 115 mm, 88 W.f. Kennedy & w.K. Christensen FOSSILS AND STRATA 44 (1997) corresponding to an estimated diameter of 440 mm. Coil Cobban 1988 (p. 595, Figs. 2; 4:1-3), from northern Mex ing is a liule more evolute than in the previous speeimen, ico, with a fastigate venter but no well-differentiated keel the umbilicus comprising 26% of the diameter, with 51% as is present in Hauericeras. of the previous whorl covered, the umbilical wall low, flat Occurrence. - Middle Santonian to Lower Campanian of ten ed, outward-inclined, with a sharp umbilical shoulder. the Miinster Basin and Braunschweig, Germany; Lyckås, The whorl section is compressed, lanceolate, with a whorl Eriksdal and Ignaberga, southern Sweden, plus borehole breadth to height ratio of 0.46, the greatest breadth below records from Kopingsberg (Kennedy Christensen 1993) mid-flank. The venter is blunt on the mould but sharp & and Hollviken (0dum 1953). and acute where replaced shell is present, the keel sepa rated fromthe venter by a marked groove. There are nine constrictions on the outer whorl, narrow, moderately deep, and preceded by a blunt, coarse collar-rib. Ribs and Family Pachydiscidae Spath, 1922 constriction are prorsiradiate, fe ebly concave across the inner flank, fe ebly convex across mid-flank and feebly Genus and subgenus concave on the outer flank, sweeping forwards and pro jected markedly across the ventro lateral shoulder to fo rm Pachydiscus Zittel, 1884 an acute chevron across the siphonal keel. The shell sur (=Parapachydiscus Hyatt, 1900, p. 570; Joaquinites Anderson, 1958, p. face between constrictions is variably preserved, but on 218; Pseudomenuites Matsumoto, 1955, p. 169). the firsthalf whorl it bears low irregular ribs and striae of Type speeies. -Ammonites neubergicus Hauer, 1858, p. 12, variable length and strength. They are weak, straight and Pl. 2:1-4, by subsequent designation of de Grossouvre prorsiradiate on the inner flank, but strengthen, sweep (1894, p. 177). fo rwards and are markedly prorsiradiate on the outer flank and ventrolateral shoulder. These large dis es are interpreted as macroconch phragmocones. Interpretation of the Braunschweig and Diilmenspeci Pachydiscus (Pachydiscus) colligatus mens as a probable macroconch and microconch pair (Binkhorst, 1861) must be regarded as tentative. From the material available Figs. 7-10 it seems that internal moulds may be virtually smooth (but for constrictions), whereas the shell surface and Synonymy. - 01861 Ammonites colligatus, Nobis - composite moulds are much more markedly ribbed. But Binkhorst, p. 25 (pars) Pl. 8 only. 01986a Pachydiscus even some composite moulds are virtually smooth, (Pachydiscus) colligatus (Binkhorst) - Kennedy, p. 36, although this might be due to post-mortem effects. If this Figs. 13-14. 01987 Pachydiscus (Pachydiscus) colligatus interpretation is accepted, the variety nodatum ofSchliiter (Binkhorst) - Kennedy, p. 162, PIs. 1:1-2; 2:1-2; 3; 4:4-5 (1899) and Hauericeras buszii Wegner, 1905 (p. 209, Pl. (with full synonymy). 01993 Pachydiscus (Pachydiscus) 8:1a-b), are synonyms of pseudogardeni. The best previ cf. colligatus (Binkhorst) - Hancock & Kennedy, p. 162. ous illustrations of the speeies are those of Miiller& Wol Types. - Lectotype, by the subsequent designation of lemann (1906) of material from Braunschweig, which Kennedy (1987, p. 162), is the original ofBinkhorst (1861, included constricted ribbed/nodate phragmocones up to Pl. 8), MNB unregistered, from the lower Upper Campa 160 mm diameter (1906, PIs. 4:1; 8:3) as well as smooth, nian of Jauche, Brabant, Belgium (Kennedy 1987, PIs. delicately constricted juveniles. The Hauericeras cf. pseu l: 1-2; 2:1-2). The paralectotypes belong to severai differ dogardeni of Mikhailov (1951, p. 81, Pl. 12:50) has con ent species as discussed by Kennedy (1987, pp. 162-163). cave constrictions and se ems far toa evolute to be referred to the present speeies. Description. - Internal moulds of the earliest growth Matsumoto in Matsumoto et al. (1990, p. 451) thought stages seen have whorl heights of 22 and 40 mm (Fig. 7A Ammonites mengedensis Schliiter, 1876 (p. 154, Pl. 40:9) B, E-F). At this size, coiling is moderately involute, the might be the microconch of H. (H.) pseudogardeni, but umbilicus small, deep, with a feebly convex wall and this is a significantly older speeies (Kaplan & Kennedy broadly rounded umbilical shoulder. The whorl section is 1994) and does not co-occur with H. (H.) pseudogardeni depressed reniform, with the greatest breadth just outside in any of the collections we have studied. the umbilical shoulder and whorl breadth to height ratios Hauericeras (H.) antiquum Collignon (1961, p. 75, Fig. of up to 1.31. Low, blunt, narrow primary ribs ar ise at the 12) from the Lower Coniacian of Madagascar appears to umbilical seam and strengthen into feeble to incipient represent the stock ancestrai to H. (H.) pseudogardeni, bullae on the umbilical shoulder. These give rise to one or having the shell shape of Hauericeras plus delicate ventrai two ribs, with single intercalated ribs between. Ribs are ribs, but no constrictions. It is transitional to the Middle straight and prorsiradiate on the inner flank,then flexing Turonian Puzosia (Puzosia) serratocarinata Kennedy & forwards and concave on the outer flankand ventrolateral FOSSILS AND STRATA 44 (1997) Santonian to Maastrichtian ammonites 89 Fig. 7. Pachydiscus (Pachydiscus) colligatus (Binkhorst, 1861), uppermost Lower Campanian, lvii K1ack. DA-B. SGU unregistered. De. SGU unregis tered. DD. Silicone squeeze froman external mould of a juvenile, SGU unregistered. DE-F. SGU unregistered. All x l. 90 W,f. Kennedy & W.K. Christensen FOSSILS AND STRA TA 44 (1997) Fig. 8. Pachydiscus (Pachydiscus) colligatus (Binkhorst, 1861), uppermost Lower Campanian, lvo K1ack. SGU unregistered. All xl. shoulder where they weaken, and cross the venter in a on the inner half of the flank, strengthening only on the very shallow convexity. A partial external mould of a juve outer flank and venter, which they cross in a broad con nile with an estimated diameter of77 mm shows the orna vexity. The ribs are subdued, and separated by very broad ment of the outer shell surface to have been much sharper interspaces, as on the only sector of the lectotype where (Fig. 7D). Eleven ribs per whorl arise at the umbilical the outer shell surface is preserved (Kennedy 1987, Pl. seam, and strengthen across the umbilical wall, most 1:1). Beyond this diameter, the unfigured side of this developing into small, sharp bullae. These give rise to specimen, which is septate to the maximum preserved pairs of narrow sharp ribs, with single intercalated ribs diameter of 210 mm, has only obscure distant low flank between. In some larger moulds (Fig. 8), this alternation ribs on the last half whorl (Figs. 9-10). Other, larger spec of primary and intercalated ribs extends to whorl heights imens show such distant ribs extending up to 330 mm of up to 70 mm. Beyond this, inner flankorna ment weak diameter. ens. Fig. 7e shows the ornament of the inner whorls of a specimen with replaced shell at a diameter of 140 mm. Discussion. - P. (P. ) colligatus from Ivo Klack is repre Inner flankor nament is very weak. There are no umbilical sented by numerous specimens in museum collections. bullae, and very weak riblets are prorsiradiate and distant The species was revised and discussed by Kennedy (1987), FOSSILS AND STRATA 44 (1997) Santonian to Maastrichtian ammonites 91 Fig. 9. Pachydiscus (Pachydiscus) colligatus (Binkhorst, 1861), uppermost Lower Campanian, lvii Klack. SGU unregistered (see also Fig. 10). xO.85. who illustrated the type material, and diseussed differ speeies occurs in the uppermost Lower Campanian rocks ences from other closely allied speeies. at Ivo Klack. It is also known from the Upper Campanian of Royan (Charente-Maritime), possibly Tercis (Landes), Occurrence. - The lectotype is from the lower Upper Campanian of Jauche, Brabant, Belgium. In Sweden the France, and, perhaps, Madagascar. 92 W,f. Kennedy & w.K. Christensen FOSSILS AND STRATA 44 (1997) Fig. 10. Pachydiscus (Pachydiscl-ls) colligatus (Binkhorst, 1861), upperm ost Lower Campanian, lvii Klack. SGU unregistered (see also Fig. 9). xO.8. Pachydiscus (Pachydiscus) haldemsis Sharpe - Moberg, p. 23, Pl. 3:1. 01889 Paehydiseus gali eianus Favre - Griepenkerl, p. 101. 01894 Paehydiseus (Schhiter, 1867) koeneni - de Grossouvre, p. 178. 01902 Paehydiseus gali Fig. II eianus (Schluter) - Wollemann, p. 103. 01913 Paehydis Synonymy. - 01867 Ammonites haldemsis - Schluter, p. eus oldhami Sharpe - Nowak, p. 362, Pls. 41:16; 43:31; 19, Pl. 3:1. 01872 Ammonites haldemsis - Schltiter, p. 70. 45:43. 0?1913 Paehydiseus kaliszanensis - Nowak, p. 359, 01872 Ammonites cf. auritoeostatus - Schluter, p. 70, Pl. Pl. 40:8. 01913 Paehydiseus haldemensis Schltiter - 22:6-7. 01872 Ammonites Galieianus Favre - Schltiter, p. Nowak, p. 349. 01925 Paehydiseus koeneni Grossouvre 63, Pl. 19:3-5, Pl. 20:9. 01885 Ammonites Oldhami Diener, p. 106. 01935 Paehydiscus ex. aff. galieianum FOSSILS AND STRATA 44 (1997) Santonian to Maastrichtian ammonites 93 Fig. 11. Pachydiscus (Pachydiscus) haldemsis (Schliiter, 1867). DA-B. LO 72lt, the original ofAmmonites oldhami Sharpe ofMoberg (1885, Pl. 3:1), prob ably middle Upper Campanian of Kopinge. De. Macroconch phragmocone, IPB 50b, the original of Schliiter (1872, Pl. 19:3-4), fromHaldem, West phalia, Germany. All x l. 94 W,J. Kennedy & W.K. Christensen FOSSILS AND STRATA 44 (1997) Favre - Brinkmann, p. 5. 01951 Paehydiseus cf. Koeneni koeneni most closely resembles P. (P.) oldhami (Sharpe, Grossouvre - Mikhailov, p. 60, Pl. 10:47. 01951 Paehydis 1855) (p. 32, Pl. 14:2; see revision in Kennedy 1986a, p. eus oldhami (Sharpe) - Wright & Wright, p. 20 (pars). 40, PIs. 3; 4:4-5; 5:1-3; Figs. 4A, 15-16 (holotype), 18). 01954 Paehydiseus (Parapaehydiscus) oldhami (Sharpe) - Microconchs ofP. (P. ) oldhami have yet to be recognized; Hagg, p. 57. 01955 Paehydiseus haldemensis Schhiter - macroconchs differ in the generally coarser ornament of Matsumoto, p. 168. 01955 Paehydiseus ambiguus Gros haldemsis, with well-developed, rather than incipient bul souvre - Matsumoto, p. 169. 01957 Pseudomenuites lae when young. Adults differ in the persistence of orna ambiguus (de Grossouvre) - Wright, p. 380. 01959 Paeh ment in haldemsis in middle growth. ydiseus koeneni Grossouvre - Naidin & Shimanskij, p. 185, Oecurrence. - Upper Upper Campanian, Kbpinge, Swe Pl. 9: 1. 01964 Paehydiseus koeneni Grossouvre - Giers, p. den, MiinsterBasin and elsewhere in Germany; Gschlief 263, Pl. 5:1, non 2; non Fig. 5. 01974 Menuites ambiguus graben, Austria; Norfolk, England; Northern Ireland; (Grossouvre) - Naidin, p. 182, Pl. 63:2. 01974 Paehydis Poland; Ukraine and Turkmenia. eus koeneni Grossouvre, 1894 - Naidin, p. 186, Pl. 65:2-3. 01980 Paehydiseus koeneni Grossouvre - Blaskiewicz, p. 42, PIs. 26: 1-2; 27: 1-4; 28: 1-4; 34:3-4. 01984 Paehydiscus (Paehydiseus) haldemsis (Schliiter) - Kennedy & Summes Pachydiscus (Pachydiscus) cf. subrobustus berger, p. 158, PIs. 4:1-5; 5:1; 6:2; 7:1-11; 13:1 (with addi Seunes, 1892 tional synonymy) . 01986a Paehydiseus (Paehydiseus) hal Fig. 12 demsis (Schliiter) - Kennedy, p. 45, PIs. 4:1-3; 5:7-14; Figs. 11A-D, F-G, 17. Synonymy.. - compare: 01892 Paehydiseus subrobustus - Seunes, p. 15, Pl. l3 (4):1. 01894 Paehydiseus subrobustus Typ es. - The lectotype of Ammonites haldemsis Schliiter, Seunes - de Grossouvre, p. 200, Pl. 36:2. 01910 Paehydis 1867 (p. 19), by subsequent designation of Kennedy & eus subrobustus Seunes - Frech, p. 4, Pl. 1:1; Figs. 2-3. Summesberger (1984, p. 158), is an unregistered speci 01913 Paehydiseus subrobustus Seunes - Nowak, p. 357, men in the Schliiter Collection (IPB), the original of Pl. 41: 15. 01925 Paehydiseus subrobustus Seunes - Diener, Schliiter (1867, Pl. 3:1), refiguredby Kennedy (1986a) as p. 108. 01951 Paehydiseus subrobustus Seunes - Fig. llA-B. The lectotype of P. koeneni de Grossouvre, Mikhailov, p. 70, Pl. 9:43-44. 01952 Paehydiseus subro 1894 (p. 178) is in the same collection and was refigured bustus Seunes - Collignon, p. 92. 01955 Paehydiseus sub by Kennedy (1986a) as Fig. lID. Both are fromthe Upper robustus Seunes - Collignon, p. 83. 01964 Paehydiseus Campanian of Haldem, Westphalia. subrobustus Seunes - Giers, p. 265, Pl. 5:3 (pars). O non 1971 Paehydiseus subrobustus Seunes - Collignon, p. 34, Deseription. - The original of Ammonites Oldham i Sharpe Pl. 454:2411. 01974 Paehydiseus subrobustus Seunes - of Moberg (1885, Pl. 3:1; Fig. 11A-B herein), from Naidin, p. 185, Pl. 65:1; Fig. 33. 01984 Paehydiseus cf. sub Kbpinge, is LO 72 It. It is a somewhat worn and crushed robustus Seunes - Kennedy Summesberger, p. 161, Pl. phragmocone with the following dimensions: D=86.0 & 8:4. 01993 Paehydiseus (Paehydiseus) cf. subrobustus (100); Wb =26.3 ( 30.6); Wh =38.4 (44.7); Wb:Wh=0.68; Seunes - Kennedy, p. 103, PIs. 1:1, 8-9; 2:l3-14. 01993 U =25.0 (34.4). Coiling is evolute, umbilicus shallow, Pachydiseus (Pachydiscus) subrobustus Seunes - Hancock with outward-inclined wall and broadly rounded shoul Kennedy, p. 161; ? Pl. 3:2-3. der. Whorl section is compressed with whorl breadth to & height ratio 0.68, the greatest breadth below mid-tlank. Typ e. - Lectotype, by subsequent designation of Kennedy Flanks are fe ebly convex, ventrolateral shoulders and ven & Summesberger (1984, p. 161), is the original of Seunes ter broadly rounded. Primary ribs arise at the umbilical (1892, Pl. 13 (4): 1), from Tercis, Landes, France. The seam and strengthen across the umbilical wall and shoul specimen has not been traced. der. They are straight and prorsiradiate on the inner to middle tlank, tlexingfo rwards and feebly concave on the Deseription. - LO 7135t (Fig. 12) from Tosterup is a outermost tlank and ventrolateral shoulder, and feebly wholly septate composite mould, distorted into an convex on the venter. One or two intercalated ribs arise ellipse, with a maximum preserved diameter of 119 mm. either low or high on the tlank and strengthen to match Coiling appears to have been moderately involute, the the primary ribs on the outermost tlank, ventrolateral umbilicus deep, with a feebly convex subvertical wall, the shoulders and venter. Suture is typical fo r genus. umbilical shoulder broadly rounded. The umbilicus comprises 22% approximately of the diameter. Whorl Diseussion. - The lectotype of Ammonites haldemsis section compressed oval, with greatest breadth just out Schliiter, 1867, is a microconch, of which Paehydiscus side the umbilical shoulder; the whorl breadth to height koeneni de Grossouvre, 1894, is the macroconch, as dem ratio is 0.75, but degree of compression may have been onstrated by Kennedy & Summesberger (1984). The modified by post-mortem compaction. On the earlier present specimen is a small, juvenile macroconch. P. (P. ) parts of the outer whorl ribs arise at the umbilical seam, FOSSILS AND STRATA 44 (1997) Santonian to Maastrichtian ammonites 95 Fig. 12. Pachydiscus (Pachydiscus) cf. subrobustus Seunes, 1892. LO 7135t, from the uppermost Lower Campanian -lower Upper Campanian ofTosterup. All xl. sweep back across the umbilical wall, strengthen on the Genus Patagiosites Spath, 1953 umbilical shoulder, and appear to develop into sharp umbilical bullae (preservation is defective) which give Type speeies. - Ammonites patagiosus Schluter, 1867, p. 22, rise to pairs of ribs, with occasional intercalated ribs Pl. 4:4-5, by original designation of Spath (1953, p. between; there are long intercalated ribs. On the last part 38);=Ammonites stobaei Nilsson, 1827, p. 5, Pl. 1:1-2. of the outer whorl most ribs are primaries, with an esti mated total of 46 ribs per whorl. The ribs are sharp, nar row, and separated by wide interspaces, straight and Patagiosites stobaei (Nilsson, 1827) prorsiradiate on the inner and middle tlank, then tlexing Figs. 13-17 fo rwards and concave on the outer tlankand tlexedback Synonymy. - 01827 Ammonites stobaei - Nilsson, p. 5, Pl. and broadly convex across the venter. 1:1-2. 01867 Ammonites patagiosus - SchlUter, p. 22, Pl. 4:4-5. O?l872 Ammonites stobaei Nilsson - Schluter, p. 56 Occurrence. - The locality Tosterup ranges from upper (pars) Pl. 17:6-7; non Pl. 17:4-5 [=P. (P. ) lundgreni de most Lower to lower Upper Campanian. The type mate Grossouvre, 1894]; non Pl. 18:10-11 [=P. (P.) pseudosto rial of P. (P. ) subrobustus is from the Upper Campanian baei (Moberg, 1885)]. 01872 Ammonites patagiosus - of Tercis, Landes, France, and there are also records Schluter, p. 66, Pl. 20:7-8. 01885 Ammonites stobaei Nils from the Upper Campanian of the Gschliefgraben, Aus son - Moberg, p. 18, Pl. 2:1-5 (with additional early syn tria; Pontus, Turkey; Poland, Ukraine, and the Munster onymy). 0?1 889 Ammonites (Pachydiscus) stobaei Nilsson Basin, Germany. - Griepenkerl, p. 100. 01894 Desmoceras stobaei Nilsson, 96 W.f. Kennedy & w.K. Christensen FOSSILS AND STRATA 44 (1997) Fig. 13. Patagiosites stobaei (Nilsson, 1827). Copy ofMoberg's original figureof the lectotype (1885, Pl. 2:1), LO 14T, probably middle Upper Campanian of Kopinge. xO.25. sp. - de Grossouvre, p. 170. 01907 Pachydiscus patagiosus b. 01993 Parapuzosia (Parapuzosia) stobaei (Nilsson) - Schliiter; Paulcke, p. 62. 01913 Pachydiscus patagiosus Kennedy, p. 104. Schliiter - Nowak, p. 344. 01925 Parapuzosia stobaei Typ e. - Lectotype, here designated, is the original of Nils Nilsson - Diener, p. 130. 01947 Puzosia (Parapuzosia) son (1827, p. 5, Pl. 1:1-2), LO 14T, from the middle stobaei (Nilsson) - Hagg, p. 93. 01953 Patagiosites patagi Upper Campanian of Kopinge, Sweden, refigured by osus (Schliiter) - Spath, p. 38. 01954 Pachydiscus (Para Moberg (1885, Pl. 2:1), Regnell (1983, Fig. 4) and he rein pachydiscus) egertoni (Forbes) - Hagg, p. 56, Pl. 9:98. as Figs. 13-14. 01954 Puzosia (Parapuzosia) stobaei (Nilsson) - Hagg, p. 56. 01964 Pachydiscus stobaei (Nilss.) - Giers, p. 258, Pl. Description. - Early growth stages, to a diameter of 67 mm 4:1-4; Fig. 4. 01964 Pachydiscus koeneni Grossouvre - approximately, are represented by the original of Moberg Giers, p. 263 (pars), non Pl. 5:1 [=P. (P. ) haldemsis (1885, Pl. 2:5), from Kopinge, LO 720t (Fig. 15C-D). The (Schliiter)]; Pl. 5:2. 01964 Pachydiscus patagiosus speeimen is a crushed, wholly septate composite internal (Schliit.) - Giers, p. 267, Pl. 5:4; Fig. 6. 01974 Pachydiscus mould. Coiling is fairly involute, with 70% of the previous stobaei (Nilsson); Naidin, p. 184, PIs. 67:2; 68:2. 01983 whorl covered. The umbilicus comprises 27% of the Parapuzosia stobaei (Nilsson) - Regnell, p. 61; Fig. 4. diameter and is shallow, with a low flattenedwall and nar 01988 Pachydiscus stobaei sensu Giers - Jagt, PIs. lb; 2a- rowly rounded umbilical shoulder. The whorl section is FOSSILS AND STRATA 44 (1997) Santonian to Maastrichtian ammonites 97 Fig. 14. Patagiosites stobaei (Nilsson, 1827). Copy of Nilsson's original figureof the lectotype (1827, Pl. 1:1-2), LO 14T, probably middle Upper Campa nian of Kopinge. Approximately xO.25. compressed oval, with the greatest breadth just outside this specimen is almost totally abraded, but for traces of a the umbilical shoulder and whorl breadth to height ratio single constriction. The fragmentaryouter whorl appears 0.6, although this may have been modified by post-mor to be body chamber, and has a maximum preserved whorl tem crushing. The inner flanks are feebly convex, the height of 35 mm. The whorl breadth to height ratio is 0.8, outer flanks convergent, the venter narrowly rounded. the whorl section oval, with greatest breadth just below There are an estimated eight narrow straight prorsira mid-flank. Two strong constrictions are present on the diate constrictions per whorl, deeply incised at the umbil fragment, with strong adapical bullate constrictions and ical shoulder, and feebly convex across the venter. All are weak non-bullate adaperturaI constrictions. There are preceded by a narrow, markedly bullate collar rib, and two distant coarse ribs on the adapical part of the frag succeeded by a subequal but non-bullate collar rib. The ment. Moberg (1885, Pl. 2:2; LO 719t) figureda very worn shell surface is smooth between constrictions, but fo r but undeform ed phragmocone from Kopinge (Fig. 16A obscure traces of ribs, most obvious across the venter. D) with the following dimensions: D= 105.5 (100); Somewhat larger is the specimen figuredby Hagg (1954, Wb =41.2 (39.1); Wh =55.0 (52.1); Wb:Wh=(0.75); Pl. 9:98) from Tosterup, as Pachydiscus (Parapachydiscus) U=26.4 (25.0). The inner whorls show five constrictions egertoni (Fig. 16E-F herein). The fragmentary nucleus of per half whorI; there are traces of two constrictions on the 98 W.f, Kennedy & W. K. Christensen FOSSILS AND STRATA 44 (1997) Fig. 15. Patagiosites stobaei (Nilsson, 1827), probably middle Upper Campanian, Kopinge. DA-B. LO 7136t, xO.8. DC-D. LO nOt, the original of Moberg (1885, Pl. 2:5), xl. adapical part of the outer whorl, but no other traces of Discussion. - We regard Pachydiscus patagiosus, P. koeneni ornament survive. A further phragmocone, LO 7136t and P. stobaei of Giers as synonyms of Nilsson's Ammo from Kopinge, lacks a nucleus and is distorted, but pre nites stobaei, and also regard Schltiter's Ammonites patagi serves ornament as a composite mould (Fig. 15A-B). osus as a further synonym, following revision of their There are three prorsiradiate, straight constrictions on the material from the Miinster Basin (Kaplan, Kennedy & adapical 60° of the outer whorl, flankedby a bullate adapi Ernst, unpublished). cal and weaker, non-bullate adaperturai collar-rib. There Speeimens referred to P. patagiosus by Giers are all are obscure traces of ribs between the constrictions, while composite moulds, mostly crushed. Early phragmocone most of the outer whorl is ribbed. Ribs arise as mere striae whorls show coiling to have been fairly involute, with on the inner flank and are markedly prorsiradiate. They 70% of the previous whorl covered. The umbilicus is of strengthen markedly on the outermost flank, where they moderate width and depth, with a broadly rounded are feebly concave, sweeping fo rwards and strengthening umbilical wall and shoulder. The whorls expand slowly across the ven tro lateral shoulder and crossing the venter with, in the least-deformed specimens, a whorl breadth to in a broad convexity. This ornament extends to the largest height ratio of 0.96, the inner flanks very broadly preserved diameter, 140 mm. rounded, the outer flanks flattened and convergent, the The lectotype has the following dimension: D=425 ventrolateral shoulders broadly rounded, the venter (100); WB =110 (25.9), Wh 172 (40.5); WB:WH=0.64; broad and feebly convex. There are typically fo ur to five U = 130 (30.6). It and other large specimens in the narrow, distant, straight, fe ebly prorsiradiate constric Lund Collections are smooth, with neither ribs nor tions per half whorl. These are flankedby a narrow, weak, constrictions. prominently bullate adapical, and non-bullate adaper- FOSSILS AND STRATA 44 (1997) Santonian to Maastrichtian ammonites 99 Fig. 16. Patagiosites stobaei (Nilsson, 1827). DA-O. LO 719t, the original of Moberg (1885, Pl. 2:2), probably middle Upper Campanian ofKopinge. DE F. RM unregistered, the original of Pachydiscus (Parapachydiscus) egertoni Forbes of Hagg (1954, Pl. 9:98), Ioc. CV5 of Tosterup, uppermost Lower lower Upper Campanian. All xl. 100 W.]. Kennedy & W.K. Christensen FOSSILS AND STRATA 44 (1997) Fig. 17. Patagiosites stobaei (Nilsson, 1827). DA-B. Paralectotype of Patagiosites patagiosus (Schliiter, 1867), IPB 22c, the original of Schliiter (1867, Pl. 4:5), fromthe Upper Campanian of Coesfeld, Westphalia, Germany. DC-E. Lectotype of Patagiosites patagiosus (Schliiter, 1867), IPB 22b, the original of Schliiter (1867, Pl. 4:4), from the Upper Campanian of Coesfeld, Westphalia, Germany. DF-H. Paralectotype of Patagiosites patagiosites (Schliiter, 1867), IPB 22a, from the Upper Campanian ofSiikerhookbei Coesfeld, Westphalia, Germany. DI-J. Copy of Schliiter (1872, Pl. 20:7-8), the original of which was from the Upper Campanian ofDarup, Westphalia, Germany. All xl. tural collar-rib. The shell between the ribs is smooth in of distant, coarse, bullate, straight prorsiradiate ribs the early whorIs of the smallest specimens, but as size flankedby feeble constrictions adapically, with or without increases, occasional low flank ribs develop. During the associated adaperturai ribs and up to four long and short later growth stage of this gro up of specimens, up to three ribs between them, the short ribs commonly extending long or short ribs appear between constrictions. They are across the ventrolateral shoulders and venter only. Such bullate or not, straight and prorsiradiate on the flanks, individuals are up to 100 mm in diameter, and some seem may weaken at mid-flank, then strengthen on the outer to be adult and thus possible microconchs, as with Hagg's flank and cross the venter in a broad convexity. Collar P. (P. ) egertoni (Fig. 16E-F). ribs strengthen, and constrictions become less prominent Specimens identified as Pachydiscus koeneni by Giers in the largest specimens of this group, where ornament is have inn er whorIs like those ofhis P. patagiosus, with con- FOSSILS AND STRAT A 44 (1997) Santonian to Maastrichtian ammonites 101 strictions and associated collar-ribs, of which traces diameter of 53 mm it corresponds to the smaller paralec remain on the inner whorls of larger individuals, as with totype, with, on the last half whorl, prominent bullate the Kopinge specimen shown as Fig. 15A-B. These are in adapical ribs associated with marked constrictions. general poorly preserved and very crushed. Coiling is Beyond 53 mm, the bullae become less conspicuous, involute, the umbilicus small, shallow (accentuated by ornament consisting of widely spaced feeble constrictions crushing) with a broadly rounded wall and umbilical flanked by weak, equal collar ribs with weak bullae and shoulder. The original whorl section appears to have been traces of primary ribs in between that are conspicuous compressed, with flattened, subparallel flanks and only on ventrolateral shoulders and venter. The lectotype broadly rounded ventrolateral shoulders and venter, is the original of Schliiter, 1867, Pl. 4:4; IPB 22b, shown which are crossed by ribs and constrictions in a broad here as Fig. 17C-E, from Coesfeld, Westphalia. It is pre convexity. There are 38 ribs per halfwhorl in the best-pre served in hard grey limestone, as are Giers' specimens. It served specimen, and ornament of this type extends to is a composite mould, distorted into an ellipse, with a around 200 mm diameter. The largest specimens are sep maximum preserved diameter of 52 mm. Coiling is fairly tate to around 350 mm diameter. BadIy crushed, they bear involute, with 80% approximately of the previous whorl distant coarse primary ribs, straight on the inner to mid covered, the umbilicus comprising an estimated 31% of dIe flank,and feebly concave on the outer, with generally the diameter, quite deep, with a flattened wall. The origi two short intercalated ribs between. nal whorl section cannot be determined being in some Specimens identified as Pachydiscus stobaei by Giers places compressed, in others dep res sed , with fe ebly con include large, wholly septate fragments with whorl vex flanks,bro adly rounded ventrolateral shoulders and a heights of up to 180 mm, bearing distant bar-like straight, feebly convex venter. The first half of the outer whorl, to prorsiradiate ribs that arise at umbilical bullae, and a diameter of 45 mm, corresponds to the previous speci weaken and effaceon the ventrolateral shoulders and ven men, with five constrictions and corresponding bullate ter. The largest complete Miinster Basin specimen is the adapical primary collar ribs. On the last half whorl there original of Giers (1964, Pl. 4:1), with 14-15 ribs on the are also fivecons trictions, broader and more conspicuous outer whorl (which is part body chamber) at a diameter of than in the early growth stages, flankedby narrow, sharp 800 mm. ribs, with no, or weak bullae. The constrictions are in The smallest of the Swedish specimens included in this some cases doubled on the outer flanks, ventrolateral species (Figs. 15C-D, 16E-F) differ in no significant shoulders and venter, with traces of pairs of ribs and respects from the surviving type material of Patagiosites incipient constrictions between successive fully developed patagiosus (Schliiter, 1867). The smallest ofthese, paralec constrictions. This ribbing becomes progressively more totype IPB 22c, is the original of Schliiter (1867, Pl. 4:5), conspicuous towards the adapertural end of the speci from Coesfeld, Westphalia (Fig. 17A-B ). It is a crushed men. Ribs and constrictions are straight and prorsiradiate composite mould, with the following dimensions: on the flanksand near-straight and transverse on the ven D=58.9 (100); Wb =11.7 (19.8); Wh =21.1 (35.8); ter. Patagiosites griffithi(S harpe, 1855) (p. 28, Pl. Il:3; Fig. Wb:Wh=0.55; U = 5.3 (9.0). Coiling is fairly involute, the 18 herein) is characterized by relatively high, ovoid umbilicus comprising 9% of the diameter. The original whorls with a narrowly arched venter and only fiveto six proportions cannot be reconstructed (because of crush deep constrictions per whorl, lacking conspicuous associ ing), but flanks and venter appear to have been broadly ated collar ribs and being smooth between constrictions rounded. There are an estimated eight strong adapical at diameters where P. stobaei is ribb ed. It is known from collar ribs per whorl. These arise at the umbilical seam, the Upper Campanian of Northern Ireland and Norwich, are strong on the umbilical wall, and strengthen into England. Patagiosites amarus (Paulcke, 1907) (p. 227, Pl. sharp bullae on the umbilical shoulder. They give rise to a 20:5, 7) from Patagonia has depressed subcircular whorls, narrow, rounded rib, straight and prorsiradiate on the evolute coiling and strong constrictions with severai inner flank, weakening, flexed fo rward and feebly concave intercalated ribs to 65 mm diameter, but thereafter lost, on the outer flank,and crossing the venter in a near-trans according to Paulke's figure; the Patagiosites aff. amarus verse course. They are succeeded by narrow constrictions of Spath (1953, p. 39, Pl. 10:7) from Graham Land, Ant on the last half of the outer whorl; constrictions are not arctica, has similarly massive whorls with very coarse col obvious on the firsthalf of the outer whorl. Constrictions lar ribs and numerous weaker intercalatories to a larger may be succeeded by a much weaker adaperturai collar size. Patagiosites arbucklensis (Anderson, 1958) (see revi rib, and there are traces of occasional non-bullate ribs sion in Matsumoto (1959, p. 60, Pls. 16:1; 17:1-2) is from between the constrictions. Paralectotype IPB 22a (Fig. the Campanian of California. It has numerous long ribs 17F-H), from Siikerhook at Coesfeld, Westphalia is a between the collar ribs at a very small size, whereas P. sto paralectotype, again a crushed composite mould, with the baei has few or none, is more evolute during later growth following dimensions: D=67.1 (100); Wb = 16.2 (24.1); with subparallel flanks and very delicate flexuous flank Wh=25.5 (38.0); Wb:Wh=0.63; U= 18.8 (28.0). To a ribbing. Patagiosites alaskensis Jones, 1963 (p. 45, Pls. 38- 102 W.f, Kennedy & w.K. Christensen FOSSILS AND STRATA 44 (1997) Fig. 18. Patagiosites grijjithi (Sharpe, 1855). Lectotype, GSM 37238, from the Upper Campanian White Limestone of Derry, Northern 1reland. All xl. 40; 41:1, 3, 7, 9; Figs. 24-25) from the Maastrichtian of Grossouvre]. 01885 Ammonites sp. - Moberg, p. 25, Pl. Alaska has markedly concave constrictions, five to seven 2:6. 01922 Tragodesmoceras hernensis (Schliiter) - Spath, per whorl on the early whorls, which are smooth or p. 128. 01925 Tragodesmoceras hernense (Schliiter) - ribb ed, the constrictions persisting to a larger size than in Diener, p. 131. O non 1953 Tragodesmoceras hernense P. stobaei. The coiling is more evolute in middle growth, Schliit. - 0dum, p. 24, Pl. 4:3 [? = Tragodesmoceras the whorls slower expanding, to give a quite different shell clypeale (Schliiter, 1872)]. 01979 Nowakites hernensis shape. (Schliiter) - Matsumoto, p. 37. O? 1991 Nowakites cf. hernensis (Schliiter) - Kennedy & Christensen, p. 210, PI. Oecurrence. - The types are from the lower Upper Cam 1:3. 01991 Nowakites hernensis (Schliiter) - Kennedy & panian of southern Sweden. The speeies has also been Christensen, p. 210, Fig. 4. recorded from Donbass (Naidin 1974); Mons Basin, Bel gium; Liege, Belgium, and southern Limburg, The Neth Ty pe. - Holotype, by monotypy, is IPB 27, from the erlands. In Germany the spe eies ranges from uppermost 'Untersenonen grauen MergeIn des Schachtes von der Lower Campanian into the upper Upper Campanian. Heydt bei Herne in Westphalen' (Fig. 19A-B). Description. - There are a series ofspeeimens from depths of 1328-l364 m in the Svedala borehole. The best-pre Genus Nowakites Spath, 1922 served ofthese, from a depth of 1364 m, is a crushed com posite mould 62 mm in diameter (Fig. 19C). Coiling is Ty pe speeies. - Paehydiseus carezi de Grossouvre, 1894, p. moderately involute, with the apparently rather shallow 190, PIs. 25:3; 37:5, by original designation of Spath umbilicus compr ising 35% of the diameter. There are (1922, p. 124). four prominent, very distant umbilical bullae on the 2700 of the outer whorl preserved; these give rise to markedly concave prorsiradiate collar ribs that are succeeded by Nowakites hernensis (Schliiter, 1867) prominent, parallel constrictions. Between constrictions are numerous shorter, concave ribs, most obvious on the Fig. 19 outer half of the flank,ventrolateral shoulders and venter. Synonymy. - 01867 Ammonites Hernensis - Schliiter, p. The Ammonites sp. of Moberg (1885, p. 25, Pl. 2:6) is 35, Pl. 6:4. O non 1872 Ammonites Hernensis Schliiter - based on SGU Type 3874 (Fig. 19D-F). Hagg (1930) Schliiter, p. 40, Pl. 11:l3-14 [=Puzosia (Puzosia) muelleri referred it to Tragodesmoceras clypeale (Schliiter, 1867), FOSSILS AND STRA T A 44 (1997) Santonian to Maastrichtian ammonites 103 Fig. 19. Nowakites hernensis (Schliiter, 1867). DA-B. Holotype, IPB 27, the original of Schliiter (1867, p. 35, Pl. 6:4), from the 'Untersenonen grauen Mergein des Schachtes von der Heydt bei Herne in Westphalen'. De. SGU unregistered, from the Svedala borehole at a depth of 1364 m. DD-F. SGU Type 3874a, the original of Ammonites sp. of Moberg (1885, Pl. 2:6), from the upper Middle Santonian ofEriksdal. All xl. but the venter is broadly rounded with widely separated strictions, weaker bullae and much coarser ribs between constrictions and associated adapical collar ribs. A pro the collar ribs. Nowakites carezi (de Grossouvre, 1894) (p. nounced ridge, displaced to one side of the mid-venter, is 190, Pl. 25:3) and N. savini (de Grossouvre, 1894) (p. 152, due to crushing. Only the outer flanksand venter are pre Pl. 25:4) are stouter shells with much coarser ribs, less served, with well-developed concave ribbing. markedly differentiated collar-ribs, and less prominent constrictions. Nowakites paillettieanus (d'Orbigny, 1841) Discussion. - The sparse constrictions, prominent bullae (p. 339, Pl. 102:2) has near-even, coarser ribb ing, without and very weak ribs between collar ribs readily separate conspicuous bullae. Nowakites hernensis from the other member of the genus, as do its seemingly compressed, slowly expanding whorls, Occurrence. - Santonian of Herne, Westphalia and else reminiscent of certain Kossmaticeratinae. Nowakites where in the Miinster Basin, Germany, Eriksdal and the lemarchandi (de Grossouvre, 1894) (p. 173, Pl. 22:5) from Svedala borehole, Sweden, and, possibly Bornholm, the Santonian of the Corbieres, France, has fewer con- Denmark. 104 W,J. Kennedy & w.K. Christensen FOSSILS AND STRATA 44 (1997) Farnily Muniericeratidae Wright, 1952 Discussion. - 0dum (1953, p. 23, Pl. 3:2) figured a erushed fragment that may belong to this species from the Upper Campanian of the Hollviken-2 borehole at a depth Genus Tragodesmoceras Spath, 1922 of 611.12 m. We have not seen this speeimen. Ty pe speeies. - Desmoceras clypealoides Leonhard, 1897, p. Occurrence. - H. (H.) coesfeldiense is known from the 57, Pl. 6:2, by original designation ofSpath (1922, p. 127). Upper Campanian of the Miinster Basin in Germany, Ukraine, Russia, Turkrnenia, Limburg, The Netherlands, and northern Aquitaine, Franee. Tragodesmoceras clypeale (Schliiter, 1867) Synonymy. - 01905 Tragodesmoceras clypeale (Sehliiter) Wegner, p. 207. 01922 Tragodesmoceras clypeale Suborder Ancyloceratina (Sehliiter) - Spath, p. 128. 01925 Tragodesmoceras Wiedrnann, 1966 clypeale Sehliiter- Diener, p. 131 (with additional synon ymy). 0?1953 Tragodesmoceras hernense Sehliit.- 0dum, SuperfarnilyTu rrilitaceae Gill, 1871 p. 24, Pl. 4:3. Farnily Nostoceratidae Hyatt, 1894 Typ es. - Leetotype, here designated, is the original of Sehliiter (1872, Pl. 15:9-10, 13), from Salzberg bei Genus Bostrychoceras Hyatt, 1900 Quedlinburg, as are the figuredparaleetot ypes, whieh are in the eolleetions of the Museum fiir Naturkunde, Berlin. Type speeies. - Turrilites polyplocus Roemer, 1841, p. 92, Pl. 14: 1-2, by original designation of Hyatt (1900, p. 588). Discussion. - 0dum (1953) reeorded this spe eies fromthe Lower Santonian of the Hollviken-2 borehole. Bostrychoceras polyplocum SuperfarnilyHop litaceae H. (Roerner, 1841) Douville, 1890 Fig. 20 Farnily Placenticeratidae Meek, 1876 Synonymy. - 01841 Turrilites polyp locus - Roemer, p. 92, Pl. 14:1 only, non 2 [=Eubostrychoceras saxonicum (Sehliiter, 1876)]. O non 1953 Bostrychoceras polyplo Genus and subgenus Hoplitoplacenticeras cum Roem. - 0dum, p. 22, Pl. 1:8 [?=Nostoceras jun Paulcke, 1907 ior (Moberg) ]. O 1986a Nostoceras (Bostrychoceras) ICZN name no. 1345 (=Dechenoceras Kayser, 1924, p. 174) polyplocum (Roemer); Kennedy, p. 92, Pl. 6:1; Pl. 15:1-3, 5-8; Figs. 32A-B; 33A-E; 34A-H; 35A-D (with Typ e speeies. - Hoplites-Placenticeras plasticus Paulcke, full synonymy). 1907, p. 186; ICZN Opinion 555, 1959: name no. 1629. Discussion. - See Kennedy (1986a, p. 63) fo r a diagnosis and diseussion of Hoplitoplacenticeras and its subgenera, plus a preliminary aeeount of the German speeies of H. (Hoplitoplacenticeras) . Hoplitoplacenticeras (Hoplitoplacenticeras) cf. coesfeldiense (Schliiter, 1867) Synonymy. - compare: 01867 Ammonites Coesfeldiensis - Sehliiter, p. 14 (pars), Pl. 1:1, 4 only [non 2-3, ?=H. vari; non 5, ?=H. dolbergense (Sehliiter, 1876)]. 01953 Dechenoceras coesfeldiense Sehliit. - 0dum, p. 23, Pl. 3:2. 01986a Hoplitoplacenticeras coesfeldiense (Sehliiter) - Kennedy, p. 73, Pl. 9:9-10; Fig. 27B-C, F (with full Fig. 20. Bostrychoceraspol yplocum (Roemer, 1841). SGU unregistered, synonymy). Upper Campanian, Trelleborg borehole, 580 m. xl. FOSSILS AND STRATA 44 (1997) Santonian to Maastrichtian ammonites 105 Type. - Lectotype, by subsequent designation of Kennedy (l986a, p. 95), is the original of Roemer (1841, Pl. 14:1) from an unspecifiedlocality in northern Germany. Description and discussion. - A fragment from the Trelle borg borehole at a depth of 580 m (Fig. 20) shows the crushed base and part of the flank of most of a whorl of speeimen with an estimated whorl height of 22.5 mm. Fig. 21. Nostoceras junior (Moberg, 1885). Holotype, LO 732T, the orig Ornament is of dense, narrow prorsiradiate ribs on the inal of Moberg (1885, Pl. 3:14), probably middle Upper Campanian of outer flank, linked at and intercalating between two rows Kopinge. xL of small tuberdes (cf. Kennedy 1986a, Pl. 15:3-6; Fig. 33A, etc.). The spe eimen from the 'Lower' Campanian of the Hollviken -1 borehole at a depth of 804 m referred to Nostoceras junior (Moberg, 1885) this speeies by 0dum (1953, Pl. 1:8) is a small spire that Fig. 21 has constrictions and seems to lack tuberdes on the rela tively few ribs, and may be doser to the Lower Campanian Synonymy. - 01885 Turrilites junior - Moberg, p. 31, Pl. Nostoceras junior (Moberg, 1885) (see below). 3:14. 0?1913 Heteroceras polyplocum Roemer - Nowak, p. 385, Pl. 40:6; Pl. 45:48. O? 1953 Bostrychoceras polyplocum Roem. - 0dum, p. 22, Pl. 1:8. 01965 Bostrychoceras Occurrence. - Upper Upper Campanian, Hollviken-1 borehole, Sweden, Northern Ireland, Norfolk, England, (Mobergoceras) junior (Moberg) - Schrnid & Ernst, p. 342, Pl. 4:4. northern Aquitaine, France, Toral10la, Montesqui and elsewhere in Catalonia; Navarra, Spain; Germany; Typ e. - Holotype,by monotypy, is the original of Moberg Poland, Russia, Ukraine, Armenia, Kazakstan, Bulgaria, (1885, Pl. 3:4), LO 732T, fromKopinge, Sweden (Fig. 21 Iran, North Africa, Texas and northern Mexico. herein). Description. - The holotype consists of 1.3 whorls of body chamber with the adult aperture preserved, the Genus Nostoceras Hyatt, 1894 whorls in contact throughout, and the maximum pre served whorl height 14.5 mm. The upper whorl face is concave, but toa poorly preserved to reveal details of Ty pe speeies. - Nostoceras stantoni Hyatt, 1894, p. 469, by ornament. The outer whorl face is convex, with 16-18 original designation. coarse, narrow prorsiradiate straight to feebly sinuous ribs that extend across the lower whorl face, where they Discussion. - Genus Mobergoceras Schrnid Ernst, 1975, & are straight and radial, dedining in strength towards the has as type species Turrilites junior Moberg, 1885 (p. 31, umbilicus. There are three, perhaps fo ur constrictions on Pl. 3:14), by original designation. Schrnid & Ernst (1975) the fragment, each preceded by a markedly strengthened regarded Mobergoceras as a subgenus of Bostrychoceras rib. The aperture is prolonged into a rostrum and Hyatt, 1900. The illustrated material of Mobergoceras is appears somewhat contracted. limited to the holotype (Fig. 21), a speeimen figured by Discussion. - The coarse, distant ribbing and constric Nowak (1913, Pl. 40:6) as Heteroceras polyplocum, a third ti(Jnsreadily distinguish N. junior from other, nontuber figuredby Schrnid Ernst (1975, Pl. 4:4), and a possible & culate Nostoceras such as N. saundersorum Stephenson, crushed individual figuredby 0dum (1953, Pl. 1:7). This 1941 (p. 46, Pl. 83:6-8), which has flatterwhorls, a shal material differs from Bostrychoceras polyplocum and low interwhorl suture, and numerous delicate ribs. Nos allied speeies in its small size, coarse ribb ing, prominent toceras platycostatum Kennedy & Cobban, 1993a (p. 131, flared constrictions, coiling with whorls in tight contact, Pl. 2:16-17; Pl. 4:1-13, 33, 34, 37A) has a much wider api apparent lack of an extended uncoiled early growth stage cal angle, with numerous near-transverse ribs, and only and of a loosely coiled, recurved adult body chamber. In ill-defined constrictions. Most other Nostoceras species these respects it resembles certain Campanian Nostoceras are tuberculate. Hyatt, 1894, such as N. saundersorum Stephenson, 1941 Occurrence. - Upper Campanian of Kopinge, Sweden; (p. 46, Pl. 83:6-8). Accordingly, Mobergoceras is here 'Lower' Campanian of the Hollviken-1 borehole, Sweden; regarded as a synonym of Nostoceras rather than a sub uppermost Lower Campanian, Misburg, Germany; Cam genus of Bostrychoceras. panian of Kaliszanay, Poland. 106 W.f, Kennedy & w.K. Christensen FOSSILS AND STRA T A 44 (1997) Fig. 22. Lewyiteselegans (Moberg, 1885). DA-B. Holotype, LO 731T, the original of Moberg (1885, Pl. 3:10), probably middle Upper Campanian of Kiipinge. DC-D. LO 733t, the original of Helico ceras? sp. ofMoberg (1885, p. 33), prob ably middle Upper Campanian of Kiipinge, Sweden.DE -G. SGU unregis tered, uppermost Lower - lower Up per Campanian ofTosterup. All xl. Family Diplomoceratidae Spath, 1926 are narrow and transverse across the venter. A second fragment from Tosterup is in the SGU Collections (Fig. Subfamily Diplomoceratinae Spath, 1926 22E-G). This has a whorl breadth to height ratio of 0.7 approximately with a maximum preserved whorl height Genus Lewyites Matsumoto & of 17 mm. The rib index is 12, with delicate ventrai tuber Miyauchi, 1984 des on some ribs, and up to three nontuberculate ribs between. A somewhat larger fragment of venter is the Ty pe speeies. - Idiohamites(?) oronensis Lewy, 1969, p. original of Moberg's Helieoceras sp. from Kopinge (1885, 127, Pl. 3:10-11, by original designation by Matsumoto & p. 33; LO 733t; Fig. 22C-D herein); well-preserved, it has Miyauchi (1984, p. 64). sharp tuberdes, without dear ribs linking them across the venter, and one or two nontuberculate ribs separating the tubereulate ones. The largest fragment, also from Kopinge, is the original of Moberg's Seaphites (1885, p. Lewyites elegans (Moberg, 1885) 30; Pl. 3: 11; LO 730t; Fig. 23 herein) . This consists of part Figs. 22-23 of the flank, with one row of ventrai tuberdes present. Synonymy. -01885 Seaphites? - Moberg, p. 30, Pl. 3:11. The maximum preserved whorl height is 34.5 mm, the rib 01885 Aneyloceras? elegans - Moberg, p. 30, Pl. 3:10. index 11, with pairs of ribs linked at coarse tuberdes, with 01885 Helicoceras? sp. Moberg, p. 33. 01913 (?)Ani one to three nontuberculate ribs between tubereulate soceras elegans - Moberg sp. - Nowak, p. 384, Pl. 40:7. groups. 01986a Neoerioceras (Sehlueterella)? elegans (Moberg) - Kennedy, p. 102, Pl. 17:3-6. Discussion. - There is some variation in the rib index of the present material, which is tre at ed as belonging to a Ty pe. - Holotype, by monotypy, is the original ofMoberg single speeies. Lewyites oronensis (Lewy, 1969) (p. 127, Pl. (1885, Pl. 3:10), LO 73lT (Fig. 22A-B herein), probably 3: 10-11) from the Upper Campanian Mishash Formation fr om the middle Upper Campanian of Kopinge, Sweden. of Israel is a dose ally, with ribs joined in pairs at ventrai Deseription. - The holotype is a curved sector with part of tuberdes but only a single nontuberculate rib between. a straight shaftof a phragmocone 41.5 mm long, with a With further and better material from Sweden it may weU maximum preserved whorl height of 16 mm. The whorl prove to be junior synonym of elegans. Lewyites eireularis section is compressed oval in intercostal section, the ven (Lewy, 1969) (p. 128, Pl. 3:9; Pl. 4:3 only), also from the ter more narrowly rounded than the dorsum, with a Upper Campanian Mishash Formation ofIsrael, has a cir whorl breadth to height ratio of 0.84. The dorsum is very cular whorl section, similar flankor nament, but ribs that worn but appears to have been ornamented by delicate weaken or disappear on the venter. Lewyites clinensis fe ebly convex ribs. These strengthen markedly across the (Adkins, 1929) (p. 208, Pl. 6; 10-11) from the Upper dorsolateral margin, and are narrow and markedly rur Campanian Anacacho Limestone near Cline, Uvalde siradiate on the flankof the curved sector, less so on the County, Texas, has a nearly circular intercostal section, shaft, where the rib index is eight. Ribs join in pairs at rib index of seven, flank ribs linked in pairs at ventrai coarse ventrai tuberdes that are linked across the venter tuberdes, and tuberdes linked across the venter by pairs by a pair of delicate loop ed ribs. One, two, and perhaps of weakened ribs, on the adapical of the holotype these three nontuberculate ribs separate tubereulate ribs and support a low siphonal node. A single nontuberculate rib FOSSILS AND STRATA 44 (1997) Santonian to Maastriehtian ammonites 107 Fig. 23. Lewyites e1egans (Moberg, 1885). LO 730t, the original of Scaphites? of Moberg (1885, Pl. 3:11), probably middle Upper Campa nian of Kopinge. All xl. Fig. 24. Glyptoxoceras crispatum (Moberg, 1885). DA. Lectotype, original separates tuberculate ribs. Lewyites taylorensis (Adkins, of Moberg (1885, Pl. 3:12), SGU Type 3877. DB. Paralectotype, SGU 1929) (p. 209, Pl. 6:12-13), from the Upper Campanian Type 3876. Both speeimens are from the upper Middle Santonian of Pecan Gap Chalk of northeast Texas, is known from poor Eriksdal. All x l. phosphatic fragments only with pairs of strong ribs linked at ventrai tuberdes with two weaker, annular, nontuber culate ribs between. It remains inadequately known. Oeeurrence. - Upper Campanian of Kopinge and Toste rup, Sweden, northern Aquitaine, France, and Wagano Diseussion. - Kennedy (1986a) discussed this species in wice, Poland. the context of Glyptoxoceras retrorsum (Schliiter, 1872) (p. 97, Pl. 30:5-10) of the Upper Campanian, noting, however, that it was much older. Small specimens of ret Genus Glyp toxoceras Spath, 1925 rorsum (Schliiter, 1872, Pl. 30:8; Kennedy 1986a, Fig. 38D; Type speeies. - Hamites rugatus Forbes, 1846, p. 117, by Kennedy 1993, p. 108, Pl. 4:1-9, 11-19, 25, 26; Fig. 4) all original designation. have ribs that are fe ebly concave on the outer tlank,while larger specimens suggest an even, elliptical or circular coil. Glyptoxoceras aquisgranense (Schliiter, 1872) (p. 102, Pl. Glyptoxoceras crispatum (Moberg, 1885) 31:6-9; Kennedy et al. 1992, p. 274, PIs. 1:6-7, 11-12, 14- 19; 2:1-5, 9-15; 3:1-9) of the Lower Campanian has a Fig. 24A-B complex coiling ontogeny, although at the size repre Synonymy. - 01885 Anisoceras (Hamites?) erispatus - sented by the lectotype of G. erispatum it has a circular to Moberg, p. 32, Pl. 3:12-13. elliptical coil with a rib index of up to nine, but decreasing Typ es. - Lectotype, here designated, is the original of to fiveor six on some body chambers, the ribs rursiradi Moberg (1885, Pl. 3:12), fromthe Santonian ofEriksdal, ate, straight on the dorsal tlank but tlexed forwards in Sweden, SGU Type 3877 (Fig. 24A). Paralectotype is SGU many specimens and fe ebly concave on the ventrai part of Type 3876 (Fig. 24B). the tlank. Glyptoxoceras souqueti Collignon, 1983 (p. 186, Description. - The lectotype is a crushed, partially septate Pl. 1:4) from the Upper Santonian of the Corbieres has a composite mould 77 mm long and gently curved, with a variable rib index, from seven in the holotype to up to ten maximum preserved whorl height of 18 mm. The rib in other, as yet undescribed specimens, and is a dose ally, index is eight, the ribs narrow, sharp, feebly convex on the the ribs being straight to feebly convex and prorsi- to rur phragmocone, less so on the adapical body chamber, and siradiate (Fig. 25). near-transverse, straight, and feebly rursiradiate at the adaperturai end. Oeeurrence. - As for type. 108 W,J. Kennedy & w.K. Christensen FOSSILS AND STRATA 44 (1997) Fig. 25. Holotypeof Glyptoxoceras sou queti Collignon, 1983, from the Upper Santonian, 'chemin de Sougraine aux Croutets', Corbieres, Aude, France. M. Bilotte Collection, Toulouse. xl. Family Baculitidae Gill, 1871 Diseussion. - The holotype has so few diagnostic features that it cannot be usefully compared with other feebly ornamented Baeulites species. In the absence of knowl Genus Baculites Lamarek, 1799 edge of intraspecific variation and original, undeformed whorl section, sueeieus is best treated as a nomen dubium. Ty pe speeies. - Baeulites vertebralis Lamarck, 1801, p. 103, through subsequent designation by Meek (1876, p. 391). Baculites incurvatus Moberg, 1885, non Baculites suecicus Moberg, 1885 Dujardin, 1837 Fig. 26A Fig. 26C-E Synonymy. - 01885 Baeulites sueeieus - Moberg, p. 34, Pl. Synonymy. - 01885 Baeulites ineurvatus Dujardin 4:1. Moberg, p. 36, Pl. 4:2-4. Diseussion. -Baeulites ineurvatus Dujardin, 1837, is a Ty pe. - Holotype, by monotypy, is LO 734T, the original Coniacian species, revised by Kennedy (1984, p. 143, PIs. ofMoberg, 1885, Pl. 4:1 (Fig. 26A herein). Moberg noted 32:12, 15-19; 33:1-22; Figs. 41, 42F-M). Moberg (1885) that the specimen was labelled Kopinge, an Upper Cam referred a series of specifically indeterminate fragments to panian locality, but suggested on the basis of the matrix the species. His Pl. 4:2, SGU Type 3878, is fr om the upper that it may have come from Kåseberga, a Lower Campa Middle Santonian of Eriksdal (Fig. 26C). It is 37.5 mm nian locality. long with a maximum preserved whorl height of 13 mm, Description. - The holotype is a very crushed composite it has two prominent dorsolateral nodes preserved, and mould of a phragmocone with a maximum preserved prominent markedly prorsiradiate lateroventral growth whorl height of 53 mm. The original whorl proportions lines striae and riblets. The original of his Pl. 4:3, LO cannot now be determined. There is an obscure orna 735T, shown here as Figure 26D, is from the lower Lower ment of growth lines, striae and low undulations, but no Campanian of Kåseberga. It is 34 mm long, with a maxi clearly differentiated ribs. The ornament de fines a tran mum preserved whorl height of 9 mm. Ornament is of sient aperture with short dorsal and long ventral ros two widely separated dorsolateral nodes which give rise to trum, and markedly concave at mid-flank. Only traces of single concave crescentic ridges, projected markedly on the suture line survive, with quite deeply incised bifid the ventrolateral margin. The specimen may belong to the lobes and saddles. widely occurring group of Baculites eapensis W oods, 1906 FOSSILS AND STRATA 44 (1997) Santonian to Maastrichtian ammonites 109 Fig. 26. DA. Baculites suecicus Moberg, 1885. Holotype, LO 734T, from the lower Lower Campanian, Kåseberga?, the original of Moberg (1885, Pl. 4:1). DB. Baculites sp., SGU Type 3881, the original ofMoberg (1885, Pl. 4:7), from the upper Middle Santonian ofEriksdal. De. Baculites incurvatus Moberg, 1885 non Dujardin, 1837. SGU Type 3878, the original of Moberg (1885, Pl. 4:2), from Eriksdal. OD. Baculites incurvatus Moberg, 1885 non Dujardin, 1837. LO 735t, the original ofMoberg (1885, Pl. 4:3), from the lower Lower Campanian, Kåseberga. DE. Baculites incurvatus Moberg, 1885 non Dujardin, 1837. LO 736t, the original of Moberg (1885, Pl. 4:4). OF. Baculites cf. aquilaensis Reeside, 1927. MGUH 24152, Tosterup conglomerate, Ioc. CV2, Rod molla, uppermost Lower Campanian, OG Baculites cf. aquilaensis Reeside, 1927. MGUH 24153, Tosterup conglomerate, Ioc. CV2, Rodmolla, uppermost Lower Campanian. OH. Baculites cf. aquilaensis Reeside, 1927. RM Mo 135639, the original of Moberg (1885, Pl. 12:3), probably middle Up per Cam panian of Kopinge. DI. Baculites vertebralis Moberg, 1885 non Lamarek, 1801. LO 737t, the original of Moberg (1885, Pl. 13:9), Åhus. DJ. Baculites cf. aquilaensis Reeside, 1927. MGUH 24154. Tosterup conglomerate, Ioc. CV2, Rodmolla, uppermost Lower Campanian. DK. Baculites cf. aquilaensis Ree side, 1927. MGUH 24155. Tosterup conglomerate, Ioc. CV2, Rodmolla, upermost Lower Campanian. DL-M. Baculites vertebralis Moberg, 1885 non Lamarek, 1801. LO 738t, the original ofMoberg (1885, Pl. 4:9), Balsberg, probably lower Upper Campanian. All xl. (p. 342, Pl. 44:6-7). The original of Moberg's Pl. 4:4, Discussion. - Moberg (1885) compared two fragments shown here as Fig. 26E, is from Eriksdal. It is 38 mm long, from Eriksdal, a Santonian locality, to Schhiter's brevi with a maximum preserved whorl height of 7.5 mm, the costa, a Coniacian species. They fall within the variation whorl section markedly trigonal as a result ofpost -mortem range of the South African Baculites capensis group. crushing. There are six delicate crescentic dorsolateral nodes and associated ribs. The fragment is again a specif ically indeterminate member of the capensis group. Baculites Sp. Moberg, 1885 Fig. 26B Baculites brevicosta? Moberg, 1885 non Synonymy. - 01885 Baculites sp. - Moberg, p. 37, Pl. 4:7. Schhiter, 1876 Discussion. - This badly crushed fragment, SGU Type Synonymy. - 01885 Baculites brevicosta Schhiter? - 3881, from the upper Middle Santonian of Eriksdal, is Moberg, p. 37, Pl. 4:5-6. specifically indeterminate. 110 W.f. Kennedy & w.K. Christensen FOSSILS AND STRATA 44 (1997) Baculites cf. aquilaensis Reeside, 1927 section compressed oval. No ornament is preserved. The two specimens are specificaUyindeterminate. Fig. 26F-G, H, J-K Synonymy. - compare: 01885 Baculites anceps auctorum (Lamarck, d'Orbigny, Schluter, cet.) - Moberg, p. 37, Pl. 4: 1l-?12. 01927 Baculites aquilaensis - Reeside, p. 12, PIs. Baculites angustus Moberg, 1885 6:1 1-13; 8:1-4; var. separatus, p. 12, PIs. 8:15-2 1; 9:6-15; Fig. 27A-E 45:5-6; ?var. obesus, p. 12, Pl. 10:1-8. 01993 Baculites Synonymy. - 01885 Baculites angustus - Moberg, p. 39, aquilaensis Reeside - Kennedy, p. 110, Pl. 4:22, 24. Pl. 4:10. Description and discussion. - We have only seen one of Typ e. - Holotype, by monotypy, is the original ofMoberg Moberg's specimens of Baculites anceps auctorum (Fig. (1885, Pl. 4:10), LO 739T, from K6pinge. 26H), but they certainly do not belong to the Upper Maastrichtian Baculites anceps Lamarck, 1822 (see revi Description. - The holotype is a dark brown phosphatic sion in Kennedy 1986b, p. 58, PIs. 11:12-14; 12:7-11; Figs. intern al mould of parts of four chambers. The fragment is 3E-H, 7 A-C). Rather, they seem to belong to the same 21.8 mm long, with a maximum preserved whorl height gro up as a series of poorly preserved fragments from the of 15.3 mm and whorl breadth to height ratio 0.67. The uppermost Lower Campanian Tosterup conglomerate whorl section is ovoid, the venter more narrowly rounded shown in Figures 26F-G, J-K. These have a compressed than the feebly convex dorsum (Fig. 27D). Ornament is of ovoid whorl section when undeformed, the venter more delicate growth lines and riblets, ne ar effaced on the dor narrowly rounded than the dorsum, the expansion rate sum, feebly concave across the dorsal half of the flanks low. Ornament is of feebly concave strong ribs, two or and projected strongly forwards and effacing on the ven three in a distance equal to the whorl height, and extend tral half (Fig. 27B). The venter is worn. Suture line mod ing across the dorsal two-thirds of the flanks. They are erately incised, with broad-stemmed elements (Fig. 27E). strongly projected forwards on the ventrai third, weaken Discussion. - The locality K6pinge ranges from upper markedly and break down into riblets and striae, which Lower to middle Upper Campanian, while this phos may also intercalate between. Ornament effaces and phatic fragment may be older. In the absence of better cross es the venter in a narrow convexity. material it cannot be determined if angustus is a feebly Despite poor preservation, these specimens compare ribb ed species or a variant of a nodose species. It is best weU with the variable Baculites aquilaensis, originaUy treated as a nomen dubium, being based on inadequate described from the Lower Campanian of the United material. States Western Interior, but also recorded fr om the Mons Basin, Belgium. Occurrence. - Tosterup conglomerate, K6pinge, upper most Lower - lower Upper Campanian. Baculites vertebralis Moberg, 1885, non Lamarek, 180 l Fig. 26I, L-M Synonymy. - 01885 Baculites vertebralis Lamarck - Moberg, p. 38, Pl. 4:8-9. Discussion. - Moberg illustrated two specimens as Bacu lites vertebralis Lamarck, an Upper Maastrichtian species revised by Kennedy (1986b) and Birkelund (1993); nei ther ofhis specimens belong to the species. The original of his Pl. 4:8, LO 737t, from Ahus (Fig. 261 herein), is a single worn chamber with a maximum preserved whorl height of 20.8 mm. Better preserved is the original of his Pl. 4:9, LO 738T; Figs. 26L-M herein. It shows the adapical part Fig. 27. OA-E. Baculites angustus Moberg, 1885. LO 739T, original of of the specimen, a phosphatized fragment from Balsberg, Moberg (1885, Pl. 4:10), a phosphatic fragment from Kiipinge,probably middle Upper Campanian. OF. Baculites schlueteri Moberg, 1885. LO preservation suggesting a lower Upper Campanian hori 74lT, original of Moberg (1885, Pl. 4:14), a phosphatic fragment from zon. It is part-septate, 62 mm long, with a maximum pre Kiipinge, probably middle Upper Campanian. Figs. A-D and F are xl. served whorl height of 18 mm approximately, the whorl Bar scale ofsuture line ofB. angustus is 10 mm. FOSSILS AND STRATA 44 (1997) Santonian to Maastrichtian ammonites III Fig. 28. Bacu/ites anceps Lamarek, 1822. LO 740t, the original of Baculites sch/u teri? ofMoberg (1885, Pl. 4:13), from the Upper Maastrichtian Calcaire a Bacu/ites of Picouville, Manche, France. All xL Baculites schlueteri Moberg, 1885 Baculites knorrianus (Desmarest, 1817) Fig. 27F Fig. 29 Synonymy. - 01817 Baculites knorrianus - Desmarest, p. Synonymy. - 01885 Baculites schlUteri - Moberg, p. 40, Pl. 48, Pl. 1:3. 01987 Baculites knorrianus Desmarest - 4:14 0nly. Kennedy & Summesberger, p. 32, Pls. 4:4-6; 5:1-15; Fig. 2 (with full synonymy). 01993 Baculites knorrianus Des Typ e. - Lectotype, here designated, is the original of marest - Kennedy, p. 109, Pls. 5:13-22; 6:1 1-13; 18-23; Moberg (1885, p. 40, Pl. 4:14), LO 74IT, a phosphatized Fig. 5A-C. 01993 Baculites knorrianus Desmarest - Birke fragment from Kopinge. lund, p. 52, Pl. 13:12-14 (with additional synonymy). Description. - The lectotype is a 14.5 mm long fr agment Typ e. - Neotype, by subsequent designation of Kennedy with a maximum preserved whorl height of 12.4 mm and & Summesberger (1987, p. 33), is No. 7459a in the collec whorl breadth to height ratio of 0.74; parts ofthree cham tions of the Naturhistorisches Museum Vienna, from the bers are preserved. The whorl section is markedly ovoid, Lower Maastrichtian of Nagoryany, near Lvov in the with venter more narrowly rounded than dorsum (Fig. Ukraine (Kennedy & Summesberger, 1987, Pl. 5:5, 7-8). 27F). There are suggestions of delicate, concave flank Description. - MGUH 241 56, from Bjarnum, is a com ornament, but preservation is poor. Suture with moder posite mould of phragmocone 82 mm long, with parts of ately incised elements. six camerae preserved, the maximum whorl height 44.5 mm, the whorl breadth to height ratio 0.6, the whorl sec Discussion. - The fragment lacks the diagnostic fe atures tion compressed ovoid with venter more narrowly necessary to characterize the species, and in the absence of rounded than dorsum. The surface of the mould is more adequate material should be regarded as a nomen smooth but fo r traces of markedly prorsiradiate riblets dubium. and grooves on the ventraI part of the flanks. Suture line complexly incised with broad lobes and saddles (Fig. The specimen from Picouville, Manche, France, figured 29C). as a questionable Baculites schlueteri by Moberg (1885, Pl. 4: 13), LO 740T, is shown as Fig. 28. It is a typical Baculites Discussion. - Large size, compression, feeble ornament anceps Lamarck, 1822, from the Upper Maastrichtian (see and sutural complexity show this fragmentto be Baculites revision in Kennedy 1986b, p. 58, Pls. 11:12-14; 12:7-11; knorrianus, as revised by Kennedy & Summesberger Figs. 3E-H, 7A- C). (1987), Kennedy (1993) and Birkelund (1993). 112 W.]. Kennedy & w.K. Christensen FOSSILS AND STRAT A 44 (1997) Fig. 29. DA-C. east of Baculites knorri anus Desmarest, 1817. MGUH 24156, fromthe basal Lower Maastrichtian, Bjarnum. A-B are natural size. Bar scale of suture line is 10 mm. Occurrence. - Lower Maastrichtian, Sweden, Denmark, solaterai ribs and may be a separate speeies or a variant Poland, Czechoslovakia, Ukraine; lower Upper Maas (Fig. 30D). Most ofthese specimens resemble the smooth trichtian of Rørdal, Denmark (Birkelund 1993). Baculites known from the Campanian of the U.S. Western Interior (e.g., Cobban 1962, p. 714, Pl. 108:1-4; Fig. lI-J). A seeond, younger assemblage, probably from the mid Baculites Spp. dIe Upper Campanian of Kopinge (Fig. 30I-L), has indi viduals with whorl heights of up to 39 mm, one with the Fig. 30 aperture seemingly preserved (Fig. 30L). The dorsum is Discussion. - Two further assemblages of Baculites are broad and flattened and the venter narrowly arehed when represented by a small number of fragments, both unfor eompared with the Ivo assemblage, and ornament weak tunately laeking sufficient criteria for specific determina to absent. tion. The older assemblage is from the uppermost Lower Campanian ofIvo Klack (Fig. 30A-H). This includes frag ments with whorl heights of up to 40 mm. Most are Aptychi of Baculites smooth, with a compressed ovoid whorl section; a few Fig. 31 have growth lines, shallow furrows and riblets that define a transient aperture with short dorsal, and long ventrai Discussion. - The middle Upper Campanian at Kopinge rostrum (Fig. 30E). A single fragment has crescentic dor- has yielded a series of eomplete and fragmentary ammo- FOSSILS AND STRATA 44 (1997) Santonian to Maastrichtian ammonites 113 Fig. 30. DA-H. Baculites sp. from the uppermost Lower Campanian of Ivii Klack. DA-B. MGUH 24157; De. SGU Collections. DO. SGU unregistered. DE. SGU unregistered. DF-G. MGUH 24158. OH. SGU unregistered. DI-L. Baculites sp. from Kiipinge,probably middle Upper Campanian. 01-]. SGU unregistered. DK-L. SGU unregistered. All x L 114 W,J. Kennedy & w.K. Christensen FOSSILS AND STRATA 44 (1997) Fig. 31. Rugaptychus.DA. Syntype ofRuga ptychusrugosus (Sharpe, 1857, Pl. 26:9), Norwich Castle Museum unregistered, from the Upper Chalk of Nor wich, England. DB. Rugaptychusf1exus (Moberg, 1885), LO 748T, the original of Moberg (1885, Pl. 1:19), from the uppermost Lower Campanian or lower Upper Campanian of Balsberg. OG-H. Rugaptychus rugosus Sharpe of Moberg (1885), Kopinge, probably middle Upper Campanian. De. LO 744t, original of Moberg (1885, Pl. 1:16). OD. LO 746t, original of Moberg (1885, Pl. 11:18). DE. LO 743t, original ofMoberg (1885, Pl. 1:15). OF. LO 747t, original ofMoberg (1885, Pl. 6:26). OG. LO 745t, original of Moberg (1885, Pl. 1:17). OH. LO 742t, original of Moberg (1885, Pl. 1:14). A, E and H, approximately xl; B-O, F-G x2. nite jaws, referable to the fo rm genus Rugaptychus Trauth, also introduced a new taxon, Aptychusflexus, fo r material 1927. Schltiter (1876, Pl. 39:16) figured a specimen of from Kdpinge (Pl. 1: 19; Fig. 3lB herein), also referring to Baculites from the Lower Maastrichtian Mucronaten his new species the original of Schliiter (1876, Pl. 40:8), a kre ide of Liineburg,Ge rmany with Rugaptychus in place, further specimen from Kdpinge. Moberg (1885, p. 41) demonstrating the association. The type species of Rug also regarded Aptychus stobaei of Lundgren (1874, p. 73, ap tychusby the subsequent designation of Moore (1957, Pl. 3:14-16) as a synonym of rugosus ofSharpe, and lepto p. 468) is Aptychus rugosus Sharpe (1857, p. 57, Pl. 24:8- phyllus of Sharpe (1857, p. 57, Pl. 24:8-9) as a possible 9), from the Upper Campanian of Norwich, England, the synonym. Trauth (1927) renamed the original ofSchliiter more complete syntype of which is shown as Fig. 31A. As (1876, Pl. 39:16) Rugaptychus knorrianus. Sharpe (1857, p. 58) noted, Hebert (1856, Pl. 28:6) The only assemblage of Rugaptychus illustrated to date described, as Aptychus insignis, an aptychus of the same is that from the Campanian of Folx -les Caves, Brabant, typebut with coarser ornament. Moberg (1885, p. 14; PIs. Belgium (de Grossouvre 1908, Pl. 1O:7-l3; Kennedy 1987, 1:14-18; 6:26; see Fig. 31C-H) describedAptychus rugosus p. 192, Pl. 16:1-22), which includes fragmentsthat match from Kdpinge, his specimens encompassing both in morphology rugosus, insignis, and flexus of previous Sharpe's rugosus (Pl. 1:14-18; Fig. 31C-E, G-H herein) authors, suggesting that these may be jaws ofbut a single and Hebert's insignis (Pl. 6:26; Fig. 31F herein). Moberg Baculites species. FOSSILS AND STRATA 44 (1997) Santonian to Maastrichtian ammonites 115 Superfamily Scaphitaceae Gill, 1871 Family Scaphitidae Gill, 1871 Subfamily Scaphitinae Gill, 1871 Genus and subgenus Scaphites Parkinson, 1811 Type speeies. - Scaphites equalis J. Sowerby, 1813, p. 53, Pl. 18:1-3, by the subsequent designation of Meek (1876, p. 413). Fig. 32. DA-D. Scaphites (Scaphites) hippocrepis (DeKay, 1828) form III ofCobban (1969). RM Mo 6327a, the original ofMoberg (1885, Pl. 3:2), from the lower Lower Campanian ofKåseberga. A-C Xl; D x2. Scaphites (Scaphites) hippocrep is (DeKay, 1828) fo rm III of Cobban, 1969 Fig. 32 from the upper Lower Campanian conica/gracilis Zone Synonymy. - 01828 Ammonites hippocrepis - DeKay, p. (= conica/papillosa Zone) and gracilis/mucronata Zone, 273, Pl. 5:5. 01885 Scaphites cf. aquisgranensis Schliiter and noted records down to lingua/quadrata Zone of the Moberg, p. 26, Pl. 3:2. 01969 Scaphites hippocrepis lower Lower Campanian (Fig. 2). (DeKay) III - Cobban, p. 21, Pls. 3:1-25, 4:35-49, 5:36- 40; Figs. 2, 4, 10-11 (with full synonymy). 01975 Scaphites hippocrepis (DeKay) - Schrnid & Ernst, p. 332, Scaphites (Scaphites) binodosus Pl. 1:1-2; Fig. 1. 01993b Scaphites (Scaphites) hippocrepis (DeKay) - Kennedy & Cobban, p. 845, Figs. 15.1, 17. 1-32. Roemer, 1841 Typ e. - Neotype, designated by Kennedy (1986a, p. 118) is Synonymy. - 01841 Scaphites binodosus - Roemer, p. 90, no. 19483 in the collections of the Academy of Naturai Pl. 13:6. 01841 Scaphites injlatus - Roemer, p. 90, Pl. 14:3. Sciences of Philadelphia, the holotype of Scaphites cuvieri 01865 Scaphites binodosus Roemer - Roemer, p. 197, Pl. Morton (1829, p. 109, Pl. 71:1) from the deep cut of the 32:6. 01872 Scaphites binodosus A. Rom. - Schliiter,p. 79, Chesapeake and Delaware Canal, USA. Pl. 24:4-6. 01872 Scaphites injlatusA. Rom. - Schliiter,p. 78, Pls. 24:1-3; 27:1. O non 1897 Scaphites binodosus Rom. Discussion. - Moberg (1885, p. 26, Pl. 3:2) described and - Fritsch, p. 36, Fig. 20 (= S. kieslingswaldensis Langenhan illustrated as Scaphites cf. aquisgranensis Schliiter, a Grundey). 0?1899 Scaphites binodosus Ad. Rom. - microconch from the Kåseberga. This specimen is RM & Schliiter, p. 414. 01902 Scaphites injlatusRoemer - Ravn, 6327a (Fig. 32). It is badly crushed and distorted, and only p. 252. 01902 Scaphites binodosus Roemer - Ravn, p. 253. 22 mm long. The spire is 16 mm in diameter with coarse 01905 Scaphites injlatusRo emer - Wegner, p. 211. 01905 ribs, the body chamber has flattened, subparallel flanks, Scaphites binodosus Roemer - Wegner, p. 211. O non 1906 weak umbilical and six strong ventro lateral clavi, with Scaphites binodosus A. Roemer - Miiller Wollemann, p. well-developed ventrai ribs, all of which characters sug & 16, Pls. 9:4-6; 10:4 (=S. fisheri Riedel, 1931). 01911 gest fo rm III of hippocrepis as recognized by Cobban Hoploscaphites injlatus Romer - Nowak, p. 565. 01915 (1969), or possibly a transition to his form Il. Scaphites binodosus A. Roemer - Frech, p. 560, Fig. 6. Occurrence. - Scaphites (S.) hippocrepis III is widespread 01915 Scaphites injlatus F.A. Roemer - Frech, p. 561, Fig. in the Lower Campanian; in the United States it is known 8. 01920 Scaphites binodosus A. Roemer - Koplitz, p. 67, from Montana, Wyoming, South Dakota, Utah, Colorado Fig. 21. 01920 Scaphites injlatus A. Roemer - Koplitz, p. and New Mexico, Travis and Lamar Counties in Texas, 68, Figs. 22-23. 01925 Scaphites injlatus Roemer - perhaps Russel County in Alabama, and is common in Diener, p. 199. 01925 Discoscaphites binodosus Roemer - New Jersey and Delaware. In Europe, S. hippocrepis III or Diener, p. 209. 01930 Ammonite spee. 1-Hagg, p. 60, Pl. passage fo rms to S. hippocrepis Il are known from Aqui 5:9. 01931 Scaphites binodosus A. Rom. - Riedel, p. 700 taine and Alpes-Maritimes in France, southern England, (pars). 01935 Scaphites binodosus A. Roemer - Hagg, p. Germany, Belgium, The Netherlands, plus the present 60. 01943 Scaphites (Discoscaphites) binodosus A. Roemer record fromso uthern Sweden. The successive subspecies - Hagg, p. 78, figure on p. 79. 01947 Scaphites (Disco mark a series of Lower Campanian Zones in the United scaphites) binodosus A. Roemer - Hagg, p. 94. O? 1951 Dis States; in Europe, Schrnid and Ernst (1973) recorded it coscaphites cf. binodosus (Roemer) - Mikhailov, p. 95, Pl. 116 W,J. Kennedy & w.K. Christensen FOSSlLS AND STRATA 44 (1997) 9:45. 01986a Scaphites binodosus (Roemer) - Kennedy, p. Trachyscaphites spiniger spiniger 116, Fig. 39A-E. 01987 Scaphites binodosus Roemer - (Schhiter, 1872) Wright & Kennedy in Smith, Pl. 38:7, Il. Figs. 33-37 Na me of the sp eeies. - Kennedy (1986a, p. 116) stated that Synonymy. - 01841 Scaphites pulcherrimus - Roemer, p. he believed S. (5.) binodosus Roemer, 1841 (p. 90, Pl. 13:6) 91 (pars), Pl. 14:4. 01872 Scaphites spiniger - Schliiter, p. to be the microconch, and s. (5.) inflatus Roemer, 1841 82, Pl. 25:1-7. 01885 Scaphites spiniger Schliiter - (p. 90, Pl. 14:3) the macroconch of a single species, for Moberg, p. 28, Pl. 3:6-8. 01889 Scaphites spiniger which he selected the name binodosus. Schliiter - Griepenkerl, p. 405. 01894 Scaphites spiniger Schliiter- de Grossouvre, p. 252. O non 1908 Scaphites cf. Typ es. - The lectotype of s. (5.) binodosus, by the subse spiniger Schliiter- de Grossouvre, p. 38, Pl. 10:6; Fig. 13 quent designation of Kennedy (1986a, p. 116), is the orig [=Hoploscaphites constrictus (J. Sowerby)J. 01915 Sca inal of Roemer (1841, Pl. 13:6), refiguredby Frech (1915, phites spiniger Schliiter - Frech, p. 564, Fig. 13. 01916 Fig. 6), from Diilmen, Westphalia, in the collections of Acanthoscaphites spiniger Schliit. - Nowak, p. 67, figure opposite p. 66. 01925 Scaphites (Acanthoscaphites) spini the Geological Museum, Wroclaw, Poland. Frech (1915, ger Schliiter - Diener, p. 207. 01927 Acanthoscaphites Fig. 8) also refiguredthe original of Scaphites inflatusRo e spiniger (Schliiter) - Reeside, p. 34. 01951 Acantho mer, 1841, Pl. 141:3, but this specimen, from Diilmen, is scaphites spiniger (Schliiter) - Mikhailov, p. 100, Pl. 19:92. now lost. 01952 Scaphites spiniger Schliiter- Basse, p. 612. 01954 Scaphites (Acanthoscaphites) spiniger Schliiter - Hagg, p. Discussion. - Hagg (1943, figure on p. 79) illustrated the 58. 01954 Scaphites sp. 2 - Hagg, p. 58, Pl. 9:97. 01964 venter of a typical individual fr om lvo Klack, which we Trachyscaphites spiniger (Schliiter) - Cobban & Scott, p. have not traced. E8. 01974 Trachyscaphites sp iniger (Schliiter) - Naidin, p. 171, Pl. 59:2. 01975 Scaphites spiniger Schliiter- Schrnid Occurrence. - Lower Campanian, with the record from & Ernst, p. 330, Pls. 2:1-4; 3:3. 01976 Trachyscaphites lvo Klack confirmed here; Hagg (1935) notes other local spiniger spiniger (Schliiter) - Atabekian & Khakhimov, p. ities. The species is als o known from the Miinster Basin 66, Pls. 9:1-4; 12:2, 4. 01980 Trachyscaphites sp iniger and Braunschweig, Germany, Yorkshire, England, and spiniger (Schliiter) - Blaskiewicz, p. 30, Pl. 13:1-3, 5-7. 01980 Trachyscaphites spiniger posterior Blaskiewicz, p. Ukraine. 31, Pls. 13:4, 14:1-7; 15:2-3; 30:2. 01986a Trachyscaphites spiniger (Schliiter) - Kennedy, p. 130, Pl. 22:4; Fig. 42. 01992 Trachyscaphites spiniger spiniger (Schliiter) - Cob Scaphites (Scaphites) Spp. ban & Kennedy, p. 86, Pls. 1:2-3; 7:1-2, 5, 9; 8:1-9; Fig. 4A. 01993 Trachyscaphites cf. spiniger (Schliiter) - Kennedy, p. 113, Pl. 7:13. Discussion. - The Ammonites spee. 1 ofHagg (1930, p. 60, Pl. 5:9) from Eriksdal is the flank of a large, specifically Types. - Lectotype, by subsequent designation of Blaskie indeterminate Scaphites of the gro up of S. (5.) kieslings wicz (1980, p. 31), is the original of Schliiter (1872, Pl. waldensis fisheri Riedel, 1931 - S. (5. ) binodosus Roemer, 25:1-3), a macroconch, from the Upper Campanian of 1841, but is toa fragmentary for precise determination. Damp, Westphalia, an unregistered specimen in the lnsti The Ammonites spee. 3 ofHagg (1930, p. 61, Pl. 5: Il), also tut fiir Palaontologie der Universitat, Bonn (Fig. 35A-D). from Eriksdal (LO 3156) is part of the body chamber and Paralectotypes are lPB 61a, the original of Schliiter(1 872, Pl. 25:4), from the Hiigelgruppe of Haldem, an adult finalhook of a Scaphites (Scaphites) with a maximum pre microconch (Kennedy 1986a, Fig. 42A-B), and lPB 61b, served whorl height of only 6 mm. from the same horizon and locality, the original of Schliiter (1872, Pl. 25:6) (Kennedy 1986a, Fig. 42F). Genus Trachyscaphites Cobban & Scott, 1964 Fig. 33. OA-K. Trachyscaphites spiniger spiniger (Schlater, 1872), proba bly middle Upper Campanian, Kiipinge. DA. LO 7137t, microconch. Ty pe speeies. - Trachyscaphites redbirdensis Cobban & DB-C. LO 7138t, macroconch. OD-E. LO 7139t, macroconch. OF-G. Scott, 1964, p. E7, Pl. 1:1-7; Fig. 3, by original designa LO 727t, the original of Moberg (1885, Pl. 3:7). OH-K. LO 7140t, small tion. macroconch. All xL FOSSILS AND STRATA 44 (1997) Santonian to Maastrichtian ammonites 117 118 W.f, Kennedy & w.K. Christensen FOSSILS AND STRAT A 44 (1997) Fig. 34. DA-F. Trachyscaphites spiniger spiniger (Schluter, 1872), probably middle Upper Campanian, K6pinge. DA. LO 7141 t. DB-C. L0726t, the orig inal of Moberg (1885, Pl. 3:6), macroconch. OD. LO 728t, the original of Moberg (1885, Pl. 3:8), a macroconch fragment. DE-F. LO 7142t. All xl. FOSSILS AND STRATA 44 (1997) Santonian to Maastrichtian ammonites 119 Fig. 35. DA-F. Trachyscaphites spiniger spiniger (Schliiter, 1872). DA-D. Lectotype, IPE Collections, the original ofSchliiter (1872, Pl. 25:1-3), from the Upper Campanian of Darup, Westphalia, Germany. DE-F. LO 7143t, Kopinge, probably middle Upper Campanian. All xl. 120 W.j. Kennedy & w.K. Christensen FOSSILS AND STRATA44 (1997) Description. - We studied 43 fragmentary specimens, mostly from Kopinge. The species is strongly dimorphic. Phragmocones are very involute, with a tiny umbilicus. The whorl section varies from compressed (Fig. 33H-K) to depressed (Fig. 34D). Numerous delicate, straight, prorsiradiate primary ribs arise singly or in pairs at the umbilical shoulder. They are generally straight and pror siradiate, and may increase by branching and intercala tion. They link to a small outer lateral tubercle either sin gly or in pairs, or intercalate. The outer lateral tubercles give rise to 1-3 ribs, and these and the intercalated ribs loop, zigzag to, or intercalate between small conical inner ventrolateral tubercles, and connect in pairs or intercalate between small conical outer ventrolateral tubercles which alternate on either side of the venter; ribs loop, alternate or zig-zag between the outer ventrolateral tubercles (Figs. 33H-K, 36A-C). Some phragmocones develop small umbilicolateral/inner lateral tubercles at the adaperturai end (Fig. 36A-C). Microconch body chambers (Fig. 33A) are slender, with a concave umbilical wall and concave profIle to the umbilical seam, not occluding the umbilicus of the phrag mocone. Whorl sections vary from compressed to depressed. Delicate ribs arise at the umbilical seam and are concave across the umbilical wall. They link in groups at, or intercalate between seven coarse umbilical bullae, connected by delicate looped ribs to up to nine outer lat eral tubercles, with other delicate ribs intercalated between. There are 9-1 1 inner and outer ventrolateral clavi that alternate and are linked by looped and zig-zag ribs, with delicate ribs intercalating. There is some varia tion in the relative strength of tubercle rows, and tubercu lation may be strong or weak and may decline towards the adult aperture. The largest microconch in the collection is an estimated 60 mm long. Macroconchs (Figs. 33B-K, 34A-C, E-F) have phrag mocones from 35 mm (Fig. 33H-K) to 50 mm diameter (Figs. 33B-C, 34B-C) , and are both compressed and slightly depressed, as in the only undeformed fragment (Fig. 34D). The largest complete macroconch is 115 mm long. The body chamber has high whorls compared with that of the microconch, with a convex umbilical wall and a straight profIle to the umbilical se am that occludes and conceals part of the umbilicus of the phragmocone (Figs. 33I-K, 34B). The innermost row of tubercles is displaced out to an umbilicolateral position rather than being perched on the umbilical shoulder. Outer lateral, inner and outer ventrolateral tubercles are coarse, and linked by and separated by delicate ribs. There are both coarsely Fig. 36. DA-G. Trachyscaphites spiniger spiniger (Schliiter, 1872) from (Figs. 33B-C, D-E, 35E-F) and finely (Figs. 33H-K, the Kopinge sandstone, probably middle Upper Campanian. DA-C. RM 34B-C) tuberculate specimens; tuberculation weakens unregistered, the original of Scaphites sp. 2 ofHagg (1954, Pl. 9:97), Fre markedly towards the adult aperture (Figs. 34B-C, E-F), driksberg. DD-E. LO 7144t, Kopinge. DF-G. LO 7145t, Kopinge. DH L LO 7146t, juvenile scaphitid, cf. Trachyscaphites spiniger spiniger, especially the umbilicolateral and outer lateral rows (Fig. Kopinge. All x l. 33F-G). FOSSILS AND STRATA 44 (1997) Santonian to Maastrichtian ammonites 121 Fig. 37. Trachyscaphites spiniger spiniger (Schhiter, 1872). The holotype of Trachyscaphites spiniger posterior Blaskiewicz, 1980, the original ofBlaskiewicz (1980, Pl. 14:5-7), no. 1,310.11.10 in the Institute of Geology Collections, Warsaw, from the Upper Campanian of Sulejow, Poland. All xl. Discussion. - The type material of Trachyscaphites spiniger 1969 (p. 132, Pl. 4:1), from the Upper Campanian of comes from localities in the MiinsterBa sin and from Hal Israel, is based on a microconch and is a synonym of dem to the north. They have undergone substantial post porehi. mortem crushing and their very compressed whorl section Trachyscaphites pulcherrimus (Roemer, 1841) (see revi is in part illusory; the present material and that from sion in Kennedy & Summesberger 1984, p. In, PIs. 11:1- sideritic concretions described by Cobban & Kennedy 2, 10-22, 13:2-6) is easily distinguished from T. spiniger (1992) shows most speeimens to have slightly dep res sed by the presence of five rows of flanktubercles as weU as a whorls, although compressed individuals also occur. siphonal row in some individuals. Trachyscaphites spiniger posterior Blaskiewicz, 1980 (p. Occurrence. - Upper Campanian of Germany, The Neth 31, PIs. 13:4; 14:1-7; 15:2-3; 30:2), from the Upper Cam erlands, Belgium(?), Kopinge, Tosterup and Fredriksberg, panian of the Vistula Valley, Poland, was differentiated Sweden; Ukraine, Armenia, Turkmenia, Poland; the spe from the nominotypical subspecies because of the eies is restricted to the lower Upper Campanian where 'smaUer number of ribs runn ing between the tuberdes of precisely dated. In the United States it is best known from the same row on the exposed part of normal spiral and the the Ozan Formation in Fannin County, Texas. presence of lateroumbilical tuberculation on earlier sec tors of the exposed, normal spiral. It also differs on the whole in a smaller degree of freeing the shaftfr om phrag Genus Hoploscaphites Nowak, 1911 mocone and in a frequent lackof ribs between the tuber des on the same row or shaft.' The holotype is shown in Type speeies. - Ammonites constrictus - J. Sowerby, 1817, Fig. 37; we regard the name as a synonym of sp iniger sensu p. 189, Pl. Al, by original designation. strieto. In contrast, Trachyscaphites sp iniger porchi (Adkins, 1929) (p. 205, Pl. 5:1-3), of which Scaphites aricki (Adkins, 1929) (p. 206, Pl. 5:7-8) is a synonym (see Hoploscaphites ikorfatensis Cobban & Scott 1964, p. E10, PIs. 2:1-23; 3:1-11; Fig. 4), Birkelund, 1965 differsfr om the nominotypical subspecies in having fewer Fig. 38 tubercles in all rows on the body chamber and generally lacking the den se ribbing so well-displayed by the present Synonymy. - 01872 Scaphites Romeri d'Orbigny - specimens. Trachyscaphites spiniger levantinensis Lewy, Schliiter, p. 89, (pars), Pl. 27:1-3, non 4 (=Hoploscaphites 122 W.]. Kennedy & W.K. Christensen FOSSILS AND STRATA 44 (1997) greenlandieus Donovan). 01885 Seaphites Roemeri d'Orbigny - Moberg, p. 29, Pl. 3:9. 01911 Acantho seaphites Rbmeri d'Orbigny - Nowak, p. 585. 01916 Aeanthoscaphites Rbmeri d'Orbigny - Nowak, p. 62. 01953 Seaphites (Aeanthoseaphites) roemeri d'Orbigny - 0dum, p. 25, Pl. 2:2. 01954 Seaphites (Acanthoseaphites) roemeri d'Orbigny - Hagg, p. 57. 01954 Scaphites green landieus Donovan - Donovan, p. 8 (pars), Pl. 2:4; non Pl. 2:5, Fig. l ( = H. greenlandieus). 01965 Seaphites (Hoplo seaphites) ikorfatensis - Birkelund, p. 102, PIs. 24:1-4; 25:1-2; 26:1; Figs. 59, 93, 121 (3). 01965 Seaphites (Hoploseaphites) ravni - Birkelund, p. 106 (pars), PIs. 26:2-4; 27:1-4; ? non Pl. 28:1; non Pl. 29:1 ( = H. greenlan dicus); Figs. 95-96, ? 97, 100 (pars), 121 (4), ? non 121 (5). Name of the sp eeies. - We regard ikorfatensis of Birkelund (1965) as the macroconch, of which co-occurring speci mens of ravni of Birkelund (1965) (including the holo type) is the microconch. As firstrev ising authors we select the name ikorfatensis for the species for those who accept this view. Typ e. - Holotype is MGUH 9815, the original of Birke lund (1965, Pl. 26:1) from the Upper Campanian of Brud kløft at lkorfat, West Greenland, 550 m above sea level. The holotype of H. ravni is MGUH 1819, the original of Birkelund (1965, Pl. 27:1, Fig. 95), from the same horizon and locality. Description. - A phragmocone from Kopinge is 17 mm in diameter with the adapical part of the body chamber pre served (Fig. 38H). It is ornamented by dense prorsiradiate branching and intercalated ribs. A fragment from this locality with a whorl height of 17 mm (Fig. 38D-E) has delicate concave ribs at the umbilical shoulder that sweep forwards across to the mid-flank, where they increase by branching and intercalation and pass across the venter in a very shallow convexity. There are three delicate ventrai tubercles on the fragment, which link up to three ribs, with four or more ribs intercalated between. A further fragmentmay be from dose to the adult aperture, with an estimated whorl height of 18 mm (Fig. 38F-G). It has del icate ribs and lacks tubercles. Much larger is Moberg's specimen from Tosterup (1885, Pl. 3:9), LO 729t (Fig. 381-K herein). This is a fragmentof a body chamber with a maximum preserved whorl height of 31 mm, lacking the umbilical region. The whorl section is compressed, with flattened, convergent flanks, narrowly rounded ventrola teral shoulders and a feebly convex venter. Ornament is of Fig. 38. OA-K. Hoploscaphites ikorfatensis Birkelund, 1965. OA-B. SGU unregistered, the original ofØdum (1954, Pl. 1:7), silicone squeeze, Upper Cam panian ofHiillviken-l borehole, depth 643.4 m. De. SGU unregistered, the original ofØdum (1954, Pl. 2:2), silicone squeeze, Upper Campanian, Hiill viken-l borehole, depth 639.25 m. OD-E. MGUH 241 59, silicone mould, Kiipinge,probably middle Upper Campanian. OF-G. MGUH 24160, silicone mould, Kiipinge,probably middle Upper Campanian. OH. MGUH 24161, silicone mould, Kiipinge, probably middle Up per Campanian. OI-K. LO 729t, the original of Moberg (1885, Pl. 3:9), probably lower Upper Campanian, Tosterup. All xl. FOSSILS AND STRATA 44 (1997) Santonian to Maastrichtian ammonites 123 delicate, prorsiradiate, feebly flexuous ribs/lirae which Moberg (1885) recorded the Maastrichtian Scaphites appear to increase by branching low on the flank, coarsen constrictus from Kopinge; these records may well be based across the flanks and are near-transverse on the venter. on fragments, or microconchs, of the present species. Up to seven ribs link at coarse ventrai clavi, with a similar number of ribs intercalated between. Occurrence. - Upper Campanian of West Greenland, The crushed specimens fromthe Hollviken-l borehole Kopinge, Tosterup, and the Hollviken-l borehole, Swe are shown as Fig. 38A-C, and differ in no significant den; the Miinster Basin and Haldem, Westphalia, Ger respects from the better-preserved Kopinge material. many. Discussion. - Birkelund (1965) studied several hundred well-preserved and precisely localized Upper Campanian Hoploscaphites from West Greenland and clarified the Hoploscaphites constrictus (J. relationships of the northwest European Scaphites roemeri Sowerby, 1817) group of authors. She recognized three species. H. ikorfa t ensis was based on macroconchs, with a single row of Fig. 39A-E three to ten ventrolateral tubercles on the body chamber, or rarely nodeless. H. greenlandicus comes from a higher Synonymy. - 01817 Ammonites constrictus - J. Sowerby, horizon and is also based on macroconchs and generally p. 189, Pl. Al. 01986b Hoploscaphites constrictus (J. Sow lacks tubercles, or may have only a few ventro lateral erby) - Kennedy, p. 64, Pls. 13:1-13; 16-24; 14:1-38; tubercles on the body chamber. Hoploscaphites ravni is 15:1-31; Figs. 9, 11A-B (with full synonymy). 01993 based on microconchs; the holotype is from the same level Hoploscaphites constrictus (J. Sowerby) - Birkelund, p. 57, as H. ikorfatensis, but Birkelund also referred to it speci Pls. 14:1-7, 12; 15:1-14; 16:6-16; 17:5-23 (with addi mens that co-occur with H. greenlandicus. It was diag tional synonymy). 01993 Hoploscaphites constrictus (J. nosed as having a single row of 0-10 ventrolateral tuber Sowerby) - Kennedy, p. 113, Pl. 7:1-11, 14-16. 01993 cles on the body chamber, the tubercles rarely extending Hoploscaphites constrictus (J. Sowerby) - Hancock & onto the phragmocone. The co-occurring macro-and Kennedy, p. 166, Pl. 20:1-4 (with additional synonymy). microconch assemblages represent a dimorphic pair in our view, with ravni of Birkelund encompassing micro Ty pes. - Lectotype, by the subsequent designation of conchs ofboth H. ikorfatensis and H. greenlandicus. Kennedy (1986b, p. 68), is BMNH C36733, the original of The Scaphites compressus of Roemer, 1841, was J. Sowerby (1817, Pl. Al); paralectotypes are BMNH regarded as a homonym of Scaphites compressus of C70645-70647, all from the Upper Maastrichtian Cal d'Orbigny by Giebel (1849), who renamed it Scaphites caire a Baculites of the Cotentin Peninsula, Manche, tuberculatus, and also by d'Orbigny (1850) who, unaware France. of Giebel's action, renamed it Scaphites roemeri, to which species the present material was fo rmerly referred. The Discussion. - Birkelund (1993) fully documented Swedish date of compressus of Roemer is 1841, but although occurrences of this species and the closely related H. ten d'Orbigny (1850, p. 214) gives the date ofhis compressus uistriatus (Kner, 1848). We illustrate here Moberg's fig as 1841 and accords it priority over compressus ofRoemer, ured specimens. the careful analysis of Sherborn (1889) shows that livrai sons 33-42 (pp. 431-662) of d'Orbigny (1840-1842) date Occurrence. - Hoploscaphites constrictus ranges through from 1842, so that compressus of Roemer has priority. out almost all the Maastrichtian. At Kronsmoor in north Kennedy (1984, p. 148) discusses the affinities of ern Germany, the firstspecimen appears 3.5-5.0 m above d'Orbigny's compressus, and Birkelund (1965) discusses the base of the Belemnella lanceolata Zone, while at Stevns those of roemeri as interpreted by Schliiter (1872). The Klint in Denmark it is common in the topmost dense ribbing that covers the adult phragmocone and hardground of the Maastrichtian White Chalk, which is body chamber of Scaphites compressus Roemer, 1841, is immediately overlain by the Palaeocene Fish Clay. It is highly distinctive and, together with the sto ut whorl sec known from the Biscay region of France and Spain, tion and bituberculation, distinguish it from the present Petites Pyrenees (Haute Garonne), Tercis (Landes) and species and other Old-World scaphites. It is a Jeletzkytes the Calcaire a Baculites of the Cotentin Peninsula (Man and resembles J. nodosus (Owen, 1852) (p. 581, Pl. 8:4) in che) (all in France), the Nekum and Meerssen Chaiks of general arrangement of ribs and tubercles, but the ribbing the Maastricht region in Belgium and The Netherlands, is finer even than that of the holotype of Owen's species, Germany, Denmark, southern Sweden, Poland, Austria and the tubercles are weaker. Scaphites elegans Tate, 1865 (Styria), the Czech Republic, Bulgaria, Ukraine, Donbass, (p. 37, Pl. 3:3; see Riccardi, 1983, Pl. 10:22-23) is no more Carpathians, Transcaspia, Kazakhstan, and Kopet Dag than an uncrushed example of Jeletzkytes compressus. (Turkmenia) . 124 W.f, Kennedy & W.K. Christensen FOSSILS AND STRAT A 44 (1997) Occurrence. - High Lower to low Upper Maastrichtian. There are records from Denmark, Sweden, Germany, The Netherlands, Poland, the Czech Republic, Ukraine and southern Russia. Genus Acanthoscaphites Nowak, 1911 Type speeies. - Scaphites tridens Kner, 1848, p. 10, Pl. 2:1, by subsequent designation by Diener (1925, p. 205). Acanthoscaphites Sp. Fig. 40 Synonymy. - 01953 Scaphites (Acanthoscaphites) tridens Kner - 0dum, p. 24, Pl. 1:4. Fig. 39. DA-E. Hoploscaphites constrictus (J. Sowerby, 1817), upper Description. - The specimen is a composite mould, 37 Lower Maastrichtian, Ulricelund near Nasbyholm. DA. LO 725t, the mm in diameter, and may indude part of the body cham original of Moberg (1885, Pl. 3:5). DB-C. LO 724t, the original of ber. Co iling is involute, the umbilicus small, shallow, with Moberg (1885, Pl. 3:4). DD-E. LO 723t, the original of Moberg (1885, Pl. 3:3). OF. Hoploscaphites tenuistriatus (Kner, 1848). SGU Collections, a flattened umbilical wall and narrowly rounded umbili the original of 0duffi (1954, Pl. 1:6), Upper Maastrichtian, Hollviken- 1 cal shoulder. Widely-separated narrow ribs are concave borehole, depth 375.57 ffi. All xl. on the innermost flankbut become straight and prorsira diate and split in to pairs of delicate riblets or lirae on the inner flankwith numerous delicate growth lines between. Ribs are feebly concave on the outer flankand terminate in delicate conical ventrai tuberdes. The ventrai view is partially concealed (Fig. 40A), but delicate growth lines link to a second row of tuberdes, while traces of a third row, alternating in position with the firstrow are also vis ible. These tuberde rows may be inner and outer ventro lateral or ventrai, siphonal and ventrai; it cannot be deter mined if there were three or fiverows of tuberdes across the venter. Discussion. - This specimen may be a microconch of Acanthoscaphites dose to A. varians (Lopuski, 1911) (pp. 120, 137, Pl. 4:1-3; Fig. l; see revision in Birkelund 1993, Fig. 40. Acanthoscaphites sp. SGU unregistered, original of Scaphites tri dens Kner of 0dum (1954, Pl. 1:4), from the Maastrichtian of the Holl p. 56; PIs. 9:3-7; 10:2-3), but uncertainty as to ventrai viken-1 borehole, depth 314.5 ffi.All x l. tuberculation predudes dose comparisons. Occurrence. - Upper Maastrichtian, Hollviken-1 bore hole, 314.5 m. Hoploscaphites tenuistriatus (Kuer, 1848) Acknowledgements. - We thank G. Kjellstrom, Geological Survey of Fig. 39F Sweden, Uppsala, S. Laufeld, formerly at the Geological Survey of Swe Synonymy. - 01848 Scaphites tenuistriatus - Kner, p. 10, den, Uppsala, J. Bergstrom, Swedish Museum of Natural History, Sec tion of Palaeozoology, Stockholm, and K. Larsson, Geological Institute, Pl. l :5. 01987 Hoploscaphites tenuistriatus (Kner) - University of Lund, for placing ammonites at our disposal. We also Kennedy, p. 210, Pl. 31:2-7 (with synonymy). 01993 thank H.C. Klinger (Cape Town), A.V. Dhondt (Brussels), M. Sander Hoploscaphites tenuistriatus (Kner) - Birkelund, p. 59, Pl. (Bonn), A. Blaskiewicz (Warsaw), W.A. Cobban (Denver), U. Kaplan 14:8-1 1, 13-16. (Giitersloh), U. Scheer (Essen), the late H. jaeger (Berlin), T. Stuart (Norwich), H. Gauthier (Paris), and M. Bilotte (Toulouse) for advice, Ty pe. - Kner's specimen from Lemberg (Lvov) (1848, Pl. discussion, and access to collections in their care. P. Bengtson (Heidel berg) reviewed the manuscript. Kennedy acknowledges the financial 1:5) should be designated lectotype if still in existence. support of the Naturai Environment Research Council (U.K.), and the technical assistance of the staffof the Geological Collections, University Discussion. - Birkelund (1993) revised Danish OCCUf Museum, Oxford and Department of Earth Sciences, Oxford. Thomas rences. A juvenile from the Hollviken-1 borehole at a Bredsdorff, Geological Museum, Copenhagen prepared the photo depth of375.75 m is shown as Fig. 39F. graphic illustrations. FOSSILS AND STRATA 44 (1997) Santonian to Maastrichtian ammonites 125 References Christensen, W.K. & Schulz, M.-G. 1976: First record of Belemnello camax balsvikensis (Brotzen, 1960) from NW Germany. Neues Jahr Adkins, W.S. 1929: Some Upper Cretaceous Taylor ammonites from buch fu r Geologie und Paliiontologie, Monatshefte 9, 522-531. Texas. Universityof Texas Bulletin 2901, 203-211, 220-223. Christensen, W.K. & Schulz, M.-G. 1997: Coniacian and Santonian be Anderson, F.M. 1958: Upper Cretaceous of the PacificCoa st. Geological lemnite faunas from Bornholm, Denmark. Fossils and Strata 44, 1-73. Societyof America Memoir 71, 378 pp. Cobban, W.A. 1 962: Baculites from the lower part ofthe Pierre Shale and Arnold, H. 1964: Die Erforschung der westfaJischenKreide und zur Def equivalent rocks in the Western Interior. Journal ofPaleontology 36, inition der Oberkreidestufen und -zonen. Fortschritte in der Geologie 704--71 8. von Rheinland und Westfalen 7, 1-14. Cobban, W.A. 1969: The Late Cretaceous ammonites Scaphites leei Ree Atabekian, A.A. & Khakhimov, F.Kh. 1976: Kampanskie i maastrikhtskie side and Scaphites hippocrepis (DeKay) in the Western Interior of the ammonity srednej Azii. [Campanian and Maastrichtian ammonites United States. United States Geological Survey Professional Paper 619, from central Asia.] 146 pp. Donish, Dushanbe. [In Russian.] 1-27. Baily, W.H. 1855: Descriptions of some Cretaceous fo ssils from South Cobban, W.A. & Kennedy, W.J. 1992: Campanian Trachyscaphites sp ini Africa. Quarterly Journal of the Geological Society of London 11, ger ammonite fauna in north-east Texas. Palaeontology 35, 63-93. 454-465. Cobban, W.A. & Scott, G.R. 1964: Multinodose scaphitid cephalopods Basse, E. 1952: Ammonoides. In Piveteau, 1. (ed.): Traite de Paleontologie from the lower part of the Pierre Shale and equivalent rocks in the 2, 522-555, 581-688. Masson, Paris. conterminous United States. United States Geological Survey Profes Binkhorst, 1.T. 1861: Monographie des Gastropodes et des Cephalopodes sional Paper 483-E, E1-E13. de la Craie Superieur du Limbourg. 1 7 + 83 pp. (gastropods) +44 pp Collignon, M. 1952: Ammonites neocretacees du Menabe (Madagas (cephalopods), Muquardt, Maastricht; Muller Freres, Brussels. car). Il - Les Pachydiscidae. Travaux du Bureau Geologique du Haut Birkelund, T. 1957: Upper Cretaceous belemnites from Denmark. Biol Commissariat de Madagascar et Dependances 41. 114 pp. ogiske Skrifter. Det Kongelige Danske Videnskabernes Selskab 9. 69 pp. Collignon, M. 1955: Ammonites neocretacees du Menabe (Madagas Birkelund, T. 1965: Ammonites from the Upper Cretaceous of West car). IL Les Pachydiscidae. Annales Geologiques du Service des Mines Greenland. Meddelelser om Grønland 179. 192 pp. de Madagascar 21. 98 pp. Birkelund, T. 1973: A note on Pseudopuzosia sp. from Siirdal. In Berg Collignon, M. 1961: Ammonites neocretacees du Menabe (Madagas stram, J., Christensen, W.K., Johansson, C. & Norling, E.: An exten car). VII. Les Desmoceratidae. Annales Geologiques du Service des sion of Upper Cretaceous rocks to the Swedish west coast at Sardal. Mines de Madagascar 31. 115 pp. Bulletin of the Geological Societyof Denmark 22, 141-142. Collignon, M. 1971: Atlas des fo ssiles characteristiques de Madagascar Birkelund, T. 1993: Ammonites fromthe Maastrichtian White Chalk in (Ammonites) XVII (Maestrichtien). Service Geologique, Tananarive. Denmark. Bulletin of the Geological Societyof Denmark 40, 33-8 1. 44 pp. Birkelund, T. & Bromley, R.G. 1979: Hauericeras cf. pseudogardeni in the Collignon, M. 1983: Les faun es d'ammonites du Santonien. Documents Upper Cretaceous of Ignaberga, Sweden. Geologiska Foreningens i des Laboratoires de Geologie de Lyon H. S. 6 (fo r 1981), 184-224. Stockholm Forhandlingar 101, 1 73-176. DeKay, J.E. 1828: Report on severai fo ssil multilocular shells from the state of Delaware: with observations on a second specimen of the new Blaskiewicz, A. 1980: Campanian and Maastrichtian ammonites of the fo ssil genus Eurypterus. Annals of the Lyceum of Natural History of Middle Vistula River Valley, Poland: a stratigraphic-paleontologic New York 2, 273-279. study. Prace Instytutu Geologicznego 92. 63 pp. Desmarest, A.G. 1817: Memoire sur deux genres coquilles fossiles ciois Boule, M., Lemoine, P. & Thevenin, A. 1906-1907: Paleontologie de onnees et siphon. Journal de Physique de Chimie, d'Histoire Na Madagascar Ill. Cephalopodes cretaces des environs de Diego-Sua it tureIles et des Arts 85, 42-51. rez. Annales de Paleontologie 1, 173-192 [ 1-20]; 2, l-56 [21-76]. Diener, C. 1925: Ammonoidea neocretacea. Fossilium Catalogus (1: An Brinkmann, R. 1935: Die Ammoniten der Gosau und des Flysch in den imalia) 29. 244 pp. nardlichen Ostalpen. Mitteilungen aus dem Geologischen Staatsinstitut Donovan, D.T. 1954: Upper Cretaceous fossils fr om Traill and Geo im Hamburg 15, 1-14. graphical Society 0er, East Greenland. Meddelelser om Grønland 72. Brotzen, F. 1945: De geologiska resultaten från borrningarne vid Hall 33 pp. viken. !. Kritan. Sveriges Geologiska Undersokning C465. 64 pp. Douville, H. 1890: Sur la classificationdes Ceratites de la Craie. Bulletin Brotzen, F. 1958: [Cretaceous.] In Magnusson, N.H. (ed.): Lexique de la Societe Geologique de France (3) 18, 275-292. Stratigraphique International. 1. Europe. Fase. 2c: Sweden. Paris. Dujardin, F. 1837: Memoire sur les couches du sol en Touraine et Brotzen, F. 1960: The Mesozoic of Scania, Southern Sweden. Interna description des coquilles et la craie et des faluns. Memoire de la Societe tional Geological Conress, XXI Session, Norden 1960. Guide to Excur Geologique de France 2, 211-311. sions A21 and C16. 15 pp. Erlstram, M. & Gabrielson, J. 1992: Petrology, fo ssil composition and Christensen, W.K. 1970: Actinocamax primus Arkhangelsky from depositional history of the Ignaberga limestone, Kristianstad Basin, Scania, Sweden. Stockholm Contributions in Geology 21, 69-76. Scania. Sveriges Geologiska Undersokning Ca80. 30 pp. Christensen, W.K. 1973: The belemnites and their stratigraphic signifi Ernst, G. 1964: Ontogenie, Phylogenie und Stratigraphie der Belem cance. In Bergstram, J., Christensen, W.K., Johansson, C. & Norling, nitengattung Gonioteuthis Bayle aus dem nordwestdeutschen Santon/ E.: An extension of Up per Cretaceous rocks to the Swedish west coast Campan. Fortschritte in der Geologie von Rheinland und Westfalen 7, at Sardal. Bulletin of the Geological Societyof Denmark 22, 113-140. 113-174. Christensen, W.K. 1975: Upper Cretaceous belemnites from the Kris Ernst, G., Schrnid,F. & Klischies, G. 1979: Multistratigraphische Unter tianstad area in Scania. Fossils and Strata 7. 69 pp. suchungen in der Oberkreide des Raumes Braunschweig-Hannover. Christensen, W.K. 1985 [date of imprint 1984]: The Albian to Maas In Wiedmann, J. (ed.): Aspekte der Kreide Europas. International Un trichtian of southern Sweden and Bornholm, Denmark: a review. ion of Geological Sciences A6, 1 1-46. Cretaceous Research 5, 313-327. Forbes, E. 1846: Report on the fossil invertebrata fromsouthern India, Christensen, W.K. 1986: Up per Cretaceous belemnites fromthe Vomb collected by Mr. Kaye and Mr. Cunliffe. Transactions of the Geological Trough in Scania. Sveriges Geologiska Undersokning Ca57. 57 pp. Societyof London (2) 7,97-174. Christensen, W.K. 1993: Upper Cretaceous belemnitellids from the Bå Frech, F. 1910: Geologische Beobachtungen im pontischen Gebirge. stad Basin, southern Sweden. Geologiska Foreningens i Stockholm Neues Jahrbuch forMinera logie, Geologie und Paliiontologie 1910 (1), Forhandlingar 115, 39-57. 1-24. Christensen, W.K. 1995: Belemnitella fromthe Upper Campanian and Frech, F. 1915: tiberScaphites. 1. Die Bedeutung von Scaphites fur die Lower Maastrichtian Chalk of Norfolk, England. Special Papers in Gliederung der Oberkreide. CentralbIattfu r Mineralogie, Geologie und Palaeontology 51. 84 pp. Paliiontologie 1915, 553-568. 126 W,J. Kennedy & w.K.Christensen FOSSILS AND STRATA 44 (1997) Fritsch, A. 1897: Studien im Gebiete der bohmischen Kreideformation. }eletzky, }.A. 1951: Die Stratigraphie und Belemnitenfauna des Ober Palaeontologische Untersuchungen der einzelnen Schichten. VI. Die campan und Maastricht Westfalens, Nordwestdeutschlands und Chlomeker Schichten. Archiv fu r die Naturwissenschaftliche Landes Danemarks, sowie einige allgemeine Gliederungs-Probleme der durchforschung von Bohmen 10. 84 pp. jiingeren borealen Oberkreide Eurasiens. Beihefte Geologisches Jahr Fritsch, A. & Kafka, }. 1887: Die Crustaceen der bohmischen Kreideforma buch 1. 142 pp. tion. 53 pp. Selbstverlag, Prague. Jones, B. 1963: Upper Cretaceous (Campanian and Maastrichtian) am Giebel, c.G. 1849: Sitzung am 14. November. Jahresbericht des Natur monites from southern Alaska. United States Geological Survey Profes wissenschaftlichen Vereinsin Halle 1849, 18-20. sional Paper 432. 55 pp. Giers, R. 1964: Die Grossfauna der Mukronatenkreide (unteres Ober Kaplan, U. & Kennedy, W.}. 1994: Ammoniten des Westfalischen campan) im ostlichen Miinsterland. Fortschritte in der Geologie von Coniac. Geologie und Palaontologie in Westfalen 31. 155 pp. Rheinland und Westfalen 7, 213-194. Kayser, E. 1924: Lehrbuch der Geologie 11, Geologischen Formations Gill, T. 1871: Arrangement of the families ofMollusks. Smithsonian Mis kunde. 7 Ed. 657 pp. Ferdinand Enke, Stuttgart. cellaneous Collections 227. 49 pp. Kennedy, W.}. 1984: Systematic palaeontology and stratigraphical dis Griepenkerl, O. 1889: Die Versteinerungen der Senonen Kreide von tribution of the ammonite faun as of the French Coniacian. Special Koenigslutter im Herzogthum Braunschweig. Palaeontologische Ab Papers in Palaeontology 31. 160 pp. handlungen 4, 305-4 19 (3-1 16). Kennedy, W.}. 1986a: Campanian and Maastrichtian ammonites Grossouvre, A. de 1894 [date of imprint 1893]: Recherches sur la from northern Aquitaine, France. Special Papers in Palaeontology craie superieur, 2, Paleontologie. Les ammonites de la craie supe 36. 145 pp. rieur. Memoires du Service de la Carte Geologique detaillee de la Kennedy, W.}. 1986b: The ammonite fauna of the Calcaire il. Baculites France. 264 pp. (Upper Maastrichtian) of the Cotentin Peninsula (Manche, France). Grossouvre, A. de 1908: Description des ammonites du Cretace Palaeontology29, 25-83. superieur du Limbourg beige et hollandais et du Hainault. Memoires Kennedy, W.}. 1987 [date ofimprint 1986]: The ammonite faunas of the du Musee Royal d'Histoire Naturelle de Belgique 4, 1-39. type Maastrichtian, with a review of Ammonites colligatus Binkhorst, 1861. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Hagg, R. 1930: Die Mollusken und Brachiopoden der schwedischen Sciences de la Terre 56, 51-267. Kreide. 1. Eriksdal. Sveriges Geologiska Undersokning C363. 94 pp. Kennedy, W.}. 1993: Campanian and Maastrichtian ammonites from Hagg, R. 1935: Die Mollusken und Brachiopoden der schwedischen the Mons Basin (Belgium). Bulletin de I'Institut Royal des Sciences Na Kreide. 2. Kullemolla, Lyckås, Kåseberga und Grasryd. Sveriges Geol turelIes de Belgique, Sciences de la Terre 63, 99-131. ogiska Undersokning C385. 94 pp. Kennedy, W.}. & Christensen, W.K. 1991: Coniacian and Santonian am Hagg, R. 1943: Scaphites (Discoscaphites) binodosus A. Roemer från Blak monites fr om Bornholm, Denmark. Bulletin of the Geological Society sudden på Ivo. Geologiska Foreningens i Stockholm Forhandlingar 65, of Denmark 38, 203-226. 78-79. Kennedy, W.}. & Christensen, W.K. 1993: Santonian ammonites from Hagg, R. 1947: Die Mollusken und Brachiopoden der schwedischen the Kopingsberg- 1 borehole, Sweden. Bulletin of the Geological Society Kreide. 3. Das Kristianstadgebiet. Sveriges Geologiska Undersokning of Denmark 40, 149-156. C485. 143 pp. Kennedy, W.}. & Cobban, W.A. 1988: Mid-Turonian ammonite faunas Hagg, R. 1954: Die Mollusken und Brachiopoden der schwedischen from northern Mexico. Geological Magazine 125, 593-612. Kreide. 4. Die Mammillaten- und Mucronatenkreide des Ystadgebi Kennedy, W.}. & Cobban, W.A. 1993a: Ammonites from the Sara toga etes. Sveriges Geologiska Undersokning C535. 72 pp. Chalk (Upper Cretaceous) of Arkansas, U.S.A. Journal ofPaleontology Hancock, }.M. & Kennedy, W.}. 1993: The high Cretaceous ammonite 67, 404-434. fauna from Tercis, Landes, France. Bulletin de I'Institut Royal des Sci Kennedy, W.}. & Cobban, W.A. 1993b: Lower Campanian (Upper Cre ences Naturelles de Belgique, Sciences de la Terre 63, 149-209. taceous) ammonites fr om the Merchantville Formation of New }er Hauer, F. von 1858: Ueber die Cephalopoden der Gosauschichten. Bei sey, Maryland and Delaware. Journal of Paleontology67, 828-849. trage zur Palaontographie von Osterreich 1, 7-14. Kennedy, W.}. & Hancock, }.M. 1993: Upper Maastrichtian ammonites Hauer, F. von 1866: Neue Cephalopoden aus dem Gosau-gebilden der from the Marnes de Nay between Gan and Rebenacq (Pyrenees-At- Alpen. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften 1antique), France. Geobios 26, 575-594. in Wien, Mathematisch-naturwissenschaftlicheKlasse 53, 1-9. Kennedy, W.}., Hansotte, M., Bilotte, M. & Burnett, }. 1992: Ammonites Hebert, E. 1856: Tableau des fo ssiles de craie de Meudon et description and nannofossils from the Campanian of Nalzen (Ariege, France). de quelques especes nouvelles. Memoire de la Sociere Geologique de Geobios 25, 263-278. (2) (2), France 5 345-374. Kennedy, W.}. & Henderson, R.A. 1992: Heteromorph ammonites from Henderson, R.A. & McNamara, K.A. 1985: Maastrichtian non-hetero the Upper Maastrichtian of Pondicherry, South India. Palaeontology morph ammonites from the Mira Formation, Western Australia. Pal 35, 693-731. aeontology 28, 35-88. Kennedy, W.}. & Klinger, H.C. 1977: Cretaceous faunas from Zululand Hyatt, A. 1889: Genesis of the Arietidae. Smithsonian Contributions to and Natal, South Africa. The ammonite family Tetragonitidae Hyatt, Knowledge 673. 239 pp. 1900. Annals of the South African Museum 73:7, 149-197. Hyatt, A. 1894: Phylogeny of an acquired characteristic. Proceedings of Kennedy, W.}. & Summesberger, H. 1984: Upper Campanian ammo the American Philosophical Society 32, 349-647. nites from the Gschliefgraben (Ultrahelvetic, Up per Austria). Beitrage Hyatt, A. 1900: Cephalopoda. In Zittel, K.A. von 1896-1 900: Textbook of zur Palaontologie von Osterreich 11, 149-206. Palaeontology, 502-592. Macmillan, London. [English translation.] Kennedy, W.}. & Summesberger, H. 1987: Lower Maastrichtian ammo lagt, }.M.W. 1988: Some stratigraphical and faunal aspects of the Up nites from Nagoryany (Ukrainian SSR). Beitrage zur Palaontologie von per Cretaceous of southern Limburg (The Netherlands) and contig Osterreich 13, 25-78. uous areas. In Streel, M. & Bless, M.}.M. (eds.): The Chalk District of Keutgen, N. 1995: Late Campanian Belemnitella from Zeven Wegen the Euregio Meuse-Rhine, 25-39. Natuurhistorisch Museum, Maas Member (Gulpen Formation) at the CPL quarry (Haccourt, NE Bel tricht, and Laboratoires de Paleontologie de I'Universite d'Etat il. gium). Second International Symposium on Cretaceous Stage Bounda Liege, Liege. ries, Abstract Volume, p. 176 only. Institut Royal des Sciences Na }eletzky, }.A. 1948: Zur Kenntnis der Oberkreide der Dnepr-Donez turelles de Belgique, Brussels. Senke und zum Vergleich der russischen borealen Oberkreide mit Kilian, W. & Reboul, P. 1909: Les cephalopodes neocretaces des !les Sey derjenigen Polens und Nordwesteuropas. Geologiska Foreningens i mour et Snow Hill. WissenschaftlicheErg ebnisse der schwedischen Sud Stockholm Forhandlingar 70, 583-602. polar-Expedition 3:6. 75 pp. FOSSILS AND STRA T A 44 (1997) Santonian to Maastrichtian ammonites 127 Kner, R. 1848: Versteinerungen des Kreidemergels von Lemberg und ceous ammonites from the southern part of European Russia and sein er Umgebung. Haidingers Naturwissenschaftliche Abhandlungen their importance for lOnal stratigraphy (Campanian, Maastrich 2. 42 pp. tian).J Trudy Instituta Geologicheskikh Nauk 129, Geology Series 50, [Koplitz, W. 1920: Uber die Fauna des oberen Untersenon im Seppen 143 pp. [In Russian.J rade - Dwmener Hohenzuge. Dissertation, Westfalia Wilhelms-U ni Moberg, J.c. 1885: Cephalopoderna i Sveriges kritsystem. Il. Arts versitat Miinster. 78 pp. J beskrifning. Sveriges Geologiska Undersokning C73, 63 pp. Kossmat, F. 1895-1898: Untersuchungen iiber die siidindische Kreide Moore, R.C. (ed.) 1957: Treatise on Invertebrate Palaeontology. Part L. formation. Beitriige zur Paliiontologie Osterreich-Ungarns und des Mollusca 4. Cephalopoda Ammonoidea. 490 pp. Geological Society of Orients 9, 97-203 [1-107J (1895); 11, 1-46 [108-153J (1897); 11, 89- America and University of Kansas Press, Lawrence, Kansas. 152 [154-217J (1898). Morton, S.G. 1829: Description of two species of fossil shells of the gen Lamarck, J.P.B.A. de M. de 1799: Prodome d'une nouvelle c1assification era Scaphites and Crepidula; with some observations on the fe rrugi des coquilles. Memoire de la Societe d'Histoire Naturelle de Paris nous sand fo rmation, plastic c1ay and upper marine fo rmations of the (1799), 63-90. United States. Journal of the Academy ofNatural Sciences of Philadel Lamarck, J.P.B.A. de M. de 1801: Systeme des Animaux sans vertebres. phia 6, 107-ll9. 432 pp. Deterville, Paris. Miiller, G. & Wollemann, A. 1906: Die Molluskenfauna des Unterse Lamarck, J.P.B.A. de M. de 1822: Histoire naturelles des Animaux sans non von Braunschweig und Ilsede. Il. Die Cephalopoden. Abhand vertebres 7. 7ll pp. Verdiere, Paris. lungen der Koniglichen Preussischen Geologischen Landesanstalt, N.F. Leonhard, R. 1897: Die Fauna der Kreideformation in Oberschlesien. 47, 30 pp. Palaeontographica 44, ll-70. Naidin, D.P. 1974: Ammonoidea. In Krymgolts, G.Ya. (ed.): Atlas Lewy, Z. 1969: Late Campanian heteromorph ammonites from southern verkhnemelevoj fa uny Donbassa [Atlas of the Upp er Cretaceous Fauna Israel. Israel Journal of Earth Sciences 18, 109-135. of Donbass], pp. 158-195. Nedra, Moscow. [In Russian.J Lopuski, C. 19l1: Przyczunki do znajmosci fauny kredowej gub, Lubel Naidin, D.P. & Shimanskij, V.N. 1959: Cephalopoda. In Moskvin, M.M. skiej. Compte Rendu des Seances et la Societe Scientifique de Varsovie 4, (ed.): Atlas verkhnemelovoj fa uny severnogo Kavkaza i Kryma [Atlas of 104-140. the Upp er Cretaceous fa una of the northern Caucasus and Crimea], Lundegren, A. 1932: Om fo rekomsten av Cenoman i Båstads-området 166-220. Moscow. [In Russian.J och dess betydeIse for dateringen av uppkomsten av Hallandsås. Ge Neumayr, M. 1875: Die Ammoniten der Kreide und die systematik der ologiska Foreningens i Stockholm Forhandlingar 54, 500-504. Ammonitiden. Zeitschrift der Deutschen Geologischen Gesellschaft27, Lundegren, A. 1933: Kopingesandstenen i sydostra Skåne. Geologiska 854-942. Foreningens i Stockholm Forhandlingar 55, 163-165. Nilsson, S. 1826: Om de mångrummiga snackor som fo rekomma i krit Lundegren, A. 1935: Die stratigraphischen Ergebnisse der Tiefbohrung fo rmationen i Sverige. Kongliga Vetenskaps-Akademiens Handlingar bei Kullemolla im siidostlichen Schonen. Sveriges Geologiska Under 1825, 329-343. sokning C386, 18 pp. Nilsson, S. 1827: Petrificata Suecana: Formationis Cretaceae. 39 pp. Ber Lundgren, B. 1874: Om en Comaster och en Aptychus från Kopinge. ling, Londini Gothorum. Ofv ersigt afKongliga Vetenskaps-Akademiens Forhandlingar 3, 61-73. Nowak, I. 19l1: Untersuchungen iiber die Cephalopoden der oberen Lundgren, B. 1881: Om Scaphites binodosus Roem. från Kåseberga. Kreide in Polen. Il. Teil. Die Skaphiten. Bulletin international de Ofversigt af Kongliga Vetenskaps-Akademiens Forhandlingar 10, I'Academie des Sciences de Cracovie. Classe des Sciences Mathematiques 23-28. et Naturelles. Serie B. Sciences Naturelles, 547-589. Mariani, E. 1898: Ammoniti del Senoniano Lombardo. Memoire del Re Nowak, J. 1913: Untersuchungen iiber die Cephalopoden der oberen ale Instituto Lombardi di Scienze e Lettere, Classe di Scienze Matemat Kreide in Polen. III Teil. Bulletin de I'Academie des Sciences de Cra iche e Naturali (3) 18, 51-58 (1-18). covie. Classe des Sciences Mathematiques et Naturelles. Serie B. Sciences Marshall, P. 1926: The Upper Cretaceous ammonites of New Zealand. Naturelles, 335-415. Transactions of the New Zealand Institute 56, 129-210. Nowak, J. 1916: Zur Bedeutung von Scaphites fiir die Gliederung der Matsumoto, T. 1938: A biostratigraphic study on the Cretaceous depos Oberkreide. Verhandlungen der Geologischen Reichsanstalt (Staatsan its of the Naibuchi Valley, South Karahuto. Proceedings ofthe Imperial stalt-Landesanstalt) fo r 1916, 55-67. Academy ofJapan 14, 190-194 0dum, H. 1953: De geologiska resultaten från borrningarna vid HoIIvik Matsumoto, T. 1955: The bituberculate pachydiscids from Hokkaido en. Sveriges Geologiska Undersokning C527. 37 pp. and Saghalien. Memoirs of the Facultyof Science, Ky ushu University, Orbigny, A. d' 1840--1842: Paleontologie fran�aise: Terrains cretaces 1. Series D, Geology 5, 153-184. Cephalopodes. 1-120 (1840); 121-430 (1841); 431-662 (1842). Mas Matsumoto, T. 1959: Upper Cretaceous ammonites of California. Part son, Paris. 1. Memoirs of the Facultyof Science, Ky ushu University, Series D, Geol Orbigny, A. d' 1850: Prodrome de Paleontologie stratigraphique uni ogy 8, 91-171. verselle des animaux mollusques et rayonnes 2. 428 pp. Masson, Paris. Matsumoto, T. 1979: Notes on Lewesiceras and Nowakites (pachydiscid Owen, D.D. 1852: Report of a Geological Survey of Wisconsin, Iowa and ammonites) fromthe Cretaceous of Hokkaido. Transactions and Pro Minnesota, and incidentally of a portion of Nebraska Territory: made ceedings of the Palaeontological Societyof Ja pan, N.S. 113, 30-44. under the direction of the US TreasuryDepartment, Philadelphia. 195 Matsumoto, T. & Miyauchi, T. 1984: Some Campanian ammonites pp. Lippincott, Grambo & Co., Philadelphia, Pa. from the Soya area. Special Paper, Palaeontological Society of Japan Parkinson, I. 18ll: Organic Remains of a Former World, 3. 479 pp. Rob 27, 33-91. son, London. Matsumoto, T. & Obata, I. 1955: Some Upper Cretaceous desmoceratids Paulcke, W. 1907: Die Cephalopoden der oberen Kreide Siidpatago from Hokkaido and Saghalien. Memoirs of the Facultyof Science, Ky niens. Bericht der Naturforschenden Gesellschaftzu Freiburg i. Breisgau ushu University, Series D, Geology5, ll9-1 51. 15,[or 1905, 167-248. Matsumoto, T., Toshimitsu, S.A. & Kawashita, Y. 1990: On Hauericeras Ravn, I.P.I. 1902-1903: Molluskerne i Danmarks Kritaflejringer. de Grossouvre, 1894, a Cretaceous ammonite. Transactions and Pro Kongelige Danske Videnskabernes Selskabs Skrifter (6) 11. 1, Lamelli ceedings of the Palaeontological Society ofJa pan, N.S. 158, 439-458. branchiater, 73-138 [5-70J (1902); 2, Scaphopoder, Gastropoder og Meek, F.B. 1876: A report on the invertebrate Cretaceous and Tertiary Cephalopoder, 209-269 [5-65J (1902); 3, Stratigrafiske Under fo ssils of the upper Missouri country. In Hayden, F.V.: Report of the søgelser, 339-433 [5-99J (1903). United States Geological Survey of the Territories 9. 629 pp. RegneIl, G. 1983: Zoologen och arkeologen som var geolog. In RegneIl, Mikhailov, N.P. 1951: Verkhnemelovye ammonity yuga evropejskoj G. (ed.): Sven Nilsson. En lard i 1800-talets Lund. Studier utgivna av chasti SSSR i ikh znachenie dlya lOnal'noj stratigrafii. [Upper Creta- Kungliga FysiografiskaSiillskapet i Lund, 1983, 23-83. 128 W,f. Kennedy & w.K.Christensen FOSSILS AND STRATA 44 (1997) Reeside, J.B. Jr. 1927: Cephalopods from the lower part of the Cody Spath, L.F. 1922: On the Senonian ammonite fauna of Pondoland. Shale of Oregon Basin, Wyoming. United States Geological Survey Transactions of the Royal Societyof South Africa 10, 1 13-147. Professional Paper 150-A, 1-19. Spath, L.F. 1925: On Senonian Ammonoidea from Jamaica. Geological Riccardi, A.C. 1983: Scaphitids from the Upper Campanian - Lower Magazine 62, 28-32. Maastrichtian Bearpaw Formation of the western interior of Canada. Spath, L.F. 1926: On new ammonites from the English Chalk. Geological Geological Survey of Canada Bulletin 354. 103 pp. Magazine 63, 77-83. Riedel, L. 1931: Zur Stratigraphie und Faciesbildung im Oberemscher Spath, L.F. 1927: Revision of the Jurassic Cephalopod fauna of Kachh und Untersenon am Siidrande des Beckens von Miinster. Jahrbuch (Cutch). Memoirs of the Geological Survey ofIndia, Palaeontologia In der Preussischen Geologischen Landesanstalt zu Berlin 51, 605-713. dica, N.S. 11, 1-71. Roemer, A. 1840-1841: Die Versteinerungen des norddeutschen Kreidege Spath, L.F. 1953: The Upper Cretaceous cephalopod fauna of Graham birges. 145 pp. (1--48, 1840; 49-145, 1841). Hahn'schen, Hannover. Land. Scientific Reparts, Falkland Islands Dependencies Survey 3, 1-60. Roemer, F.A. 1865: Die Quadraten-Kreide des Sudmerberges bei Goslar. Stephenson, L.W. 1941: The larger invertebrate fossiIs of the Navarro Palaeontographica 13:4, 193-199. Group of Texas (exclusive of corals and crustaceans and exclusive of Rollier, L. 1922: Phylogenie des Ammonoides. Eclogae Geologicae Helve the fa una of the Escondido Formation). University of Texas Publica tiae 17, 358-360. tion 4101. 641 pp. Roman, F. 1938: Les ammonites jurassiques et mitacees. Essai de genera. Stobaeus, K. 1732: Opuscula in quibus petrefactorum, numismatum et 554 pp. Masson, Paris. antiquitatum historia illustratur, in unum volumen collecta. 327 pp. Dantiscum. Schhiter, C. 1867: Beitrag zur Kenntniss der jilngsten Ammoneen Nord Stolley, E. 1896: Einige Bemerkungen iiber die obere Kreide insbeson deutschlands. 36 pp. Henry, Bonn. dere von Liineburg und Lagerdorf. Archiv filrAn thropologie und Ge Schliiter, C. 1870: Bericht iiber eine geognostisch-paHiontologische ologie Schleswig-Hoisteins 1, 139-176. Reise im siidlichenSchweden. Neues Jahrbuch fu r Mineralogie, Geolo Stolley, E. 1897: Ueber die Gliederung des norddeutschen und gie und Paliiontologie, Jahrgang 1870, 929-969. baltischen Sen on sowie die dasselbe characterisierenden Belemniten. Schliiter, C. 1871-1876: Cephalopoden der oberen deutschen Kreide. Archiv fu r Anthropologie und Geologie Schleswig-Holsteins 2, 216-302. Palaeontographica 21, 1-24 (1871); 21, 25-120 (1872); 24, 123-204, Stolley, E. 1930: Einige Bemerkungen iiberdie Kreide Siidskandinavi 207-263 36-55 (1876). ens. Geologiska Foreningens i Stockholm Forhandlingar 52, 157-190. Schliiter, C. 1899: Podocrates im Senon von Braunschweig und Verb rei Stolley, E. 1932: Die Belemniten De Geer's von Båstad. Geologiska FOre tung und Benennung der Gattung. Zeitschrift der Deutschen Geolo ningens i Stockholm Forhandlingar 54, 498--499. gischen Gesellschaft51, 409--430. Surlyk, F. 1984: The Maastrichtian Stage in NW Europe and its bra Schrnid, F. & Ernst, G. 1975: Ammoniten aus dem Campan der Lehrter chiopod zonation. Bulletin of the Geological Society ofDenmar k 33, Westmulde und ihre stratigraphische Bedeutung. I. Tei!. Scaphites, 217-233. Bostrychocerasund Hoplitoplacenticeras. Bericht der Naturhistorischen Tate, R. 1865: On the correlation of the Cretaceous fo rmations of the Gesellschaftzu Hannover 119, 315-359. northeast ofIreland. Quarterly Journal ofthe Geological Societyof Lon Schulz, M.-G. 1878: Zur Litho- und Biostratigraphie des Obercampan don 21, 15-44. Untermaastricht von Lagerdorf und Kronsmoor (SW-Holstein). Tomlin, J.R.B. 1930: Some preoccupied generic names. Il. Proceedings of Newsletters in Stratigraphy 7, 73-89. the Malacological Society19 , 22-24. Schulz, M.-G. 1979: Morphometrisch-variationsstatistische Untersuch Wahlenberg, G. 1818: Petrificata telluris Svecanae. 116 pp. Uppsala. ungen zur Phylogenie der Belemniten-Gattung Belemnella im Unter Ward, P.D. & Kennedy, W.J. 1993: Maastrichtian ammonites from maastricht NW-Europas. Geologisches Jahrbuch A47, 157 pp. the Biscay Region (France, Spain). Paleontological Society Memoir Schulz, M.-G. 1996: Macrofossil biostratigraphy. In Schonfeld, J. & 34. 58 pp. Schulz, M.-G. (Coord.) et al.: New results on biostratigraphy, palaeo Wegner, T. 1905: Die Granulatenkreide des westlichen Miinsterlandes. magnetism, geochemistry and correlation from the standard section Zeitschrift der Deutschen Geologischen Gesellschaft57, 112-232. for the Up per Cretaceous White Chalk of northem Germany (Lager Wiedmann, J. 1966: Stammesgeschichte und System der posttriadischen dorf-Kronsmoor-Hemmoor). Mitteilungen aus dem Geologisch-Palii Ammonoideen; ein Oberblick. Neues Jahrbuch fu r Geologie und Palii ontologischen Institut der UniversitiitHa mburg 77, 548-550. 13-81. ontologie, Abhandlungen 125, 49-79; 127, Schulz, M.-G., Ernst, G., Ernst, H. & Schrnid, F. 1984: Coniacian to Wollemann, A. 1902: Die Fauna der Luneburger Kreide. Abhandlungen Maastrichtian stage boundaries in the standard section for the Upper der Preussischen geologischen Landesanstalt N.F. 37. III pp. Cretaceous white chalk of NW Germany (Lagerdorf-Kronsmoor Woods, H. 1906: The Cretaceous fa una of Pondoland. Annals of the Hemmoor): Definitionsand proposals. Bulletin of the Geological Soci South African Museum 4, 275-350. etyof Denmark 33, 203-215. Worm, O. 1655: Museum Wormianum. 389 pp. Elsevier, Amsterdam. Seunes, J. 1890-1892: Contributions it l'etude des cephalopodes du Wright, C.W. 1952: A classificationof the Cretaceous Ammonites. Jour Cretace Superieur de France. I. Ammonites du Calcaire it Baculites nal of Paleontology26, 213-222. du Cotentin. Mt!moiresde la Sociht! Gt!ologiquede France, Palt!ontolo Wright, C.W. 1957: [Cretaceous Ammonoidea.] In Moore, R.C. (ed.): gie 1, Mt!moire2, 1-7; 2, Mt!moire2, 2-22. Treatise on Invertebrate Paleontology. Part L, Mollusca 4, Cephalopoda Sharpe, D. 1853-57: Description of the fo ssil remains of Mollusca fo und Ammonoidea. 490 pp. Geological Society of America and University in the Chalk of England. I. Cephalopoda. Palaeontographical Society of Kansas Press, Lawrence, Kansas. Monographs 68, 1-26 (1853); 27-36 (1855); 37-68 (1857). Wright, C.W. 1979: The ammonites of the English Chalk Rock (Upper Sherborn, C.D. 1889: On the dates of the 'Paleontologie Fran�aise' of Turonian). Bulletin of the British Museum (Natural History), Geology d'Orbigny. Geological Magazine (iv) 6, 223-225. Series 31, 281-332. Shimizu, S. 1934: [Ammonites. ] In Shimizu, S. & Obata, T.: [Cephalop Wright, C.W. & Matsumoto, T. 1954: Some doubtful Cretaceous am oda. Iwanami's lecture series of Geology and Palaeontology.] 137 pp. monite genera from Japan and Saghalien. Memoirs of the Facultyof [In Japanese.] Science, Ky ushu University, Series D, Geology 4, 107-134. Siverson, M. 1992: Biology, dental morphology and taxonomy oflamni Wright, C.W. & Wright, E.V. 1951: A survey of the fo ssil Cephalopoda fo rm sharks from the Campanian of the Kristianstad Basin, Sweden. of the Chalk of Great Britain. Palaeontographical Society Monographs, Palaeontology 35, 519-554. 1--40. Smith, A.S. 1987: Fossils of the Chaik. 306 pp. Palaeontological Associa Zittel, K.A. von 1884: Handbuch der Palaeontologie. I. Abt. 2; Lieferung tion, London. 3, Cephalopoda, pp. 329-522. R. Oldenbourg, Munich & Leipzig. Sowerby, J. 1812-1822: The Mineral Conchology of GreatBritain. 383 pls. Zittel, K.A. von 1895: Grundzuge der Palaeontologie (Palaeozoologie). 972 London. pp. R. Oldenbourg, Munich & Leipzig. Instructions to authors Consult the editor at an early stage regarding suitability of topie, technical Illustrations requirements, and financing. Final acceptance for publication, however, will not be made until the manuscript has been refereed. lllustrations may be either figures in the text or plates fo llowing the text. Plan the figures so that they take up either the entire width of the type area (170 mm) or the width ofone colurnn (80 mm). If an intermediate width Text has to be used, do not exceed 120 mm. Maximum height is 233 mm. Plates The text should be as brief as possible without sacrificingclarity and vital should either fillup the whole type area (170x233 mm), for eaptions to be information. Write simply and direetly, avoid unnecessary jargon, and do placed on the opposing page, or be sufficientlyreduced in height to leave not shun the personal pronouns T and 'we' (but do not overuse them room for the captions below. Use the available rectangular space economi either), except in the abstract. Do notwrite as you speak (unless your speech cally: no unneeessary space between items and no protruding text. IlIustra is consistently lucid and does not rely on voiee inflexions,stresses, pauses, tion originals should not be smaUer than the finalprint and not Iarger than etc., to be understood). lf the message is clear and the writing succinct, the 200x300 mm. Mark all originals clearly with author name and illustration editor will not tamper (much) with your style. number. Though contributions in German and Freneh may exceptionally be Photographs must be clear and sharply contrasted, but without pro aceepted (with abstract and captions for illustrations and tables also pro nounced light areas and heavy shadows. If ineident light is used for vided in English), Fossils and Strata strongly favours English as the language illumination, the light should fall consistently from the upper left. In a of international scientificcommunication. This is not a linguistie or politi composite figure, all items should be of similar tone and contrast. Mount cal preferenee but simply an acceptance of fact. the photographs on pieces of cardboard, preferably A4 or Letter Size, or Use Sl (Systeme International d'Unites) units wherever possible. submit the finalillustrations as TIFF computer files witha resolution (in final size) of at least 233 dpi and a pixel depth of at least 8 bits. Tit/e. - Keep the title short ('The relationship between conodonts and Line drawings may also be submitted as computer files (confer with the vertebrates'). Avoid titles that are assertive ('Conodonts arevertebrates') or editors about fo rmats ). Ifsubmitted on paper, they should have lines of even interrogative ('Are conodonts vertebrates?'), and do not use 'noise' words thickness and blaekening, and any screens used should be coarser than 40 or parentheses ('Some remarks about conodonts (Vertebrata)') lines/cm (or the spaces left blank to be patterned by the editors). lf the figures include text, Do Not Monumentalize the Text by Capitalizing Authors' names. - Spell out one or more ofthe author's (or authors') first Words. For metrie units, use the standard symbols - (not mm, m, names. (Do the same with all persons mentioned in the text, for example in � Il), km. Do not leave out zero before decimal points. the acknowledgment section, but not for authors ofliterature referred to.) Markingofitems in composite figures(A, B, C, etc.; nota, b, c ... ) or plates Include the current address of eaeh author. (1,2,3, etc.) should be distinet but not dominant and placed as consistently Abstraet. - The abstract should be short (about 2000 letters and spaees) and as possible in the differentitems. Use Helvetica or a similar typeface. informative, stating the results presented in the monograph rather than describing its contents. New systematie or stratigraphic names should be mentioned in the abstraet. Conclude the abstraet with a list of keywords (index entries). Manuscript proeessing Main text. - Use fo ur or fewer grades of heading. Wherever possible, incorporate tabular material and systematie listings in the running text. Submit the manuscript initially as two paper copies for refereeing. Double Literature references in the text should be given in parentheses with the space all text. On the basis of the referees' report, the editor will decide author's surname followed by the year of publieation and no comma in whether to publish the manuscript and which modifications will be re between; use ampersand ('&') for two authors and 'et a/. ' if more than two. quired. The final version of text and figures should as far as possible be Thus '(Smith 1988, 1990; Smith & Jones 1990, p. 405; Black eta/. 1992, pp. submitted as computer files; the editor will supply the neeessary instruc 310-312)'. Do not use 'ibid. ' or 'ap. cit. ' or other sueh memory burdens. tions. Failure to submit computer fileswill not be cause for rejection but may lead to longer publication time and higher costs. Computer filesand Reference list - FoUow the fo rmat in reeent issues of Fossils and Strata or illustration originals should not be submitted until the paper has been Lethaia. lf in doubt, simply be as consistent as possible. accepted for publication. ou will receive galley proofs and page proofs in the form of laser Figure and table captions. - Captions should be self-explanatory but as short y as possible. Order them separately in the manuscript. printouts. Read the proofs carefully, making only necessary corrections, and return them promptly to the editor. Note that both text and illustrations Tables. - Keep tables few and simple. Small tables might be possible to run will have been modifiedby the editor and must be checked for errors; editor in with the text, and larger ones might preferably be presented as diagrams. and publisher take no responsibility for errors in the proofs not discovered lf, nonetheless, a table is most appropriate for the data, do not use excessive by the authors. fo rmatting (heavy lines, screened eells, boldface characters, etc.) or vertical Twenty copies of the publieation will be given free of charge to the lines, and do not design it to be turned 90° on the page. author(s). FOSSILS AND STRATA Number 44 · Oslo 22nd August 1997 Recent numbers 28 Muller, K.l. & Hinz, L 1991: Upper Cambrian 37 Noe-Nygaard, N. 1995: Ecological, sedimentary, and conodonts from Sweden. 153 pp. ISBN 82-00- geochemical evolution of the late-glacial to postglacial 37475-0. USD 44.00. Åmose lacustrine basin, Denmark. 436 pp. ISBN 82- 29 Moczycllowska, M. 1991: Acritarch biostratigraphy of 00-37656-7. USD 85.00. the Lower Cambrian and the Precambrian-Cambrian 38 Nielsen, A.T. 1995: Trilobite systematics, boundary in southeastern Poland. 127 pp. ISBN 82- biostratigraphy and palaeoecology of the Lower 00-37474-2. USD 38.00. Ordovician Komstad Limestone and Huk 30 Werdelin, L. & Solounias, N. 1991: The Hyaenidae: Formations, southern Scandinavia. 374 pp. ISBN 82- taxonomy, systematics and evolution. 104 pp. 00-37658-3. USD 75.00. ISBN 82-00-37481-5. USD 31.00. 39 Baarli, B.G. 1995: Orthacean and strophomenid 31 Smith, A.B. & Bengtson, P. 1991: Cretaceous brachiopods from the Lower Silurian of the central echinoids from north-eastern Brazil. 88 pp. ISBN 82- Oslo Region. 93 pp. ISBN 82-00-37659-1. USD 29.00. 00-37480-7. USD 27.00. 40 Jarvik, E. 1996: The Devonian tetrapod Ichthyostega. 32 Walossek, D. 1993: The Upper Cambrian Rehbachiella 213 pp. ISBN 82-00-37660-5. USD 49.00. and the phylogeny of Branchiopoda and Crustacea. 41 Bornmalm, L. 1997: Taxonomy and paleoecology of 202 pp. ISBN 82-00-37487-4. USD 48.00. late Neogene benthic foraminifera from the 33 Liljedahl, L. 1994: Silurian nuculoid and Caribbean Sea and eastern equatorial Pacific Ocean. modiomorphid bivalves from Sweden. 89 pp. 96 pp. ISBN 82-00-37666-4. USD 29.00. ISBN 82-00-37648-6. USD 28.00. 42 Jensen, S. 1997: Trace fossiIs from the Lower 34 Butterfield, N.J., Knoll, A.H. & Swett, K. 1994: Cambrian Mickwitzia sandstone, south-central Paleobiology of the N eoproterozoic Svanbergfjellet Sweden. III pp. ISBN 82-00-37665-6. USD 32.00. Formation, Spitsbergen. 84 pp. ISBN 82-00-37649-4. 43 Widmark, J.G.V. 1997: Deep-sea benthic foraminifera USD 27.00. from Cretaceous-Paleogene boundary strata in the 35 Popov, L. & Holmer, L.E. 1994: Cambrian- South Atlantic - taxonomy and paleoecology. 94 pp. Ordovician lingulate brachiopods from Scandinavia, ISBN 82-00-37667-2. USD 29.00. Kazakhstan, and South Ural Mountains. 156 pp. ISBN 44 Christensen, W.K. & Schulz, M.-G. 1997: Coniacian 82-00-37651-6. USD 45.00. and Santonian belemnite faunas from Bornholm, 36 Kielan-Jaworowska, Z. & Gambaryan, P.P. 1994: Denmark. Pp. 1-73. Kennedy, W.J. & Christensen, Postcranial anatomy and habits of Asian W.K. 1997: Santonian to Maastrichtian ammonites multituberculate mammais. 92 pp. ISBN 82-00- from Scania, southern Sweden. Pp. 75-128. 37650-8. USD 29.00. ISBN 82-00-37695-8. USD 36.00. ISBN 82-00-37695-8 Composed on a Macintosh® using Adobe FrameMaker+SGMVM, PageMaker®, Photoshop®, IlIustrator"', Minion"', and Deneba Canvas'" Set to film and printed by Berlings Grafiska AB, Arlov, Sweden, 1997 • © Lethaia Foundation 1997 I II 9788200I 376958