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- THE RUSH FAMILY

Juncaceae:

Genera: (22) and (2)

Mostly freshwater (except J. roemerianus)

10 are common or frequent:

Juncus roemerianus Juncus scirpoides Juncus dichotomous Juncus repens Juncus elliottii Juncus paludosus Juncus polycephalus

Credit: floridagrasses.org Credit: John R. Gwaltney Morphological Features:

• Cespitose or rhizomatous • 3-ranked, mostly terete (round) and septate, a few species have flat leaves • Culm (stems) pithy • Nodes without joints • an open or congested; lateral or terminal; panicles or heads • Sheaths fused or overlapping • is a three-part containing many Credit: Hilton Pond Center Credit: floridagrasses.org JUNCACEAE - Rushes

Common Rushes: 1. Juncus effusus: Usually 1m tall or so. appearing lateral, sheaths lacking blades. Often found growing in depressional areas within pastures.

2. Juncus marginatus: flowers borne in glomerate clusters, blades are flat, not septate. Capsule abruptly tapered. This species is extremely variable.

3. Juncus scirpoides: flowers borne in congested, headlike lobed clusters. Leaves round, sheath closed. Found across a wide range of .

4. Juncus megacephalus: Flowers borne in congested, headlike spherical clusters. Leaves round, sheath closed.

5. Juncus paludosus: is an endemic first identified in in 2008. Superficially similar to J. polycephalus, but more closely related to J. megacephalus. Distinguished form the former by having blades of the lower leaves laterally flattened, from the latter by having > 25 heads.

6. Juncus polycephalus: Similar in appearance to J. paludosus. See description above.

1 2 3 4 5 6 Artificial Identification Key JUNCACEAE – Morphological Features

Capsule Capsule Capsule

Perianth Segment Segment

Perianth Pedicel Segment

Flowers borne singly (one pedicel – one capsule/)

1 2 3 Capsule

Capsule

Pedicel

Flowers borne in clusters (one pedicel – many capsules/flowers) Morphology

Capsule

Perianth Segment JUNCACEAE – INFLORESCENCE TYPES

Inflorescence appearing Inflorescence appearing Inflorescence appearing Inflorescence appearing lateral terminal, branching, flowers terminal, branching, flowers terminal, in tight head-like borne singly borne 2+ clusters clusters Juncus effusus Juncus dichotomous Juncus marginatus Juncus megacephalus Juncus repens Juncus paludosus Juncus elliottii Juncus polycephalus Juncus scirpoides Inflorescence appearing lateral

Juncus roemerianus Juncus effusus Juncus coriaceus

Description: Large, stout growing in Description: Freshwater wetlands and Description: Freshwater wetlands. Sheaths saline/brackish wetlands(saltmarshes). Foliage pastures; much larger than J. coriaceus; with blades. Capsules subglobose or spherical. turning dark brown. Sheaths without blades. Capsules triangular. Inflorescence appearing terminal, flowers borne singly

Juncus bufonius Juncus dichotomous

Description: Small annual, rarely more than 30 cm tall. Description: Perennial with . Usually > 35 cm Description: Perennial with basal shorts forming Inflorescence typically more than ½ the size of the tall (to 1 meter). Blades terete, channeled or flat. small tufts or tussocks. Blades sometimes reduced or plant. absent. When present, blades flat or involute. Inflorescence appearing terminal, branching, flowers borne in 2+ clusters Juncus repens Juncus marginatus Juncus diffusisimus Juncus elliottii

Description: J. repens and Description: Perennial, Description: Perennial Description: Perennial Description: Perennial, marginatus are the only two sometimes rhizomatous cespitose. Leaves basal and caespitose. Leaves basal caespitose. Leaves basal species with flat grass-like Stems erect. Plant often cauline, terete or round and and cauline, terete. and cauline, terete. leaves. Unlike J. marginatus, growing in acidic or sandy septate. Inflorescence Capsules equaling or only Capsules equaling or only species has creeping stems soil shorelines. Leaves basal, branched (diffuse). slightly exceeding perianth slightly exceeding perianth and often forms mats. flat, auriculate. Very variable Capsules much longer than bristles. Perianth segments. Perianth in appearance. perianth segments. segments 3-4mm. segments <3mm. Inflorescence appearing terminal, branching, flowers in tight, head-like clusters

Juncus polycephalus Juncus validus Juncus megacephalus Juncus scirpoides

Description: Perennial, cespitose. Description: Perennial, usually Description: Perennial, cespitose. Description: Rhizomatous Leaves basal, somewhat flattened. rhizomatous Culms round or flat Leaves basal, somewhat flattened. perennial. Culms round or flat. Heads > 12mm in diameter. Leaves mostly basal, somewhat Heads > 12mm in diameter. Leaves mostly basal, somewhat Capsule tips fused. flattened sheaths fused. Capsule Capsule tips fused flattened sheaths fused. Capsule valves free valves free.