Mark Johnston, Editor-in-Chief University of Colorado HSC-Denver Tracey DePellegrin, GENETICS Editorial Board Executive Editor

CELLULAR GENETICS Yi Rao Lars M. Steinmetz Mark A. Beaumont Deborah Andrew Peking University European Molecular Biology Laboratory & University of Bristol Johns Hopkins University School of Medicine Meera V. Sundaram Graham Coop Orna Cohen-Fix University of Pennsylvania Trisha Wittkopp University of California, Davis NIDDK, National Institutes of Health Mariana F. Wolfner University of Michigan Simon Gravel Cornell University Amy S. Gladfelter McGill University UNC Chapel Hill and Dartmouth College Yongbiao Xue STATISTICAL GENETICS AND Joachim Hermisson Chinese Academy of Sciences N. Louise Glass GENOMICS University of Vienna University of California, Berkeley Sharon Browning Guillaume Martin Bob Goldstein GENE EXPRESSION University of Washington l’Institut des Sciences de l’Evolution de University of North Carolina at Chapel Hill James A. Birchler Jeffrey Endelman Montpellier (ISEM) Barth D. Grant University of Missouri University of Wisconsin-Madison Joanna Masel Rutgers University Junbiao Dai Eleazar Eskin University of Arizona Soni Lacefield Shenzhen Institutes of Advanced Technology University of California, Los Angeles Sohini Ramachandran Chinese Academy of Sciences Indiana University Haiyan Huang Brown University Michael Freitag University of California, Berkeley Aurélien Tellier Oregon State University Neil Risch Technical University of Munich COMPLEX TRAITS Pamela Geyer University of California, San Francisco Lindi M. Wahl Elissa J. Chesler University of Iowa Chiara Sabatti Western University The Jackson Laboratory Craig Kaplan Stanford University Brian Charlesworth (Perspectives Editor) Hans Daetwyler University of Pittsburgh Mikko J. Sillanpää University of Edinburgh La Trobe University Mitzi Kuroda Dirk-Jan de Koning Harvard Medical School University of Oulu Deborah Charlesworth (Perspectives Editor) Swedish University of Agricultural Sciences Victoria Meller William Valdar University of Edinburgh Jonathan Flint Wayne State University University of North Carolina at Chapel Hill Elizabeth A. De Stasio (Primers Editor) University of California, Los Angeles Aaron P. Mitchell Noah Zaitlen Lawrence University Jennifer Gleason University of Georgia University of California, Los Angeles Jasper Rine (Reviews Editor) The University of Kansas Oliver J. Rando University of California, Berkeley Michael Allen Gore University of Massachusetts Medical School METHODS, TECHNOLOGY, AND Trudi Schüpbach (Consulting Editor) Cornell University Nathan Springer RESOURCES Princeton University Elizabeth Hauser University of Minnesota Hannes Bülow Michael Turelli (Reviews Editor) Duke University Elizabeth Tran Albert Einstein College of Medicine University of California, Davis Thomas E. Juenger Purdue University Stanley Fields Adam S. Wilkins (Perspectives Editor) University of Texas University of Washington Humboldt University of Berlin Dahlia Nielsen GENOME INTEGRITY AND Liqun Luo North Carolina State University TRANSMISSION Stanford University Andrew H. Paterson BOARD OF SENIOR EDITORS Jack Bateman Kate O'Connor-Giles University of Georgia Karen M. Arndt Bowdoin College Brown University Katie Peichel University of Pittsburgh Needhi Bhalla Norbert Perrimon University of Bern Nicholas H. Barton University of California, Santa Cruz Harvard Medical School Christine Queitsch Douglas Bishop Institute of Science and Technology Austria University of Washington Valerie Reinke University of Chicago David J. Begun Paul Scheet Yale University Anne Britt University of California, Davis MD Anderson Cancer Center Jay Shendure University of California, Davis Karl Broman Albert Tenesa University of Washington Brian R. Calvi The University of Edinburgh University of Wisconsin-Madison Indiana University Trisha Wittkopp Mario Calus Francesca Cole EMPIRICAL University of Michigan Wageningen UR Livestock Research University of Texas MD Anderson Daniel A. Barbash Audrey Gasch Jason B. Wolf Cancer Center Cornell University University of Bath University of Wisconsin-Madison JoAnne Engebrecht Santiago C. González-Martínez Jianming Yu University of California, Davis INRAE UMR BIOGECO David I. Greenstein Iowa State University University of Minnesota Eva Hoffmann Matthew W. Hahn University of Copenhagen Indiana University Oliver Hobert DEVELOPMENTAL AND Neil Hunter Charles H. Langley Columbia University BEHAVIORAL GENETICS University of California, Davis University of California, Davis Anthony D. Long University of California at Irvine Kathryn Barton Amy MacQueen Mara Lawniczak Carnegie Institution for Science Wesleyan University Wellcome Sanger Institute Mark D. Rose Hugo J. Bellen Houra Merrikh Jeffrey G. Lawrence Georgetown University Baylor College of Medicine Vanderbilt University University of Pittsburgh Jeff Sekelsky Hannes Bülow Jac A. Nickoloff Leonie C. Moyle University of North Carolina at Chapel Hill Albert Einstein College of Medicine Colorado State University Indiana University Barbara Conradt Yikang Rong Michael W. Nachman GENETICS publishes high quality, original University College London University of South China University of California, Berkeley research presenting novel findings on a Gerald B. Downes Jennifer Surtees John Novembre range of topics bearing on heredity and University of Massachusetts, Amherst University at Buffalo University of Chicago variation. GENETICS is a peer-edited Bruce W. Draper Daven Presgraves journal—all editorial decisions are made University of California, Davis GENOME AND SYSTEMS BIOLOGY University of Rochester by the authors’ peers—with a tradition of Robert J. Duronio Kirsten Bomblies Andrea Sweigart rigorous peer review. Publication is open University of North Carolina at Chapel Hill ETH Zürich University of Georgia equally to members and non-members of Elissa Hallem Alan Moses Daniel M. Weinreich the Genetics Society of America. Submit University of California, Los Angeles University of Toronto Brown University manuscripts at https://genetics.msubmit.net. Yasuhiko Kawakami Norbert Perrimon Stephen I. Wright GSA Journals Editorial Office University of Minnesota Harvard Medical School University of Toronto 6120 Executive Boulevard Scott Kennedy Valerie Reinke Suite 550 Harvard University Medical School Yale University THEORETICAL POPULATION Rockville, MD 20852 Abraham A. Palmer Jay Shendure University of Chicago University of Washington GENETICS Phone: 412-226-5930 Amy Ralston Jason Stajich Aneil F. Agrawal Fax: 412-226-5931 Michigan State University University of California, Riverside University of Toronto E-mail: [email protected]