Letter from the Chairman

For our inaugural annual report, I am thrilled to describe the tremendous progress that eLife Sciences has achieved during its first year of operation.

In June 2011, the Howard Hughes Medical Institute, the Max Planck Society, and the Wellcome Trust announced eLife Sciences – a disruptive, refreshing new model for the publishing and dissemination of cutting-edge scientific research. It is built on digital technologies, and driven by the needs of academic science, the research community, and the broader population interested in science.

The mission of eLife Sciences is to establish a leading journal for the publication of advancements in the life and biomedical sciences, using active researchers to assess the science efficiently and quickly, and leveraging the power of digital media to bring the work to life and make it openly accessible in multiple dimensions to a global audience.

The Board is very proud of the eLife Sciences team’s accomplishments in 2012, including the initial development and operation of the journal and the! warm and open reception from the research community.

Critical to eLife Sciences’ success has been the driving force of our talented academic editorial team. Led by Editor-in-Chief Randy Schekman (University of California at Berkeley, USA), and Deputy Editors Fiona Watt (King’s College, London, UK) and Detlef Weigel (Max Planck Institute for Developmental Biology, Germany) the team has devised an open and transparent editorial process that has been welcomed by the scientific community. This process ensures that new submissions are assessed swiftly and fairly, and that the editorial feedback after peer review is focused on essential revision requests. The eLife Sciences staff, based in Cambridge, England, have also put together a leading-edge publishing platform that delivers a rich, intuitive digital content experience and was launched with eLife’s new website in December 2012.

It has been eLife’s goal to quickly attract the highest calibre of content and to build a reputation for excellence in editorial standards. Given that eLife is competing with established and influential venues for the most important scientific work, this is a considerable challenge. It is, therefore, hugely gratifying that well over 100 research articles that meet our high standards of excellence have already been published mid-way through 2013. Subjects range from structural biology to human perception, from plant-predator interactions to public health, and each article is accompanied by an eLife Digest to make these stories more accessible for the broader public interested in science.

eLife Sciences Publications, Ltd is a limited liability non-profit non-stock corporation incorporated in the State of Delaware, USA, with company number 5030732, and is registered in the UK with company number FC030576 and branch number BR015634 at the address Suite 203, Sheraton House, Castle Park, Cambridge CB3 0AX. It is important to remember, however, that the eLife Sciences project is about more than launching a journal. It is about exploiting the potential of a new medium to communicate and share the results of scientific research. The digital medium has had profound and widespread effects on the way we conduct our lives, and yet has had limited impact on journals and research communication in general. Indeed, this area has been severely limited by the paradigm of the printed medium and outmoded print-based business models. The eLife Sciences Board will also therefore assess the progress of the initiative in terms of how journals and publishing practices change more broadly.

This Annual Report includes the audited accounts of eLife Sciences Publications Limited. Consistent with the far-reaching ambitions of the project, the founders have committed more than £15 million to its launch and development.

We would like to acknowledge the important contributions to the eLife Sciences Board from two colleagues who have moved on to other positions: Sir Mark Walport, who led the W ellcome Trust for 10 years and was instrumental in the conception and early life of the project; and Elizabeth Marincola, who chaired the Board for our first year. Sir Mark is currently the UK Government’s Chief Science Advisor, and Elizabeth has become the CEO of the open-access publisher, PLOS. We wish them both well in their new endeavours.

The Board could not be happier with the start that eLife Sciences has made, and we congratulate all involved, most especially the researchers who have taken part – as editors, reviewers or authors – in this unique opportunity to transform the dissemination of knowledge and stimulate scientific innovation.

Toby Coppel Chairman, Board of Directors eLife Sciences

To view the complete eLife Sciences 2012 Annual Report, please visit 2012.elifesciences.org. ELIFE SCIENCES PUBLICATIONS, LTD.

Form 8868 Application for Extension of Time To File an (Rev. January 2013) Exempt Organization Return OMB No. 1545-1709 Department of the Treasury I Internal Revenue Service File a separate application for each return. m m m m m m m m m m m m m m m m m % If you are filing for an Automatic 3-Month Extension, complete only Part I and check this box I X % If you are filing for an Additional (Not Automatic) 3-Month Extension, complete only Part II (on page 2 of this form). Do not complete Part II unless you have already been granted an automatic 3-month extension on a previously filed Form 8868.

Electronic filing (e-file). You can electronically file Form 8868 if you need a 3-month automatic extension of time to file (6 months for a corporation required to file Form 990-T), or an additional (not automatic) 3-month extension of time. You can electronically file Form 8868 to request an extension of time to file any of the forms listed in Part I or Part II with the exception of Form 8870, Information Return for Transfers Associated With Certain Personal Benefit Contracts, which must be sent to the IRS in paper format (see instructions). For more details on the electronic filing of this form, visit www.irs.gov/efile and click on e-file for Charities & Nonprofits. Part I Automatic 3-Month Extension of Time. Only submit original (no copies needed). A corporatimonm rmeqm umirem dm tmo mfilme mFom rm m 9m90m -Tm manm dm rem qm umesm tinm gm am nm amutmom am tmic m6-m mm omntmh mexm tem nmsimonm -m cmhem cmk mthmism bmoxm amndm cm ommmplmetme m m m m m Part I only I All other corporations (including 1120-C filers), partnerships, REMICs, and trusts must use Form 7004 to request an extension of time to file income tax returns. Enter filer's identifying number, see instructions Name of exempt organization or other filer, see instructions. Employer identification number (EIN) or Type or print ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 File by the Number, street, and room or suite no. If a P.O. box, see instructions. Social security number (SSN) due date for filing your SUITE 203 SHERATON HOUSE CB3 OAX return. See City, town or post office, state, and ZIP code. For a foreign address, see instructions. instructions. CASTLE PARK CAMBRIDGE UNITED KINGDOM m m m m m m m m m m m m Enter the Return code for the return that this application is for (file a separate application for each return) 0 1

Application Return Application Return Is For Code Is For Code Form 990 or Form 990-EZ 01 Form 990-T (corporation) 07 Form 990-BL 02 Form 1041-A 08 Form 4720- (individual) 03 Form 4720 09 Form 990-PF 04 Form 5227 10 Form 990-T (sec. 401(a) or 408(a) trust) 05 Form 6069 11 Form 990-T (trust other than above) 06 Form 8870 12

% The books are in the care of I PAUL KELLY I I % Telephone No. FAX No. m m m m m m m m m m m m m m m I % If the organization does not have an office or place of business in the United States, check this box X If this is for a Group Return, enter tm hme mormgam nmization's four digit Group Exemption Number (GEN) m m m m m m m . If this is for the whole group, check this box I . If it is for part of the group, check this box I and attach a list with the names and EINs of all members the extension is for. 1 I request an automatic 3-month (6 months for a corporation required to file Form 990-T) extension of time until 08/15 , 20 13 , to file the exempt organization return for the organization named above. The extension is for the organization's return for: I X calendar year 2012 or I tax year beginning , 20 , and ending , 20 .

2 If the tax year entered in line 1 is for less than 12 months, check reason: Initial return Final return Change in accounting period

3a If this application is for Form 990-BL, 990-PF, 990-T, 4720, or 6069, enter the tentative tax, less any nonrefundable credits. See instructions. 3a $ b If this application is for Form 990-PF, 990-T, 4720, or 6069, enter any refundable credits and estimated tax payments made. Include any prior year overpayment allowed as a credit. 3b $ c Balance due. Subtract line 3b from line 3a. Include your payment with this form, if required, by using EFTPS (Electronic Federal Tax Payment System). See instructions. 3c $ Caution. If you are going to make an electronic fund withdrawal with this Form 8868, see Form 8453-EO and Form 8879-EO for payment instructions. For Privacy Act and Paperwork Reduction Act Notice, see Instructions. Form 8868 (Rev. 1-2013)

JSA

2F8054 2.000 8195FH 2502 5/9/2013 10:07:41 AM V 12-4.6F 2885408 PAGE 1 ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Form 990 (2012) Page 2 Part III Statement of Program Service Accomplishments mmmmmmmmmmmmmmmmmmmmmmmm Check if Schedule O contains a response to any question in this Part III X 1 Briefly describe the organization's mission: ELIFE IS A UNIQUE COLLABORATION BETWEEN FUNDERS AND PRACTITIONERS OF RESEARCH TO COMMUNICATE INFLUENTIAL DISCOVERIES IN THE LIFE AND BIOMEDICAL SCIENCES IN THE MOST EFFECTIVE WAY.

2 Did the organization undertakemmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmm any significant program services during the year which were not listed on the prior Form 990 or 990-EZ? YesX No If "Yes," describe these new services on Schedule O. 3 Did the organizationmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmmm cease conducting, or make significant changes in how it conducts, any program services? YesX No If "Yes," describe these changes on Schedule O. 4 Describe the organization's program service accomplishments for each of its three largest program services, as measured by expenses. Section 501(c)(3) and 501(c)(4) organizations are required to report the amount of grants and allocations to others, the total expenses, and revenue, if any, for each program service reported.

4a (Code: ) (Expenses $ 2,726,966. including grants of $ ) (Revenue $ ) ATTACHMENT 1

4b (Code: ) (Expenses $ including grants of $ ) (Revenue $ )

4c (Code: ) (Expenses $ including grants of $ ) (Revenue $ )

4d Other program services (Describe in Schedule O.) (Expenses $ including grants of $ ) (Revenue $ ) 4e Total program service expenses I 2,726,966. JSA 2E1020 2.000 Form 990 (2012) 8195FH 2502 6/17/2013 3:14:36 PM V 12-5F 2885408 PAGE 2 ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Form 990 (2012) Page 3 Part IV Checklist of Required Schedules Yes No 1 Is the organization desm cmribm emd minm sm emctmiomn m5m01m (cm )(m 3m) morm 4m 9m47m (am )m(1m) m(omthmemr tmham nm am mprmivmatme mfom umndm amtiom nm)?m Ifm "mYmesm,"m m complete Schedule A m m m m m m m m m 1 X 2 Is the organization required to complete Schedule B, Schedule of Contributors (see instructions)? 2 X 3 Did the organization engage in direct or indirect political campamigmn macm timvimtiem sm omn mbmehm amlf mofm omr minm om pmpom smitimonm tmom m candidates for public office? If "Yes," complete Schedule C, Part I 3 X 4 Section 501(c)(3) organizations. Did the organization engage in lobbyingm am cmtivmitimesm , morm hm amvem am sm emctmiomn m50m 1m (hm )m m election in effect during the tax year? If "Yes," complete Schedule C, Part II 4 X 5 Is the organization a section 501(c)(4), 501(c)(5), or 501(c)(6) organization that receives membership dues, assessmmenm tsm , momr msim milamr mam om umnmts m ams m dmefminmedm minm Rm emvem nmuem mPrmocm emdum rem m98m -m19m ?m Imf m"Ymesm ,"m cmompm lem tme mSmchmedm umlem Cm ,m m Part III 5 X 6 Did the organization maintain any donor advised funds or any similar funds or accounts for which donors have the right to provide advice onm tmhem mdimstmribm umtiom nm om r minmvem smtmm emntm om f m am moum nmtsm inm msum cmh mfumndm sm om r macm cmoum nmtsm? mIf m m "Yes," complete Schedule D, Part I 6 X 7 Did the organization receive or hold a conservation easement, including easements to preservme mopm emnm smpamcem ,m m the environment, historic land areas, or historic structures? If "Yes," complete Schedule D, Part II 7 X 8 Did the organization maintain mcom llmecm timonm sm omf wm om rkm sm om f marmt, m hmistmomricm aml tmremasm umrems,m omr motmhem r m smimm ilam rm amssmetms?m Imf "mYmesm,"m m complete Schedule D, Part III 8 X 9 Did the organization report an amount in Part X, line 21, for escrow or custodial account liability; serve as a custodian for amounts not listed in Part X; or provide credit coum nmsem limngm , mdmebm t mmmamnam gmemm em nmt, mcrmemdimt rmempam irm, om rm m debt negotiation services? If "Yes," complete Schedule D, Part IV 9 X 10 Did the organization, directly or through a related organization, hold assets in temporarily rmesm trmicmtemdm m endowments, permanent endowments, or quasi-endowments? If "Yes," complete Schedule D, Part V 10 X 11 If the organization’s answer to any of the following questions is "Yes," then complete Schedule D, Parts VI, VII, VIII, IX, or X as applicable. a Did the organization report am nm am mm om umntm fmorm mlamndm , m bmuimldmingm sm, m amndm meqm umipmmmenm t m imn m Pmarmt mXm, mlinmem 1m 0m? m Ifm "mYmesm,"m m complete Schedule D, Part VI 11a X b Did the organization report an amount for investments-other securities in Part X, mlinme m 1m2 m thm amt mis m5%m m omr m mormem m of its total assets reported in Part X, line 16? If "Yes," complete Schedule D, Part VII 11b X c Did the organization report an amount for investments-program related in Part X, mlinm em 1m3 m thm amt mis m 5m%m omr m mormem m of its total assets reported in Part X, line 16? If "Yes," complete Schedule D, Part VIII 11c X d Did the organization report an amount for other assets in Part X, limnem 1m 5m tmham t m ism 5m%m om r mmm omrem omf mitsm tmotmalm amssmemtsm m reported in Part X, line 16? If "Yes," complete Schedule D, Part IX 11d X e Did the organization report an amount for other liabilities in Part X, line 25? If "Yes," complete Schedule D, Part X 11e X f Did the organization’s separate or consolidated financial statements for the tax year include a footnote that admdrmesmsem sm m the organization's liability for uncertain tax positions under FIN 48 (ASC 740)? If "Yes," complete Schedule D, Part X 11f X 12 a Did the organization obtain separate, m inm dmepm emndm emntm maum dmitem dm fminmanm cmiaml sm tam tmemm em nmtsm fmorm tmhem mtamx m ymeamr?m mIf m "Ym ems,m" m complete Schedule D, Parts XI and XII 12a X b Was the organization included in consolidated, independent audited financial statements for the mtaxm ym eam r?m Imf "mYem s,m" manmd m ifm m the organization answered "No" to line 12a, then completing Schedule D, Parts XI and XII is optional m m m m m m m m m m 12b X 13 Is the organization a school described in section 170(b)(1)(A)(ii)? If "Yes," complete Schedulme mE m m m m m m m m m m m 13 X 14 a Did the organization maintain an office, employees, or agents outside of the United States? 14a X b Did the organization have aggregate revenues or expenses of more than $10,000 from grantmaking, fundraising, business, investment, and program service activities outside the United Statems,m om r m amggm rmegm amtem m foreign investments valued at $100,000 or more? If "Yes," complete Schedule F, Parts I and IV 14b X 15 Did the organization report on Part IX, column (A), line 3, more than $5,000 of grants or assistancme m tom manm ym m organization or entity located outside the United States? If "Yes," complete Schedule F, Parts II and IV 15 X 16 Did the organization report on Part IX, column (A), line 3, more than $5,000 of aggregate granm tsm om r masm smismtamncmem m to individuals located outside the United States? If "Yes," complete Schedule F, Parts III and IV 16 X 17 Did the organization report a total of more than $15,000 of expenses for professional fundmram isminmg m smermvimcemsm m on Part IX, column (A), lines 6 and 11e? If "Yes," complete Schedule G, Part I (see instructions) 17 X 18 Did the organization report more than $15,000 total of fundramismingm mevm emntm gmromssm imncm ommme m amndm cm omntm rimbum timonm sm omnm m Part VIII, lines 1c and 8a? If "Yes," complete Schedule G, Part II 18 X 19 Did the organization report more thanm m$1m 5m,0m 0m0 m omf gm rmosms m inmcom mm em fmrom mm gm ammminmg m amctmivimtiem sm om nm Pmarmt mVImII,m limnem m9am?m m If "Yes," complete Schedule G, Part III m m m m m m m m m m m m m 19 X 20 a Did the organization operate one or more hospital facilities? If "Yes," complete Schedule H m m m m m m 20a X b If "Yes" to line 20a, did the organization attach a copy of its audited financial statements to this return? 20b

JSA Form 990 (2012)

2E1021 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 3 ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Form 990 (2012) Page 4 Part IV Checklist of Required Schedules (continued) Yes No 21 Did the organization report more than $5,000 of grants and other assistance to any governmm emntm omr mormgam nmizmatmiomnm in the United States on Part IX, column (A), line 1? If "Yes," complete Schedule I, Parts I and II 21 X 22 Did the organization report more than $5,000 of grants and other assistanm cme mtom inm dmivmidmuam lsm inm tmhem Um nm item dm Sm tam tem sm m on Part IX, column (A), line 2? If "Yes," complete Schedule I, Parts I and III 22 X 23 Did the organization answer "Yes" to Part VII, Section A, line 3, 4, or 5 about compensation of the organization's current and former officerms,m dm irmecm tom rms,m tmrumstmeem sm, mkem ym em mm pm lomyem ems,m am nmd m hm igmhem smt m cmomm pm emnsm amtemdm employees? If "Yes," complete Schedule J 23 X 24 a Did the organization have a tax-exempt bond issue with an outstanding principal amount of more than $100,000 as of the last day of the year, that was issued amftem rm Dmemcem mm bmerm 3m1m, 2m 0m0m2?m Imf m"Ymesm ,"m amnsm wmerm lminmesm 2m 4mbm through 24d and complete Schedule K. If “No,” go to line 25 m m m m m m m 24a X b Did the organization invest any proceeds of tax-exempt bonds beyond a temporary period exception? 24b c Did the organization maintain an emscmrom wm am cmcom umntm om thm emr mthmanm ma mrem fum nmdimngm em smcrmomw m amt am nmy m tim me mdmurminmg mthm em ymeam rm to defease any tax-exempt bonds? m m m m m m m 24c d Did the organization act as an "on behalf of" issuer for bonds outstanding at any time during the year? 24d 25 a Section 501(c)(3) and 501(c)(4) organizations. Did the organization engage minm amn m emxcmesm sm bmenm emfitm tmramnmsamctmiomnm with a disqualified person during the year? If "Yes," complete Schedule L, Part I 25a X b Is the organization aware that it engaged in an excess benefit transaction with a disqualified person in a prior year, and that the transaction has nm omt bm emenm mrempom rmtemd monm manm y m omf tmhem mormgam nm izmatmiomn'ms mprmiomr mFom rmms m99m 0m om r m 9m90m-Em Zm?m If "Yes," complete Schedule L, Part I 25b X 26 Was a loan to or by a current or former officer, director, trustee, key employee, highly compensated employee, orm disqualified person outstanding as of the end of the organization's tax year? If "Yes," complete Schedule L, Part II 26 X 27 Did the organization provide a grant or other assistance to an officer, director, trustee, key employee, substantial contributor or employee thereof, a grant selection committee member,m omr m tom ma m3m5%m m cmonm trmomllemdm entity or family member of any of these persons? If "Yes," complete Schedule L, Part III 27 X 28 Was the organization a party to a business transaction with one of the following parties (see Schedule L, Part IV instructions for applicable filing thresholds, conditions, and exceptions): m m m m m m m m a A current or former officer, director, trustee, or key employee? If "Yes," complete Schedule L, Part IV 28a X b A family member m omf ma m cmurm rem nmt m omr mfomrmm em r m om ffmicmerm, mdimremctmomr, m trm umstmeem , m omr m kmeym em mm pm lomyem em? m Ifm m"Ymesm ,"m cm ommmplmetmem Schedule L, Part IV 28b X c An entity of which a current or former officer, director, trustee, or key employee (or a family mem mm bmerm thm em rem omf)m was an officer, director, trustee, or direct or indirect owner? If "Yes," complete Schedule L, Part IV 28c X 29 Did the organization receive more than $25,000 in non-cash contributions? If "Yes," complete Schedule M 29 X 30 Did the organization receive contributions of art, historicm aml mtrmeam smurmesm, m omr motmhem rm sm imm ilmarm masmsem tsm , m omr mqmuam lifmiemdm conservation contributions? If "Yes," complete Schedule M 30 X 31 Did them momrgmanm izm amtiom nm limqum idm amtem, mtemrmm imnam tem , m omr mdimssm omlvem manm dm cm emasm em ompmermatmiomnsm? m Ifm "mYmesm," m cmo m pm lem tem mScm hmedm umlem Nm ,m Part I 31 X 32 Did the organization sell, mexmchm amngm em, m dmispm omsem momf, m omr m trmanm smfemr mmm omrem tmham nm m25m %m momf mitsm mnem t m am ssm emtsm? m Ifm "mYmesm,"m complete Schedule N, Part II 32 X 33 Did the organization own 100% of an entity disregarded as separate from m thm em omrgm amnimzam timonm um nmdem rm Rm emgum lam timonmsm sections 301.7701-2 and 301.7701-3? If "Yes," complete Schedule R, Part I 33 X 34 Was the organization mremlamtemd m tom manm ym tmaxm-em xmemm pm t momr mtamxam bmle m emntmitym ?m Imf "m Ymesm,"m cmompm lem tme mSmchmedm umlem Rm , m Pmarmt mII,m ImII,m or IV, and Part V, line 1 m m m m m m m m m m m m m m 34 X 35 a Did the organization have a controlled entity within the meaning of section 512(b)(13)? 35a X b If "Yes" to line 35a, did the organization receive any payment from or engage in any transactionm mwmithm mam controlled entity within the meaning of section 512(b)(13)? If "Yes," complete Schedule R, Part V, line 2 35b 36 Section 501(c)(3) organizations. Did the organization makem am nm y m tmramnsm fem rsm mtom manm memxemmm pmt mnom nm-cmhmarmitam bmlem related organization? If "Yes," complete Schedule R, Part V, line 2 36 X 37 Did the organization conduct more than 5% of its activities through an entity that is not a related organization and thatm ism mtrem amtemdm ams ma m pmarmtnmemrsmhimp mfomr mfemdmermalm inm cmomm em mtamx m pmurmpom sem sm? m Ifm "mYem sm," mcom mm pmlemtem Sm cmhem dmulme m Rm, m m m m m Part VI 37 X 38 Did the organization complete Schedule O and provide explanationms minm Sm cmhem dmumle m Om fmorm Pm amrt m VmI, mlinm ems m1m1bm am nm dm 19? Note. All Form 990 filers are required to complete Schedule O 38 X Form 990 (2012)

JSA

2E1030 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 4 ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Form 990 (2012) Page 5 Part V Statements Regarding Other IRS Filings and Tax Compliance m m m m m m m m m m m m m m m m m m m m m m m Check if Schedule O contains a response to any question in this Part V X m m m m m m m m m m Yes No 1a Enter the number reported in Box 3 of Form 1096. Enter -0- if not applicable m m m m m m m m m 1a 20 b Enter the number of Forms W-2G included in line 1a. Enter -0- if not applicable 1b 0 c Did the organization comply with backup withholdminmg m rmulmesm mfom r m rmepm omrtmabm lem mpam ymmm emntms m tom m vmenm dmorm s m am nmdm reportable gaming (gambling) winnings to prize winners? 1c X 2a Enter the number of employees reported on Form W-3, Transmittal of Wage and Taxm Statements, filed for the calendar year ending with or within the year covered by this return 2a 11 b If at least one is reported on line 2a, did the organization file all required federal employment tam x m rem tmurmnsm?m 2b X Note. If the sum of lines 1a and 2a is greater than 250, you may be required to e-file (see instructimonm sm) m m m m m m m 3a Did the organization have unrelated business gross income of $1,000 or more during the yemarm?m m m m m m m m m m m 3a X b If "Yes," has it filed a Form 990-T for this year? If "No," provide an explanation in Schedule O 3b 4a At any time during the calendar year, did the organization have an interest in, or a signature or other authority over, a finam nmciam l m amccm omunm t m inm am mfomrem igmn m cmoum nmtrmy m(sm umchm am sm am bm amnkm am cmcom umntm, msem cmurmitimesm am cmcom umntm, morm om thm emr m fimnam nmciam lm account)? 4a X b If “Yes,” enter the name of the foreign country: I UNITED KINGDOM See instructions for filing requirements for Form TD F 90-22.1, Report of Foreign Bank and Financial mAcm cmoum nmtsm. m m 5a Was the organization a party to a prohibited tax shelter transaction at any time during the tax year? 5a X b Did any taxable party notify the organization that it was or is m am pmamrtym tmo m am pm rom hmibmitem dm tmaxm mshm emltem r mtrmanm smacm tiom nm?m 5b X c If "Yes" to line 5a or 5b, did the organization file Form 8886-T? 5c 6a Does the organization have annual gross receipts that are normally greater than $100,0m 0m0,m am nm dm dm idm mthmem organization solicit any contributions that were not tax deductible as charitable contributions? 6a X b If "Yes," did the organization imncm lum dme m wmithm memvemrym sm omlicm itmatmiomn m amn mexm pmremssm mstmatmemm em nmt mthmatm sm umchm cm om nmtrimbum timonm sm om rm gifts were not tax deductible? 6b 7 Organizations that may receive deductible contributions under section 170(c). a Did the organization receive a paymm emntm imn m emxcmesm sm omf m$7m 5m m madm em pm amrtlmy masm ma mcom nmtrmibmutmiomn m amndm pm amrtmlym fom rm gmoom dmsm and services provided to the payor? m m m m m m m m m m m m 7a X b If "Yes," did the organization notify the donor of the value of the goods or services provided? 7b c Did the organization sell, exm cmham nmgem , m omr m omthmermwmisme m dmismpom sme m omf mtamngm ibm lem mpmermsom nmalm pm rom pmerm tym mfomr mwmhimchm mit m wm amsm required to file Form 8282? m m m m m m m m m m m m m m m m 7c X d If "Yes," indicate the number of Forms 8282 filed during the year 7d m m m e Did the organization receive any funds, directly or indirectly, to pay premiums on a personal benefit contract? 7e X f Did the organization, during the year, pay premiums, directly or indirectly, on a personal benefit contramctm?m 7f X g If the organization received a contribution of qualified intellectual property, did the organization file Form 8899 as required? 7g h If the organization received a contribution of cars, boats, airplanes, or other vehicles, did the organization file a Form 1098-C? 7h 8 Sponsoring organizations maintaining donor advised funds and section 509(a)(3) supporting organizations. Did the supporting organization, or a donor advisemd m fmunm dm mmmaimntmaimnem dm mbym m a m sm pmonm smorminmgm organization, have excess business holdings at any time during the year? 8 X 9 Sponsoring organizations maintaining donor advised funds. m m m m m m m m m m m m m m m m m m m m m m m a Did the organization make any taxable distributions under section 4966? m m m m m m m m m m m m m m m m 9a b Did the organization make a distribution to a donor, donor advisor, or related person? 9b 10 Section 501(c)(7) organizations. Enter: m m m m m m m m m m m m m m a Initiation fees and capital contributions included on Part VIII, line 12 m m m m 10a b Gross receipts, included on Form 990, Part VIII, line 12, for public use of club facilities 10b 11 Section 501(c)(12) organizations. Enter: m m m m m m m m m m m m m m m m m m m m m m m m m m a Gross income from members or shareholders 11a b Gross income from other sources (Do not m nmetm mamm om umntms m dmuem momr mpam idm mtom motmhem r m smomurmcemsm against amounts due or received from them.) 11b 12 a Section 4947(a)(1) non-exempt charitable trusts. Is the organization filing Form 9m9m0 m inm lmieu of Form 1041? 12a b If "Yes," enter the amount of tax-exempt interest received or accrued during the year 12b 13 Section 501(c)(29) qualified nonprofit health insurance issuers. m m m m m m m m m m m m m m m m m m a Is the organization licensed to issue qualified health plans in more than one state? 13a Note. See the instructions for additional information the organization must report on Schedule O. b Enter the amount of reserves the organization is required tom m am inm tam inm bm ym thm em smtamtems minm wm hmicmh m m the organization is licensed to issue qumalmifiem dm hm emalmthm pmlamnsm m m m m m m m m m m m m m m m m m m m m 13b c Enter the amount of reserves on hand m m m 1m3cm m m m m m m m m 14 a Did the organization receive any payments for indoor tanning services during the tax year? m m m m m m 14a X b If "Yes," has it filed a Form 720 to report these payments? If "No," provide an explanation in Schedule O 14b JSA 2E1040 1.000 Form 990 (2012) 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 5 Form 990 (2012) ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Page 6 Part VI Governance, Management, and Disclosure For each "Yes" response to lines 2 through 7b below, and for a "No" response to line 8a, 8b, or 10b below, describe the circumstances, procemssmesm , morm cmham nmgem sm inm Sm cmhem dmulme mO.m Sm eem imnsm trm umctimonm s. Check if Schedule O contains a response to any question in this Part VI X Section A. Governing Body and Management m m m m m m m m m m m Yes No 1a Enter the number of voting members of the governing body at the end of the tax year. 1a 5 If there are material differences in voting rights among members of the governing body, or if the governing body delegated broad authority to an executive committee or similar committee, explain in Schedumle mOm. m m m b Enter the number of voting members included in line 1a, above, who are independent 1b 5 2 Did any officer, director, trustee, or key employee hmavme m am fmamm imly m rem lam timonm smhimp m omr ma mbum sminmesm sm rmelmatmiomnsmhmip m wm itmh m any other officer, director, trustee, or key employee? 2 X 3 Did the organization delegate control over management duties customarily performed by or under the dirmecm tm supervision of officers, directors, or trustees, or key employees to a management company or other pemrsmonm ?m m m m 3 X 4 Did the organization make any significant changes to its governing documents since the prior Form 990 was filed? m m m m m 4 X 5 Did the organization become aware during the year ofm am sm igmnimficm amntm dmivmermsimonm om f mthme mormgam nmizam timonm 'sm amssm emtsm? m m m m m 5 X 6 Did the organization have members or stockholders? 6 X 7a Did the organization have members, stockholmdem rsm , m omr motmhem r m pmerm som nms m wmhom mham dm tmhem mpom wm emr mtom em lem cmt morm am pmpom inm tm one or more members of the governing body? 7a X b Are any governance decisions of the organization rem smerm vem dm mtom m(omr m smubm jem cmt m tom mamppm rom vmalm bm ym) m mm emmmbem rms,m stockholders, or persons other than the governing body? 7b X 8 Did the organization contemporaneously document the meetings held or written actions undertaken during the year by the followminmg:m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m a The governing body? m m m m m m m m m m m m m m m m m m m m m m m 8a X b Each committee with authority to act on behalf of the governing body? 8b X 9 Is there any officer, director, trustee, or key employee listed in Part VII, Section A, who camnnm omt mbem mremacm hmedm am tm the organization's mailing address? If "Yes," provide the names and addresses in Schedule O 9 X Section B. Policies (This Section B requests information about policies not required by the Internal Revenue Code.) m m m m m m m m m m m m m m m m m m m m m m m m m m m Yes No 10a Did the organization have local chapters, branches, or affiliates? 10a X b If "Yes," did the organization have written policies and procedures governing the activities of such chapm tem rsm ,m affiliates, and branches to ensure their operations are consistent with the organization's exempt purposes? m m 10b 11a Has the organization provided a complete copy of this Form 990 to all members of its governing body before filing the form? 11a X b Describe in Schedule O the process, if any, used by the organization to review this Fm om rmm m99m 0m. m m m m m m m m m m m 12a Did the organization have a written conflict of interest policy? If "No," go to line 13 12a X b Were officers, direm cmtom rsm , morm tmrumstmeem sm, am nmd m kem ym em mm pmlomyemesm mremqum irmedm mtom dmismclmosm em amnnm umalmly m inm tem rem smtsm thm am t mcom umldm gm ivmem rise to conflicts? 12b X c Did the organization regularly and consism tem nmtlym m monm itmorm manm dm em nmfomrcme m cmom pm limanm cme m wm itmh m tmhem mpom limcym? m Ifm "mYmesm,"m describe in Schedule O how this was done m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m 12c X 13 Did the organization have a written whistleblower policy? m m m m m m m m m m m m m m m m m m m 13 X 14 Did the organization have a written document retention and destruction policy? 14 X 15 Did the process for determining compensation of the following persons include a review and approval by independent persons, comparability data, and contemporaneous substanm timatmiomn mofm thm em dmelmibmermamtiom nm amndm dm emcimsiom nm? m a The organization's CEO, Executive Director, or top mm amnam gmem em nmt om ffmicmialm m m m m m m m m m m m m m m m m m m m m m m m 15a X b Other officers or key employees of the organization 15b X If "Yes" to line 15a or 15b, describe the process in Schedule O (see instructions). 16a Did the organization invest in, contrmibmutme m amssmetms m tom , m omr mpam rtmicmipmatme m inm ma mjom inmt mvem nmtumrem om rm sm imm ilmarm am rmramngm emmmenm tm with a taxable entity during the year? 16a X b If "Yes," did the organization follow a written policy or procedure requiring the organization to evaluate its participation in joint venture arrangements under applicable fedemraml mtamx mlamwm, am nmd m tam kme mstmepm sm tmo msam fem gmuam rmd mthmem organization's exempt status with respect to such arrangements? 16b Section C. Disclosure 17 List the states with which a copy of this Form 990 is required to be filed I CA,DE, 18 Section 6104 requires an organization to make its Forms 1023 (or 1024 if applicable), 990, and 990-T (Section 501(c)(3)s only) available for public inspection. Indicate how you made these available. Check all that apply. X Own website Another's website X Upon request Other (explain in Schedule O) 19 Describe in Schedule O whether (and if so, how), the organization made its governing documents, conflict of interest policy, and financial statements available to the public during the tax year. 20 State the name, physical address, and telephone number of the person who possesses the books and records of the organization: I PAUL KELLY SUITE 203 SHERATON HOUSE CB3 OAX CASTLE PARK CAMBRIDGE UK JSA Form 990 (2012) 2E1042 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 6 Form 990 (2012) ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Page 7 Part VII Compensation of Officers, Directors, Trustees, Key Employees, Highest Compensated Employees, and Independent Contractors m m m m m m m m m m m m m m m m m m m m Check if Schedule O contains a response to any question in this Part VII Section A. Officers, Directors, Trustees, Key Employees, and Highest Compensated Employees 1a Complete this table for all persons required to be listed. Report compensation for the calendar year ending with or within the organization's tax year. % List all of the organization's current officers, directors, trustees (whether individuals or organizations), regardless of amount of compensation. Enter -0- in columns (D), (E), and (F) if no compensation was paid. % List all of the organization's current key employees, if any. See instructions for definition of "key employee." % List the organization's five current highest compensated employees (other than an officer, director, trustee, or key employee) who received reportable compensation (Box 5 of Form W-2 and/or Box 7 of Form 1099-MISC) of more than $100,000 from the organization and any related organizations. % List all of the organization's former officers, key employees, and highest compensated employees who received more than $100,000 of reportable compensation from the organization and any related organizations. % List all of the organization's former directors or trustees that received, in the capacity as a former director or trustee of the organization, more than $10,000 of reportable compensation from the organization and any related organizations. List persons in the following order: individual trustees or directors; institutional trustees; officers; key employees; highest compensated employees; and former such persons. Check this box if neither the organization nor any related organization compensated any current officer, director, or trustee. (C) (A) (B) Position (D) (E) (F) Name and Title Average (do not check more than one Reportable Reportable Estimated hours per box, unless person is both an compensation compensation from amount of week (list any officer and a director/trustee) from related other

hours for o I I O K e H F compensation

n n the organizations m r e o i f d s g

f y r d

p from the t i i h m c

related v organization (W-2/1099-MISC) i i e l t e r e o i u e d e m s r y organization t r u c i organizations t (W-2/1099-MISC) e o p

t a c o e n l and related l o o

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below dotted a t y m r l e u organizations t p e s line) r u e t e s n e t s e a e t e d

(1) ELIZABETH MARINCOLA 5.00 DIRECTOR AND CHAIR X 0 0 0 (2) SIR MARK WALPORT 2.00 DIRECTOR X 0 0 0 (3) ROBERT TJIAN 2.00 DIRECTOR X 0 0 0 (4) PETER GRUSS 2.00 DIRECTOR X 0 0 0 (5) TOBY COPPEL 2.00 DIRECTOR X 0 0 0 (6) DR. RANDY SCHEKMAN 20.00 EDITOR IN CHIEF X 87,500. 0 0 (7) DR. MARK PATTERSON 35.00 EXEC DIR, SECRETARY & TREAS X 116,211. 0 11,621. (8)

(9)

(10)

(11)

(12)

(13)

(14)

JSA Form 990 (2012)

2E1041 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 7 ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Form 990 (2012) Page 8 Part VII Section A. Officers, Directors, Trustees, Key Employees, and Highest Compensated Employees (continued) (A) (B) (C) (D) (E) (F) Name and title Average Position Reportable Reportable Estimated hours per (do not check more than one compensation compensation from amount of week (list any box, unless person is both an from related other hours for officer and a director/trustee) compensation

o I I O K e H F the organizations n n m r e o i f d s g from the

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m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m I 1b Sub-total m m m m m m m m m m m m m I 203,711. 0 11,621. c Total from continuation shem etm sm tom Pm am rtm Vm IIm, Sm em ctm iom nm Am m m m m m m m m m m m m m 0 0 0 d Total (add lines 1b and 1c) I 203,711. 0 11,621. 2 Total number of individuals (including but not limited to those listed above) who received more than $100,000 of reportable compensation from the organization I 1 Yes No 3 Did the organization list any former officer, director, or trusteem , m km emy m emmmplmoym eme,m morm mhimghm emstm mcommm pmenm smatmedm employee on line 1a? If "Yes," complete Schedule J for such individual 3 X 4 For any individual listed on line 1a, is the sum of reportable compensation and other compensation from the organizatiom nm manm dm mrem lam tem dm mormgam nmizmatmiomnsm m gmrematmerm m thm amn m $m 1m5m0,m00m 0m?m mIf m “mYem sm,” m cmom mplmetme m Sm cmhem dmulme m Jm m fom r m sm umchm individual 4 X 5 Did any person listed on line 1a receive or accrue compensation from any unrelatedm mormgam nm izmatmiomn m omr minmdimvimdum aml for services rendered to the organization? If “Yes,” complete Schedule J for such person 5 X Section B. Independent Contractors 1 Complete this table for your five highest compensated independent contractors that received more than $100,000 of compensation from the organization. Report compensation for the calendar year ending with or within the organization's tax year.

(A) (B) (C) Name and business address Description of services Compensation

2 Total number of independent contractors (including but not limited to those listed above) who received more than $100,000 in compensation from the organization I 0 JSA Form (2012) 2E1055 3.000 990 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 8 Form 990 (2012) ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Page 9 Part VIII Statement of Revenue m m m m m m m m m m m m m m m m m m m m m m m m m Check if Schedule O contains a response to any question in this Part VIII (A) (B) (C) (D) Total revenue Related or Unrelated Revenue exempt business excluded from tax function revenue under sections m m m m m m m m revenue 512, 513, or 514 s s t t 1a Federated campaigns 1a n n a

u m m m m m m m m m r

o b Membership dues 1b G

m m m m m m m m m m , s A c Fundraising events 1c

t f r

i m m m m m m m m a l

G 1d

i d Related organizations

, m m m s i 1e

n e Government grants (contributions) S

o i r t

e f All other contributions, gifts, grants, u

h m b t i 1f 5,140,020. r and similar amounts not included above O t

n d

o g Noncash contributions included in lines 1a-1f: $ n m m m m m m m m m m m m m m m m m m m C a I h Total. Add lines 1a-1f 5,140,020. e

u Business Code n e

v 2a e R b e c i

v c r e

S d

m

a e

r m m m m m g f All other program service revenue o

r m m m m m m m m m m m m m m m m m m m I

P g Total. Add lines 2a-2f 0 3 Investment income (inmclum dmingm dm ivmidmenm dsm , mintmermesmt, manm dm m m m I other similar amounts) m m m I 0 4 Income frommm inm vmesmtmm emntm ofm tam xm-emxemmmpt mbom nmd pm rom cem emdsm m m m I 0 5 Royalties 0 m m m m m m m m (i) Real (ii) Personal 6a Gross rents m m m b Less: rental expenses m m c Rental income or (loss) m m m m m m m m m m m m m m m m m I d Net rental income or (loss) 0 (i) Securities (ii) Other 7a Gross amount from sales of assets other than inventory b Less: cost or other bamsims m m and sales expemnsmesm m m m m c Gain or (loss) m m m m m m m m m m m m m m m m m m m m m I d Net gain or (loss) 0

e 8a Gross income from fundraising u

n events (not including $ e

v of contributions reported on line 1c). e m m m m m m m m m m m R

See Part IV, line 18 a r m m m m m m m m m m e b Less: direct expenses b h m m m m m m m m t I c Net income or (loss) from fundraising events 0 O 9a Gross income from gm am imngm amctimvitmiems. m m m See Part IV, line 19 m m m m m m m m m m a b Less: direct expenses m bm m m m m m m m I c Net income or (loss) from gaming activities 0 10a Gross sales of invmenm tom rym, m mlesm sm m returns and allowances m m m m m m m m m a b Less: cost of goods sold m bm m m m m m m m I c Net income or (loss) from sales of inventory 0 Miscellaneous Revenue Business Code

11a OTHER INCOME 900099 1,791. 1,791. b c m m m m m m m m m m m m m d All other revenue m m m m m m m m m m m m m m m m m I e Total. Add lines 11a-11d m m m m m m m m m m m m m m I 1,791. 12 Total revenue. See instructions 5,141,811. 1,791. Form (2012) JSA 990 2E1051 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 9 Form 990 (2012) ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Page 10 Part IX Statement of Functional Expenses Section 501(c)(3) and 501(c)(4) organizations must complete all columns. All other omrgmanm izmatmiomnsm mm um smt cmompm lem tem cm omlum mn m(Am). m m m m m Check if Schedule O contains a response to any question in this Part IX Do not include amounts reported on lines 6b, 7b, (A) (B) (C) (D) Total expenses Program service Management and Fundraising 8b, 9b, and 10b of Part VIII. expenses general expenses expenses 1 Grants and other assistance to governments andm organizations in the United States. See Part IV, line 21 0 2 Grants and other assistance to indivm idm umalsm minm the United States. See Part IV, line 22 0 3 Grants and other assistance to governments, organizations, and individuals outsidem mthme m United States. See Part IV, lines 1m 5m amndm 1m6m m m m 0 4 Benefits paid to or for members 0 5 Compensation of current omffimcemrsm, m dmiremctmorms,m trustees, and key employees 409,272. 285,751. 123,521. 6 Compensation not included above, to disqualified persons (as defined under section 4958m(f)m(1m)) m anm dm persons described in section 4m95m 8(mc)m(3m)(Bm) m m m m m m 0 7 Other salaries and wages 386,486. 343,921. 42,565. 8 Pension plan accruals and contributions (incmludm e msemctimonm 401(k) and 403(b) employer cmonm trmibum tiom nsm ) m m m m m m 42,163. 31,946. 10,217. 9 Other employmeem bmenm efmitsm m m m m m m m m m m m m 683. 683. 10 Payroll taxes 72,595. 56,661. 15,934. 11 Fees for servicesm (mnomn-memm pmlomyemesm):m m m m m m m m a Managem mmenm t m m m m m m m m m m m m m m m m m m 0 b Legal m m m m m m m m m m m m m m m m m m 333,106. 333,106. c Accountingm m m m m m m m m m m m m m m m m m m 84,101. 84,101. d Lobbying 0 e Professional fundraising services. Seme mPamrt mIVm, lminem 1m 7 m 0 f Investment management fees 0 g Other. (If line 11g amount exceeds 10% of line 25, column m m m m m m 386,212. 255,292. 130,920. (A) amount, list line 11g expenses on mScmhemdulme Om .) m m m m m m 12 Advertising and pmrom motimonm m m m m m m m m m m m 331,450. 331,450. 13 Office expenses m m m m m m m m m m m m m 0 14 Informatiom nm tem chm nmolmogmy m m m m m m m m m m m m m 0 15 Royalties m m m m m m m m m m m m m m m m m m 0 16 Occupamncm ym m m m m m m m m m m m m m m m m m m 114,352. 86,643. 27,709. 17 Travel 128,180. 107,622. 20,558. 18 Payments of travel or entertainment expenses for any federal, state, or local public ofm ficm iamlsm 0 19 Conferenmcems,m cmonmvem ntmiomnsm, am ndm mm emetminmgsm m m m m 0 20 Interest m m m m m m m m m m m m m m 0 21 Payments to affiliates m m m m 0 22 Depreciatiomn,m dem pmlemtiomn,m amndm ammmormtizmatmionm m m m m 7,149. 5,417. 1,732. 23 Insurance 21,032. 8,666. 12,366. 24 Other expenses. Itemize expenses not covered above (List miscellaneous expenses in line 24e. If line 24e amount exceeds 10% of line 25, column (A) amount, list line 24e expenses on Schedule O.) a EDITORIAL COSTS 940,810. 940,810. b RECRUITMENT 125,165. 89,440. 35,725. c WEBSITE & DEVELOPMENT 155,005. 155,005. d MISCELLANEOUS 28,086. 27,660. 426. e All other expenses 25 Total functional expenses. Add lines 1 through 24e 3,565,847. 2,726,967. 838,880. 26 Joint costs. Complete this line only if the organization reported in column (B) joint costs from a combined educational caImpaign and fundraising solicitation. Check here m m m m m m ifm following SOP 98-2 (ASC 958-720) 0 JSA Form 990 (2012) 2E1052 1.000

8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 10 ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Form 990 (2012) Page 11 Part X Balance Sheet m m m m m m m m m m m m m m m m m m m m m Check if Schedule O contains a response to any question in this Part X (A) (B) Beginning of year End of year 1 Cash - non-interest-bearing m m m m m m m m m m m m m m m m m m m m m m m m m m m 0 1 3,022,470. 2 Savings and temporary cash investments m m m m m m m m m m m m m m m m m m m m 0 2 0 3 Pledges and grants receivable, net m m m m m m m m m m m m m m m m m m m m m m m 0 3 0 4 Accounts receivable, net m m m m m m m m m m m m m m m m m m m m m m m m m m m m 0 4 0 5 Loans and other receivables from current and former officers, directors, trustees, key employees, and highest compensated employees. Complete Part II of Schedule L m m m m m m m m m m m m m m m m m m m m m m m m m 0 5 0 6 Loans and other receivables from other disqualified persons (as defined under section 4958(f)(1)), persons described in section 4958(c)(3)(B), and contributing employers and sponsoring organizations of section 501(c)(9) voluntary employees' beneficiary organizations (see instructions). Complete Part II of Schedule L m m m m m m m m m m m 0 6 0 s

t m m m m m m m m m m m m m m m m m m m m m m m m m

e 7 Notes and loans receivable, net 0 7 0 s m m m m m m m m m m m m m m m m m m m m m m m m m m m m s 8 Inventories for sale or use 0 8 0 A m m m m m m m m m m m m m m m m m m m m 9 Prepaid expenses and deferred charges 0 9 0 10 a Land, buildings, and equipment: cost or other basis. Complete Part VI of mScm hmedm umlem Dm m m m 10a 35,881. b Less: accumulated depreciation m m m m m m10m bm m m m m m m m 7,274.m m m m m 0 10c 28,607. 11 Investments - publicly traded securities m m m m m m m m m m m m m m m 0 11 0 12 Investments - other securities. See Part IV, line 11 m m m m m m m m m m m m m m 0 12 0 13 Investments - promgrmam -mrem lamtemd.m Sm eme mPam rtm IVm ,m linm em 1m1 m m m m m m m m m m m m m m m 0 13 0 14 Intangible assets m m m m m m m m m m m m m m m m m m m m m m m m 0 14 0 15 Other assets. See Part IV, line 11 m m m m m m m m m m 0 15 325,061. 16 Total assets. Add lines 1 through 15 (mumstm emqum aml lminme m34m )m m m m m m m m m m m 0 16 3,376,138. 17 Accounts payabmlem amndm macm crm umedm em xpm emnsm ems m m m m m m m m m m m m m m m m m m m m 0 17 884,034. 18 Grants payable m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m 0 18 0 19 Deferred revenue m m m m m m m m m m m m m m m m m m m m m m m m m m m 0 19 884,479. 20 Tax-exempt bond liabilities m m m m 0 20 0 s 21 Escrow or custodial account liability. Complete Part IV of Schedule D 0 21 0 e i t

i 22 Loans and other payables to current and former officers, directors, l i

b trustees, key employees, highest compensated employees, and a i

L m m m m m m m m m m m m m m disqualified persons. Complete Part II of Schedule L m m m m m m m 0 22 0 23 Secured mortgages and notes payable to unrelated third partimesm m m m m m m m 0 23 0 24 Unsecured notes and loans payable to unrelated third parties 0 24 0 25 Other liabilities (including federal income tax, payables to related third parties, and othmerm limabm ilmitimesm nm omt minmclmudm emd momn mlinm ems m17m -m24m ).m Cm ommmpmletme m Pmarmt mX m of Schedule D m m m m m m m m m m m m m m m m m m m m 0 25 24,260. 26 Total liabilities. Add lines 17 through 25 0 26 1,792,773. Organizations that follow SFAS 117 (ASC 958), check here I X and

s complete lines 27 through 29, and lines 33 and 34. e

c m m m m m m m m m m m m m m m m m m m m m m m m m m m m m n 27 Unrestricted net assets 0 27 1,583,365. a l

a 28 Temporarily restricted net assets m m m m m m m m m m m m m m m m m m m m m m m m 0 28 0 B

m m m m m m m m m m m m m m m m m m m m m m m m

d 29 Permanently restricted net assets 0 29 0 n

u I Organizations that do not follow SFAS 117 (ASC 958), check here and F

r complete lines 30 through 34. o

s m m m m m m m m m m m m m m m m

t 30 Capital stock or trust principal, or current funds 30 e m m m m m m m m s 31 Paid-in or capital surplus, or land, building, or equipment fund 31 s

A m m m m 32 Retained earnings, endowment, accumulated income, or other funds 32 t e 33 Total net assets or fund balances m m m m m m m m m m m m m m m m m m m m m m m m 0 1,583,365. N m m m m m m m m m m m m m m m m m m 33 34 Total liabilities and net assets/fund balances 0 34 3,376,138. Form 990 (2012)

JSA 2E1053 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 11 ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Form 990 (2012) Page 12 Part XI Reconciliation of Net Assets m m m m m m m m m m m m m m m m m m Check if Schedule O contains a response to amnmy mqum em smtiom nm inm mthmism Pmamrt mXmI m m m m m m m X 1 Total revenue (must equal Part VIII, column (A), line 12) m m m m m m m m m m m m m m m m m m m m m m m 1 5,141,811. 2 Total expenses (must equal Part IX, column (A), linem 2m5)m m m m m m m m m m m m m m m m m m m m m m m m 2 3,565,847. 3 Revenue less expenses. Subtract line 2 from line 1 m m m m m 3 1,575,964. 4 Net assets or fund balances at beginning of ymeam r m(mm um smt em qmuam l Pm am rtm Xm, lminme m33m , mcomlummmn m(Am))m m m m m m 4 0 5 Net unrealized gains (losses) on investm em nmtsm m m m m m m m m m m m m m m m m m m m m m m m m m m m m m 5 0 6 Donated services andm umsem om f mfamcilmitiem sm m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m 6 0 7 Investment expenses m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m 7 0 8 Prior period adjustments m m m m m m m m m m m m m m m m 8 0 9 Other changes in net assets or fund balances (explain in Schedule O) 9 7,401. 10 Net assets or funm dm bmalmanm cmesm am t m emndm om f myem amr.m Cmommbimnem lminem sm 3m thm rom umghm 9m (mmmusm t meqm umalm Pmarmt Xm ,m linm em m m 33, column (B)) 10 1,583,365. Part XII Financial Statements and Reporting m m m m m m m m m m m m m m m m m Check if Schedule O contains a response to any question in this Part XII Yes No 1 Accounting method used to prepare the Form 990: Cash X Accrual Other If the organization changed its method of accounting from a prior year or checked "Other," explain in Schedule O. m m m m m m 2a Were the organization's financial statements compiled or reviewed by an independent accountant? 2a X If "Yes," check a box below to indicate whether the financial statements for the year were compiled or reviewed on a separate basis, consolidated basis, or both: Separate basis Consolidated basis Both consolidated and sepamramtem bmasm ism m m m m m m m m b Were the organization's financial statements audited by an independent accountant? 2b X If "Yes," check a box below to indicate whether the financial statements for the year were audited on a separate basis, consolidated basis, or both: X Separate basis Consolidated basis Both consolidated and separate basis c If "Yes" to line 2a or 2b, does the organization have a committee that assumes responsibility for oversight of the audit, review, or compilation of its financial statements and selection of an independent accountant? 2c X If the organization changed either its oversight process or selection process during the tax year, explain in Schedule O. 3a As a result of a federal award, was the organimzamtiom nm rmeqm umirem dm tmo m umndm emrgm om amn m amudm itm omr maum dmitsm masm sm emt fmorm thm imnm the Single Audit Act and OMB Circular A-133? 3a X b If "Yes," did the organization undergo the required audit or audits? If the organization did not undergo the required audit or audits, explain why in Schedule O and describe any steps taken to undergo such audits 3b Form 990 (2012)

JSA 2E1054 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 12 SCHEDULE A Public Charity Status and Public Support OMB No. 1545-0047 (Form 990 or 990-EZ) À¾µ¶ Complete if the organization is a section 501(c)(3) organization or a section 4947(a)(1) nonexempt charitable trust. Open to Public Department of the Treasury I I Internal Revenue Service Attach to Form 990 or Form 990-EZ. See separate instructions. Inspection Name of the organization Employer identification number ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Part I Reason for Public Charity Status (All organizations must complete this part.) See instructions. The organization is not a private foundation because it is: (For lines 1 through 11, check only one box.) 1 A church, convention of churches, or association of churches described in section 170(b)(1)(A)(i). 2 A school described in section 170(b)(1)(A)(ii). (Attach Schedule E.) 3 A hospital or a cooperative hospital service organization described in section 170(b)(1)(A)(iii). 4 A medical research organization operated in conjunction with a hospital described in section 170(b)(1)(A)(iii). Enter the hospital's name, city, and state: 5 An organization operated for the benefit of a college or university owned or operated by a governmental unit described in section 170(b)(1)(A)(iv). (Complete Part II.) 6 A federal, state, or local government or governmental unit described in section 170(b)(1)(A)(v). 7 X An organization that normally receives a substantial part of its support from a governmental unit or from the general public described in section 170(b)(1)(A)(vi). (Complete Part II.) 8 A community trust described in section 170(b)(1)(A)(vi). (Complete Part II.) 9 An organization that normally receives: (1) more than 331/3 % of its support from contributions, membership fees, and gross receipts from activities related to its exempt functions - subject to certain exceptions, and (2) no more than 331/3% of its support from gross investment income and unrelated business taxable income (less section 511 tax) from businesses acquired by the organization after June 30, 1975. See section 509(a)(2). (Complete Part III.) 10 An organization organized and operated exclusively to test for public safety. See section 509(a)(4). 11 An organization organized and operated exclusively for the benefit of, to perform the functions of, or to carry out the purposes of one or more publicly supported organizations described in section 509(a)(1) or section 509(a)(2). See section 509(a)(3). Check the box that describes the type of supporting organization and complete lines 11e through 11h. a Type I b Type II c Type III-Functionally integrated d Type III-Non-functionally integrated e By checking this box, I certify that the organization is not controlled directly or indirectly by one or more disqualified persons other than foundation managers and other than one or more publicly supported organizations described in section 509(a)(1) or section 509(a)(2). f If the organization received a written determination from the IRS that it is a Type I, Type II, or Type III supporting organization, check this box m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m g Since August 17, 2006, has the organization accepted any gift or contribution from any of the following persons? (i) A person who directly or indirectly controls, either alone or together with persons described in (ii) Yes No and (iii) below, the governing body of the supported organization? m m m m m m m m m m m m m m m m m m m m m 11g(i) (ii) A family member of a person described in (i) above? m m m m m m m m m m m m m m m m m m m m m m m m m m m m m 11g(ii) (iii) A 35% controlled entity of a person described in (i) or (ii) above? m m m m m m m m m m m m m m m m m m m m m m 11g(iii) h Provide the following information about the supported organization(s). (i) Name of supported (ii) EIN (iii) Type of organization (iv) Is the (v) Did you notify (vi) Is the (vii) Amount of monetary organization (described on lines 1-9 organization in the organization organization in support above or IRC section col. (i) listed in in col. (i) of col. (i) organized your governing (see instructions)) document? your support? in the U.S.? Yes No Yes No Yes No

(A)

(B)

(C)

(D)

(E)

Total For Paperwork Reduction Act Notice, see the Instructions for Schedule A (Form 990 or 990-EZ) 2012 Form 990 or 990-EZ.

JSA 2E1210 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 13 ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Schedule A (Form 990 or 990-EZ) 2012 Page 2 Part II Support Schedule for Organizations Described in Sections 170(b)(1)(A)(iv) and 170(b)(1)(A)(vi) (Complete only if you checked the box on line 5, 7, or 8 of Part I or if the organization failed to qualify under Part III. If the organization fails to qualify under the tests listed below, please complete Part III.) Section A. Public Support Calendar year (or fiscal year beginning in) I (a) 2008 (b) 2009 (c) 2010 (d) 2011 (e) 2012 (f) Total

1 Gifts, grants, contributions, and membership fees received. m (mDmo m nm omt include any "unusual grants.") 0 0 0 0 5,140,020. 5,140,020.

2 Tax revenues levied for the organization's benefit and m emithm emr m pam idm to or expended on its behalf 0

3 The value of services or facilities furnished by a governmentaml um nmit m tom tmhem organization without charge m m m m m m m 0 4 Total. Add lines 1 through 3 5,140,020. 5,140,020. 5 The portion of total contributions by each person (other than a governmental unit or publicly supported organization) included on line 1 that exceeds 2% of m thm em am mmoum nmt shown on line 11, column (f) 2,217,174. 6 Public support. Subtract line 5 from line 4. 2,922,846. Section B. Total Support I Calendar year (or fiscal year mbem gminnm inm gm inm ) m m m (a) 2008 (b) 2009 (c) 2010 (d) 2011 (e) 2012 (f) Total 7 Amounts from line 4 5,140,020. 5,140,020. 8 Gross income from interest, dividends, payments received on securities loans, rents, romyam ltimesm am nmd minmcom mme m frmomm msim milamr sources 0

9 Net income from unrelated business activities, whether or m nmotm tmhem mbum simnem ssm is regularly carried on 0

10 Other income. Do not include gain or loss from the salem m ofm mcam pmitaml m asm smetsm (Explain in Part IV.) ATCH 1 m m 1,791. 1,791. 11 Total support. Add lines 7 through 10 m m m m m m m m m m m m m m m m m m m m m m m m m m 5,141,811. 12 Gross receipts from related activities, etc. (see instructions) 12 13 First five years. If the Form 990 is for m thm em om rgm amnimzamtiomn'ms m fimrstm, msem com nmd,m tmhimrdm, mfomurmth,m mor m fmiftmh m tam x m ymeam r m asm ma m smecm tiom nm 5m 0m1(mc)m(3) organization, check this box and stop here I X Section C. Computation of Public Support Percentage m m m m m m m m 14 Public support percentage for 2012 (line 6, column (f) divided by lminme m11m ,m cmolum mm nm (fm))m m m m m m m m m 14 % 15 Public support percentage from 2011 Schedule A, Part II, line 14 15 % 16a 33 1/3 % support test - 2012. If the organization did not check the box on line 13, anm dm limnem 1m 4m ims m3m31m /3m %m om rm mm omrem , mchm emck this box and stop here. The organization qualifies as a publicly supported organization I b 33 1/3 % support test - 2011. If the organization did not check a box on line 13 or 16a, amndm mlinm em 1m5m ism 3m 3m 1m/3m%m omr mmmomre, check this box and stop here. The organization qualifies as a publicly supported organization I 17a 10%-facts-and-circumstances test - 2012. If the organization did not check a box on line 13, 16a, or 16b, and line 14 is 10% or more, and if the organization meets the "facts-and-circumstances" test, check this box and stop here. Explain in Part IV how mthme m omrgmamnizm amtiom nm mm em emtsm tmhem "m fam cmtsm-amndm -cm irmcum mm stmanm cmesm ” m tem smt. mTmhem om rgm am nmizam timonm mqum amlifmiems masm am mpum bmlicm lym sm umppm omrtmed organization I b 10%-facts-and-circumstances test - 2011. If the organization did not check a box on line 13, 16a, 16b, or 17a, and line 15 is 10% or more, and if the organization meets the "facts-and-circumstances" test, check this box and stop here. Explain in Part IV how mthme m omrgmamnizm amtiom nm mm em emtsm tmhem "m fam cmtsm-amndm -cm irmcum mm smtamncmesm " m tem smt. m Tmhem om rmgam nmizmatmiomn mqum amlifmiems masm ma mpum bm licm ly supported organization I 18 Private foundm amtiom nm . mIf mthme mormgmanm izmatmiomn mdmid m nmotm cmhem cmk ma mbom xm omn mlinm em 1m3,m 1m 6ma,m 1m6bm , m1m7am , morm 1m 7mb,m cmhem cmk mthmis mbom xm amndm sm eme m m m m instructions I Schedule A (Form 990 or 990-EZ) 2012

JSA

2E1220 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 14 ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Schedule A (Form 990 or 990-EZ) 2012 Page 3 Part III Support Schedule for Organizations Described in Section 509(a)(2) (Complete only if you checked the box on line 9 of Part I or if the organization failed to qualify under Part II. If the organization fails to qualify under the tests listed below, please complete Part II.) Section A. Public Support I Calendar year (or fiscal year beginning in) (a) 2008 (b) 2009 (c) 2010 (d) 2011 (e) 2012 (f) Total 1 Gifts, grants, contributions, and membership fees received. (Do not include any "unusual grants.") 2 Gross receipts from admissions, merchandise sold or services performed, or facilities furnished in any activity that is relmatem d m tom thm em organization's tax-exempt purpose 3 Gross receipts from activities that are not anm unrelated trade or business under section 513 4 Tax revenues levied for the organization's benefit and meimthmerm pm amidm to or expended on its behalf 5 The value of services or facilities furnished by a governmentalm umnimt tmo m thm em organization without charge m m m m m m m 6 Total. Add lines 1 through 5 7a Amounts included on lines 1, 2, am nmd m3m received from disqualified persons b Amounts included on lines 2 and 3 received from other than disqualified persons that exceed the greater of $5,000 or 1% of the amount omn mlinme m13m fmor m thme myemarm c Add lines 7a and 7b 8 Public smupm pmormt m(Sm umbtrmacm t m linm em 7m cm fmrom m line 6.) Section B. Total Support I Calendar year (or fiscal ymeam r mbem gminmninm gm inm )m m (a) 2008 (b) 2009 (c) 2010 (d) 2011 (e) 2012 (f) Total 9 Amounts from line 6 10 a Gross income from interest, dividends, payments received on securities loans, rents, romyamltimesm amndm imncm om em mfrom mm sm imm ilmarm sources b Unrelated business taxable income (less section 511 taxes) from bm usm inmesm sem sm acquired after June 30, 1m97m 5m m m m m m m c Add lines 10a and 10b 11 Net income from unrelated business activities not included in line 10b, whether or nm omt tmhem bm umsinm emssm ism rmegm umlarmlym carried on 12 Other income. Do not include gain or loss from the salem mof m cm ampitmalm am smsemtsm (Explain in Part IV.) 13 Total supmpomrtm. (m Amddm mlinmesm 9m , m 1m0cm , m 11m ,m and 12.) 14 First five years. If the Form 990 is for m thm em om rgm anm izm amtiomn'sm mfirmst,m sm emcomndm , mthmirdm , m fom umrthm , morm fmiftmh m tam x m ymeamr masm ma msem ctmiomn m 5m01m(cm)(3m )m m organization, check this box and stop here I Section C. Computation of Public Support Percentage m m m m m m m m m m m m m m 15 Public support percentage for 2012 (line 8, column (f) divided by lminem 1m 3m, cmolum mm nm (fm)) m m m m m m m m m m m m m m 15 % 16 Public support percentage from 2011 Schedule A, Part III, line 15 16 % Section D. Computation of Investment Income Percentage m m m m m m m m m m 17 Investment income percentage for 2012 (line 10c, column (f) divided bym limnem 1m3,m com lummmn m(f)m) m m m m m m m m m m 17 % 18 Investment income percentage from 2011 Schedule A, Part III, line 17 18 % 19 a 33 1/3 % support tests - 2012. If the organization did not check the box on line 14, and line 15 is more than 331/3 %, and line I 17 is not more than 331/3 %, check this box and stop here. The organization qualifies as a publicly supported organization b 33 1/3 % support tests - 2011. If the organization did not check a box on line 14 or line 19a, and line 16 is more than 331/3 %, and I line 18 is not more than 331/3 %, check this box and stop here. The organization qualifies as a publicly supported organization I 20 Private foundation. If the organization did not check a box on line 14, 19a, or 19b, check this box and see instructions JSA Schedule A (Form 990 or 990-EZ) 2012 2E1221 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 15 ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Schedule A (Form 990 or 990-EZ) 2012 Page 4 Part IV Supplemental Information. Complete this part to provide the explanations required by Part II, line 10; Part II, line 17a or 17b; and Part III, line 12. Also complete this part for any additional information. (See instructions). ATTACHMENT 1 SCHEDULE A, PART II - OTHER INCOME

DESCRIPTION 2008 2009 2010 2011 2012 TOTAL

OTHER INCOME 1,791. 1,791.

TOTALS 1,791. 1,791.

JSA Schedule A (Form 990 or 990-EZ) 2012

2E1225 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 16 Schedule B Schedule of Contributors OMB No. 1545-0047 (Form 990, 990-EZ, I or 990-PF) Attach to Form 990, Form 990-EZ, or Form 990-PF. À¾µ¶ Department of the Treasury Internal Revenue Service Name of the organization Employer identification number ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Organization type (check one):

Filers of: Section:

Form 990 or 990-EZ X 501(c)( 3 ) (enter number) organization

4947(a)(1) nonexempt charitable trust not treated as a private foundation

527 political organization

Form 990-PF 501(c)(3) exempt private foundation

4947(a)(1) nonexempt charitable trust treated as a private foundation

501(c)(3) taxable private foundation

Check if your organization is covered by the General Rule or a Special Rule. Note. Only a section 501(c)(7), (8), or (10) organization can check boxes for both the General Rule and a Special Rule. See instructions.

General Rule

X For an organization filing Form 990, 990-EZ, or 990-PF that received, during the year, $5,000 or more (in money or property) from any one contributor. Complete Parts I and II.

Special Rules

For a section 501(c)(3) organization filing Form 990 or 990-EZ that met the 33 1/3 % support test of the regulations under sections 509(a)(1) and 170(b)(1)(A)(vi) and received from any one contributor, during the year, a contribution of the greater of (1) $5,000 or (2) 2% of the amount on (i) Form 990, Part VIII, line 1h, or (ii) Form 990-EZ, line 1. Complete Parts I and II.

For a section 501(c)(7), (8), or (10) organization filing Form 990 or 990-EZ that received from any one contributor, during the year, total contributions of more than $1,000 for use exclusively for religious, charitable, scientific, literary, or educational purposes, or the prevention of cruelty to children or animals. Complete Parts I, II, and III.

For a section 501(c)(7), (8), or (10) organization filing Form 990 or 990-EZ that received from any one contributor, during the year, contributions for use exclusively for religious, charitable, etc., purposes, but these contributions did not total to more than $1,000. If this box is checked, enter here the total contributions that were received during the year for an exclusively religious, charitable, etc., purpose. Do not complete any of the parts unless the General Rule applies to this organization because it received nonexclusively religious, charitable, etc., contributions of $5,000 or more during the year m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m I $

Caution. An organization that is not covered by the General Rule and/or the Special Rules does not file Schedule B (Form 990, 990-EZ, or 990-PF), but it must answer "No" on Part IV, line 2 of its Form 990; or check the box on line H of its Form 990-EZ or on Part I, line 2 of its Form 990-PF, to certify that it does not meet the filing requirements of Schedule B (Form 990, 990-EZ, or 990-PF).

For Paperwork Reduction Act Notice, see the Instructions for Form 990, 990-EZ, or 990-PF. Schedule B (Form 990, 990-EZ, or 990-PF) (2012)

JSA

2E1251 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 17 Schedule B (Form 990, 990-EZ, or 990-PF) (2012) Page 2 Name of organization ELIFE SCIENCES PUBLICATIONS, LTD. Employer identification number 45-3588477 Part I Contributors (see instructions). Use duplicate copies of Part I if additional space is needed.

(a) (b) (c) (d) No. Name, address, and ZIP + 4 Total contributions Type of contribution

1 HOWARD HUGHES MEDICAL INSTITUTE Person X Payroll 4000 JONES BRIDGE ROAD $ 2,320,010. Noncash

CHEVY CHASE, MD 20815 (Complete Part II if there is a noncash contribution.)

(a) (b) (c) (d) No. Name, address, and ZIP + 4 Total contributions Type of contribution

2 WELLCOME TRUST Person X Payroll 215 EUSTON ROAD NW1 2BE $ 2,320,010. Noncash

LONDON (Complete Part II if there is a noncash contribution.) UNITED KINGDOM (a) (b) (c) (d) No. Name, address, and ZIP + 4 Total contributions Type of contribution

3 MAX PLANCK SOCIETY Person X Payroll HOFGARTENSTR 8 80593 $ 500,000. Noncash

MUNICH (Complete Part II if there is a noncash contribution.) GERMANY (a) (b) (c) (d) No. Name, address, and ZIP + 4 Total contributions Type of contribution

Person Payroll $ Noncash (Complete Part II if there is a noncash contribution.)

(a) (b) (c) (d) No. Name, address, and ZIP + 4 Total contributions Type of contribution

Person Payroll $ Noncash (Complete Part II if there is a noncash contribution.)

(a) (b) (c) (d) No. Name, address, and ZIP + 4 Total contributions Type of contribution

Person Payroll $ Noncash (Complete Part II if there is a noncash contribution.)

JSA Schedule B (Form 990, 990-EZ, or 990-PF) (2012)

2E1253 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 18 Schedule B (Form 990, 990-EZ, or 990-PF) (2012) Page 3 Name of organization ELIFE SCIENCES PUBLICATIONS, LTD. Employer identification number 45-3588477 Part II Noncash Property (see instructions). Use duplicate copies of Part II if additional space is needed.

(a) No. (c) (b) (d) from FMV (or estimate) Description of noncash property given Date received Part I (see instructions)

$

(a) No. (c) (b) (d) from FMV (or estimate) Description of noncash property given Date received Part I (see instructions)

$

(a) No. (c) (b) (d) from FMV (or estimate) Description of noncash property given Date received Part I (see instructions)

$

(a) No. (c) (b) (d) from FMV (or estimate) Description of noncash property given Date received Part I (see instructions)

$

(a) No. (c) (b) (d) from FMV (or estimate) Description of noncash property given Date received Part I (see instructions)

$

(a) No. (c) (b) (d) from FMV (or estimate) Description of noncash property given Date received Part I (see instructions)

$

JSA Schedule B (Form 990, 990-EZ, or 990-PF) (2012)

2E1254 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 19 Schedule B (Form 990, 990-EZ, or 990-PF) (2012) Page 4 Name of organization ELIFE SCIENCES PUBLICATIONS, LTD. Employer identification number 45-3588477 Part III Exclusively religious, charitable, etc., individual contributions to section 501(c)(7), (8), or (10) organizations that total more than $1,000 for the year. Complete columns (a) through (e) and the following line entry. For organizations completing Part III, enter the total of exclusively religious, charitable, etc., contributions of $1,000 or less for the year. (Enter this information once. See instructions.) I $ Use duplicate copies of Part III if additional space is needed. (a) No. from (b) Purpose of gift (c) Use of gift (d) Description of how gift is held Part I

(e) Transfer of gift

Transferee's name, address, and ZIP + 4 Relationship of transferor to transferee

(a) No. from (b) Purpose of gift (c) Use of gift (d) Description of how gift is held Part I

(e) Transfer of gift

Transferee's name, address, and ZIP + 4 Relationship of transferor to transferee

(a) No. from (b) Purpose of gift (c) Use of gift (d) Description of how gift is held Part I

(e) Transfer of gift

Transferee's name, address, and ZIP + 4 Relationship of transferor to transferee

(a) No. from (b) Purpose of gift (c) Use of gift (d) Description of how gift is held Part I

(e) Transfer of gift

Transferee's name, address, and ZIP + 4 Relationship of transferor to transferee

JSA Schedule B (Form 990, 990-EZ, or 990-PF) (2012)

2E1255 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 20 OMB No. 1545-0047 SCHEDULE D Supplemental Financial Statements (Form 990) À¾µ¶ I Complete if the organization answered "Yes," to Form 990, Part IV, line 6, 7, 8, 9, 10, 11a, 11b, 11c, 11d, 11e, 11f, 12a, or 12b. Open to Public Department of the Treasury I I Internal Revenue Service Attach to Form 990. See separate instructions. Inspection Name of the organization Employer identification number ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Part I Organizations Maintaining Donor Advised Funds or Other Similar Funds or Accounts. Complete if the organization answered "Yes" to Form 990, Part IV, line 6. m m m m m m m m m m m (a) Donor advised funds (b) Funds and other accounts 1 Total number at end of year m m m m 2 Aggregate contributions to (during yem amr)m m m m m 3 Aggregate grants from (during ym emarm) m m m m m m m 4 Aggregate value at end of year 5 Did the organization inform all donors and donor advisors in writing that the assets held inm mdom nmorm am dmvism emdm m funds are the organization’s property, subject to the organization's exclusive legal control? Yes No 6 Did the organization inform all grantees, donors, and donor advisors in writing that grant funds can be used only for charitable purposes and not for thme mbmenm emfitm omf tmhem mdom nmorm om r mdom nmorm am dmvimsom r,m omr fm omr am nmy motmhem rm pmurmpom smem m conferring impermissible private benefit? Yes No Part II Conservation Easements. Complete if the organization answered "Yes" to Form 990, Part IV, line 7. 1 Purpose(s) of conservation easements held by the organization (check all that apply). Preservation of land for public use (e.g., recreation or education) Preservation of an historically important land area Protection of natural habitat Preservation of a certified historic structure Preservation of open space 2 Complete lines 2a through 2d if the organization held a qualified conservation contribution in the form of a conservation easement on the last day of the tax year. m m m m m m m m m m m m m m m m m m m m m m m m m m m m Held at the End of the Tax Year a Total number of conservation easements m m m m m m m m m m m m m m m m m m m m m m 2a b Total acreage restricted by conservation easements m m m m m m 2b c Number of conservation easements on a certified historic structure included in (a) 2c d Number of conservation easements included inm (cm ) macm qmuimrem dm amftem rm 8m/1m7/m06m , m amndm nm omt om nm am m m historic structure listed in the National Register 2d 3 Number of conservation easements modified, transferred, released, extinguished, or terminated by the organization during the tax year I 4 Number of states where property subject to conservation easement is located I 5 Does the organization have a written policy regarding the periodic mom nmitom rimngm , minmspm emctmiomn,m hmanm dmlinm gm omf m m m m m violations, and enforcement of the conservation easements it holds? Yes No 6 SItaff and volunteer hours devoted to monitoring, inspecting, and enforcing conservation easements during the year 7 AImount of expenses incurred in monitoring, inspecting, and enforcing conservation easements during the year $ 8 Does each conservation easemm emntm rmepm omrtmedm om nm limnem 2m (dm )m ambom vme msam timsfmy mthme mremqum irmem em nmtsm omf sm emctimonm 1m 7m0(mh)m(4m )(mB)m (i) and section 170(h)(4)(B)(ii)? Yes No 9 In Part XIII, describe how the organization reports conservation easements in its revenue and expense statement, and balance sheet, and include, if applicable, the text of the footnote to the organization’s financial statements that describes the organization’s accounting for conservation easements. Part III Organizations Maintaining Collections of Art, Historical Treasures, or Other Similar Assets. Complete if the organization answered "Yes" to Form 990, Part IV, line 8. 1a If the organization elected, as permitted under SFAS 116 (ASC 958), not to report in its revenue statement and balance sheet works of art, historical treasures, or other similar assets held for public exhibition, education, or research in furtherance of public service, provide, in Part XIII, the text of the footnote to its financial statements that describes these items. b If the organization elected, as permitted under SFAS 116 (ASC 958), to report in its revenue statement and balance sheet works of art, historical treasures, or other similar assets held for public exhibition, education, or research in furtherance of public service, provide the following amounts relatingm tom mthmesm em itmemm sm: m m m m m m m m m m m m m m m m m m m m I (i) Revenues included in Form 990, Part Vm ImII, mlinm em 1m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m $ (ii) Assets included in Form 990, Part X I $ 2 If the organization received or held works of art, historical treasures, or other similar assets for financial gain, provide the following amounts required to be reported under mSFm AmSm 1m16m (m AmSCm m95m 8m) mremlamtinm gm tom tmhmesme mitem mm sm: m m m m m m I a Revenues included in Form 990, Part Vm ImII, mlinm em 1m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m $ b Assets included in Form 990, Part X I $ For Paperwork Reduction Act Notice, see the Instructions for Form 990. Schedule D (Form 990) 2012 JSA 2E1268 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 21 ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Schedule D (Form 990) 2012 Page 2 Part III Organizations Maintaining Collections of Art, Historical Treasures, or Other Similar Assets (continued)

3 Using the organization's acquisition, accession, and other records, check any of the following that are a significant use of its collection items (check all that apply): a Public exhibition d Loan or exchange programs b Scholarly research e Other c Preservation for future generations 4 Provide a description of the organization's collections and explain how they further the organization's exempt purpose in Part XIII. 5 During the year, did the organization solicit or receive donations of art, historical treasures, or other smimm ilam rm m m assets to be sold to raise funds rather than to be maintained as part of the organization's collection? Yes No Part IV Escrow and Custodial Arrangements. Complete if the organization answered "Yes" to Form 990, Part IV, line 9, or reported an amount on Form 990, Part X, line 21. 1a Is the organization an agent, trum smteme,m cum smtomdimanm om rm omthmerm inm tem rm medm iam rym fmorm cm omntmribm umtiom nms morm om thm emr am smsem tsm nm omt m m m included on Form 990, Part X? Yes No b If "Yes," explain the arrangement in Part XIII and complete the following table: m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m Amount c Beginning balance m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m 1c d Additions during the year m m m m m m m m m m m m m m m m m m m m m m m m m m m m m 1d e Distributions durimngm tmhem myem amr m m m m m m m m m m m m m m m m m m m m m m m m m m m m m 1e f Ending balance m m m m m m m 1mf m m m m m m m m m m m m m m 2a Did the organization include an amount on Form 990, Part X, line 21? m m m m m m m Ym em s No b If "Yes," explain the arrangement in Part XIII. Check here if the explanation has been provided in Part XIII Part V Endowment Funds. Complete if the organization answered "Yes" to Form 990, Part IV, line 10. m m m m (a) Current year (b) Prior year (c) Two years back (d) Three years back (e) Four years back 1a Beginning of yem amr mbamlamncm em m m m m b Contributions c Net investmm emntm em amrnminmgsm, mgaminms,m m and losses m m m m m m d Grants or scholarships e Other expenditmurmesm fmorm fmacm ilmitiem sm m and programs m m m m m f Administrative expenm smesm m m m m m g End of year balance 2 Provide the estimated percentage of the current year end balance (line 1g, column (a)) held as: a Board designated or quasi-endowment I % b Permanent endowment I % c Temporarily restricted endowment I % The percentages in lines 2a, 2b, and 2c should equal 100%. 3a Are there endowment funds not in the possession of the organization that are held and administered for the organization by: m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m Yes No (i) unrelated organizationms m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m 3a(i) (ii) related organizations m m m m m m m m m m m m m m m m m m 3a(ii) b If "Yes" to 3a(ii), are the related organizations listed as required on Schedule R? 3b 4 Describe in Part XIII the intended uses of the organization's endowment funds. Part VI Land, Buildings, and Equipment. See Form 990, Part X, line 10. Description of property (a) Cost or other basis (b) Cost or other basis (c) Accumulated (d) Book value m m m m m m m m m m m m m m m m m m m m m (investment) (other) depreciation 1a Land m m m m m m m m m m m m m m m m m m b Buildings m m m m m m m m m m c Leasehold imm pmromvem mm emntms m m m m m m m m m m d Equipmmenm tm m m m m m m m m m m m m m m m m m 35,881. 7,274. 28,607. e Other m m m m m m Total. Add lines 1a through 1e. (Column (d) must equal Form 990, Part X, column (B), line 10(c).) I 28,607. Schedule D (Form 990) 2012

JSA 2E1269 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 22 ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Schedule D (Form 990) 2012 Page 3 Part VII Investments - Other Securities. See Form 990, Part X, line 12. (a) Description of security or category (b) Book value (c) Method of valuation: (including nm ammme mof msem cum rimty)m m m m m m m m m Cost or end-of-year market value (1) Financial derivatives m m m m m m m m m m m m m (2) Closely-held equity interests (3) Other (A) (B) (C) (D) (E) (F) (G) (H) (I) I Total. (Column (b) must equal Form 990, Part X, col. (B) line 12.) Part VIII Investments - Program Related. See Form 990, Part X, line 13. (a) Description of investment type (b) Book value (c) Method of valuation: Cost or end-of-year market value (1) (2) (3) (4) (5) (6) (7) (8) (9) (10) I Total. (Column (b) must equal Form 990, Part X, col. (B) line 13.) Part IX Other Assets. See Form 990, Part X, line 15. (a) Description (b) Book value (1) OTHER CURRENT ASSETS 325,061. (2) (3) (4) (5) (6) (7) (8) (9) (10) m m m m m m m m m m m m m m m m m m m m m m m m m m Total. (Column (b) must equal Form 990, Part X, col. (B) line 15.) I 325,061. Part X Other Liabilities. See Form 990, Part X, line 25. 1. (a) Description of liability (b) Book value (1) Federal income taxes (2) OTHER LIABILITIES 24,260. (3) (4) (5) (6) (7) (8) (9) (10) (11) Total. (Column (b) must equal Form 990, Part X, col. (B) line 25.) I 24,260. 2. FIN 48 (ASC 740) Footnote. In Part XIII, provide the text of the footnote to the organization's financial statements thatm rmepm orm tsm tmhem omrgmanm izm ation's liability for uncertain tax positions under FIN 48 (ASC 740). Check here if the text of the footnote has been provided in Part XIII JSA 2E1270 1.000 Schedule D (Form 990) 2012 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 23 ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Schedule D (Form 990) 2012 Page 4 Part XI Reconciliation of Revenue per Audited Financial Statemenmtsm Wm mithm Rm em vm emnum em pm emr mRmetmumrn 1 Total revenue, gains, and other support per audited financial statements 1 5,141,811. 2 Amounts included on line 1 but not on mFom rm m 9m90m , mPmarmt mVImII,m limnem 1m 2m: m m m m m m a Net unrealized gains on investments m m m m m m m m m m m m m m m m m m m m m m 2a b Donated services and use of famcilmitiem sm m m m m m m m m m m m m m m m m m m m m m m 2b c Recoveries of prior year grantms m m m m m m m m m m m m m m m m m m m m m m m m m m 2c d Other (Describe in Part XIImI.)m m m m m m m m m m m m m m m m m m m m m m m m m m m m m 2m dm m m m m m m m m m m m e Add lines 2a through 2d m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m 2e 3 Subtract line 2e from line 1 3 5,141,811. 4 Amounts included on Form 990, Part VIII, line 12, but not on line 1m :m m m m m m a Investment expenses not inclum dm emd monm Fm omrmm m99m 0m, mPam rtm Vm IImI, mlinme m7bm m m m m m m m 4a b Other (Describe in Parmt Xm IImI.)m m m m m m m m m m m m m m m m m m m m m m m m m m m m m 4m bm m m m m m m m m m m m c Add lines 4a and 4b m m m m m m m m m m m m m m 4c 5 Total revenue. Add lines 3 and 4c. (This must equal Form 990, Part I, line 12.) 5 5,141,811. Part XII Reconciliation of Expenses per Audited Financial Statements With Expenses per Return 1 Total expenses and losses per audited financial statements m m m m m m m m m m m m m m m m m m m m m m m m 1 3,565,847. 2 Amounts included on line 1 but not on mFom rm m 9m90m , mPmarmt mIXm, lminme m25m :m m m m m m m a Donated services and umsem om f mfamcilmitiem sm m m m m m m m m m m m m m m m m m m m m m m 2a b Prior year admjusm tm em nmtsm m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m 2b c Other losses m m m m m m m m m m m m m m m m m m m m m m m m m m m 2c d Other (Describe in Part XIImI.)m m m m m m m m m m m m m m m m m m m m m m m m m m m m m 2m dm m m m m m m m m m m m e Add lines 2a through 2d m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m m 2e 3 Subtract line 2e from line 1 3 3,565,847. 4 Amounts included on Form 990, Part IX, line 25, but not on line 1:m m m m m m m a Investment expenses not inclum dm emd monm Fm omrmm m99m 0m, mPam rtm Vm IImI, mlinme m7bm m m m m m m m 4a b Other (Describe in Parmt Xm IImI.)m m m m m m m m m m m m m m m m m m m m m m m m m m m m m 4m bm m m m m m m m m m m m c Add lines 4a and 4b m m m m m m m m m m m m m m 4c 5 Total expenses. Add lines 3 and 4c. (This must equal Form 990, Part I, line 18.) 5 3,565,847. Part XIII Supplemental Information Complete this part to provide the descriptions required for Part II, lines 3, 5, and 9; Part III, lines 1a and 4; Part IV, lines 1b and 2b; Part V, line 4; Part X, line 2; Part XI, lines 2d and 4b; and Part XII, lines 2d and 4b. Also complete this part to provide any additional information. ASC 740 FOOTNOTE

THE ORGANIZATION'S FINANCIAL STATEMENTS DO NOT INCLUDE ANY LIABILITY FOR

UNCERTAIN TAX POSITIONS.

Schedule D (Form 990) 2012

JSA 2E1271 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 24 Schedule D (Form 990) 2012 ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Page 5 Part XIII Supplemental Information (continued)

Schedule D (Form 990) 2012

JSA

2E1226 2.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 25 SCHEDULE F Statement of Activities Outside the United States OMB No. 1545-0047 (Form 990) I Complete if the organization answered "Yes" to Form 990, À¾µ¶ Part IV, line 14b, 15, or 16. I I Open to Public Department of the Treasury Attach to Form 990. See separate instructions. Internal Revenue Service Inspection Name of the organization Employer identification number ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Part I General Information on Activities Outside the United States. Complete if the organization answered "Yes" to Form 990, Part IV, line 14b. 1 For grantmakers. Does the organization maintain records to substantiate the amount of its grants and other assistance, the granteem sm' em limgimbimlitmy mfomr tm hme mgrmanm tsm om rm amssm ismtamncm em, am nmd mthmem smelmecm timonm cm rimtemriam um smedm tm om amwmarmd mthme grants or assistance? Yes No

2 For grantmakers. Describe in Part V the organization's procedures for monitoring the use of its grants and other assistance outside the United States.

3 Activities per Region. (The following Part I, line 3 table can be duplicated if additional space is needed.) (a) Region (b) Number of (c) Number of (d) Activities conducted in (e) If activity listed in (d) is (f) Total offices in the employees, region (by type) (e.g., a program service, expenditures for region agents, and fundraising, program services, describe specific type of and investments independent investments, service(s) in region in region contractors grants to recipients in region located in the region)

(1) EUROPE 1. 10. PROGRAM SERVICES ONLINE JOURNAL 2,726,966.

(2)

(3)

(4)

(5)

(6)

(7)

(8)

(9)

(10)

(11)

(12)

(13)

(14)

(15)

(16) (17) m m m m m m m m m m m 3a Sub-total 1. 10. 2,726,966. b Total from mcom nmtinm umatmiomn sheets to Part I c Totals (add lines 3a and 3b) 1. 10. 2,726,966. For Paperwork Reduction Act Notice, see the Instructions for Form 990. Schedule F (Form 990) 2012 JSA 2E1274 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 26 ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Schedule F (Form 990) 2012 Page 2 Part II Grants and Other Assistance to Organizations or Entities Outside the United States. Complete if the organization answered "Yes" to Form 990, Part IV, line 15, for any recipient who received more than $5,000. Part II can be duplicated if additional space is needed. (i) Method of 1 (a) Name of (b) IRS code (c) Region (d) Purpose of (e) Amount of (f) Manner of (g) Amount of (h) Description valuation cash non-cash of non-cash organization section and EIN grant cash grant (book, FMV, (if applicable) disbursement assistance assistance appraisal, other)

(1)

(2)

(3)

(4)

(5)

(6)

(7)

(8)

(9)

(10)

(11)

(12)

(13)

(14)

(15)

(16) 2 Enter total number of recipient organizations listed above that are recognized as charities by the fmormeimgnm cm omunm trm ym, rmecm omgnm izm emd masm tmaxm -em xmemIpt by the IRS, or for which the grantee or counsel has pmromvimdem dm am smecm timonm 5m 0m1(m cm)(3m )m emqum ivmalmenm cmy mlemttem rm m m m m m m m m m m m m m m m m m m m m 3 Enter total number of other organizations or entities I Schedule F (Form 990) 2012

JSA

2E1275 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 27 ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Schedule F (Form 990) 2012 Page 3 Part III Grants and Other Assistance to Individuals Outside the United States. Complete if the organization answered "Yes" to Form 990, Part IV, line 16. Part III can be duplicated if additional space is needed. (e) Manner of (f) Amount of (g) Description (h) Method of (a) Type of grant or assistance (b) Region (c) Number of (d) Amount of cash non-cash of non-cash valuation recipients cash grant disbursement assistance assistance (book, FMV, appraisal, other)

(1)

(2)

(3)

(4)

(5)

(6)

(7)

(8)

(9)

(10)

(11)

(12)

(13)

(14)

(15)

(16)

(17)

(18) Schedule F (Form 990) 2012

JSA

2E1276 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 28 ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Schedule F (Form 990) 2012 Page 4 Part IV Foreign Forms

1 Was the organization a U.S. transferor of property to a foreign corporation during the tax year? If "Yes," the organization may be required to file Fomrmm 9m 26m , m Rmetmurnm mbym am Um.Sm. Tm ram nmsfem rom r mofm Pmrompem rtmy tm o m a m Fmorem igm n Corporation (see Instructions for Form 926) Yes X No

2 Did the organization have an interest in a foreign trust during the tax year? If "Yes," the organization may be required to file Form 3520, Annual Return to Report Transactions with Foreign Trusts and Receipt of Certain Foreign Gifts, and/or Form 3520-A, Annm umal mInmfomrmm amtiomn mRem tum rnm omf Fm omremignm Tm rum smt Wm imth m a U.S. Owner (see Instructions for Forms 3520 and 3520-A) Yes X No

3 Did the organization have an ownership interest in a foreign corporation during the tax year? If "Yes," the organization may be required to file Form 5471, Informatiom nm Rmetmurmn mof m Um.Sm. Pm ermsom nsm Wm imthm Rmesm pem cmt Tm o Certain Foreign Corporations. (see Instructions for Form 5471) Yes X No

4 Was the organization a direct or indirect shareholder of a passive foreign investment company or a qualified electing fund during the tax year? If "Yes," the organization may be required to file Form 8621, Information Return by a Shareholder om f m a m Pmasm simvem Fm ormeimgnm Imnvmesm tmm emntm Cm om pm amnym om r mQmuam lifmiedm mElmecmtinm g Fund. (see Instructions for Form 8621) Yes X No

5 Did the organization have an ownership interest in a foreign partnership during the tax year? If "Yes," the organization may be required to file Form 8865,m Rm emtumrn m omf Um .Sm . m Pmersm omnsm Wm imth m Rmesmpem cmt Tm om Cmermtamin Foreign Partnerships. (see Instructions for Form 8865) Yes X No

6 Did the organization have any operations in or related to any boycotting countries during the tax year? If "Yes," the organimzamtiomn mmmaym bme mremqum irem dm tom fmilem Fm orm mm 5m71m 3m, Imntmermnam tiom nmal m Bmoymcom ttm Rmepm ormt m(sem e m Inmstmrumctmionm s for Form 5713) Yes X No

Schedule F (Form 990) 2012

JSA

2E1277 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 29 ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477 Schedule F (Form 990) 2012 Page 5 Part V Supplemental Information Complete this part to provide the information required by Part I, line 2 (monitoring of funds); Part I, line 3, column (f) (accounting method; amounts of investments vs. expenditures per region); Part II, line 1 (accounting method); Part III (accounting method); and Part III, column (c) (estimated number of recipients), as applicable. Also complete this part to provide any additional information (see instructions).

JSA Schedule F (Form 990) 2012

2E1502 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 30 OMB No. 1545-0047 SCHEDULE O Supplemental Information to Form 990 or 990-EZ (Form 990 or 990-EZ) À¾µ¶ Complete to provide information for responses to specific questions on Form 990 or 990-EZ or to provide any additional information. Open to Public Department of the Treasury I Internal Revenue Service Attach to Form 990 or 990-EZ. Inspection Name of the organization Employer identification number ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477

SCHEDULE O SUPPLEMENTAL DISCLOSURES

PART V, LINE 2A

THERE ARE 11 EMPLOYEES IN TOTAL, 10 OF WHOM ARE LOCATED IN THE UNITED

KINGDOM. THE SOLE U.S. EMPLOYEE WAS REPORTED ON THE FORM W-3.

PART VI, LINES 6 AND 7

THE MEMBERS OF ELIFE ARE HOWARD HUGHES MEDICAL INSTITUTE AND WELLCOME

TRUST. EACH MEMBER HAS THE POWER TO APPOINT ONE MEMBER OF THE BOARD OF

DIRECTORS OF ELIFE. CERTAIN DECISIONS REQUIRE THE UNANIMOUS CONSENT OF

THE BOARD MEMBERS.

PART VI, LINE 11B

FORM 990 WAS PREPARED BY ELIFE'S INDEPENDENT ACCOUNTING FIRM AND WAS

REVIEWED BY SENIOR MANAGEMENT AND THE AUDIT COMMITTEE. PRIOR TO FILING

THE FORM 990 WITH THE IRS, A COPY OF FORM 990 WAS PROVIDED TO EACH BOARD

MEMBER.

PART VI, LINE 12C

ELIFE HAS A CONFLICT OF INTEREST POLICY THAT REQUIRES DIRECTORS, OFFICERS

AND OTHER EMPLOYEES TO DISCLOSE INTERESTS THAT MAY GIVE RISE TO A

For Privacy Act and Paperwork Reduction Act Notice, see the Instructions for Form 990 or 990-EZ. Schedule O (Form 990 or 990-EZ) (2012) JSA 2E1227 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 31 Schedule O (Form 990 or 990-EZ) 2012 Page 2 Name of the organization Employer identification number ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477

CONFLICT. ELIFE MONITORS THE POLICY BY REQUIRING INDIVIDUALS TO COMPLETE

AN ANNUAL QUESTIONNAIRE THAT REQUIRES DISCLOSURE OF ANY POTENTIAL

CONFLICTS OF INTEREST.

PART VI, LINE 15

COMPENSATION DECISIONS ARE MADE BY INDIVIDUALS THAT ARE INDEPENDENT WITH

RESPECT TO THE INDIVIDUAL RECEIVING THE COMPENSATION. EFFECTIVE WITH

COMPENSATION PAID IN 2013, THE BOARD WILL OBTAIN AND RELY ON

COMPARABILITY DATA IN ESTABLISHING THE COMPENSATION FOR ITS OFFICERS.

COMPENSATION WILL BE APPROVED IN ADVANCE OF PAYMENT, AND THE DECISION

WILL BE DOCUMENTED IN THE BOARD MINUTES.

PART VI, LINE 19

ELIFE DOES NOT MAKE ITS GOVERNING DOCUMENTS AND CONFLICT OF INTEREST

POLICY AVAILABLE TO THE GENERAL PUBLIC UNLESS THERE IS A LEGAL OBLIGATION

TO DO SO.

OUR AUDITED FINANCIAL STATEMENTS WILL BE AVAILABLE FROM THE UK COMPANIES

REGISTRY.

PART XI, LINE 5

FOREIGN CURRENCY TRANSLATION ADJUSTMENT $7,401

JSA Schedule O (Form 990 or 990-EZ) 2012

2E1228 1.000 8195FH 2502 6/21/2013 1:09:46 PM V 12-5F 2885408 PAGE 32 Schedule O (Form 990 or 990-EZ) 2012 Page 2 Name of the organization Employer identification number ELIFE SCIENCES PUBLICATIONS, LTD. 45-3588477

ATTACHMENT 1

FORM 990, PART III - PROGRAM SERVICE, LINE 4A

ELIFE WAS FOUNDED IN RESPONSE TO AN INITIATIVE TO DRIVE

IMPROVEMENTS IN RESEARCH COMMUNICATION FROM THREE INTERNATIONALLY

PROMINENT, NONPROFIT ORGANISATIONS OPERATING IN THE PUBLIC

INTEREST: HOWARD HUGHES MEDICAL INSTITUTE, MAX PLANCK SOCIETY FOR

THE ADVANCEMENT OF SCIENCE, AND WELLCOME TRUST.

ELIFE OPERATES AN ONLINE, OPEN-ACCESS JOURNAL FOR THE MOST

IMPORTANT ADVANCES IN LIFE SCIENCE AND BIOMEDICINE. THE EDITORIAL

CONTENT OF ELIFE IS THE RESPONSIBILITY OF AN INDEPENDENT EDITORIAL

TEAM COMPRISING ACTIVE, PRACTICING SCIENTISTS. THE ELIFE JOURNAL

IS ALSO A PLATFORM FOR EXTENDING THE REACH AND INFLUENCE OF NEW

DISCOVERIES, AND TO SHOWCASE NEW APPROACHES TO THE PRESENTATION,

USE AND ASSESSMENT OF RESEARCH.

JSA Schedule O (Form 990 or 990-EZ) 2012

2E1228 1.000 8195FH 2502 6/17/2013 3:01:25 PM V 12-5F 2885408 PAGE 33