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The Skull of the Upper Cretaceous Snake Dinilysia Patagonica Smith-Woodward, 1901, and Its Phylogenetic Position Revisited
Zoological Journal of the Linnean Society, 2012, 164, 194–238. With 24 figures The skull of the Upper Cretaceous snake Dinilysia patagonica Smith-Woodward, 1901, and its phylogenetic position revisited HUSSAM ZAHER1* and CARLOS AGUSTÍN SCANFERLA2 1Museu de Zoologia da Universidade de São Paulo, Avenida Nazaré 481, Ipiranga, 04263-000, São Paulo, SP, Brasil 2Laboratorio de Anatomía Comparada y Evolución de los Vertebrados. Museo Argentino de Ciencias Naturales ‘Bernardino Rivadavia’, Av. Angel Gallardo 470 (1405), Buenos Aires, Argentina Received 23 April 2010; revised 5 April 2011; accepted for publication 18 April 2011 The cranial anatomy of Dinilysia patagonica, a terrestrial snake from the Upper Cretaceous of Argentina, is redescribed and illustrated, based on high-resolution X-ray computed tomography and better preparations made on previously known specimens, including the holotype. Previously unreported characters reinforce the intriguing mosaic nature of the skull of Dinilysia, with a suite of plesiomorphic and apomorphic characters with respect to extant snakes. Newly recognized plesiomorphies are the absence of the medial vertical flange of the nasal, lateral position of the prefrontal, lizard-like contact between vomer and palatine, floor of the recessus scalae tympani formed by the basioccipital, posterolateral corners of the basisphenoid strongly ventrolaterally projected, and absence of a medial parietal pillar separating the telencephalon and mesencephalon, amongst others. We also reinterpreted the structures forming the otic region of Dinilysia, confirming the presence of a crista circumfenes- tralis, which represents an important derived ophidian synapomorphy. Both plesiomorphic and apomorphic traits of Dinilysia are treated in detail and illustrated accordingly. Results of a phylogenetic analysis support a basal position of Dinilysia, as the sister-taxon to all extant snakes. -
Birds, Reptiles, Amphibians, Vascular Plants, and Habitat in the Gila River Riparian Zone in Southwestern New Mexico
Birds, Reptiles, Amphibians, Vascular Plants, and Habitat in the Gila River Riparian Zone in Southwestern New Mexico Kansas Biological Survey Report #151 Kelly Kindscher, Randy Jennings, William Norris, and Roland Shook September 8, 2008 Birds, Reptiles, Amphibians, Vascular Plants, and Habitat in the Gila River Riparian Zone in Southwestern New Mexico Cover Photo: The Gila River in New Mexico. Photo by Kelly Kindscher, September 2006. Kelly Kindscher, Associate Scientist, Kansas Biological Survey, University of Kansas, 2101 Constant Avenue, Lawrence, KS 66047, Email: [email protected] Randy Jennings, Professor, Department of Natural Sciences, Western New Mexico University, PO Box 680, 1000 W. College Ave., Silver City, NM 88062, Email: [email protected] William Norris, Associate Professor, Department of Natural Sciences, Western New Mexico University, PO Box 680, 1000 W. College Ave., Silver City, NM 88062, Email: [email protected] Roland Shook, Emeritus Professor, Biology, Department of Natural Sciences, Western New Mexico University, PO Box 680, 1000 W. College Ave., Silver City, NM 88062, Email: [email protected] Citation: Kindscher, K., R. Jennings, W. Norris, and R. Shook. Birds, Reptiles, Amphibians, Vascular Plants, and Habitat in the Gila River Riparian Zone in Southwestern New Mexico. Open-File Report No. 151. Kansas Biological Survey, Lawrence, KS. ii + 42 pp. Abstract During 2006 and 2007 our research crews collected data on plants, vegetation, birds, reptiles, and amphibians at 49 sites along the Gila River in southwest New Mexico from upstream of the Gila Cliff Dwellings on the Middle and West Forks of the Gila to sites below the town of Red Rock, New Mexico. -
Snakes of the Siwalik Group (Miocene of Pakistan): Systematics and Relationship to Environmental Change
Palaeontologia Electronica http://palaeo-electronica.org SNAKES OF THE SIWALIK GROUP (MIOCENE OF PAKISTAN): SYSTEMATICS AND RELATIONSHIP TO ENVIRONMENTAL CHANGE Jason J. Head ABSTRACT The lower and middle Siwalik Group of the Potwar Plateau, Pakistan (Miocene, approximately 18 to 3.5 Ma) is a continuous fluvial sequence that preserves a dense fossil record of snakes. The record consists of approximately 1,500 vertebrae derived from surface-collection and screen-washing of bulk matrix. This record represents 12 identifiable taxa and morphotypes, including Python sp., Acrochordus dehmi, Ganso- phis potwarensis gen. et sp. nov., Bungarus sp., Chotaophis padhriensis, gen. et sp. nov., and Sivaophis downsi gen. et sp. nov. The record is dominated by Acrochordus dehmi, a fully-aquatic taxon, but diversity increases among terrestrial and semi-aquatic taxa beginning at approximately 10 Ma, roughly coeval with proxy data indicating the inception of the Asian monsoons and increasing seasonality on the Potwar Plateau. Taxonomic differences between the Siwalik Group and coeval European faunas indi- cate that South Asia was a distinct biogeographic theater from Europe by the middle Miocene. Differences between the Siwalik Group and extant snake faunas indicate sig- nificant environmental changes on the Plateau after the last fossil snake occurrences in the Siwalik section. Jason J. Head. Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, P.O. Box 37012, Washington, DC 20013-7012, USA. [email protected] School of Biological Sciences, Queen Mary, University of London, London, E1 4NS, United Kingdom. KEY WORDS: Snakes, faunal change, Siwalik Group, Miocene, Acrochordus. PE Article Number: 8.1.18A Copyright: Society of Vertebrate Paleontology May 2005 Submission: 3 August 2004. -
Language Kinship Between Mandarin, Hokkien Chinese and Japanese (Lexicostatistics Review)
LANGUAGE KINSHIP BETWEEN MANDARIN, HOKKIEN CHINESE AND JAPANESE (LEXICOSTATISTICS REVIEW) KEKERABATAN ANTARA BAHASA MANDARIN, HOKKIEN DAN JEPANG (TINJAUAN LEXICOSTATISTICS) Abdul Gapur1, Dina Shabrina Putri Siregar 2, Mhd. Pujiono3 1,2,3Faculty of Cultural Sciences, University of Sumatera Utara Jalan Universitas, No. 19, Medan, Sumatera Utara, Indonesia Telephone (061) 8215956, Facsimile (061) 8215956 E-mail: [email protected] Article accepted: July 22, 2018; revised: December 18, 2018; approved: December 24, 2018 Permalink/DOI: 10.29255/aksara.v30i2.230.301-318 Abstract Mandarin and Hokkien Chinese are well known having a tight kinship in a language family. Beside, Japanese also has historical relation with China in the field of language and cultural development. Japanese uses Chinese characters named kanji with certain phonemic vocabulary adjustment, which is adapted into Japanese. This phonemic adjustment of kanji is called Kango. This research discusses about the kinship of Mandarin, Hokkien Chinese in Indonesia and Japanese Kango with lexicostatistics review. The method used is quantitative with lexicostatistics technique. Quantitative method finds similar percentage of 100-200 Swadesh vocabularies. Quantitative method with lexicostatistics results in a tree diagram of the language genetics. From the lexicostatistics calculation to the lexicon level, it is found that Mandarin Chinese (MC) and Japanese Kango (JK) are two different languages, because they are in a language group (stock) (29%); (2) JK and Indonesian Hokkien Chinese (IHC) are also two different languages, because they are in a language group (stock) (24%); and (3) MC and IHC belong to the same language family (42%). Keywords: language kinship, Mandarin, Hokkien, Japanese Abstrak Bahasa Mandarin dan Hokkien diketahui memiliki hubungan kekerabatan dalam rumpun yang sama. -
The Sclerotic Ring: Evolutionary Trends in Squamates
The sclerotic ring: Evolutionary trends in squamates by Jade Atkins A Thesis Submitted to Saint Mary’s University, Halifax, Nova Scotia in Partial Fulfillment of the Requirements for the Degree of Master of Science in Applied Science July, 2014, Halifax Nova Scotia © Jade Atkins, 2014 Approved: Dr. Tamara Franz-Odendaal Supervisor Approved: Dr. Matthew Vickaryous External Examiner Approved: Dr. Tim Fedak Supervisory Committee Member Approved: Dr. Ron Russell Supervisory Committee Member Submitted: July 30, 2014 Dedication This thesis is dedicated to my family, friends, and mentors who helped me get to where I am today. Thank you. ! ii Table of Contents Title page ........................................................................................................................ i Dedication ...................................................................................................................... ii List of figures ................................................................................................................. v List of tables ................................................................................................................ vii Abstract .......................................................................................................................... x List of abbreviations and definitions ............................................................................ xi Acknowledgements .................................................................................................... -
Status and Protection of Globally Threatened Species in the Caucasus
STATUS AND PROTECTION OF GLOBALLY THREATENED SPECIES IN THE CAUCASUS CEPF Biodiversity Investments in the Caucasus Hotspot 2004-2009 Edited by Nugzar Zazanashvili and David Mallon Tbilisi 2009 The contents of this book do not necessarily reflect the views or policies of CEPF, WWF, or their sponsoring organizations. Neither the CEPF, WWF nor any other entities thereof, assumes any legal liability or responsibility for the accuracy, completeness, or usefulness of any information, product or process disclosed in this book. Citation: Zazanashvili, N. and Mallon, D. (Editors) 2009. Status and Protection of Globally Threatened Species in the Caucasus. Tbilisi: CEPF, WWF. Contour Ltd., 232 pp. ISBN 978-9941-0-2203-6 Design and printing Contour Ltd. 8, Kargareteli st., 0164 Tbilisi, Georgia December 2009 The Critical Ecosystem Partnership Fund (CEPF) is a joint initiative of l’Agence Française de Développement, Conservation International, the Global Environment Facility, the Government of Japan, the MacArthur Foundation and the World Bank. This book shows the effort of the Caucasus NGOs, experts, scientific institutions and governmental agencies for conserving globally threatened species in the Caucasus: CEPF investments in the region made it possible for the first time to carry out simultaneous assessments of species’ populations at national and regional scales, setting up strategies and developing action plans for their survival, as well as implementation of some urgent conservation measures. Contents Foreword 7 Acknowledgments 8 Introduction CEPF Investment in the Caucasus Hotspot A. W. Tordoff, N. Zazanashvili, M. Bitsadze, K. Manvelyan, E. Askerov, V. Krever, S. Kalem, B. Avcioglu, S. Galstyan and R. Mnatsekanov 9 The Caucasus Hotspot N. -
Controlled Animals
Environment and Sustainable Resource Development Fish and Wildlife Policy Division Controlled Animals Wildlife Regulation, Schedule 5, Part 1-4: Controlled Animals Subject to the Wildlife Act, a person must not be in possession of a wildlife or controlled animal unless authorized by a permit to do so, the animal was lawfully acquired, was lawfully exported from a jurisdiction outside of Alberta and was lawfully imported into Alberta. NOTES: 1 Animals listed in this Schedule, as a general rule, are described in the left hand column by reference to common or descriptive names and in the right hand column by reference to scientific names. But, in the event of any conflict as to the kind of animals that are listed, a scientific name in the right hand column prevails over the corresponding common or descriptive name in the left hand column. 2 Also included in this Schedule is any animal that is the hybrid offspring resulting from the crossing, whether before or after the commencement of this Schedule, of 2 animals at least one of which is or was an animal of a kind that is a controlled animal by virtue of this Schedule. 3 This Schedule excludes all wildlife animals, and therefore if a wildlife animal would, but for this Note, be included in this Schedule, it is hereby excluded from being a controlled animal. Part 1 Mammals (Class Mammalia) 1. AMERICAN OPOSSUMS (Family Didelphidae) Virginia Opossum Didelphis virginiana 2. SHREWS (Family Soricidae) Long-tailed Shrews Genus Sorex Arboreal Brown-toothed Shrew Episoriculus macrurus North American Least Shrew Cryptotis parva Old World Water Shrews Genus Neomys Ussuri White-toothed Shrew Crocidura lasiura Greater White-toothed Shrew Crocidura russula Siberian Shrew Crocidura sibirica Piebald Shrew Diplomesodon pulchellum 3. -
WHO Guidance on Management of Snakebites
GUIDELINES FOR THE MANAGEMENT OF SNAKEBITES 2nd Edition GUIDELINES FOR THE MANAGEMENT OF SNAKEBITES 2nd Edition 1. 2. 3. 4. ISBN 978-92-9022- © World Health Organization 2016 2nd Edition All rights reserved. Requests for publications, or for permission to reproduce or translate WHO publications, whether for sale or for noncommercial distribution, can be obtained from Publishing and Sales, World Health Organization, Regional Office for South-East Asia, Indraprastha Estate, Mahatma Gandhi Marg, New Delhi-110 002, India (fax: +91-11-23370197; e-mail: publications@ searo.who.int). The designations employed and the presentation of the material in this publication do not imply the expression of any opinion whatsoever on the part of the World Health Organization concerning the legal status of any country, territory, city or area or of its authorities, or concerning the delimitation of its frontiers or boundaries. Dotted lines on maps represent approximate border lines for which there may not yet be full agreement. The mention of specific companies or of certain manufacturers’ products does not imply that they are endorsed or recommended by the World Health Organization in preference to others of a similar nature that are not mentioned. Errors and omissions excepted, the names of proprietary products are distinguished by initial capital letters. All reasonable precautions have been taken by the World Health Organization to verify the information contained in this publication. However, the published material is being distributed without warranty of any kind, either expressed or implied. The responsibility for the interpretation and use of the material lies with the reader. In no event shall the World Health Organization be liable for damages arising from its use. -
2013 - Year of the Snake
2013 - YEAR OF THE SNAKE The black water yin snake ! As ominous as the year of the Black Water Snake may sound, having survived the year of the Black Water Dragon and the end of the Mayan calendar(!), surely we can slither through 2013 unscathed?! A Snake year rides on the glory of the Dragon and can bring many great achievements and successes. Of course, it can also be a slippery fellow so be prepared to shed your Ophidiophobia and embrace the New Year! The first cycle of the zodiac started in 2637 B.C., introduced by the Emperor Huang Ti. Every twelfth year is a Snake year and they are the sixth animal in the zodiac cycle. A complete cycle of the Chinese lunar calendar takes 60 years and is made up of five cycles of 12 years, 12 animal signs and the 5 elements, meaning that each animal and element only combine once during each full cycle. The last time we had Water and Snake ensemble was in 1953. Each of the animal signs also has its own fixed element, the Snake’s is Fire. Finally, we throw Yin and Yang into the mix as well and you start to see just how intricate an art Chinese Astrology is! Black is the colour of the Water element and the ancient Chinese regarded it as the king of colours, also said to be the colour of heaven and space. Of course, it also makes up one half of the Tajii symbol characterising the unity of Yin and Yang. In Chinese Astrology, odd years are Yin years and even years are Yang years. -
THE ORIGIN and EVOLUTION of SNAKE EYES Dissertation
CONQUERING THE COLD SHUDDER: THE ORIGIN AND EVOLUTION OF SNAKE EYES Dissertation Presented in Partial Fulfillment for the Requirements for the Degree of Doctor of Philosophy in the Graduate School of The Ohio State University By Christopher L. Caprette, B.S., M.S. **** The Ohio State University 2005 Dissertation Committee: Thomas E. Hetherington, Advisor Approved by Jerry F. Downhower David L. Stetson Advisor The graduate program in Evolution, John W. Wenzel Ecology, and Organismal Biology ABSTRACT I investigated the ecological origin and diversity of snakes by examining one complex structure, the eye. First, using light and transmission electron microscopy, I contrasted the anatomy of the eyes of diurnal northern pine snakes and nocturnal brown treesnakes. While brown treesnakes have eyes of similar size for their snout-vent length as northern pine snakes, their lenses are an average of 27% larger (Mann-Whitney U test, p = 0.042). Based upon the differences in the size and position of the lens relative to the retina in these two species, I estimate that the image projected will be smaller and brighter for brown treesnakes. Northern pine snakes have a simplex, all-cone retina, in keeping with a primarily diurnal animal, while brown treesnake retinas have mostly rods with a few, scattered cones. I found microdroplets in the cone ellipsoids of northern pine snakes. In pine snakes, these droplets act as light guides. I also found microdroplets in brown treesnake rods, although these were less densely distributed and their function is unknown. Based upon the density of photoreceptors and neural layers in their retinas, and the predicted image size, brown treesnakes probably have the same visual acuity under nocturnal conditions that northern pine snakes experience under diurnal conditions. -
CITES Appendices I, II and III Valid from 10 March 2016
CONVENTION ON INTERNATIONAL TRADE IN ENDANGERED SPECIES OF WILD FAUNA AND FLORA Appendices I, II and III valid from 2 January 2017 Interpretation 1. Species included in these Appendices are referred to: a) by the name of the species; or b) as being all of the species included in a higher taxon or designated part thereof. 2. The abbreviation “spp.” is used to denote all species of a higher taxon. 3. Other references to taxa higher than species are for the purposes of information or classification only. The common names included after the scientific names of families are for reference only. They are intended to indicate the species within the family concerned that are included in the Appendices. In most cases this is not all of the species within the family. 4. The following abbreviations are used for plant taxa below the level of species: a) “ssp.” is used to denote subspecies; and b) “var(s).” is used to denote variety (varieties). 5. As none of the species or higher taxa of FLORA included in Appendix I is annotated to the effect that its hybrids shall be treated in accordance with the provisions of Article III of the Convention, this means that artificially propagated hybrids produced from one or more of these species or taxa may be traded with a certificate of artificial propagation, and that seeds and pollen (including pollinia), cut flowers, seedling or tissue cultures obtained in vitro, in solid or liquid media, transported in sterile containers of these hybrids are not subject to the provisions of the Convention. -
Watchable Wildlife Form
U.S. Fish & Wildlife Service Watchable Wildlife Bitter Lake National Wildlife Refuge Welcome Roundnose minnow Dionda episcopa Amphibians Bitter Lake National Wildlife Refuge is one Speckled chub Extrarius aestivalis Family Ambystomatidae – Mole Salamanders of New Mexico’s most important sanctuaries Plains minnow Hybognathus placitus Tiger salamander Ambystoma tigrinum Arkansas River shiner Notropis girardi and breeding grounds for migratory birds and Family Leptodactylidae – Tropical Frogs Rio Grande shiner Notropis jemezanus other wildlife. Established in 1937, the 24,500- Eastern barking frog Eleutherodactylus augusti Pecos bluntnose shiner Notropis simus pecosensis acre refuge is strategically located along the latrans Pecos River where the Chihuahuan Desert Fathead minnow Pimephales promelas Family Pelobatidae – Spadefoot Toads meets the Great Plains. The convergence of Family Catostomidae – Suckers Couch’s spadefoot toad Scaphiopus couchii these vastly different terrains has produced a River carpsucker Carpoides carpio diverse range of habitats, providing a home to New Mexico spadefoot toad Spea multiplicata a rich array of plant and animal life, including Family Ictaluridae – Catfishes Plains spadefoot toad Spea bombifrons Channel catfish Ictalurus punctatus a number of rare species. Family Bufonidae – Toads Bitter Lake NWR is best known for its Family Cyprinodontidae – Pupfish Woodhouse’s toad Bufo woodhousii spectacular variety of birds, particularly the Pecos pupfish Cyprinodon pecosensis Red-spotted toad Bufo punctatus Great Plains toad Bufo cognatus large migrations of ducks, geese, and cranes Family Fundulidae – Killifishes Texas toad Bufo speciosus during the fall and winter months. Many of Plains killifish Fundulus zebrinus Western green toad Bufo debilis insidior these creatures are drawn by the refuge’s Rainwater killifish Lucania parva wetlands, which offer an abundance of food as Family Hylidae – Treefrogs Family Poeciliidae – Livebearers well as ideal nesting habitat for some species.