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The Lower Cretaceous Flora of the Gates Formation from Western Canada
The Lower Cretaceous Flora of the Gates Formation from Western Canada A Shesis Submitted to the College of Graduate Studies and Research in Partial Fulfillment of the Requirements for the Degree of Doctor of Philosophy in the Department of Geological Sciences Univ. of Saska., Saskatoon?SI(, Canada S7N 3E2 b~ Zhihui Wan @ Copyright Zhihui Mian, 1996. Al1 rights reserved. National Library Bibliothèque nationale 1*1 of Canada du Canada Acquisitions and Acquisitions et Bibliographic Services services bibliographiques 395 Wellington Street 395. rue Wellington Ottawa ON KlA ON4 Ottawa ON K1A ON4 Canada Canada The author has granted a non- L'auteur a accordé une licence non exclusive licence allowing the exclusive permettant à la National Libraxy of Canada to Bibliothèque nationale du Canada de reproduce, loan, distribute or sell reproduire, prêter, distribuer ou copies of this thesis in microfom, vendre des copies de cette thèse sous paper or electronic formats. la fome de microfiche/nlm, de reproduction sur papier ou sur foxmat électronique. The author retains ownership of the L'auteur conserve la propriété du copyright in this thesis. Neither the droit d'auteur qui protège cette thèse. thesis nor substantial extracts fiom it Ni la thèse ni des extraits substantiels may be printed or otherwise de celle-ci ne doivent être imprimés reproduced without the author's ou autrement reproduits sans son permission. autorisation. College of Graduate Studies and Research SUMMARY OF DISSERTATION Submitted in partial fulfillment of the requirernents for the DEGREE OF DOCTOR OF PHILOSOPHY ZHIRUI WAN Depart ment of Geological Sciences University of Saskatchewan Examining Commit tee: Dr. -
Jurassic Flora of Cape 1,Isburne Alaska
DEPARTMENT OF THE INTERIOR UNITED STATES GEOLOGICAL SURVEY GEORGE OTIS SMITH, DIRECTOR THE JURASSIC FLORA OF CAPE 1,ISBURNE ALASKA F. H. KNOWLTON Publishecl January 28, 1914 PART D OF PROFESSIONAL PAPER 85, "CONTRIBUTIONS TO GENERAL GEOLOGY 1913" WASHINGTON GOVERNMENT PRINTING OFFICE 1914 CONTENTS. Page . Introduction ............................................................................................ 39 The Corwin formation ..................................................................................... 39 Plant collections ......................................................................................... 40 Age of the plant-bearing beds ............................................................................. 41 Distribution of Jurassic floras ............................................................................. 43 Geographicrange .................................................................................... 43 Means of dispersal ................................................................................... 45 Avenues of dispersal ................................................................................. 45 Probable climatic conditions .......................................................................... 46 .The flora ................................................................................................ 46 ILLUSTRATIONS. Page . PLATESV-VIII . Jurassic flora of Cape Lisburne, Alaska .................................................... 57-64 THE' JUR,ASSIC FLORA OF CAPE -
Fern Classification
16 Fern classification ALAN R. SMITH, KATHLEEN M. PRYER, ERIC SCHUETTPELZ, PETRA KORALL, HARALD SCHNEIDER, AND PAUL G. WOLF 16.1 Introduction and historical summary / Over the past 70 years, many fern classifications, nearly all based on morphology, most explicitly or implicitly phylogenetic, have been proposed. The most complete and commonly used classifications, some intended primar• ily as herbarium (filing) schemes, are summarized in Table 16.1, and include: Christensen (1938), Copeland (1947), Holttum (1947, 1949), Nayar (1970), Bierhorst (1971), Crabbe et al. (1975), Pichi Sermolli (1977), Ching (1978), Tryon and Tryon (1982), Kramer (in Kubitzki, 1990), Hennipman (1996), and Stevenson and Loconte (1996). Other classifications or trees implying relationships, some with a regional focus, include Bower (1926), Ching (1940), Dickason (1946), Wagner (1969), Tagawa and Iwatsuki (1972), Holttum (1973), and Mickel (1974). Tryon (1952) and Pichi Sermolli (1973) reviewed and reproduced many of these and still earlier classifica• tions, and Pichi Sermolli (1970, 1981, 1982, 1986) also summarized information on family names of ferns. Smith (1996) provided a summary and discussion of recent classifications. With the advent of cladistic methods and molecular sequencing techniques, there has been an increased interest in classifications reflecting evolutionary relationships. Phylogenetic studies robustly support a basal dichotomy within vascular plants, separating the lycophytes (less than 1 % of extant vascular plants) from the euphyllophytes (Figure 16.l; Raubeson and Jansen, 1992, Kenrick and Crane, 1997; Pryer et al., 2001a, 2004a, 2004b; Qiu et al., 2006). Living euphyl• lophytes, in turn, comprise two major clades: spermatophytes (seed plants), which are in excess of 260 000 species (Thorne, 2002; Scotland and Wortley, Biology and Evolution of Ferns and Lycopliytes, ed. -
Phytochrome Diversity in Green Plants and the Origin of Canonical Plant Phytochromes
ARTICLE Received 25 Feb 2015 | Accepted 19 Jun 2015 | Published 28 Jul 2015 DOI: 10.1038/ncomms8852 OPEN Phytochrome diversity in green plants and the origin of canonical plant phytochromes Fay-Wei Li1, Michael Melkonian2, Carl J. Rothfels3, Juan Carlos Villarreal4, Dennis W. Stevenson5, Sean W. Graham6, Gane Ka-Shu Wong7,8,9, Kathleen M. Pryer1 & Sarah Mathews10,w Phytochromes are red/far-red photoreceptors that play essential roles in diverse plant morphogenetic and physiological responses to light. Despite their functional significance, phytochrome diversity and evolution across photosynthetic eukaryotes remain poorly understood. Using newly available transcriptomic and genomic data we show that canonical plant phytochromes originated in a common ancestor of streptophytes (charophyte algae and land plants). Phytochromes in charophyte algae are structurally diverse, including canonical and non-canonical forms, whereas in land plants, phytochrome structure is highly conserved. Liverworts, hornworts and Selaginella apparently possess a single phytochrome, whereas independent gene duplications occurred within mosses, lycopods, ferns and seed plants, leading to diverse phytochrome families in these clades. Surprisingly, the phytochrome portions of algal and land plant neochromes, a chimera of phytochrome and phototropin, appear to share a common origin. Our results reveal novel phytochrome clades and establish the basis for understanding phytochrome functional evolution in land plants and their algal relatives. 1 Department of Biology, Duke University, Durham, North Carolina 27708, USA. 2 Botany Department, Cologne Biocenter, University of Cologne, 50674 Cologne, Germany. 3 University Herbarium and Department of Integrative Biology, University of California, Berkeley, California 94720, USA. 4 Royal Botanic Gardens Edinburgh, Edinburgh EH3 5LR, UK. 5 New York Botanical Garden, Bronx, New York 10458, USA. -
The Stele – a Developmental Perspective on the Diversity and Evolution of Primary Vascular 2 Architecture 3 4 Alexandru M.F
Preprints (www.preprints.org) | NOT PEER-REVIEWED | Posted: 31 May 2020 doi:10.20944/preprints202005.0472.v1 1 The stele – a developmental perspective on the diversity and evolution of primary vascular 2 architecture 3 4 Alexandru M.F. Tomescu 5 6 Department of Biological Sciences, Humboldt State University, Arcata, California 95521, USA 7 (+1) 707‐826‐3229 [email protected] 8 9 Abstract 10 The stele concept is one of the oldest enduring concepts in plant biology. This paper reviews the 11 concept and its foundations, and builds an argument for an updated view of steles and their evolution. 12 The history of studies of stelar organization has generated a widely ranging array of definitions of the 13 stele that determine the way we classify steles and construct scenarios about the evolution of stelar 14 architecture. Because at the level of the organism biological evolution proceeds by, and is reflected in, 15 changes in development, concepts of structure need to be grounded in development in order to be 16 relevant in an evolutionary perspective. For the stele, most of the traditional definitions that 17 incorporate development have viewed it as the totality of tissues that either originate from procambium 18 – currently the prevailing view – or are bordered by a boundary layer (e.g., endodermis). A definition of 19 the stele that would bring consensus between these perspectives recasts the stele as a structural entity 20 of dual nature. Here, I review briefly the history of the stele concept, basic terminology related to stelar 21 organization, and traditional classifications of the steles. -
Mesozoic Lycopods and Ferns from the Bureja Basin
Palaeontographica Abt. В 166 Lfg. 1-3 16—29 Stuttgart, August, 1978 MESOZOIC LYCOPODS AND FERNS FROM THE BUREJA BASIN BY VALENTIN KRASSILOV, Vladivostok*) With 11 plates and 1 figure in the text Abstract Lycopods Lycopodites macrostomus sp. nov. and Synlycostrobus tyrmensis gen. et sp. nov. (leafy shoots, cuticles, strobili, spores) and ferns Osmunda diamensis (SEW.) comb, nov., Klukia tyganensis KRASSIL., Dicksonia concinna HEER (= Coniopteris burejensis (ZAL.) SEW.), D. arctica (PRYN.) comb. nov. and Cyatbea tyrmensis (SEW.) comb. nov. (leaf morphology, indusia, sporangia, spores) are de- scribed from the Late Jurassic — Early Cretaceous strata of the Bureja basin. Thermophilous ferns came from the southern localities about 50° North suggesting a rather steep temperature gradient. Fertile shoots of Synlycostrobus resembled the compound strobilus of conifers evidencing the high evolutionary potential of the Mesozoic lycopods. Osmunda probably originated in the Siberian pro- vince. Most species of Coniopteris had bivalvate (not cup-shaped) indusia as in Dicksonia or Cibotium. In the Cyatheaceae, indusia probably evolved from hemitelioid to cyatheoid. Contents Introduction 16 Stratigraphy 17 Description 17 Lycopodites macrostomus sp. nov 17 Synlycostrobus tyrmensis gen. et sp. nov 18 Osmunda diamensis (SEWARD) comb, nov 19 Klukia tyganensis KRASSILOV 20 Dicksonia concinna HEER 21 Dicksonia arctica (PRYNADA) comb, nov 22 Cyathea tyrmensis (SEWARD) comb, nov 23 Palaeoecology 23 Evolutionary significance 24 Summary 26 References 26 Explanation of Plates 27 Introduction 1976 was a centennial year for paleobotany in Siberia and the Amurland. Initial work of OSWALD HEER (1876) based on the collections furnished by F. B. SCHMIDT in 1859, not only brought to light the fossil floras of northern Asia, but also challenged a widely held belief in universal homogeneity of Jurassic vegetation and climate. -
USGS Open-File Report 2007-1047, Short Research Paper 081, 4 P.; Doi:10.3133/Of2007-1047.Srp081
U.S. Geological Survey and The National Academies; USGS OF-2007-1047, Short Research Paper 081; doi:10.3133/of2007-1047.srp081 Paleobotany of Livingston Island: The first report of a Cretaceous fossil flora from Hannah Point M. Leppe,1 W. Michea,2 C. Muñoz,3 S. Palma-Heldt,3 and F. Fernandoy3 1 Scientific Department, Chilean Antarctic Institute-INACH, Plaza Muñoz Gamero 1055, Punta Arenas, Chile ([email protected]) 2 Departamento de Geología, Universidad de Chile, Plaza Ercilla 803, Casilla 13518, Correo 21, Santiago, Chile ([email protected]) 3 Departamento Ciencias de La Tierra, Universidad de Concepción, Casilla 160-C, Concepción, Chile ([email protected]). Abstract This is the first report of a fossil flora from Hannah Point, Livingston Island, South Shetland Islands, Antarctica. The fossiliferous content of an outcrop, located between two igneous rock units of Cretaceous age are mainly composed of leaf imprints and some fossil trunks. The leaf assemblage consists of 18 taxa of Pteridophyta, Pinophyta and one angiosperm. The plant assemblage can be compared to other Early Cretaceous floras from the South Shetland Islands, but several taxa have an evidently Late Cretaceous affinity. A Coniacian-Santonian age is the most probable age for the outcrops, supported by previous K/Ar isotopic studies of the basalts over and underlying the fossiliferous sequence. Citation: Leppe, M., W. Michea, C. Muñoz, S. Palma-Heldt, and F. Fernandoy (2007), Paleobotany of Livingston Island: The first report of a Cretaceous fossil flora from Hannah Point, in Antarctica: A Keystone in a Changing World – Online Proceedings of the 10th ISAES, edited by A. -
Historical Reconstruction of Climatic and Elevation Preferences and the Evolution of Cloud Forest-Adapted Tree Ferns in Mesoamerica
Historical reconstruction of climatic and elevation preferences and the evolution of cloud forest-adapted tree ferns in Mesoamerica Victoria Sosa1, Juan Francisco Ornelas1,*, Santiago Ramírez-Barahona1,* and Etelvina Gándara1,2,* 1 Departamento de Biología Evolutiva, Instituto de Ecología AC, Carretera antigua a Coatepec, El Haya, Xalapa, Veracruz, Mexico 2 Instituto de Ciencias/Herbario y Jardín Botánico, Benemérita Universidad Autónoma de Puebla, Puebla, Mexico * These authors contributed equally to this work. ABSTRACT Background. Cloud forests, characterized by a persistent, frequent or seasonal low- level cloud cover and fragmented distribution, are one of the most threatened habitats, especially in the Neotropics. Tree ferns are among the most conspicuous elements in these forests, and ferns are restricted to regions in which minimum temperatures rarely drop below freezing and rainfall is high and evenly distributed around the year. Current phylogeographic data suggest that some of the cloud forest-adapted species remained in situ or expanded to the lowlands during glacial cycles and contracted allopatrically during the interglacials. Although the observed genetic signals of population size changes of cloud forest-adapted species including tree ferns correspond to predicted changes by Pleistocene climate change dynamics, the observed patterns of intraspecific lineage divergence showed temporal incongruence. Methods. Here we combined phylogenetic analyses, ancestral area reconstruction, and divergence time estimates with climatic and altitudinal data (environmental space) for phenotypic traits of tree fern species to make inferences about evolutionary processes Submitted 29 May 2016 in deep time. We used phylogenetic Bayesian inference and geographic and altitudinal Accepted 18 October 2016 distribution of tree ferns to investigate ancestral area and elevation and environmental Published 16 November 2016 preferences of Mesoamerican tree ferns. -
Fern Genomes Elucidate Land Plant Evolution and Cyanobacterial Symbioses
ARTICLES https://doi.org/10.1038/s41477-018-0188-8 Fern genomes elucidate land plant evolution and cyanobacterial symbioses Fay-Wei Li 1,2*, Paul Brouwer3, Lorenzo Carretero-Paulet4,5, Shifeng Cheng6, Jan de Vries 7, Pierre-Marc Delaux8, Ariana Eily9, Nils Koppers10, Li-Yaung Kuo 1, Zheng Li11, Mathew Simenc12, Ian Small 13, Eric Wafula14, Stephany Angarita12, Michael S. Barker 11, Andrea Bräutigam 15, Claude dePamphilis14, Sven Gould 16, Prashant S. Hosmani1, Yao-Moan Huang17, Bruno Huettel18, Yoichiro Kato19, Xin Liu 6, Steven Maere 4,5, Rose McDowell13, Lukas A. Mueller1, Klaas G. J. Nierop20, Stefan A. Rensing 21, Tanner Robison 22, Carl J. Rothfels 23, Erin M. Sigel24, Yue Song6, Prakash R. Timilsena14, Yves Van de Peer 4,5,25, Hongli Wang6, Per K. I. Wilhelmsson 21, Paul G. Wolf22, Xun Xu6, Joshua P. Der 12, Henriette Schluepmann3, Gane K.-S. Wong 6,26 and Kathleen M. Pryer9 Ferns are the closest sister group to all seed plants, yet little is known about their genomes other than that they are generally colossal. Here, we report on the genomes of Azolla filiculoides and Salvinia cucullata (Salviniales) and present evidence for episodic whole-genome duplication in ferns—one at the base of ‘core leptosporangiates’ and one specific to Azolla. One fern- specific gene that we identified, recently shown to confer high insect resistance, seems to have been derived from bacteria through horizontal gene transfer. Azolla coexists in a unique symbiosis with N2-fixing cyanobacteria, and we demonstrate a clear pattern of cospeciation between the two partners. Furthermore, the Azolla genome lacks genes that are common to arbus- cular mycorrhizal and root nodule symbioses, and we identify several putative transporter genes specific to Azolla–cyanobacte- rial symbiosis. -
Vol. XIV, No. 1,. the CLASSIFICATION of PLANTS, X
198 The Ohio Naturalist. [Vol. XIV, No. 1,. THE CLASSIFICATION OF PLANTS, X.* JOHN H. SCHAFFNER. Our knowledge of the anatomy, cytology, and life history of many of the groups of Pteridophytes is still far from satisfactory and only a tentative arrangement is at present possible. How- ever, three great lines of development are clearly marked giving three great phyla with which to begin. There may be some dis- pute as to the true relationship of a few isolated groups but in the great majority of living forms the connection is quite evident. Some of the recent speculations in respect to the Pteridophytes have very little morphological evidence for their support. The writer believes that it is best not to disturb the arrangements of the various groups as accepted in the past until there is more than a mere foundation of assumptions based on doubtful evolutionary hypotheses, many of which are all but disproven at the present time. There is a notion that external characters are less stable than internal anatomy. But there is really no evidence that this is so. We should first find out whether there is any ecological response and if so whether one set of structures responds more readily than another. Even if it could be shown that there is ecological adap- tion by direct response to environment or by natural selection this would still be inconclusive, for the internal structure would neces- sarily have to be co-ordinated with the external. A given type of vascular system may be found in a group and thus indicate rela- tionship, but the same is sometimes true of unimportant external structures like the ligule in Selaginclla. -
First Megafossil Evidence of Cyatheaceous Tree Fern from the Indian Cenozoic
First megafossil evidence of Cyatheaceous tree fern from the Indian Cenozoic Subir Bera1,∗, Sudha Gupta2, Mahasin Ali Khan1, Aniruddha De1 and Radhanath Mukhopadhyay3 1Centre of Advanced Study, Department of Botany, University of Calcutta, 35, B.C. Road, Kolkata 700 019, West Bengal, India. 2Department of Botany, University of Kalayni, Nadia 741 235, West Bengal, India. 3Department of Botany, University of Burdwan, Burdwan 713 104, West Bengal, India. ∗Corresponding author. e-mail: [email protected] A part of the compressed tree fern axis with leaf and adventitious root scars in unusual arrangement from Plio–Pleistocene sediments of Arunachal Pradesh, India is described as Cyathea siwalika sp. nov. This record suggests that Cyathea was an important component of tropical evergreen forest in the area during the Plio–Pleistocene and this group subsequently declined in importance within the local vegetation. 1. Introduction 10.5 Ma and the middle to upper Siwaliks transi- tion is dated at 2.6 Ma. The generalized Neogene– Miocene to Pleistocene strata deposited as a pied- Quaternary stratigraphy of Arunachal Pradesh is mont along the southern flank of the Himalayas given in table 1. are rich in fossil plants but pteridophytes are rare Here we describe the first evidence of Cya- and poorly preserved. Although dispersed spores of theaceae based on a part of the compressed stem various fern groups have been recorded regularly, a with leaf and adventitious root scars from the few macroscopic fern remains have been reported. upper Siwalik of Papumpare district of Arunachal They were recorded from Uttarakhand, Assam and Pradesh, India (figure 1). -
A Classification for Extant Ferns
55 (3) • August 2006: 705–731 Smith & al. • Fern classification TAXONOMY A classification for extant ferns Alan R. Smith1, Kathleen M. Pryer2, Eric Schuettpelz2, Petra Korall2,3, Harald Schneider4 & Paul G. Wolf5 1 University Herbarium, 1001 Valley Life Sciences Building #2465, University of California, Berkeley, California 94720-2465, U.S.A. [email protected] (author for correspondence). 2 Department of Biology, Duke University, Durham, North Carolina 27708-0338, U.S.A. 3 Department of Phanerogamic Botany, Swedish Museum of Natural History, Box 50007, SE-104 05 Stock- holm, Sweden. 4 Albrecht-von-Haller-Institut für Pflanzenwissenschaften, Abteilung Systematische Botanik, Georg-August- Universität, Untere Karspüle 2, 37073 Göttingen, Germany. 5 Department of Biology, Utah State University, Logan, Utah 84322-5305, U.S.A. We present a revised classification for extant ferns, with emphasis on ordinal and familial ranks, and a synop- sis of included genera. Our classification reflects recently published phylogenetic hypotheses based on both morphological and molecular data. Within our new classification, we recognize four monophyletic classes, 11 monophyletic orders, and 37 families, 32 of which are strongly supported as monophyletic. One new family, Cibotiaceae Korall, is described. The phylogenetic affinities of a few genera in the order Polypodiales are unclear and their familial placements are therefore tentative. Alphabetical lists of accepted genera (including common synonyms), families, orders, and taxa of higher rank are provided. KEYWORDS: classification, Cibotiaceae, ferns, monilophytes, monophyletic. INTRODUCTION Euphyllophytes Recent phylogenetic studies have revealed a basal dichotomy within vascular plants, separating the lyco- Lycophytes Spermatophytes Monilophytes phytes (less than 1% of extant vascular plants) from the euphyllophytes (Fig.