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THE DESERT EDGE: FLORA OF THE GUAYMAS REGION OF , . PART 1: THE CHECKLIST

Item Type Article

Authors Felger, Richard Stephen; Carnahan, Susan Davis; Sanchez- Escalante, Jose Jesus

Publisher University of (Tucson, AZ)

Journal Desert

Rights Copyright © Arizona Board of Regents. The University of Arizona.

Download date 25/09/2021 05:16:18

Link to Item http://hdl.handle.net/10150/627276 19 Felger, Carnahan, Sánchez-Escalante The Desert Edge: Flora of the Guaymas Region of Sonora, Mexico

THE DESERT EDGE: FLORA OF THE GUAYMAS REGION OF SONORA, MEXICO. PART 1: THE CHECKLIST

Richard Stephen Felger Herbarium, University of Arizona Tucson, Arizona 85721 & International Alliance Ajo, Arizona Author for correspondence: [email protected]

Susan Davis Carnahan Herbarium, University of Arizona Tucson, Arizona 85721 [email protected]

José Jesús Sánchez-Escalante Herbario USON, Universidad de Sonora-DICTUS Blvd. Luis Encinas y Hermosillo, Sonora, C.P. 83000, México [email protected]

Abstract: A checklist is provided for the vascular plants of the Guaymas region of western Sonora. This region encompasses 532,000 hectares (1,314,600 acres) where the southern Sonoran Desert transitions from subtropical thornscrub. This flora includes 820 native and non-native taxa in 113 families and 471 genera. There are 97 non-natives established in the flora area, 27 of which are grasses. Nineteen taxa are endemic to the flora area.

Resumen: Se proporciona un listado florístico de las plantas vasculares para la región de Guaymas en el occidente de Sonora. Esta región cubre 532,000 hectáreas (1,314,600 acres), donde ocurre la transición del matorral espinoso subtropical hacia la parte sur del Desierto Sonorense. Esta flora incluye 820 taxa nativas y no nativas, distribuidas en 113 familias y 471 géneros. Hay 97 taxa no nativas establecidas en el área de la flora, de las cuales 27 son pastos. Diecinueve taxa son endémicas a la región. 20 Desert Plants 33(1) October 2017

Figure 1. Guaymas region. Flora area is approximately within the dashed red line, from the vicinity of Tastiota to Río Yaqui, coastal ranges and adjacent inland ranges including parts of Sierra Libre and Sierra Bacatete. Also included are Isla San Pedro Nolasco and near-shore islands. Map by Alberto Búrquez-Montijo.

This checklist covers the vascular plants of the Guaymas re- Edward Palmer made the first major collections from the region, gion of western Sonora, Mexico. The boundaries of our floris- collecting hundreds of specimens on multiple trips beginning in tic study extend from the vicinity of Tastiota southward to the 1867 (McVaugh 1956; Watson 1889). Over the ensuing century Río Yaqui and include the Sierra El Aguaje as well as parts of the and a half, thousands of additional vouchers were acquired and nearby Sierra Libre and Sierra Bacatete. Also included are Isla San deposited at the University of Arizona Herbarium (ARIZ), the Pedro Nolasco and small islands in Guaymas Bay and near San Herbario de la Universidad de Sonora (USON), and other her- Carlos (Figure 1). This is the desert edge, the southern limit of baria, including ASU, CAS, DES, GH, HCIB, MEXU, MO, NMC, the Sonoran Desert in mainland Mexico and the northern coastal RSA, SD, TEX, UC, UCR, and US. Many of these records appear limit of subtropical thornscrub—the most arid expression of the in SEINet (2017). Despite this extensive botanical history, there tropical deciduous . The terrain is complex and encompasses coastal areas with mangroves, desert and thornscrub plains, ba- jadas, and rugged mountains with riparian canyons (Felger et al. Table I. Taxa endemic to the Guaymas region 2016). Shreve (1951) designated the vegetation of the region as Mammillaria tayloriorum part of the Sonoran segment of the Gulf Coast subdivision of the Mandevilla nacapulensis CONVOLVULACEAE Sonoran Desert. Ipomoea seaania This flora includes 820 native and non-native taxa (, Coreocarpus sanpedroensis subspecies, varieties, and hybrids) in 113 families and 471 genera. Non-natives established and reproducing in the flora area total 97 Coreocarpus sonoranus pediculifera taxa (12 percent of the flora), 27 of which are grasses (). var. libranus var. linearifolia An additional 24 non-natives are not established as reproducing Hofmeisteria crassifolia populations. The most diverse families are grasses (Poaceae, 92 Verbesina felgeri Briquetia sonorae taxa), composites (Asteraceae, 82), legumes (, 72), eu- CACTACEAE Gossypium turneri phorbs (Euphorbiaceae, 55), mallows (Malvaceae, 42), and cacti Echinocereus llanuraensis (Cactaceae, 34). The largest genera are Euphorbia (27 species), Ipo- Echinocereus websterianus alata moea (15), and Cyperus (14). Taxa based on specimens from the flora Grusonia reflexispina area total 154. Nineteen taxa are endemic to the flora area (Table Mammillaria boolii Stenaria sanchezii 1); four of these are known only from Isla San Pedro Nolasco: Mammillaria johnstonii SOLANACEAE Coreocarpus sanpedroensis, Echinocereus websterianus, Mammillaria multi- digitata, and Mammillaria tayloriorum. Mammillaria multidigitata Physalis purpurea 21 Felger, Carnahan, Sánchez-Escalante The Desert Edge: Flora of the Guaymas Region of Sonora, Mexico

exists no comprehensive floristic treatment of the greater Guay- NYMPHAEALES mas region. The unique flora of Isla San Pedro Nolasco comprises 59 species (Felger et al. 2011, 2012). A revised flora of Cañón del NYMPHAEACEAE Nacapule in the Sierra El Aguaje includes 310 taxa, encompassing Nymphaea elegans Hooker 38% of the regional flora (Felger et al. 2017). An in-depth flora of the region, including identification keys and species descriptions, is in preparation. In the following checklist, family designations follow APG III Anisacanthus thurberi (Torrey) A. Gray & IV (Angiosperm Phylogeny Group 2009, 2016) and the An- Aphanosperma sinaloensis (Leonard & Gentry) T.F. Daniel giosperm Phylogeny Website (Stevens 2012). Selected synonyms [Carlowrightia sinaloensis Leonard & Gentry] are given in brackets. Non-native taxa established as reproduc- Avicennia germinans (Linnaeus) Linnaeus ing populations in the flora area are indicated with an asterisk (*) Carlowrightia arizonica A. Gray and non-natives not established as reproducing populations are Carlowrightia pectinata Brandegee marked with two asterisks (**). Taxa described from the flora area resupinata (Vahl) de Jussieu (those with the type locality in the flora area) are indicated with (T) Elytraria imbricata (Vahl) Persoon following the scientific name. Henrya insularis Nees ex Bentham Holographis virgata (Harvey ex Bentham & Hooker) T.F. LYCOPHYTES Daniel subsp. virgata [Berginia virgata Harvey ex Bentham & SELAGINELLACEAE Hooker] Selaginella rupincola Underwood Justicia californica (Bentham) D.N. Gibson Justicia candicans (Nees) L.D. Benson [Jacobinia ovata A. Gray PTERIDOPHYTES var. subglabrata S. Watson (T)] MARSILEACEAE Justicia longii Hilsenbeck [Siphonoglossa longiflora (Torrey) A. Marsilea vestita Hooker & Greville Gray] Justicia sonorae Wasshausen PSILOTACEAE Ruellia californica (Rose) I.M. Johnston subsp. californica Psilotum nudum (Linnaeus) Palisot de Beauvois Ruellia ciliatiflora Hooker [R. nudiflora (Engelmann & A. Gray) Urban. R. nudiflora var. glabrata Leonard] PTERIDACEAE Ruellia intermedia Leonard Adiantum capillus-veneris Linnaeus Ruellia leucantha Brandegee subsp. postinsularis (Gentry) T.F. Adiantum tricholepis Fée Daniel Astrolepis sinuata (Lagasca ex Swartz) D.M. Benham subsp. Tetramerium nervosum Nees [Dianthera sonorae S. Watson (T). sinuata [Notholaena sinuata (Lagasca ex Swartz) Kaulfuss] Tetramerium hispidum Nees] Cheilanthes lozanoi (Maxon) R.M. Tryon & A.F. Tryon var. seemannii (Hooker) Mickel & Beitel ACHATOCARPACEAE Myriopteris lindheimeri (Hooker) J. Small [Cheilanthes lindheimeri Phaulothamnus spinescens A. Gray Hooker] Myriopteris pringlei (Davenport) Grusz & Windham subsp. AIZOACEAE pringlei [Cheilanthes pringlei Davenport] *Mesembryanthemum crystallinum Linnaeus Notholaena lemmonii D.C. Eaton var. lemmonii Sesuvium portulacastrum (Linnaeus) Linnaeus Notholaena standleyi Maxon [Cheilanthes standleyi (Maxon) Sesuvium verrucosum Rafinesque Mickel] Trianthema portulacastrum Linnaeus

THELYPTERIDACEAE (includes Chenopodiaceae) Thelypteris puberula (Baker) C.V. Morton var. sonorensis A.R. Achyranthes watsonii Standley Smith Allenrolfea occidentalis (S. Watson) Kuntze Amaranthus fimbriatus (Torrey) Bentham ex S. Watson EARLY ANGIOSPERMS Amaranthus obcordatus (A. Gray) Standley Amaranthus palmeri S. Watson Amaranthus watsonii Standley (T) Arthrocnemum subterminale (Parish) Standley [Salicornia watsonii Wooton & Standley subterminalis Parish] barclayana (Bentham) D. Dietrich SAURURACEAE Atriplex canescens (Pursh) Nuttall var. canescens Anemopsis californica (Nuttall) Hooker & Arnott 22 Desert Plants 33(1) October 2017

Atriplex elegans (Moquin) D. Dietrich [A. fasciculata S. Watson. Mandevilla nacapulensis (Felger & Henrickson) A.O. Simões, A. elegans subsp. fasciculata (S. Watson) H.M. Hall & Clements] Kinoshita & M.E. Endress [Telosiphonia nacapulensis Felger & Atriplex linearis S. Watson (T) [A. canescens subsp. linearis (S. Henrickson (T)] Watson) H.M. Hall & Clements] Marsdenia edulis S. Watson (T) Atriplex pacifica A. Nelson Matelea cordifolia, see Polystemma cordifolium Atriplex polycarpa (Torrey) S. Watson Matelea parvifolia (Torrey) Woodson *Atriplex wrightii S. Watson Matelea pringlei (A. Gray) Woodson *Chenopodiastrum murale (Linnaeus) S. Fuentes, Uotila & Metastelma arizonicum A. Gray [Cynanchum arizonicum (A. Borsch [Chenopodium murale Linnaeus] Gray) Shinners. Metastelma albiflorum S. Watson (T), not Grise- Chenopodium neomexicanum Standley bach. M. watsonianum Standley (T)] *Dysphania ambrosioides (Linnaeus) Mosyakin & Clemants Metastelma californicum Bentham [Chenopodium ambrosioides Linnaeus var. anthelminticum **Nerium oleander Linnaeus (Linnaeus) A. Gray] Plumeria rubra Linnaeus [P. acutifolia Poiret] Froelichia interrupta (Linnaeus) Moquin var. alata (S. Watson) Polystemma cordifolium (A. Gray) McDonnell & Fishbein R.A. McCauley [Matelea cordifolia (A. Gray) Woodson. Rothrockia cordifolia A. Gomphrena sonorae Torrey Gray] Iresine alternifolia S. Watson (T) Vallesia glabra (Cavanilles) Link var. glabra Iresine latifolia (M. Martens & Galeotti) Bentham & Hooke f. Vallesia laciniata Brandegee [V. baileyana Woodson (T)] [I. calea (Ibáñez) Standley] Monolepis nuttalliana (Schultes) Greene ASTERACEAE Salicornia bigelovii Torrey Acourtia wrightii (A. Gray) Reveal & R.M. King [Perezia wrightii *Salsola tragus Linnaeus [Kali tragus Scopoli] A. Gray] Sarcocornia pacifica (Standley) A.J. Scott [Salicornia pacifica Adenophyllum anomalum (Canby & Rose) Strother [Dyssodia Standley] anomala (Canby & Rose) B.L. Robinson] Suaeda esteroa Ferren & Whitmore Aldama glomerata (Brandegee) E.E. Schilling & Panero Suaeda nigra (Rafinesque) J.F. Macbride [S. moquinii (Torrey) [ glomerata Brandegee var. glomerata] Greene. S. ramosissima (Standley) I.M. Johnston] Ambrosia acuminata (Brandegee) W.W. Payne Tidestromia lanuginosa (Nuttall) Standley subsp. eliassoniana Ambrosia ambrosioides (Cavanilles) W.W. Payne Sánchez-del Pino & Flores-Olvera (T) [T. eliassoniana Ambrosia confertiflora de Candolle (Sánchez-del Pino & Flores-Olvera) Sánchez-del Pino] Ambrosia cordifolia (A. Gray) W.W. Payne (Torrey) W.W. Payne ANACARDIACEAE Ambrosia divaricata (Brandegee) W.W. Payne Rhus terebinthifolia Schlechtendal & Chamisso [R. hartmanii Ambrosia monogyra (Torrey & A. Gray) Strother & B.G. F.A. Barkley] Baldwin [Hymenoclea monogyra Torrey & A. Gray] **Schinus terebinthifolia Raddi Ambrosia psilostachya de Candolle Arida coulteri, see Leucosyris coulteri APIACEAE Arida crispa, see Leucosyris crispa *Cyclospermum leptophyllum (Persoon) Sprague ex Britton & Baccharis salicifolia (Ruiz & Pavón) Persoon P. Wilson var. leptophyllum Baccharis sarothroides A. Gray Daucus pusillus Michaux Bahiopsis chenopodina (Greene) E.E. Schilling & Panero Eryngium nasturtiifolium de Jussieu ex F. Delaroche [Viguiera chenopodina Greene. V. deltoidea A. Gray var. chenopo- dina (Greene) S.F. Blake] APOCYNACEAE Bahiopsis triangularis (M.E. Jones) E.E. Schilling & Panero * curassavica Linnaeus [Viguiera triangularis M.E. Jones] Asclepias leptopus I.M. Johnston (T) juncea (Bentham) Greene var. aspera Greene Asclepias subulata Decaisne laevis (Linnaeus) Britton, Sterns, & Poggenburg * R. Brown Brickellia coulteri A. Gray var. coulteri Dictyanthus altatensis (Brandegee) W.D. Stevens [Matelea Brickellia rhomboidea Greene (T) altatensis (Brandegee) Woodson] **Carthamus tinctorius Linnaeus clausum (Jacquin) Schlechter [Cynanchum clausum Chloracantha spinosa (Bentham) G.L. Nesom var. spinosa Jacquin. clausum (Jacquin) Roemer & Schultes] [Aster spinosus Bentham var. spinosus] Funastrum heterophyllum (Engelmann ex Torrey) Standley [F. Chromolaena sagittata (A. Gray) R.M. King & H. Robinson hartwegii Schlechter. Sarcostemma cynanchoides Decaisne subsp. [Eupatorium sagittatum A. Gray] hartwegii R.W. Holm. S. heterophyllum Engelmann ex Torrey] Cirsium rhaphilepis (Hemsley) Petrak Haplophyton cimicidum A. de Candolle 23 Felger, Carnahan, Sánchez-Escalante The Desert Edge: Flora of the Guaymas Region of Sonora, Mexico

Coreocarpus sanpedroensis E.B. Smith (T) [C. arizonicus (A. Pectis purpurea Brandegee var. sonorae D.J. Keil (T) Gray) S.F. Blake var. sanpedroensis (E.B. Smith) B.L. Turner] Pectis rusbyi Greene ex A. Gray [P. palmeri S. Watson (T)] Coreocarpus sonoranus Sherff var. libranus B.L. Turner (T) Perityle californica Bentham [P. deltoidea S. Watson (T)] Coreocarpus sonoranus var. sonoranus [C. johnstonii Sherff (T)] Perityle emoryi Torrey canescens A. Gray Perityle microglossa Bentham var. microglossa * prostrata (Linnaeus) Linnaeus [E. alba (Linnaeus) Perityle palmeri S. Watson (T)] [P. leptoglossa Harvey & A. Gray Hasskarl. E. erecta Linnaeus] subsp. palmeri (S. Watson) Felger & Lowe] ** viscosa (Linnaeus) Lessing [Cotula viscosa Linnaeus] Pleurocoronis laphamioides (Rose) R.M. King & H. Robinson A. Gray ex Torrey [Hofmeisteria laphamioides Rose. H. laphamioides var. pauciseta Encelia halimifolia Cavanilles (I.M. Johnston) S.F. Blake. H. pluriseta var. pauciseta I.M. *Erigeron canadensis Linnaeus var. glabratus A. Gray [Conyza Johnston (T)] canadensis (Linnaeus) Cronquist var. glabrata (A. Gray) Cron- Pluchea salicifolia (Miller) S.F. Blake [P. adnata (Humboldt & quist] Bonpland ex Willdenow) C. Mohr. P. subdecurrens Cassini] Gamochaeta sphacelata (Kunth) Cabrera [Gnaphalium Porophyllum gracile Bentham pedunculosum I.M. Johnston. G. sphacelatum Kunth] Porophyllum pausodynum B.L. Robinson & Greenman (T) [P. *Glebionis coronaria (Linnaeus) Cassini ex Spach brachypodum B.L. Robinson (T). P. seemannii S. Watson (T), not [Chrysanthemum coronarium Linnaeus] P. seemannii Schultz-Bipontinus] Gnaphalium palustre Nuttall Pseudognaphalium stramineum (Kunth) Anderberg [Gnaphali- *Gnaphalium polycaulon Persoon [G. indicum of authors, not um chilense Sprengel. G. stramineum Kunth] Linnaeus] Senecio lemmonii A. Gray Gymnosperma glutinosum (Sprengel) Lessing **Silybum marianum (Linnaeus) Gaertner Helenium laciniatum A. Gray (T) *Sonchus oleraceus Linnaeus *Helianthus annuus Linnaeus Symphyotrichum subulatum (Michaux) G.L. Nesom var. Helianthus niveus (Bentham) Brandegee var. niveus parviflorum (Nees) S.D. Sundberg [Aster subulatus Michaux] Heliopsis anomala (M.E. Jones) B.L. Turner [Heliopsis rubra T.R. Thymophylla concinna (A. Gray) Strother [Dyssodia concinna (A. Fisher. H. parvifolia A. Gray var. rubra (T.R. Fisher) Wiggins] Gray) B.L. Robinson] Hofmeisteria crassifolia S. Watson (T) **Thymophylla tenuiloba (de Candolle) Small Isocoma acradenia (Greene) Greene var. acradenia [Haplo- Trichocoronis wrightii (Torrey & A. Gray) A. Gray [Ageratum pappus acradenius (Greene) S.F. Blake. Isocoma veneta (Kunth) wrightii Torrey & A. Gray] Greene] Trixis californica Kellogg var. californica Koanophyllon palmeri (A. Gray) R.M. King & H. Robinson *Verbesina encelioides (Cavanilles) Bentham & Hooker f. ex A. [Eupatorium palmeri A. Gray] Gray *Lactuca serriola Linnaeus Verbesina felgeri B.L. Turner (T) Lagascea decipiens Hemsley var. decipiens Viguiera dentata (Cavanilles) Sprengel Leucosyris coulteri (A. Gray) Pruski & R.L. Hartman *Xanthium strumarium Linnaeus [Arida coulteri (A. Gray) D.R. Morgan & R.L. Hartman. Xylothamia diffusa (Bentham) G.L. Nesom [Ericameria diffusa coulteri (A. Gray) B.L. Turner & Horne var. Bentham. Haplopappus sonorensis (A. Gray) S.F. Blake. Linosyris coulteri. Psilactis coulteri A. Gray (T)] sonorensis A. Gray (T)] Leucosyris crispa (Brandegee) Pruski & R.L. Hartman [Arida crispa (Brandegee) D.R. Morgan & R.L. Hartman. BATACEAE Machaeranthera crispa (Brandegee) B.L. Turner & Horne. Batis maritima Linnaeus Psilactis crispa Brandegee] Logfia filaginoides (Hooker & Arnott) Morefield Filago[ californica Nuttall. Logfia californica (Nuttall) Holub] ** alata Kunth Malacothrix sonorae W.S. Davis & P.H. Raven Handroanthus impetiginosus (Martius ex de Candolle) Mattos Melampodium cupulatum A. Gray (T) [Tabebuia impetiginosa (Martius ex de Candolle) Standley. T. Palafoxia linearis (Cavanilles) Lagasca palmeri Rose] Parthenium tomentosum de Candolle var. stramonium Tecoma stans (Linnaeus) de Jussieu ex Kunth var. angustata (Greene) Rollins Rehder [T. incisa I.M. Johnston, not T. incisa Sweet] Pectis coulteri Harvey & A. Gray (T) *Tecoma stans var. stans Pectis cylindrica (Fernald) Rydberg [P. prostrata Cavanilles var. cylindrica Fernald] Pectis filipes Harvey & A. Gray var. filipes Pectis linifolia Linnaeus var. linifolia Pectis papposa Harvey & A. Gray var. papposa 24 Desert Plants 33(1) October 2017

BORAGINACEAE sensu lato CACTACEAE This formerly broadly circumscribed family is treated Carnegiea gigantea (Engelmann) Britton & Rose as 6 families in the flora area: , Cylindropuntia arbuscula (Engelmann) F.M. Knuth [Opuntia CORDIACEAE, EHRETIACEAE, HELIOTROPIACEAE, arbuscula Engelmann] HYDROPHYLLACEAE, and NAMACEAE. Cylindropuntia bigelovii (Engelmann) F.M. Knuth [Opuntia bigelovii Engelmann] BORAGINACEAE Cylindropuntia fulgida (Engelmann) Knuth var. fulgida maritima Greene [C. maritima var. pilosa I.M. [Opuntia fulgida Engelmann] Johnston] Cylindropuntia fulgida var. mamillata (Schott ex Engelmann) Johnstonella angustifolia (Torrey) Hasenstab & M.G. Simpson Backeberg [Opuntia mamillata A. Schott ex Engelmann] [Cryptantha angustifolia (Torrey) Greene] Cylindropuntia leptocaulis (de Candolle) F.M. Knuth [Opuntia Johnstonella grayi (Vasey & Rose) Hasenstab & M.G. Simp- leptocaulis de Candolle] son var. cryptochaeta (J.F. Macbride) Hasenstab & M.G. Cylindropuntia leptocaulis × C. versicolor Simpson [Cryptantha grayi (Vasey & Rose) J.F. Macbride var. Cylindropuntia versicolor (Engelmann ex J.M. Coulter) F.M. cryptochaeta I.M. Johnston] Knuth [Opuntia versicolor Engelmann ex J.M. Coulter] Johnstonella grayi var. nesiotica (I.M. Johnston) Hasenstab & Echinocereus leucanthus N.P. Taylor (T) [Wilcoxia albiflora M.G. Simpson [Cryptantha grayi var. nesiotica I.M. Johnston] Backeberg, not Echinocereus albiflorus Weingart] Pectocarya recurvata I.M. Johnston Echinocereus llanuraensis (Rutow) W. Blum & D. Felix (T) [E. engelmannii (Engelmann) Lemaire subsp. llanuraensis (Rutow) Felger. E. nicholii (L.D. Benson) B.D. Parfitt subsp.llanuraensis *Brassica kaber (de Candolle) L.C. Wheeler [Not Brassica arvensis Rutow] Linnaeus; B. arvensis Rabenhorst is illegitimate. Sinapis arvensis Echinocereus scopulorum Britton & Rose (T) Linnaeus. S. kaber de Candolle] Echinocereus websterianus G.E. Lindsay (T) *Brassica nigra (Linnaeus) W.D.J. Koch [Sinapis nigra Linnaeus] emoryi (Engelmann) Orcutt [F. covillei Britton & **Brassica rapa Linnaeus var. oleifera de Candolle Rose. F. emoryi subsp. covillei (Britton & Rose) D.R. Hunt & **Brassica rapa var. rapa Dimmitt] *Brassica tournefortii Gouan Ferocactus herrerae J. González Ortega [F. wislizeni Descurainia pinnata (Walter) Britton (Engelmann) Britton & Rose var. herrerae (J. González cuneifolia, see Tomostima cuneifolia Ortega) N.P. Taylor] Dryopetalon runcinatum A. Gray [D. runcinatum var. laxiflorum Ferocactus tiburonensis (G.E. Lindsay) Backeberg [F. wislizeni Rollins] (Engelmann) Britton & Rose var. tiburonensis G.E. Lindsay] *Eruca vesicaria (Linnaeus) Cavanilles subsp. sativa (Miller) Grusonia reflexispina (Wiggins & Rollins) E.F. Anderson Thellung [E. sativa Miller] [Corynopuntia reflexispina (Wiggins & Rollins) Backeberg. *Lepidium didymum Linnaeus [Coronopus didymus (Linnaeus) Opuntia reflexispina Wiggins & Rollins (T)] Smith] Lophocereus schottii (Engelmann) Britton & Rose var. Lepidium lasiocarpum Nuttall ex Torrey & A. Gray australis (K. Brandegee) Borg [L. schottii var. tenuis G.E. Lepidium virginicum Linnaeus Lindsay (T)] Lyrocarpa coulteri Hooker & Harvey ex Harvey var. coulteri Lophocereus schottii var. schottii [Pachycereus schottii *Raphanus sativus Linnaeus (Engelmann) D.R. Hunt] Rorippa teres (Michaux) Stuckey Mammillaria bocensis R.T. Craig *Sisymbrium irio Linnaeus Mammillaria boolii G.E. Lindsay (T) *Sisymbrium orientale Linnaeus Mammillaria grahamii Engelmann subsp. sheldonii (Britton & Tomostima cuneifolia (Nuttall ex Torrey & A. Gray) Rose) D.R. Hunt [M. inaiae R.T. Craig (T). M. swinglei (Britton Al-Shehbaz, M. Koch & Jordon-Thaden [Draba cuneifolia & Rose) Boedeker. M. swinglei var. diabloa P.C. Fischer (T). Nuttall ex Torrey & A. Gray. D. cuneifolia var. integrifolia S. Neomammillaria sheldonii Britton & Rose (T)] Watson. D. cuneifolia var. sonorae (Greene) Parish. D. sonorae Mammillaria johnstonii (Britton & Rose) Orcutt [M. johnstonii Greene. Tomostima sonorae (Greene) Al-Shehbaz, M. Koch & var. guaymensis R.T. Craig (T). M. johnstonii var. sancarlosensis Jordon-Thaden] R.T. Craig (T). Neomammillaria johnstonii Britton & Rose (T)] Mammillaria multidigitata W.T. Marshall ex G.E. Lindsay (T) BURSERACEAE Mammillaria tayloriorum Glass & R.A. Foster (T) Bursera fagaroides (Kunth) Engelmann var. elongata McVaugh Mammillaria yaquensis R.T. Craig (T) [M. thornberi Orcutt var. & Rzedowski [B. confusa (Rose) Engler, in part] yaquensis (R.T. Craig) D.R. Hunt] Bursera hindsiana (Bentham) Engler Opuntia bravoana E.M. Baxter Bursera laxifloraS. Watson (T) Opuntia gosseliniana F.A.C. Weber (T) Bursera microphylla A. Gray Pachycereus pecten-aboriginum (Engelmann) Britton & Rose 25 Felger, Carnahan, Sánchez-Escalante The Desert Edge: Flora of the Guaymas Region of Sonora, Mexico

Pachycereus pringlei (S. Watson) Britton & Rose CONVOLVULACEAE Peniocereus marianus (Gentry) Sánchez-Mejorada [Wilcoxia *Convolvulus arvensis Linnaeus marianus Gentry] Cressa truxillensis Kunth Peniocereus striatus (Brandegee) F. Buxbaum [Cereus diguetii K. Cuscuta americana Linnaeus Weber. C. striatus Brandegee. Neoevansia diguetii (F.A.C. Weber) Cuscuta corymbosa Ruiz & Pavón var. grandiflora Engelmann W.T. Marshall. N. striata (Brandegee) Sánchez-Mejorada] Cuscuta desmouliniana Yuncker [C. umbellata var. dubia Yuncker Pereskiopsis porteri (Brandegee ex F.A.C. Weber) Britton & (T)] Rose [Opuntia porterii Brandegee ex F.A.C. Weber. Pereskiopsis *Cuscuta indecora Choisy var. indecora [C. decora Choisy ex gatesii E.M. Baxter] Engelmann var. indecora (Choisy) Engelmann. Epithymum Stenocereus alamosensis (J.M. Coulter) A.C. Gibson & K.E. indecorum (Choisy) Nieuwland & Lunell] Horak [Rathbunia alamosensis (J.M. Coulter) Britton & Rose] Cuscuta legitima Costea & Stefanović (T) [C. californica Hooker Stenocereus thurberi (Engelmann) F. Buxbaum [Lemaireocereus & Arnott var. reflexa J.M. Coulter. C. umbellata var. reflexa (J.M. thurberi (Engelmann) Britton & Rose] Coulter) Yuncker] Cuscuta tuberculata Brandegee CAMPANULACEAE Cuscuta umbellata Kunth var. umbellata Lobelia cardinalis Linnaeus subsp. graminea (Lamarck) Evolvulus alsinoides (Linnaeus) Linnaeus var. angustifolia McVaugh Torrey [E. alsinoides var. acapulcensis (Willdenow) van Ooststroom] CANNABACEAE Ipomoea arborescens (Humboldt & Bonpland ex Willdenow) **Cannabis sativa Linnaeus G. Don var. arborescens Celtis pallida Torrey subsp. pallida [C. tala Gillies var. pallida *Ipomoea batatas (Linnaeus) Lamarck [Batatas edulis (Thunberg) (Torrey) Planchon] Choisy. Ipomoea triloba of authors, not Linnaeus] Celtis reticulata Torrey Ipomoea bracteata Cavanilles [Exogonium bracteatum (Cavanilles) Choisy ex G. Don] CAPPARACEAE **Ipomoea carnea Jacquin subsp. fistulosa (Martius ex Choisy) Atamisquea emarginata Miers ex Hooker & Arnott [Capparis D.F. Austin [I. fistulosa Martius ex Choisy] atamisquea Kuntze] Ipomoea cholulensis Kunth [I. cristulata Hallier. Quamoclit cholulensis (Kunth) G. Don] Ipomoea hederacea Jacquin Achyronychia cooperi A. Gray Ipomoea hederifolia Linnaeus [I. coccinea of authors, not Drymaria arenarioides Humboldt & Bonpland ex Schultes Linnaeus. Quamoclit coccinea (Linnaeus) Moench. Q. coccinea subsp. peninsularis (S.F. Blake) J.A. Duke var. hederifolia (Linnaeus) House. Q. hederifolia (Linnaeus) G. Drymaria glandulosa Bartling Don] Drymaria viscosa S. Watson *Ipomoea heptaphylla Sweet [I. pulchella of authors, not Roth. I. *Spergularia salina J. Presl & C. Presl spiralis House (T). I. wrightii A. Gray] *Ipomoea leucantha Jacquin CELASTRACEAE Ipomoea muricata (Linnaeus) Jacquin Maytenus phyllanthoides Bentham var. phyllanthoides Ipomoea nil (Linnaeus) Roth [I. hederacea of authors, not Jacquin] [Tricerma phyllanthoides (Bentham) Lundell] *Ipomoea pes-caprae (Linnaeus) R. Brown subsp. brasiliensis (Linnaeus) van Ooststroom CLEOMACEAE Ipomoea seaania Felger & D.F. Austin (T) Cleome tenuis S. Watson (T) Ipomoea ternifolia Cavanilles var. leptotoma (Torrey) J.A. *Corynandra viscosa (Linnaeus) Cochrane & Iltis subsp. McDonald [I. divergens House (T). I. leptotoma Torrey] viscosa [Arivela viscosa (Linnaeus) Rafinesque.Cleome viscosa *Ipomoea triloba Linnaeus Linnaeus] Jacquemontia agrestis (Choisy) Meisner [J. palmeri S. Watson Wislizenia palmeri A. Gray [W. mamillata Rose ex Greene (T). (T)] W. refracta Engelmann var. palmeri (A. Gray) I.M. Johnston] Jacquemontia polyantha (Schlechtendal & Chamisso) Hallier f. [J. albida Wiggins & Rollins (T)] Jacquemontia pringlei A. Gray [J. pringlei var. glabrescens A. Gray] de Candolle Merremia palmeri (S. Watson) Hallier f. [Ipomoea palmeri S. Watson (T)] COMBRETACEAE Laguncularia racemosa (Linnaeus) C.F. Gaertner CORDIACEAE Cordia parvifolia A. de Candolle Cordia sonorae Rose 26 Desert Plants 33(1) October 2017

Varronia curassavica Jacquin [Cordia brevispicata M. Martens Cnidoscolus palmeri (S. Watson) Rose [Jatropha palmeri S. & Galeotti. C. curassavica (Jacquin) Roemer & Schultes. C. Watson (T)] palmeri S. Watson (T)] Croton alamosanus Rose Croton californicus Müller Argoviensis CRASSULACEAE Croton ciliatoglandulifer Ortega Graptopetalum rusbyi (Greene) Rose Croton magdalenae Millspaugh Croton sonorae Torrey CROSSOSOMATACEAE Croton subjucundus Croizat Crossosoma bigelovii S. Watson Croton texensis (Klotzsch) Müller Argoviensis Dalechampia scandens Linnaeus var. scandens [D. scandens var. CUCURBITACEAE fimbriata (Kunth) Müller Argoviensis] Apodanthera palmeri S. Watson (T) Euphorbia abramsiana L.C. Wheeler [ abramsiana **Citrullus lanatus (Thunberg) Matsumura & Nakai subsp. (L.C. Wheeler) Koutnik. Euphorbia pediculifera var. abramsiana lanatus Ewan] *Cucumis melo Linnaeus var. dudaim (Linnaeus) Naudin [C. Euphorbia albomarginata Torrey & A. Gray [Chamaesyce dudaim Linnaeus] albomarginata (Torrey & A. Gray) Small] Echinopepon wrightii (A. Gray) S. Watson Euphorbia arizonica Engelmann [Chamaesyce arizonica Ibervillea sonorae (S. Watson) Greene [Maximowiczia insularis (Engelmann) Arthur. Euphorbia portulana S. Watson (T)] Brandegee. M sonorae S. Watson (T)] Euphorbia californica Bentham Luffa sepium (G. Meyer) C. Jeffrey [L. operculata (Linnaeus) Euphorbia capitellata Engelmann [Chamaesyce capitellata Cogniaux] (Engelmann) Millspaugh. Euphorbia capitellata var. laxiflora S. Tumamoca macdougalii Rose [Ibervillea macdougalii (Rose) Lira, Watson (T). E. gladiosa M.E. Jones (T)] Dávila & Legaspi] Euphorbia ceroderma I.M. Johnston (T) Vaseyanthus insularis (S. Watson) Rose [Echinopepon insularis Euphorbia cymosa Poiret [E. colletioides Bentham. E. plicata S. S. Watson (T). E. palmeri S. Watson. Vaseyanthus insularis var. Watson] inermis I.M. Johnston. V. insularis var. palmeri (S. Watson) Euphorbia eriantha Bentham [Poinsettia eriantha (Bentham) Rose Gentry] & Standley] Euphorbia florida Engelmann [Chamaesyce florida (Engelmann) EHRETIACEAE Millspaugh] Bourreria sonorae S. Watson (T) Euphorbia gentryi V.W. Steinmann & T.F. Daniel Pholisma culiacanum (Dressler & Kuijt) Yatskievych Euphorbia gracillima S. Watson [Chamaesyce gracillima (S. Tiquilia cuspidata (I.M. Johnston) A.T. Richardson [Coldenia Watson) Millspaugh] cuspidata I.M. Johnston] *Euphorbia hirta Linnaeus [Chamaesyce hirta (Linnaeus) Millspaugh] ELATINACEAE *Euphorbia hypericifolia Linnaeus [Chamaesyce hypericifolia Bergia texana (Hooker) Seubert (Linnaeus) Millspaugh] Euphorbia hyssopifolia Linnaeus [Chamaesyce hyssopifolia EUPHORBIACEAE (Linnaeus) Small] aliena Brandegee Euphorbia incerta Brandegee [Chamaesyce incerta (Brandegee) Acalypha californica Bentham [A. pringlei S. Watson] Millspaugh] Acalypha papillosa Rose Euphorbia leucophylla Bentham subsp. comcaacorum V.W. cinerea (Wiggins & Rollins) A. Cervantes, V.W. Steinmann & Felger Steinmann & Flores Olvera [Bernardia cinerea Wiggins & Euphorbia lomelii V.W. Steinmann [Pedilanthus macrocarpus Rollins (T)] Bentham] Adelia obovata Wiggins & Rollins Euphorbia magdalenae Bentham [Chamaesyce magdalenae adenophora A. Gray [Ditaxis adenophora (A. Gray) (Bentham) Millspaugh] Pax & K. Hoffmann] Euphorbia pediculifera Engelmann var. linearifolia S. Watson Argythamnia lanceolata (Bentham) Müller Argoviensis [A. (T) palmeri S. Watson (T). Ditaxis lanceolata (Bentham) Pax & K. Euphorbia petrina S. Watson [Chamaesyce petrina (S. Watson) Hoffmann. D. palmeri (S. Watson) Pax & K. Hoffmann] Millspaugh. Euphorbia polycarpa Bentham var. petrina (S. Argythamnia serrata (Torrey) Müller Argoviensis [A. Watson) I.M. Johnston] neomexicana Müller Argoviensis. Ditaxis gracilis Rose & Euphorbia polycarpa Bentham [Chamaesyce polycarpa (Bentham) Standley (T). D. neomexicana (Müller Argoviensis) A. Heller. Millspaugh. Euphorbia intermixta S. Watson (T). E. polycarpa D. serrata (Torrey) A. Heller] var. hirtella Boissier] Bernardia viridis Millspaugh * Aiton [Chamaesyce prostrata (Aiton) Small] 27 Felger, Carnahan, Sánchez-Escalante The Desert Edge: Flora of the Guaymas Region of Sonora, Mexico

*Euphorbia serpens Kunth [Chamaesyce serpens (Kunth) Small] Coursetia glandulosa A. Gray [Pictetia microphylla Bentham ex Euphorbia setiloba Engelmann ex Torrey [Chamaesyce setiloba Hemsley (T)] (Engelmann) Millspaugh] Dalea bicolor Humboldt & Bonpland ex Willdenow var. bicolor Euphorbia tomentulosa S. Watson [Chamaesyce tomentulosa (S. Dalea mollis Bentham Watson) Millspaugh] Dalea pringlei A. Gray var. multijuga Barneby Euphorbia trachysperma Engelmann [Chamaesyce trachysperma Dalea purpusii Brandegee (Engelmann) Millspaugh] bicornutus S. Watson Euphorbia xanti Engelmann ex Boissier Desmanthus covillei (Britton & Rose) Wiggins [Acuan Jatropha cardiophylla (Torrey) Müller Argoviensis bicornutum (S. Watson) Britton & Rose. A. subulatum Britton Jatropha cinerea (Ortega) Müller Argoviensis [J. canescens & Rose. Desmanthus subulatus (Britton & Rose) Wiggins] (Bentham) Müller Argoviensis] Desmodium procumbens (Miller) Hitchcock [D. scopulorum S. Jatropha cordata (Ortega) Müller Argoviensis Watson (T). D. wigginsii Schubert] Jatropha cuneata Wiggins & Rollins Diphysa occidentalis Rose (T) Manihot angustiloba (Torrey) Müller Argoviensis flabelliformis Kearney Manihot davisiae Croizat Erythrostemon palmeri (S. Watson) E. Gagnon & G.P. Lewis Manihot rubricaulis I.M. Johnston [Caesalpinia arida (Britton & Rose) Wiggins. C. palmeri S. Pleradenophora bilocularis (S. Watson) Esser & A.L. Melo Watson (T). Poinciana palmeri (S. Watson) Britton & Rose. P. [Sapium biloculare (S. Watson) Pax. bilocularis S. palmeri (S. Watson) Rose] Watson] Galactia wrightii A. Gray *Ricinus communis Linnaeus Galactia sp. Tragia jonesii Radcliffe-Smith & Govaerts [T. scandens M.E. Haematoxylum brasiletto H. Karsten Jones (T), not T. scandens Linnaeus] Havardia sonorae (S. Watson) Britton & Rose [Pithecellobium sonorae S. Watson (T)] FABACEAE Indigofera jamaicensis Sprengel [I. laevis Rydberg (T)] Acacia angustissima, see Acaciella angustissima *Indigofera suffruticosa Miller Acacia californica, see Vachellia californica *Leucaena leucocephala (Lamarck) de Wit subsp. glabrata Acacia cochliacantha, see Vachellia campechiana (Rose) S. Zárate Acacia constricta, see Vachellia constricta Lupinus arizonicus (S. Watson) S. Watson Acacia farnesiana, see Vachellia farnesiana Lysiloma candidum Brandegee Acacia occidentalis, see Senegalia occidentalis Lysiloma divaricatum (Jacquin) J.F. Macbride [L. microphyllum Acacia russelliana, see Mariosousa russelliana Bentham] Acacia willardiana, see Mariosousa willardiana Lysiloma watsonii Rose [L. microphyllum var. thornberi (Britton & Acaciella angustissima (Miller) Britton & Rose [Acacia Rose) Isely. L. thornberi Britton & Rose] angustissima (Miller) Kuntze. A. filicina Willdenow] Macroptilium atropurpureum (Sessé & Moçiño ex de Aeschynomene fascicularis Schlechtendal & Chamisso Candolle) Urban Albizia sinaloensis Britton & Rose Marina evanescens (Brandegee) Barneby [Dalea evanescens Astragalus nuttallianus de Candolle var. cedrosensis M.E. Brandegee. Petalostemon evanescens (Brandegee) Rose] Jones Marina parryi (Torrey & A. Gray) Barneby [Dalea parryi Torrey **Caesalpinia cacalaco, see Tara cacalaco & A. Gray] Caesalpinia palmeri, see Erythrostemon palmeri Marina vetula (Brandegee) Barneby [Dalea vetula Brandegee] *Caesalpinia pulcherrima (Linnaeus) Swartz [Poinciana Mariosousa russelliana (Britton & Rose) Seigler & Ebinger pulcherrima Linnaeus] [Acacia coulteri Bentham, in part. A. russelliana (Britton & Caesalpinia pumila, see Coulteria pumila Rose) Lundell] Calliandra californica Bentham Mariosousa willardiana (Rose) Seigler & Ebinger [Acacia Calliandra eriophylla Bentham willardiana Rose (T). Prosopis heterophylla Bentham (T)] Cercidium ×sonorae, see Parkinsonia ×sonorae *Medicago polymorpha Linnaeus Chamaecrista nictitans (Linnaeus) Moench var. leptadenia *Melilotus indicus (Linnaeus) Allioni (Greenman) Gandhi & S.L. Hatch [Cassia leptadenia Mimosa aculeaticarpa Ortega var. biuncifera (Bentham) Greenman var. mensalis Greenman. Chamaecrista nictitans var. Barneby [M. biuncifera Bentham] mensalis (Greenman) H.S. Irwin & Barneby] Mimosa distachya Cavanilles var. laxiflora (Bentham) Barneby Coulteria pumila (Britton & Rose) Sotuyo & G.P. Lewis [M. laxiflora Bentham] [Caesalpinia pumila (Britton & Rose) F.J. Hermann. Guaymasia Nissolia schottii (Torrey) A. Gray pumila Britton & Rose (T), based on Caesalpinia gracilis tesota A. Gray Bentham ex Hemsley, not C. gracilis Miquel] *Parkinsonia aculeata Linnaeus Coursetia caribaea (Jacquin) Lavin var. caribaea Parkinsonia ×carterae Hawkins 28 Desert Plants 33(1) October 2017

Parkinsonia florida (Bentham ex A. Gray) S. Watson subsp. FOUQUIERIACEAE florida [Cercidium floridum Bentham ex A. Gray subsp. Fouquieria diguetii (van Tieghem) I.M. Johnston floridum] Fouquieria macdougalii Nash Parkinsonia florida subsp. peninsulare (Rose) Hawkins & Engelmann subsp. splendens Felger [Cercidium floridum subsp. peninsulare (Rose) Carter. C. peninsulare Rose] GENTIANACEAE Parkinsonia microphylla Torrey [Cercidium microphyllum (Torrey) Zeltnera nudicaulis (Engelmann) G. Mansion [Centaurium Rose & I.M. Johnston] nudicaule (Engelmann) B.L. Robinson] Parkinsonia praecox (Ruiz & Pavón ex Hooker) Hawkins [Cercidium praecox (Ruiz & Pavón ex Hooker) Harms HELIOTROPIACEAE Parkinsonia ×sonorae (Rose & I.M. Johnston ex I.M. Johnston) Euploca convolvulacea Nuttall [E. convolvulacea subsp. californica Hawkins & Felger [Cercidium molle I.M. Johnston. C.×sonorae (Greene) Abrams. Heliotropium convolvulaceum (Nuttall) Rose & I.M. Johnston ex I.M. Johnston (T)] A. Gray. H. convolvulaceum var. californicum (Greene) I.M. Phaseolus acutifolius A. Gray var. acutifolius Johnston] Phaseolus filiformis Bentham Euploca procumbens (Miller) Diane & Hilger [Heliotropium Piscidia mollis Rose procumbens Miller] *Pithecellobium dulce (Roxburgh) Bentham Heliotropium angiospermum Murray Prosopis articulata S. Watson (T) Heliotropium curassavicum Linnaeus var. oculatum (A. Prosopis glandulosa Torrey var. torreyana (L.D. Benson) M.C. Heller) I.M. Johnston ex Tidestrom Johnston [P. juliflora (Swartz) de Candolle var. torreyana L.D. Heliotropium wigginsii I.M. Johnston (T) Benson] Myriopus volubilis (Linnaeus) Small [Tournefortia volubilis Psorothamnus emoryi (A. Gray) Rydberg var. emoryi [Dalea Linnaeus] emoryi A. Gray] Tournefortia hartwegiana Steudel Rhynchosia minima (Linnaeus) de Candolle Rhynchosia precatoria (Humboldt & Bonpland ex Willdenow) HYDROPHYLLACEAE de Candolle chrysanthemifolia (Bentham) Greene var. Senegalia occidentalis (Rose) Britton & Rose [Acacia occidentalis bipinnatifida (Torrey) Constance Rose] Eucrypta micrantha (Torrey) Heller Senna covesii (A. Gray) H.S. Irwin & Barneby [Cassia covesii A. Phacelia crenulata Torrey ex S. Watson [P. ambigua M.E. Jones. Gray] P. crenulata var. minutiflora (J.W. Voss ex Munz) Jepson] Senna pallida (Vahl) H.S. Irwin & Barneby var. shreveana H.S. Phacelia distans Bentham Irwin & Barneby Phacelia scariosa Brandegee Sesbania herbacea (Miller) McVaugh Sphinctospermum constrictum (S. Watson) Rose [Tephrosia KOEBERLINIACEAE constricta S. Watson (T)] Koeberlinia spinosa Zuccarini var. tenuispina Kearney & **Tara cacalaco (Bonpland) Molinari-Nova & Sánchez Ocharan Peebles [Caesalpinia cacalaco Bonpland] Tephrosia tenella A. Gray [T. vicioides Schlechtendal] KRAMERIACEAE Vachellia californica (Brandegee) Seigler & Ebinger [Acacia Krameria erecta Willdenow ex Schultes [K. parvifolia Bentham] californica Brandegee. A. pringlei Rose subsp. californica Krameria sonorae Britton (T) (Brandegee) Lee, Seigler & Ebinger. A. sonorensis Rose (T)] Vachellia campechiana (Miller) Seigler & Ebinger [Acacia cochliacantha Humboldt & Bonpland ex Willdenow. A. albida (Kunth) Harley & J.F.B. Pastore [ cymbispina Sprague & Riley (T)] (Torrey) Harley & J.F.B. Pastore. albida Kunth. H. Vachellia constricta (Bentham) Seigler & Ebinger [Acacia emoryi Torrey. H. emoryi var. palmeri (S. Watson) I.M. Johnston. constricta Bentham] H. palmeri S. Watson (T)] Vachellia farnesiana (Linnaeus) Wight & Arnott [Acacia Salvia lasiocephala Hooker & Arnott farnesiana (Linnaeus) Willdenow. A. minuta (M.E. Jones) Salvia misella Kunth [S. privoides Bentham. S. riparia Kunth] Beauchamp subsp. densiflora (Alexander ex Small) Salvia similis Brandegee Beauchamp. A. smallii Isely] Stachys coccinea Ortega [S. coccinea Jacquin] Zapoteca formosa (Kunth) H.M. Hernández subsp. rosei Teucrium cubense Jacquin subsp. depressum (Small) (Wiggins) H.M. Hernández [Calliandra rosei Wiggins. C. schottii McClintock & Epling [T. cubense var. densum Jepson. T. subsp. rosei (Wiggins) Felger & Lowe] depressum Small] Vitex mollis Kunth **Vitex trifolia Linnaeus 29 Felger, Carnahan, Sánchez-Escalante The Desert Edge: Flora of the Guaymas Region of Sonora, Mexico

LOASACEAE Corchorus hirtus Linnaeus [C. orinocensis Kunth. C. hirtus var. Eucnide cordata Kellogg ex Curran orinocensis (Kunth) K. Schumann] Eucnide rupestris (Baillon) H.J. Thompson & W.R. Ernst **Corchorus olitorius Linnaeus [Loasella rupestris Baillon (T). Sympetaleia rupestris (Baillon) A. Gossypium davidsonii Kellogg Gray ex S. Watson] Gossypium turneri Fryxell (T) Mentzelia adhaerens Bentham Guazuma ulmifolia Lamarck Mentzelia aspera Linnaeus Helicteres baruensis Jacquin Heliocarpus attenuatus S. Watson LORANTHACEAE Herissantia crispa (Linnaeus) Brizicky Psittacanthus calyculatus (de Candolle) G. Don Hermannia pauciflora S. Watson Psittacanthus sonorae (S. Watson) Kuijt [Loranthus sonorae S. Hibiscus biseptus S. Watson Watson (T). Phrygilanthus sonorae (S. Watson) Standley] Bentham Struthanthus palmeri Kuijt [S. haenkei (Presl) Engler sensu lato, Hibiscus phoeniceus Jacquin in part. S. haenkei var. angustatus I.M. Johnston (T)] Horsfordia newberryi (S. Watson) A. Gray Kosteletzkya hispidula (Sprengel) Garcke [K. coulteri A. Gray. K. digitata A. Gray (T)] robusta Heer & Regel *Malva parviflora Linnaeus Malvastrum bicuspidatum (S. Watson) Rose subsp. bicuspidatum Callaeum macropterum (Moçiño & Sessé ex de Candolle) Malvella leprosa (Ortega) Krapovickas D.M. Johnson [ macroptera (Moçiño & Sessé ex de Melochia tomentosa Linnaeus [M. arida Rose (T). M. speciosa Candolle) Niedenzu] S. Watson (T). M. tomentosa var. speciosa (S. Watson) A. Cottsia californica (Bentham) W.R. Anderson & C. Davis [C. Goldberg] scandens Dubard & Dop (T). Janusia californica Bentham] Pseudabutilon thurberi (A. Gray) Fryxell [ thurberi A. Cottsia gracilis (A. Gray) W.R. Anderson & C. Davis [Janusia Gray. Pseudabutilon sonorae Wiggins] gracilis A. Gray] abutifolia Miller Cottsia linearis (Wiggins) W.R. Anderson & C. Davis [Janusia Sida acuta Burman f. linearis Wiggins] *Sida ciliaris Linnaeus Echinopterys eglandulosa (A. de Jussieu) Small [Bunchosia Sida glabra Miller eglandulosa A. de Jussieu] Sida hyalina Fryxell Galphimia angustifolia Bentham [G. angustifolia var. oblongifolia Sphaeralcea ambigua A. Gray subsp. ambigua (A. Gray) S. Watson (T). Thryallis angustifolia (Bentham) Sphaeralcea coulteri (S. Watson) A. Gray Kuntze] Sphaeralcea emoryi Torrey ex A. Gray [S. arida Rose (T)] Malpighia watsonii Rose [Bunchosia parvifolia S. Watson (T), not **Talipariti tiliaceum (Linnaeus) Fryxell var. tiliaceum [Hibiscus Malpighia parvifolia A. de Jussieu. Malpighia sonorae M.E. Jones tiliaceus Linnaeus] (T)] Waltheria indica Linnaeus

MALVACEAE (includes Bombacaceae, Sterculiaceae, and MARTYNIACEAE Tiliaceae) Proboscidea altheifolia (Bentham) Decaisne Abutilon abutiloides (Jacquin) Garcke ex Hochreutiner [A. Proboscidea parviflora (Wooton) Wooton & Standley subsp. scabrum S. Watson (T)] sinaloensis (Van Eseltine) Bretting [P. sinaloensis Van Abutilon californicum Bentham Eseltine] (Link) Sweet [A. pringlei Hochreutiner] Abutilon mollicomum (Willdenow) Sweet A. Gray (T) [A. aurantiacum S. Watson, not A. diversifolius de Candolle aurantiacum Linden ex Turczaninow] S. Watson MOLLUGINACEAE Abutilon reventum S. Watson *Glinus lotoides Linnaeus Anoda crenatiflora Ortega *Glinus radiatus (Ruiz & Pavón) Rohrbach Anoda pentaschista A. Gray *Mollugo cerviana (Linnaeus) Seringe Ayenia filiformis S. Watson (T) [A. compacta Rose. A pusilla of Mollugo verticillata Linnaeus various authors, not A. pusilla Linnaeus] Ayenia jaliscana S. Watson Briquetia sonorae Fryxell (T) insipida Willdenow [F. radulina S. Watson] Ceiba acuminata (S. Watson) Rose 30 Desert Plants 33(1) October 2017

Ficus palmeri S. Watson [F. petiolaris Kunth subsp. palmeri (S. ochroleuca Sweet Watson) Felger & Lowe] Ficus pertusa Linnaeus f. [F. fasciculata S. Watson (T), not F. PASSIFLORACEAE fasciculata F. Mueller ex Bentham. F. padifolia Kunth. F. sonorae Passiflora arida (Masters & Rose) Killip [P. foetida Linnaeus var. S. Watson (T)] arida Masters & Rose (T)] Passiflora arizonica (Killip) D.H. Goldman [P. foetida var. NAMACEAE arizonica Killip] Nama hispida A. Gray var. sonorae C.L. Hitchcock Passiflora mexicana de Jussieu Nama jamaicensis Linnaeus Turnera diffusa Willdenow var. diffusa Nama stenocarpa A. Gray PEDALIACEAE NYCTAGINACEAE **Sesamum indicum Linnaeus [S. orientale Linnaeus] Abronia maritima Nuttall ex S. Watson subsp. maritima Allionia incarnata Linnaeus [A. incarnata var. nudata (Standley) PETIVERIACEAE Munz. A. incarnata var. villosa (Standley) B.L. Turner] Rivina humilis Linnaeus Boerhavia alata S. Watson (T) Boerhavia coccinea Miller [B. caribaea Jacquin] PHRYMACEAE Boerhavia erecta Linnaeus Erythranthe floribunda (Douglas ex Lindley) G.L. Nesom Boerhavia gracillima Heimerl [Mimulus floribundus Douglas ex Lindley] Boerhavia spicata Choisy [B. coulteri (Hooker f.) S. Watson. B. palmeri S. Watson, 1889 (T), not B. palmeri S. Watson, 1883. B. PHYLLANTHACEAE spicata Choisy var. palmeri S. Watson (T). B. watsonii Standley Andrachne microphylla (Lamarck) Baillon [A. ciliatoglandulosa (T). Senkenbergia coulteri Hooker f. (T)] (Millspaugh) Croizat] Boerhavia triquetra S. Watson [B. intermedia M.E. Jones. B. Phyllanthus evanescens Brandegee maculata Standley (T). B. triquetra var. intermedia (M.E. Jones) Spellenberg] PHYTOLACCACEAE, Rivina, see PETIVERIACEAE Boerhavia wrightii A. Gray Boerhavia xanti S. Watson (includes Scrophulariaceae, in part) Commicarpus scandens (Linnaeus) Standley terrestris Rafinesque C.[ deflexa A. Braun ex Pisonia capitata (S. Watson) Standley [Cryptocarpus capitatus S. Hegelmaier] Watson (T)] Gambelia juncea (Bentham) D.A. Sutton [Galvezia juncea Salpianthus macrodontus Standley (Bentham) Ball] Mecardonia procumbens (Miller) Small [Bacopa procumbens (Mill- OLACACEAE, see SCHOEPFIACEAE and XIMENIACEAE er) Greenman. Mecardonia vandellioides (Kunth) Pennell] Nuttallanthus texanus (Scheele) D.A. Sutton [Linaria canadensis OLEACEAE (Linnaeus) Dumont de Courset var. texana (Scheele) Pennell] Forestiera angustifolia Torrey Plantago ovata Forsskål var. fastigiata (Morris) S.C. Meyers Menodora scabra A. Gray [M. scoparia Engelmann ex A. Gray] & Liston [P. fastigiata E. Morris. P. insularis Eastwood var. fastigiata (E. Morris) Jepson] ONAGRACEAE Pseudorontium cyathiferum (Bentham) Rothmaler [Antirrhinum Eremothera chamaenerioides (A. Gray) W.L. Wagner & Hoch cyathiferum Bentham] [Camissonia chamaenerioides (A. Gray) P.H. Raven. Oenothera Russelia sonorensis Carlson var. pubescens Carlson chamaenerioides A. Gray] Sairocarpus costatus (Wiggins) D.A. Sutton [Antirrhinum Ludwigia octovalvis (Jacquin) P.H. Raven subsp. octovalvis costatum Wiggins (T)] Ludwigia peploides (Kunth) P.H. Raven Sairocarpus watsonii (Vasey & Rose) D.A. Sutton [Antirrhinum Oenothera arizonica (Munz) W.L. Wagner [O. avita (W.M. Klein) kingii S. Watson var. watsonii (Vasey & Rose) Munz. A. watsonii W.M. Klein subsp. arizonica (Munz) W.M. Klein. O. californica Vasey & Rose] S. Watson subsp. arizonica (Munz) W.M. Klein] Schistophragma intermedium (A. Gray) Pennell [Conobea Oenothera kunthiana (Spach) Munz intermedia A. Gray] Schistophragma polystachyum (Brandegee) B.L. Turner OXALIDACEAE [Conobea polystachya (Brandegee) Minod. Stemodia polystachya *Oxalis sp. Brandegee] Stemodia durantifolia (Linnaeus) Swartz var. durantifolia Stemodia palmeri A. Gray Argemone gracilenta Greene [A. intermedia Sweet] 31 Felger, Carnahan, Sánchez-Escalante The Desert Edge: Flora of the Guaymas Region of Sonora, Mexico

Veronica peregrina Linnaeus subsp. xalapensis (Kunth) Ziziphus amole (Sessé & Moçiño) M.C. Johnston [Z. sonorensis S. Pennell Watson (T)] Ziziphus obtusifolia (Hooker ex Torrey & A. Gray) A. Gray var. PLUMBAGINACEAE canescens (A. Gray) M.C. Johnston [Condaliopsis lycioides (A. Plumbago zeylanica Linnaeus [P. scandens Linnaeus] Gray) Suessenguth var. canescens (A. Gray) Suessenguth]

POLEMONIACEAE RHIZOPHORACEAE Dayia sonorae (Rose) J.M. Porter [Gilia sonorae Rose (T). Rhizophora mangle Linnaeus Ipomopsis sonorae (Rose) A. Grant] RUBIACEAE POLYGALACEAE Chiococca petrina Wiggins Hebecarpa macradenia (A. Gray) J.R. Abbott [Polygala macrade- Diodia teres Walter nia A. Gray] microphyllum A. Gray Galium proliferum A. Gray POLYGONACEAE Hintonia latiflora (de Candolle) Bullock [Coutarea latiflora de Antigonon leptopus Hooker & Arnott Candolle. C. pterosperma (S. Watson) Standley. Portlandia Coccoloba goldmanii Standley pterosperma S. Watson (T)] *Persicaria cf. maculosa Gray *Oldenlandia corymbosa Linnaeus *Polygonum argyrocoleon Steudel ex Kunze Randia obcordata S. Watson (T) Rumex inconspicuus Rechinger f. Randia sonorensis Wiggins Randia thurberi S. Watson (T) PORTULACACEAE Stenaria sanchezii Lorence (T) Portulaca halimoides Linnaeus [P. parvula A. Gray] Portulaca johnstonii Henrickson RUTACEAE *Portulaca oleracea Linnaeus [P. retusa Engelmann, perhaps not Amyris balsamifera Linnaeus P. retusa of western authors] Esenbeckia hartmanii B.L. Robinson & Fernald Portulaca suffrutescens Engelmann Zanthoxylum fagara (Linnaeus) Sargent [Z. sonorense Lundell] Portulaca umbraticola Kunth subsp. lanceolata (Engelmann) Zanthoxylum mazatlanum Sandwith J.F. Matthews & Ketron [P. lanceolata Engelmann] SALICACEAE PRIMULACEAE Populus mexicana de Candolle subsp. dimorpha (Brandegee) *Anagallis arvensis Linnaeus Eckenwalder [P. dimorpha Brandegee] Bonellia macrocarpa (Cavanilles) B. Ståhl & Källersjö subsp. Salix gooddingii C.R. Ball pungens (A. Gray) B. Ståhl & Källersjö [Jacquinia macrocarpa Cavanilles subsp. pungens (A. Gray) B. Ståhl. J. pungens A. SANTALACEAE Gray] Phoradendron brachystachyum (de Candolle) Oliver [P. Samolus parviflorus Rafinesque S.[ floribundus Kunth] brachystachyum (de Candolle) Nuttall. P. diguetianum van Tieghem. P. globuliferum Trelease (T)] Phoradendron californicum Nuttall drummondii Torrey & A. Gray SAPINDACEAE RESEDACEAE Cardiospermum corindum Linnaeus Forchhammeria watsonii Rose (T) Dodonaea viscosa Jacquin [D. angustifolia Linnaeus f. D. viscosa Oligomeris linifolia (Vahl) J.F. Macbride var. angustifolia (Linnaeus f.) Bentham] Paullinia sonorensis S. Watson (T) Sapindus saponaria Linnaeus californica I.M. Johnston Serjania palmeri S. Watson (T) Colubrina triflora Brongniart ex G. Don Colubrina viridis (M.E. Jones) M.C. Johnston [C. glabra S. SAPOTACEAE Watson (T)] Sideroxylon occidentale (Hemsley) T.D. Pennington [Bumelia globosa I.M. Johnston var. globosa occidentalis Hemsley] Condalia globosa var. pubescens I.M. Johnston Gouania rosei Wiggins SCHOEPFIACEAE Karwinskia humboldtiana (Roemer & Schultes) Zuccarini [K. Schoepfia shreveana Wiggins (T) parvifolia Rose] 32 Desert Plants 33(1) October 2017

SIMMONDSIACEAE THEOPHRASTACEAE, see PRIMULACEAE Simmondsia chinensis (Link) C.K. Schneider TROPAEOLACEAE SOLANACEAE **Tropaeolum majus Linnaeus Calibrachoa parviflora (de Jussieu) D’Arcy [Petunia parviflora de Jussieu] URTICACEAE Capsicum annuum Linnaeus var. glabriusculum (Dunal) Parietaria hespera B.D. Hinton var. hespera Heiser & Pickersgill [C. annuum var. aviculare (Dierbach) D’Arcy & Eshbaugh, nom. illegit. C. indicum Dierbach forma aviculare Dierbach] Aloysia sonorensis Moldenke (T) Datura discolor Bernhardi Bouchea dissecta S. Watson (T) A. Gray var. andersonii [L. andersonii var. flabellifolium S. Watson (T) deserticola (C.L. Hitchcock) Jepson. L. andersonii var. pubescens Glandularia gooddingii (Briquet) Solbrig [ gooddingii S. Watson. L. andersonii var. wrightii A. Gray] Briquet. V. gooddingii var. nepetifolia Tidestrom] Lycium berlandieri Dunal var. longistylum C.L. Hitchcock Lantana camara Linnaeus Lycium berlandieri var. peninsulare (Brandegee) C.L. Lantana hispida Kunth [L. velutina M. Martens & Galeotti] Hitchcock Lippia palmeri S. Watson (T) [L. palmeri var. spicata Rose] Lycium brevipes Bentham var. brevipes [L. palmeri A. Gray (T). Phyla nodiflora (Linnaeus) Greene [Lippia nodiflora (Linnaeus) L. richii A. Gray] Michaux. Phyla nodiflora var. incisa (Small) Moldenke] Lycium californicum Nuttall ex A. Gray subsp. carinatum (S. Verbena menthifolia Bentham [V. officinalis Linnaeus subsp. halei Watson) Felger & C.H. Lowe [L. carinatum S. Watson (T)] (Small) S. Barber. V. halei Small] Lycium exsertum A. Gray Lycium fremontii A. Gray var. fremontii VIOLACEAE Lycium macrodon A. Gray var. dispermum (Wiggins) F. Hybanthus fruticulosus (Bentham) I.M. Johnston [H. fruticulosus Chiang [L. dispermum Wiggins (T)] var. flavescens (Dowell) I.M. Johnston (T)] Nicotiana clevelandii A. Gray Hybanthus mexicanus Gingins *Nicotiana glauca Graham var. glauca Nicotiana obtusifolia M. Martens & Galeotti [N. palmeri A. Gray. N. trigonophylla Dunal. N. trigonophylla var. palmeri (A. mexicana de Candolle [C. mayoensis Gentry] Gray) M.E. Jones] Cissus trifoliata (Linnaeus) Linnaeus Petunia parviflora, see Calibrachoa parviflora Cissus verticillata (Linnaeus) Nicolson & C.E. Jarvis [C. sicyoides Physalis acutifolia (Miers) Sandwith [P. wrightii A. Gray] Linnaeus. Viscum verticillatum Linnaeus] Physalis crassifolia Bentham var. crassifolia [P. crassifolia var. versicolor (Rydberg) Waterfall. P. versicolor Rydberg (T). P. XIMENIACEAE versicolor var. microphylla Rydberg (T)] Ximenia glauca (DeFilipps) Bentouil [X. parviflora Bentham var. *Physalis pubescens Linnaeus [P. pubescens var. integrifolia (Dunal) glauca DeFilipps] Waterfall] Physalis purpurea Wiggins (T) ZYGOPHYLLACEAE *Solanum americanum Miller [S. nodiflorum Jacquin] Fagonia barclayana (Bentham) Rydberg [sensu Wiggins 1964. *Solanum elaeagnifolium Cavanilles F. californica Bentham var. barclayana Bentham, but not as to Solanum erianthum D. Don [S. verbascifolium of authors, not type] Linnaeus] Guaiacum coulteri A. Gray [G. palmeri Vail (T)] Solanum hindsianum Bentham Kallstroemia californica (S. Watson) Vail *Solanum lycopersicum Linnaeus [Lycopersicum esculentum Miller] Kallstroemia grandiflora Torrey ex A. Gray (de Candolle) Coville var. tridentata [L. STEGNOSPERMATACEAE divaricata Cavanilles subsp. tridentata (de Candolle) Felger & Stegnosperma halimifolium Bentham [S. watsonii D.J. Rogers] C.H. Lowe] *Tribulus terrestris Linnaeus TALINACEAE Viscainoa geniculata (Kellogg) Greene var. geniculata Talinum paniculatum (Jacquin) Gaertner MONOCOTS TAMARICACEAE ALISMATACEAE *Tamarix aphylla (Linnaeus) H. Karsten Echinodorus berteroi (Sprengel) Fassett [Echinodorus rostratus *Tamarix chinensis Loureiro [T. ramosissima Ledebour] (Nuttall) Engelmann] Limnocharis flava (Linnaeus) Buchenau [Alisma flava Linnaeus] 33 Felger, Carnahan, Sánchez-Escalante The Desert Edge: Flora of the Guaymas Region of Sonora, Mexico

Sagittaria longiloba Engelmann ex J.G. Smith Cyperus flavicomus Michaux [C. albomarginatus (Martius & Sagittaria montevidensis Chamisso & Schlechtendal subsp. Schrader ex Nees) Steudel] calycina (Engelmann) Bogin *Cyperus iria Linnaeus Cyperus ochraceus Vahl AMARYLLIDACEAE Cyperus odoratus Linnaeus [C. ferax Richard. Mariscus ferax Hymenocallis sonorensis Standley (Richard) C.B. Clarke. Torulinium odoratum (Linnaeus) S.S. Nothoscordum bivalve (Linnaeus) Britton Hooper] *Cyperus rotundus Linnaeus ARACEAE (includes Lemnaceae) Cyperus semiochraceus Boeckeler Lemna aequinoctialis Welwitsch Cyperus squarrosus Linnaeus [C. aristatus Rottbøll. Mariscus Lemna sp. squarrosus (Linnaeus) C.B. Clarke] Cyperus subsquarrosus (Muhlenberg) Bauters [Hemicarpha ARECACEAE micrantha (Vahl) Pax. Lipocarpha micrantha (Vahl) G.C. Tucker] Brahea brandegeei (Purpus) H.E. Moore [Brahea clara (L.H. Eleocharis geniculata (Linnaeus) Roemer & Schultes [E. capitata Bailey) Espejo & López-Ferrari. B. elegans (Franceschi ex R. Brown. E. caribaea (Rottbøll) S.F. Blake] Beccari) H.E. Moore. B. roezlii sensu Wiggins, not B. roezlii Eleocharis macrostachya Britton Linden. Erythea brandegeei Purpus. E. clara L.H. Bailey (T). E. Eleocharis montana (Kunth) Roemer & Schultes [E. montana elegans Franceschi ex Beccari] var. nodulosa (Roth) Svenson] **Phoenix dactylifera Linnaeus Eleocharis montevidensis Kunth Sabal uresana Trelease Fimbristylis complanata (Retzius) Link Washingtonia robusta H. Wendland [W. sonorae S. Watson (T)] Fimbristylis dichotoma (Linnaeus) Vahl [F. annua (Allioni) Roemer & Schultes. F. laxa Vahl] ASPARAGACEAE (includes Agavaceae and Nolinaceae) Fuirena simplex Vahl var. simplex Agave angustifolia Haworth var. angustifolia [A. owenii I.M. Schoenoplectus americanus (Persoon) Volkart ex Schinz & Johnston (T). A. pacifica Trelease. A. yaquiana Trelease] R. Keller [Scirpus americanus Persoon. S. olneyi A. Gray of Agave angustifolia × A. colorata western authors] Agave chrysoglossa I.M. Johnston (T) Agave colorata Gentry JUNCACEAE Agave felgeri Gentry (T) Juncus bufonius Linnaeus Dasylirion gentryi Bogler POACEAE ASPHODELACEAE Andropogon glomeratus (Walter) Britton, Sterns & Poggenburg **Aloe vera (Linnaeus) Burman f. var. scabriglumis C.S. Campbell Aristida adscensionis Linnaeus BROMELIACEAE Aristida californica Thurber var. californica Hechtia montana Brandegee Aristida divaricata Humboldt & Bonpland ex Willdenow [A. Tillandsia exserta Fernald palmeri Vasey] Tillandsia recurvata Linnaeus Aristida ternipes Cavanilles var. gentilis (Henrard) Allred [A. gentilis Henrard. A. hamulosa Henrard. A. ternipes var. hamulosa COMMELINACEAE (Henrard) Trent] Commelina erecta Linnaeus Aristida ternipes var. ternipes *Arundo donax Linnaeus CYMODOCEACEAE *Avena fatua Linnaeus Halodule wrightii Ascherson Bothriochloa barbinodis (Lagasca) Herter aristidoides (Kunth) Grisebach [B. aristidoides var. CYPERACEAE arizonica M.E. Jones] Bolboschoenus maritimus (Linnaeus) Palla [Scirpus maritimus Bouteloua barbata Lagasca var. barbata [B. arenosa Vasey (T). B. Linnaeus] barbata var. sonorae (Griffiths) Gould.B. sonorae Griffiths (T)] Cyperus articulatus Linnaeus Bouteloua diversispicula J.T. Columbus [Cathestecum brevifolium Cyperus canus J. Presl & C. Presl Swallen] Cyperus dentoniae G. Tucker [Mariscus pubescens J. Presl & C. Bouteloua parryi (E. Fournier) Griffiths B.[ gentryi Gould] Presl] (Kunth) Scribner & Merrill [B. filiformis (E. Cyperus elegans Linnaeus Fournier) Griffiths] Cyperus entrerianus Boeckeler Brachiaria reptans, see Urochloa reptans Cyperus esculentus Linnaeus var. leptostachyus Boeckeler *Cenchrus ciliaris Linnaeus [Pennisetum ciliare (Linnaeus) Link] 34 Desert Plants 33(1) October 2017

*Cenchrus echinatus Linnaeus [C. insularis Scribner] Eragrostis pectinacea (Michaux) Nees var. pectinacea [E. *Cenchrus longispinus (Hackel) Fernald pectinacea var. miserrima (E. Fournier) Reeder] *Cenchrus myosuroides Kunth var. myosuroides *Eragrostis pilosa (Linnaeus) P. Beauvois var. pilosa Cenchrus palmeri Vasey (T) Eragrostis spicata Vasey *Cenchrus setaceus (Forsskål) Morrone [Pennisetum ruppelii Eriochloa acuminata (J. Presl) Kunth var. acuminata [E. Steudel. P. setaceum (Forsskål) Chiovenda] acuminata var. minor (Vasey) R.B. Shaw. E. lemmonii Vasey & *Cenchrus spinifex Cavanilles Scribner var. gracilis (E. Fournier) Gould] *Chloris barbata Swartz [C. inflata Link] Eriochloa aristata Vasey var. aristata Chloris crinita, see crinita Heteropogon contortus (Linnaeus) P. Beauvois ex Roemer & Chloris virgata Swartz Schultes Cottea pappophoroides Kunth Hilaria belangeri (Steudel) Nash [H. belangeri var. longifolia * dactylon (Linnaeus) Persoon var. dactylon (Vasey) Hitchcock. H. cenchroides Kunth var. ciliata Scribner. *Dactyloctenium aegyptium (Linnaeus) Willdenow H. cenchroides var. longifolia Vasey (T). H. ciliata (Scribner) *Digitaria bicornis (Lamarck) Roemer & Schultes Nash] Digitaria californica (Bentham) Henrard var. californica Jouvea pilosa (J. Presl) Scribner [Trichachne californica (Bentham) Chase] Lasiacis ruscifolia (Kunth) Hitchcock var. ruscifolia *Digitaria ciliaris (Retzius) Koeler [D. adscendens (Kunth) Leptochloa crinita (Lagasca) P.M. Peterson & N. Snow [Chloris Henrard. D. sanguinalis (Linnaeus) Scopoli var. ciliaris (Retzius) crinita Lagasca. Trichloris crinita (Lagasca) Parodi. T. mendocina Parlatore] (Philippe) Kurtz] Digitaria insularis (Linnaeus) Mez ex Ekman [Trichachne insularis Leptochloa dubia, see Disakisperma dubium (Linnaeus) Nees] Leptochloa fusca, see fusca panicea (Retzius) P.M. Peterson & N. Snow subsp. Leptochloa panicea, see Dinebra panicea brachiata (Steudel) P.M. Peterson & N. Snow [Leptochloa *Leptochloa panicoides, see Dinebra panicoides brachiata Steudel. L. filiformis (Persoon) P. Beauvois. L. panicea Leptochloa viscida, see Dinebra viscida (Retzius) Ohwi subsp. brachiata (Steudel) N. Snow] *Lolium perenne Linnaeus [Festuca perennis (Linnaeus) Columbus *Dinebra panicoides (J. Presl) P.M. Peterson & N. Snow [Lep- & J.P. Smith] tochloa panicoides (J. Presl) Hitchcock] Luziola gracillima Prodoehl Dinebra viscida (Scribner) P.M. Peterson & N. Snow [Leptochloa *Melinis repens (Willdenow) Zizka subsp. repens [Rhynchelytrum viscida (Scribner) Beal] repens (Willdenow) C.E. Hubbard. R. roseum (Nees) Stapf & Diplachne fusca (Linnaeus) P. Beauvois ex Roemer & Schultes C.E. Hubbard] subsp. fascicularis (Lamarck) P.M. Peterson & N. Snow Monanthochloe littoralis, see Distichlis littoralis [Leptochloa fusca (Linnaeus) Kunth subsp. fascicularis (Lamarck) Muhlenbergia microsperma (de Candolle) Trinius N. Snow] Muhlenbergia rigens (Bentham) Hitchcock Diplachne fusca subsp. uninervia (J. Presl) P.M. Peterson & N. Panicum alatum Zuloaga & Morrone var. alatum Snow [Leptochloa fusca subsp. uninervia (J. Presl) N. Snow. L. Panicum alatum var. minus (Andersson) Zuloaga & Morrone uninervia (J. Presl) Hitchcock & Chase] [P. hirticaule var. minus Andersson] Disakisperma dubium (Kunth) P.M. Peterson & N. Snow [Lep- Panicum hians, see Steinchisma hians tochloa dubia (Kunth) Nees] Panicum hirticaule J. Presl var. hirticaule [P. hirticaule Distichlis littoralis (Engelmann) H.L. Bell & Columbus [Monan- var. miliaceum (Vasey) Beetle. P. hirticaule var. pampinosum thochloe littoralis Engelmann] (Hitchcock & Chase) Beetle. P. hirticaule subsp. sonorum (Beal) Distichlis spicata (Linnaeus) Greene [D. stricta (Torrey) Rydberg] Freckmann & Lelong. P. pampinosum Hitchcock & Chase. P. *Echinochloa colona (Linnaeus) Link var. colona sonorum Beal] *Echinochloa crus-galli (Linnaeus) P. Beauvois var. crus-galli Panicum hirticaule var. stramineum (Hitchcock & Chase) [E. crus-galli var. mitis (Pursh) Peterman] Beetle [P. capillare Linnaeus var. stramineum (Hitchcock & Echinochloa crus-pavonis (Kunth) Schultes var. macera Chase) Gould. P. stramineum Hitchcock & Chase (T)] (Wiegand) Gould Pappophorum philippianum Parodi Echinochloa muricata (P. Beauvois) Fernald var. microstachya Paspalidium geminatum, see Setaria geminata Wiegand Paspalum distichum Linnaeus * indica (Linnaeus) C.F. Gaertner Paspalum hartwegianum E. Fournier chlorideus (J. Presl) Clayton [Chloris brandegeei Paspalum langei (E. Fournier) Nash (Vasey) Swallen. C. chloridea (J. Presl) Hitchcock. Gouinia *Pennisetum ciliare, see Cenchrus ciliaris brandegeei (Vasey) Hitchcock] *Pennisetum setaceum, see Cenchrus setaceus *Eragrostis barrelieri Daveau Phragmites australis (Cavanilles) Trinius ex Steudel subsp. *Eragrostis cilianensis (Allioni) Vignolo-Lutati ex Janchen berlandieri (E. Fournier) Saltonstall & Hauber [P. communis Trinius] 35 Felger, Carnahan, Sánchez-Escalante The Desert Edge: Flora of the Guaymas Region of Sonora, Mexico

*Poa annua Linnaeus MALVACEAE Sporobolus airoides (Torrey) Torrey subsp. airoides Horsfordia rotundifolia S. Watson Sporobolus contractus Hitchcock Sporobolus coromandelianus (Retzius) Kunth [S. patens RUBIACEAE Swallen. S. pulvinatus Swallen. S. pyramidatus (Lamarck) Randia echinocarpa de Candolle Hitchcock] Sporobolus cryptandrus (Torrey) A. Gray POACEAE Sporobolus flexuosus (Thurber ex Vasey) Rydberg [S. cryptandrus Enneapogon desvauxii P. Beauvois var. flexuosus Thurber ex Vasey] Munroa pulchella (Kunth) L.D. Amarilla [Dasyochloa pulchella (Kunth) Sporobolus virginicus (Linnaeus) Kunth Willdenow ex Rydberg] Steinchisma hians (Elliott) Nash [Panicum hians Elliott] **Stenotaphrum secundatum (Walter) Kuntze Trichloris crinita, see Leptochloa crinita ACKNOWLEDGEMENTS Tripsacum lanceolatum Ruprecht ex E. Fournier **Triticum aestivum Linnaeus Assistance from the University of Arizona Herbarium (ARIZ) Urochloa arizonica (Scribner & Merrill) Morrone & Zuloaga and Herbario de la Universidad de Sonora (USON) has made [Brachiaria arizonica (Scribner & Merrill) S.T. Blake. Panicum this work possible. We thank the staff of herbaria that provid- arizonicum Scribner & Merrill. P. dissitiflorum Vasey (T)] ed loans or images of specimens, including CAS, DES, MEXU, Urochloa fusca (Swartz) B.F. Hansen & Wunderlin [Brachiaria MO, NMC, RSA, SD, TEX, UC, and US. Online resources from fasciculata (Swartz) Parodi] Southwest Environmental Information Network (SEINet), Trop- Urochloa reptans (Linnaeus) Stapf [Brachiaria reptans (Linnaeus) icos, and the San Diego Natural History Museum have been es- C.A. Gardner & C.E. Hubbard] pecially valuable. We thank all, including the late Daniel F. Aus- tin, Alberto Búrquez-Montijo, Thomas F. Daniel, Christopher PONTEDERIACEAE J.S. Davis, George McNeil Ferguson, Walter Frank Fertig, Mark *Eichhornia crassipes (C. Martius) Solms-Laubach [Pontederia Fishbein, Judy Ann Gibson, Wendy C. Hodgson, Matthew B. crassipes C. Martius] Johnson, Kirsten Lake, Max Licher, Michelle “Shelley” McMa- Heteranthera limosa (Swartz) Willdenow hon, Guy L. Nesom, Jon P. Rebman, Ana Lilia Reina-, Andrew Michael Salywon, Andrew C. Sanders, Silke Schneider, POTAMOGETONACEAE Curtis Latham Smith, Victor W. Steinmann, Gordon C. Tucker, Zannichellia palustris Linnaeus Thomas R. Van Devender, James (Jim) Thomas Verrier, Benja- min T. Wilder, and George Yatskievych. Richard Felger extends RUPPIACEAE gratitude to the Wallace Research Foundation for earlier support. Ruppia maritima Linnaeus

TYPHACEAE Typha domingensis Persoon

ZOSTERACEAE Zostera marina Linnaeus var. atam T.W.H. Backman

TAXA DOUBTFUL AND EXCLUDED FROM THE GUAYMAS REGION FLORA

PTERIDACEAE Notholaena californica de Candolle [Cheilanthes deserti Mickel]

ASTERACEAE Malperia tenuis S. Watson

FABACEAE Tephrosia palmeri S. Watson Vachellia pennatula (Schlechtendal & Chamisso) Seigler & Ebinger [Acacia pennatula (Schlechtendal & Chamisso) Bentham] 36 Desert Plants 33(1) October 2017

LITERATURE CITED Angiosperm Phylogeny Group. 2009. An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG III. Bot. J. Linn. Soc. 161: 105–121. DOI: 10.1111/j.1095- 8339.2009.00996 Angiosperm Phylogeny Group. 2016. An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG IV. Bot. J. Linn. Soc. 181: 1–20. DOI: 10.1111/boj.12385 Felger, R.S., S.D. Carnahan, and J.J. Sánchez-Escalante. 2017. Oasis at the Desert Edge: Flora of Cañón del Nacapule, Sonora, Mexico. Proceedings of the Desert Laboratory, Univ. of Arizona, vol. 1 (in press). Felger, R.S., S.D. Carnahan, J.J. Sánchez-Escalante, M. Bogan, A. Búrquez-Montijo, and F.S. Molina. 2016. The Desert Edge: Flora and ethnobotany of the Guaymas region of Sonora, Mexico. The Press, The Arizona Native Plant Society 39 (1): 1–6. Felger, R.S., B.T. Wilder, and J.P. Gallo-Reynoso. 2011. Floristic diversity and long- term vegetation dynamics of Isla San Pedro Nolasco, Gulf of Califor- nia, Mexico. Proceed. San Diego Soc. Nat. Hist. 43: 1–42. Felger, R.S. and B.T. Wilder in collaboration with H. Romero-Morales. 2012. Plant Life of a Desert Archipelago: Flora of the Sonoran Islands in the Gulf of . Univ. of Arizona Press, Tucson. McVaugh, R. 1956. Edward Palmer: Plant Explorer of the American West. Univ. of Oklahoma Press, Norman. Shreve, F. 1951. Vegetation of the Sonoran Desert. Publ. Carnegie Inst. Washing- ton No. 591. Reprinted as vol. 1, part 1: 1-186 + 37 plates in F. Shreve and I.L. Wiggins, Vegetation and flora of the Sonoran Desert. Stanford Univ. Press, Stanford. Southwest Environmental Information Network (SEINet). 2017. Stevens, P.F. 2012 (onward). Angiosperm Phylogeny Website, version 12, July 2012 onward. Watson, S. 1889. Upon a collection of plants made by Dr. E. Palmer, in 1887, about Guaymas, Mexico, at Muleje and Los Angeles Bay, in Lower California, and on the Island of San Pedro Martin, in the Gulf of California. Proc. Am. Acad. Arts Sci. 24: 36–87.