IV EUROPEAN WORKSHOP ON VERTEBRATE PALAEONTOLOGY

ALBARRACIN (Teruel, Spain) JUNIO DE 1999

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Programme and Abstracts

Field guide

Edited b y

José Ignacio Canudo Gloria Cuenca-Bescós Universidad de Zaragoza Zaragoza, Spain IV EUROPEAN WORKSHOP ON VERTEBRATE PALAEONTOLOGY (1999) ALBARRACIN, SPAIN 81

DINOSAUR REMAINS IN THE BARREMIAN (LOWER CRETACEOUS) OF THE TERUEL PROVINCE (IBERIAN CHAIN, NE SPAIN)

José I. RUIZ-OMEÑACA, Gloria CUENCA-BESCOS & José I. CANUDO Departamento de Ciencias de la Tierra, Paleontología, Universidad de Zaragoza, 50009 Zaragoza, SPAIN. e-mail: [email protected]

Dinosaur remains are frequent in the region of Aragon. There is fossil record in the Tithonian-Cenomanian in the Iberian Chain (provinces of Teruel a n d Zaragoza), and in the Maastrichtian in the Pyrenees (province of Huesca). Barremian dinosaurs are the most common in the province of Teruel, and they are represented by bones, isolated teeth, and eggshells. Although dinosaur tracks are best represented in other periods in the province of Teruel (Tithonian- Barremian, Hauterivian, Cenomanian), up to the moment there are only two barremian tracksites.

Lower Barremian (LB) and Upper Barremian (UB) dinosaur-bearing formations are present, W to E, in the Oliete Basin (Blesa Fm., LB and UB, Alacón Fm., UB), Galve Basin (Castellar and Camarillas Fms., LB) and Morella Basin (Mirambel Fm., LB., Artoles Fm., UB). French paleontologist A. F. de Lapparent found in the sixties 6 localities in the "weald" of Teruel, in Galve, Morella, Salzedella a n d Peñagolosa Basins, but their exact location remains unknown; two of them may be UB.

With regard to direct remains (i.e. bones and teeth), currently, we have 2 2 localities in the LB (Oliete, Galve and Morella Basins) and 5 localities in the UB (Oliete and Morella basins), five of them consist of isolated remains, and the remainder are isolated carcasses or concentrations of bones, teeth or both. The principal concentration of sites is in Galve (Galve Basin) and Castellote (Morella Basin). There are representative remains of sauropod, theropod, ornithopod a n d thyreophoran dinosaurs.

There are Sauropoda in the LB (a tibia of Sauropoda indet., isolated teeth of two different Camarasauridae indet. and the brachiosaurid cf. Pleurocoelus sp., vertebrae of Titanosauriformes indet.) and in the UB (vertebrae of two types of Titanosauriformes indet, isolated teeth of Camarasauridae/Brachiosauridae).

Among Theropoda, the most representative remains are isolated teeth of very different sizes (ranging from 1 to 34 mm of fore-aft basal length), in LB and UB. IV EUROPEAN WORKSHOP ON VERTEBRATE PALAEONTOLOGY (1999) ALBARRACIN, SPAIN 82

There are representatives of Theropoda indet. (at least five types), Baryonychidae indet., small unserrated coelurosaurians, "paronychodontids", dromeosaurs (at least six types: Dromeosauridae indet., Dromaeosaurinae indet. and Velociraptorinae indet.). Bony remains are infrequent (vertebrae of Theropoda indet in LB and UB, ungual phalanx in LB). Eggshells of the oofamily Elongatoolitidae may belong to oviraptorosaur theropods (Amo, 1998), other eggshells of ratite morphotype may represent undetermined theropods.

Ornithopods are the most frequent group, with some cranial material, abundant postcranial material, and isolated teeth. In the LB there are a partial articulated skeleton of a new Othnielia-like hypsilophodontid, an incomplete femur of a possibly driosaurid (? Valdosaurus sp.), articulated vertebrae and appendicular elements of Iguanodontidae indet., and cranial remains of Iguanodon atherfieldensis. Isolated teeth indicate the presence of at least three hypsilophodontids in the LB and one in the UB, two iguanodontids in the LB an d UB, and one Ornithopoda indet. in the LB.

Thyreophorans are only represented in the LB: a dorsal spine of a nodosaurid ankylosaur and a fragment of a dorsal plate of an stegosaur (Nodosauridae indet., Stegosauria indet.).

Amo, O. 1998. Fragmentos de cáscara de huevo de vertebrados del Cretácico Inferior de Galve (Teruel). Tesis de Licenciatura, Universidad de Zaragoza, 116p.