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Home , Adelpha cocala, Adelpha lycorias, Altinote, Anaeini, Anartia, Anthanassa

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Higher Classification1 Kingdom: Animalia, Phylum: Arthropoda, Class: Insecta, : , Superfamily: Papilionoidea

Family (F:), Subfamily (sF:) and Tribe (T:) Scientific Name1 English Name1 F: Hesperiidae (Skippers) sF: Eudaminae (Spreadwing Skippers) anaphus annetta Yellow-tipped Flasher Central American Banded- vectilucis Skipper pronus Pronus Longtail sF: Hesperiinae () T: Anthoptini Synapte salenus salenus Salenus Faceted-Skipper T: Calpodini cf. ethlius Brazilian Skipper Talides alternata Alternate Ruby- T: cf. phyleus phyleus inimica Yellow-stained Skipper Poanes cf. zabulon T: Halotus angellus Angellus Skipper accius Clouded Skipper Remella rita Rita's Remella sF: Heteropterinae () Dalla lethaea Schaus' sF: Pyrginae (Spread-wing Skippers) T: Achlyodidini Doberes anticus Dark Doberes T: lactifera lactifera Cryptic Skipper T: Erynnini cf. maimon Common Mylon F: (Gossamerwings) sF: (Hairstreaks) T: (Hairstreaks) bassania White-etched Hairstreak F: (Brushfoots) sF: (Emperors) cyane mexicana Mexican cherubina Turquoise Emperor sF: (Exotic Brushfoots) T: Callicorini anna anna Anna’s Eighty-eight Diaethria astala astala Astala Eighty-eight marchalii Widespread Eighty-eight Diaethria gabaza eupepla Smudged Eighty-eight Diaethria pandama Orange-striped Eighty-eight T: Catonephelini Cantonephele chromis godmani Sister-spotted Banner T: Epiphelini iblis plusios Plutonia Banner sF: (Leafwings & Relatives) T: electra Pearly Leafwing Consul panariste jansoni Blackened Leafwing

Page 1 of 5 Cloudbridge , Costa Rica Last Updated: February 18, 2018 Papilionoidea (Butterfly & Skipper) Species List

Family (F:), Subfamily (sF:) and Tribe (T:) Scientific Name1 English Name1 F: Nymphalidae (Brushfoots) (cont’d) sF: Charaxinae (Leafwings & Relatives) (cont’d) T: Anaeini (cont’d) glycerium glycerium Angled Leafwing Fountainea nobilis peralta Noble Leafwing ambrosia ambrosia Ambrosia Leafwing Memphis arginussa eubaena Mottled Leafwing Memphis aulica Aulica Leafwing Memphis beatrix Beatrix Leafwing Memphis elara Elara Leafwing pithyusa Pale-spotted Leafwing Memphis proserpina Proserpina Leafwing Memphis xenocles carolina Carolina Leafwing amphimachus T: Preponini White-spotted amphiktion One-spotted Prepona centralis sF: (Daggerwings) T: Cyrestini (Daggerwings) marcella valetta Purple-stained Daggerwing dentigera Waiter Daggerwing sF: (Monarch & Relatives) T: plexippus plexippus Monarch T: jemina Jemina Clearwing (Clearwings & Tigerwings) Dircenna klugii Klug's Clearwing Dircenna olyras relata Olyras Clearwing salvinia Salvin’s Clearwing Episcada salvinia opleri Salvin's Clearwing hypereia theope Crisp Tigerwing nero Nero Clearwing annette White-spotted Clearwing Greta annette championi White-spotted Clearwing Greta polissena umbrana Polissena Clearwing virginiana evanides Virginiana Clearwing heraldica heralica Heraldica Clearwing Ithomia patilla Patilla Clearwing menapis saturata Menapis Tigerwing vicina Vicina Clearwing Olyras crathis staudingeri Crathis Tigerwing artena Artena Clearwing Pteronymia fulvimargo Fulvimargo Clearwing Pteronymia simplex simplex Simple Clearwing sF: (Actinotes, Heliconians & Fritillaries) T: (Actinotes) anteas Anteas Actinote ozomene nox Lamplight Actinote T: (Longwings) juno juno Juno Longwing

Page 2 of 5 Cloudbridge Nature Reserve, Costa Rica Last Updated: February 18, 2018 Papilionoidea (Butterfly & Skipper) Species List

Family (F:), Subfamily (sF:) and Tribe (T:) Scientific Name1 English Name1 F: Nymphalidae (Brushfoots) (cont’d) sF: Heliconiinae (Actinotes, Heliconians & Fritillaries) (cont’d) T: Heliconiini (Longwings) poeyii Mexican Silverspot (cont’d) Banded Longwing procula vulgiformis Darkened Longwing charithonia vazquezae Zebra Longwing montanus Montane Longwing Heliconius cydno or pachinus Heliconius Doris Longwing Heliconius cydno or pachinus Heliconius pachinus Pachinus Longwing Heliconius petiverana Crimson-patched Longwing sF: (Sisters & Admirals) T: cocala lorzae Lorza’s Sister Adelpha erymanthis erymanthis Erymanthis Sister Adelpha leucophthalma Leucophthalma Sister leucophthalma Adelpha melanippe Adelpha tracta Tracta Sister sF: (True Brushfoots) T: acheronta acheronta Tailed Cecropian T: ardys Ardys Crescent Anthanassa atronia Brown Crescent Anthanassa crithona Orange-banded Crescent Anthanassa otanes Blackended Crescent Anthanassa otanes fulviplaga Blackened Crescent Anthanassa otanes otanes Blackened Crescent Anthanassa sosis Sosis Crescent fulgora Fulgora Crescent Chlosyne cf. janais janais Crimson-patch Checkerspot Tegosa anieta anieta Black-bordered Crescent Tegosa nigrella Nigrella Crescent Tegosa nigrella niveonotis Nigrella Crescent T: dirce dirce Dirce Beauty dione arcaei Banded Mapwing Hypanartia godmanii Godman’s Mapwing Hypanartia kefersteini Mapwing Hypanartia Orange Mapwing Hypanartia trimaculata autumna Reddish Mapwing blomfildia datis Blomfild's Beauty T: Victorinini fatima fatima Banded Peacock epaphus Rusty-tipped Page sF: (Satyrs) Satyrinae sp. Satyr T: (Owl ) dionysos Banded Owl-Butterfly

Page 3 of 5 Cloudbridge Nature Reserve, Costa Rica Last Updated: February 18, 2018 Papilionoidea (Butterfly & Skipper) Species List

Family (F:), Subfamily (sF:) and Tribe (T:) Scientific Name1 English Name1 F: Nymphalidae (Brushfoots) (cont’d) sF: Satyrinae (Satyrs) (cont’d) T: Brassolini (Owl Butterflies) sulanus Dark Owl-Butterfly (cont’d) bubocula Bubocula Owl-Butterfly staudingeri staudingeri Staudinger's Owl-Butterfly bogotanus alajuela2 Bogota Owl-Butterfly2 chiriquensis Split-banded Owl-Butterfly Opsiphanes quiteria talamancensis Scalloped Owl-Butterfly T: Morphini sp. ( & Antirrheas) T: Pronophilini Oxeoschistus hilara euriphyle Dot-banded Satyr T: confusa Confused Satyr Cissia gigas3 Cissia Cissia labe Labe Satyr argentella Ochre-patched Gemmed-Satyr Cyllopsis philodice Pale-banded Gemmed-Satyr Cyllopsis rogersi Costa Rican Gemmed-Satyr Cyllopsis sp. Drucina leonata Leonata Satyr White-dotted Satyr Harmonia Satyr Hermes Satyr Oxeoschistus cothon Lemon-patched Satyr Oxeoschistus crothonides Rust-patched Satyr metaleuca White-banded Satyr dejecta dejecta Dejected Satyr Pedaliodes manis Manis Satyr timanthes Timanthes Satyr Pseudomaniola phaselis rogersi Fire-spotted Satyr Satyrotaygetis satyrina Wide-bordered Satyr uzza Uzza Satyr renata Renata Satyr F: Papilionidae (Swallowtails) sF: Papiloninae T: calliste olbius Yellow Kite-Swallowtail (Kite Swallowtails) T: Papilionini garamas syedra Magnificent Swallowtail (Fluted Swallowtails) Papilio isidorus rhodostictus Ruby-patched Swallowtail F: (Whites & Yellows) sF: (Sulphurs & Yellows) daira eugenia Barred Yellow Salome Yellow Eurema salome jamapa Salome Yellow

Page 4 of 5 Cloudbridge Nature Reserve, Costa Rica Last Updated: February 18, 2018 Papilionoidea (Butterfly & Skipper) Species List

Family (F:), Subfamily (sF:) and Tribe (T:) Scientific Name1 English Name1 F: Pieridae (Whites & Yellows) (cont’d) sF: Coliadinae (Sulphurs & Yellows) (cont’d) Dark Apricot Sulphur marcellina Cloudless Sulphur nelphe Mimosa Yellow sF: (Mimic-Whites) crisia lubina Crisia Mimic-White Dismorphia zathoe pallidula Zathoe Mimic-White Frosted Mimic-White atthis Frosted Mimic-White Lieinix viridifascia Greenish Mimic-White sF: (Whites) T: (Whites) ctemene actinotis Darkened Dartwhite Catasticta flisa melanisa Narrow-banded Dartwhite Catasticta hegemon hegemon Frosted Dartwhite bryson Mexican Dartwhite Catasticta sisamnus smalli Whitened Dartwhite Catasticta teutila flavomaculata Pure-banded Dartwhite Catasticta theresa Small Dartwhite dysoni Dyson’s White Leodonta dysoni dysoni Dyson’s White aripa elodia Mountain White Leptophobia caesis tenuicornis Bluish White charops Darkened White F: (Metalmarks) sF: T: cypria paphia Orange-striped Emesis Falcate Emesis T: Mesosemia asa asa Deep-blue Eyed Metalmark Mesosemia grandis Giant Eyed-Metalmark T: cf. glauca Glauca Metalmark

NOTES: Short-forms: sp. = one species of the given taxon identified; cf. = indicates the individual was difficult to identify and the full species identification is uncertain. 1, Classification, scientific and English names based on current classifications as found on Butterflies of America (http://butterfliesofamerica.com/). 2, identified from a . Could also potentially be Opsiphanes tamarindi tamarindi. Final ID based on location as Opsiphanes bogotanus alajuela is found in habitat in Costa Rica, while Opsiphanes tamarindi tamarindi is found more generally from E & W to Columbia (http://butterfliesofamerica.com/). 3, Identified using: DeVries, P.J. 1987. The Butterflies of Costa Rica and their Natural History: Papilionidae, Pieridae, Nymphalidae. Oxford: Princeton University Press. Have been unable to confirm species name from other sources.

Contributors: Alex Acott, Brittany Beagle, Jeff Boone, Tim Carlson, Neil Hancart, Timo Jäger, Spencer Kane, Sarah Rathbone, Ian Rowntree, Frank Spooner, and Bob Yukich.

Page 5 of 5 Cloudbridge Nature Reserve, Costa Rica Last Updated: February 18, 2018