ARTÍCULOS

Botanica Complutensis ISSN-e: 1988-2874

http://dx.doi.org/10.5209/BOCM.63970

Diversity of Albanian Cladoniaceae

Ana Rosa Burgaz1, Sandra Luna-González1, Blanca Gutiérrez-Larruga1, Raquel Pino-Bodas2, László Lőkös3 & Edit Farkas4

Recibido: 2019 / Aceptado:

Abstract. As a result of collections made during the years 2009-2017, the number of Cladoniaceae species is enlarged to 27 species, providing fifteen new records to Albanian Cladonia. The distribution of many of the previously mentioned taxa is extended and the chemical variability for many of the species is discussed. Cladonia borealis, C. caespiticia, C. cervicornis, C. coccifera, C. conista, C. cryptochlorophaea, C. cyathomorpha, C. humilis, C. merochlorophaea, C. parasitica, C. peziziformis, C. polycarpoides, C. ramulosa, C. rei and C. subulata are new for . Key words. Cladonia; Albania; phytogeography.

[es] Diversidad de las Cladoniaceae de Albania

Resumen. Como consecuencia de las recolecciones efectuadas en varios periodos durante los años 2009-2017, se amplía a 27 especies el número de Cladoniaceae, además se aportan 15 nuevas especies de Cladonia para Albania. Se amplía la distribución de muchos de los taxones citados con anterioridad y se discute la variabilidad química en la mayoría de las especies encontradas. Cladonia borealis, C. caespiticia, C. cervicornis, C. coccifera, C. conista, C. cryptochlorophaea, C. cyathomorpha, C. humilis, C. merochlorophaea, C. parasitica, C. peziziformis, C. polycarpoides, C. ramulosa, C. rei y C. subulata son nuevas para Albania. Palabras clave. Cladonia; Albania; fitogeografía.

Introduction nean climate, and further east the mountains act as a barrier to the penetration of temperate Albania is a Mediterranean country located in air masses. The climate of the mountainous re- SE Europe and surrounded by the gions is alpine, with strong changes in precip- and in the NW, by the Ionian Sea itation and temperature depending on height and Greece in the SW and North Macedonia in and exposure. The temperature differences the East. Despite its small extension, 28,748 between the coast and the interior regions km2, it is topographically very diverse. Al- are more marked in winter, while the precip- though Albania lies within the temperate zone itations are more intense in summer (Tatli & the topographic heterogeneity and the combi- Türkeş 2014). nation of Mediterranean and continental influ- The karstic soils (limestones) are predom- ences confer a great climatic and vegetation inant, although there are locations with ser- diversity (Reed et al. 2004). pentine substrates containing a certain amount The main mountain ranges in the country, of silicates, yeilding slightly acid soils (Bego the Dinaric Alps, run parallel to the coast and et al. 2018). they present the highest peak at 2753 m (Mt. Most of the Albanian territory is included Korab). The coastal region a Mediterra- in the Illyrian province of the Circumboreal

1 Department of Biodiversity, Ecology and Evolution (Botany Unit), Faculty of Biology, Complutense University, E-28040 Madrid, Spain. Corresponding author, e-mail: [email protected] 2 Comparative Plant and Fungal Biology Department, Royal Botanic Gardens Kew, TW9 3AB, Richmond, Surrey, UK. [email protected] 3 Department of Botany, Hungarian Natural History Museum, H-1431 Budapest, Pf. 137, Hungary. [email protected] 4 Institute of Ecology and Botany, MTA Centre for Ecological Research, Hungarian Academy of Sciences, H-2163 Vácrátót, Alkotmány u. 2-4, Hungary. [email protected]

Bot. complut. 43, 2019: 15-40 15 16 Burgaz, A. R. et al. Bot. complut. 43, 2019: 15-40

Region with predominant deciduous forest Materials and Methods of Quercus ssp. (Q. pubescens, Q. frainetto and Q. cerris) and with an important pres- After several recollections in the Albanian terri- ence of Carpinus orientalis (Polunin & Wal- tory at different periods during 2009-2017, we ters 1989). In the higher altitudinal zones the studied and analysed 335 Cladonia specimens, grasslands and beech forests are dominant. which are listed in Annex 1. The new collections The Mediterranean vegetation belongs to the were deposited in Madrid (MACB), Helsinki (H) Adriatic province and is usually well devel- or Budapest (BP). Most of the specimens were oped. It occupies a small coastal strip and analysed by means of thin-layer chromatography has sub-Mediterranean and Mediterranean (TLC, HPTLC) to identify the major lichen sub- species of evergreen and maquis shrubs (Sa- stances according to standardized methods (Or- dori et al. 2015). ange et al. 2001, 2010; Arup et al. 1993). The main The lichenological studies in Albania are secondary metabolites of Albanian Cladonia quite scarce as confirmed by Hafellner & Kash- are listed in Table 1. Comments on world dis- ta (2003) and Hafellner (2007). Later Svoboda tribution of species are from Ahti & Stenroos et al. (2012) increased the number of species (2013). The distribution maps are presented ac- found up to 333 taxa of lichenized fungi and cording to the administrative divisions in coun- 11 of them represent Cladonia. The aim of this ties of Albania (Figure 1). The ITS rDNA region contribution is to summarize results from new was sequenced in some specimens to confirm the collections in the framework of a project on identifications and the GeneBank numbers are the family Cladoniaceae in the Mediterranean provided (Annex 1). The primers and protocols area of Eurasia. used are described in Pino-Bodas et al. (2015).

A B C

Figure 1. A- Map of Albania (http://www.lahistoriaconmapas.com/atlas/albania-map/tirana-albania-map. htm). B- Albanian counties (https://www.revolvy.com/page/Counties-of-Albania Albania_Counties_Gray.svg: Joshbaumgartner). C- : 1 , 2 Bulqizë, 3 Delvinë, 4 , 5 Dibër, 6 Durrës, 7 , 8 , 9 Gjirokastër, 10 Gramsh, 11 Has, 12 Kavajë, 13 Kolonjë, 14 Korçë, 15 Krujë, 16 Kuçovë, 17 Kukës, 18 , 19 Lezhë, 20 , 21 Lushnjë, 22 Malësi e Madhe, 23 Mallakastër, 24 , 25 Mirditë, 26 , 27 Përmet, 28 , 29 Pukë, 30 Sarandë, 31 Shkodër, 32 , 33 Tepelenë, 34 , 35 Tropojë and 36 Vlorë (https://en.wikipedia.org/wiki/Districts_of_Albania). Burgaz, A. R. et al. Bot. complut. 43, 2019: 15-40 17

Results and discussion conista, C. cryptochlorophaea, C. cyathomorpha, C. humilis, C. merochlorophaea, C. parasitica, Cladonia is the only genus of Cladoniaceae C. peziziformis, C. polycarpoides, C. ramulosa, represented in Albania. Of the 27 analysed species C. rei and C. subulata. The species studied occur the following 15 are newly reported to the former between the sea level and 1800 m of altitude based lichen checklist of Albania (*): Cladonia borealis, on our records and the information published by C. caespiticia, C. cervicornis, C. coccifera, C. other authors (Figure 2).

Figure 2. Species altitudinal rank in Albania (except for C. carneola in the lack of data).

The commonest species are: C. cariosa, C. to , and C. borealis, C. fimbriata, C. foliacea, C. furcata, C. pyxidata and cryptochlorophaea, C. humilis, C. merochlorophaea C. rangiformis, while C. caespiticia, C. coccifera, and C. parasitica have a southern or a south-eastern C. digitata, C. macilenta, C. peziziformis, C. distribution (Figure 3). polycarpoides and C. ramulosa are restricted 18 Burgaz, A. R. et al. Bot. complut. 43, 2019: 15-40

Cladonia borealis Cladonia caespiticia

Cladonia cariosa Cladonia cervicornis

Cladonia chlorophaea Cladonia coccifera

Cladonia coniocraea Cladonia conista

Cladonia cryptochlorophaea Cladonia cyathomorpha Burgaz, A. R. et al. Bot. complut. 43, 2019: 15-40 19

Cladonia digitata Cladonia fimbriata

Cladonia foliacea Cladonia furcata

Cladonia humilis Cladonia macilenta

Cladonia merochlorophaea Cladonia parasitica

Cladonia peziziformis Cladonia polycarpoides 20 Burgaz, A. R. et al. Bot. complut. 43, 2019: 15-40

Cladonia pyxidata Cladonia ramulosa

Cladonia rangiformis Cladonia rei

Cladonia subulata Cladonia symphycarpa Figure 3. Distribution of Cladonia species in Albania (except of C. carneola in the lack of exact locality data). Bibliographic reports: ●, new records: ▲.

Nine species are chemically variable C. polycarpoides atranorin and norstictic acid, (Table 1) C. cariosa, C. foliacea, C. furcata, C. and one specimen containing only norstictic acid. polycarpoides, C. pyxidata, C. rangiformis, C. rei Cladonia pyxidata contains fumarprotocetraric and C. symphycarpa. Cladonia cariosa specimens acid and additionally physodalic acid, and also contain three chemotypes: atranorin; atranorin and some specimens have fumarprotocetraric and norstictic acid; atranorin and rangiformic acid, psoromic acids. The commonest chemotype in C. and additional zeorin. Most of the specimens rangiformis contains atranorin, fumarprotocetraric of C. foliacea contain fumarprotocetraric and and rangiformic acids sometimes with additional usnic acids, additionally physodalic acid; but a physodalic acid, the other chemotype with few contain fumarprotocetraric, psoromic and atranorin and rangiformic acid is very rare in the usnic acids. In C. furcata most of the specimens Albanian specimens. The specimens of C. rei contain fumarprotocetraric acid, occasionally contain fumarprotocetraric and homosekikaic with physodalic acid; specimens with atranorin acids and other with only homosekikaic acid. and fumarprotocetraric acid or atranorin Specimens of C. symphycarpa have atranorin and and bourgeanic acid were also found. Two norstictic acid; and other also atranorin, norstictic chemotypes were detected in the specimens of and fumarprotocetraric acids. Burgaz, A. R. et al. Bot. complut. 43, 2019: 15-40 21

Table 1. Main secondary metabolites of the Cladonia species chemically analysed from Albania. ATR: atranorin, BAR: barbatic acid, BOU: bourgeanic acid, CRY: cryptochlorophaeic acid, FUM: fumarprotocetraric acid, HSEK: homosekikaic acid, MER: merochlorophaeic acid, NST: norstictic acid, PHY: physodalic acid, PSO: psoromic acid, RAN: rangiformic acid, THAM: thamnolic acid, USN: usnic acid, ZEO: zeorin. Species ATR BAR BOU CRY FUM HSEK MER NST PHY PSO RAN THAM USN ZEO C. borealis + C. caespiticia + C. cariosa 1 + C. cariosa 2 + + C. cariosa 3 + + C. cariosa 4 + + + C. cervicornis + C. chlorophaea + + C. coccifera + + C. coniocraea + C. conista + + C. cryptochlorophaea + C. cyathomorpha + C. fimbriata + C. foliacea 1 + + C. foliacea 2 + + + C. foliacea 3 + + + C. furcata 1 + C. furcata 2 + + C. furcata 3 + + C. furcata 4 + + + C. furcata 5 + + + C. humilis + + C. macilenta + + C. merochlorophaea + + C. parasitica + + C. peziziformis + C. polycarpoides 1 + C. polycarpoides 2 + + C. pyxidata 1 + C. pyxidata 2 + + C. pyxidata 3 + + C. ramulosa + C. rangiformis 1 + + C. rangiformis 2 + + + C. rangiformis 3 + + + + C. rei 1 + C. rei 2 + + C. subulata + C. symphycarpa 1 + + C. symphycarpa 2 + + + C. symphycarpa 2 + + + 22 Burgaz, A. R. et al. Bot. complut. 43, 2019: 15-40

List of species tha 2003); from Dibër and Tiranë districts by Markgraf (1931, 1932) (Hafellner 2007). It is a *1. Cladonia borealis S. Stenroos, Ann. Bot. cosmopolitan species (Ahti & Stenroos 2013). Fenn. 26: 160 (1989) This terricolous species was only found in *7. Cladonia coccifera (L.) Willd., Fl. Berol. Korçë County growing on sandstone in slopes Prodr.: 361 (1787) of beech forest at 1550 m altitude. It is distrib- This is a very rare species only collected in uted in arctic/antarctic-temperate and circum- Shkodër County at 735 m altitude on serpen- boreal regions (Ahti & Stenroos 2013). tine substrate. It has a cosmopolitan distribu- tion (Ahti & Stenroos 2013). *2. Cladonia caespiticia (Pers.) Flörke, Clad. Comm.: 8 (1828) 8. Cladonia coniocraea (Flörke) Spreng., It is a very rare species only found in Lezhë Syst. Veg., Edn 16, 4(1): 272 (1827) County on limestone in slopes of Carpinus syn. C. ochrochlora Flörke orientalis formations at 40 m altitude. This is It was collected from Durrës, Lezhë and Vlorë mainly a temperate species with oceanic ten- counties from sea level until 828 m altitude, on dencies, not continuously circumpolar (Ahti & limestone and sandy substrates or stumps of Stenroos 2013). Castanea sativa. It was reported of Dibër and Shkodër districts by Svoboda et al. (2012) and 3. Cladonia cariosa (Ach.) Spreng., Syst. Hafellner & Kashta (2003). All of the Albani- Veg., Edn 16, 4(1): 272 (1827) an specimens belong to C. coniocraea s.str. the A very common species found in Elba- more oceanic morphotype often recognized as san, Korçë, Kukës, Shkodër and Tiranë coun- C. ochrochlora, which is not distinguishable in ties. It was reported by Svoboda et al. (2012) DNA analyses (Pino-Bodas et al. 2011) is not from Dibër County on calcareous soil. This present here. is a terricolous species growing on bare soils of limestone, sandstone, serpentine and slate *9. Cladonia conista (Nyl.) Robbins, in Allen, substrates in a wide variability of formations Rhodora 32: 92 (1930) at 120-1550 m altitude. It is a Holarctic spe- A common species collected in Durrës, El- cies with an outlier in southern South Ameri- basan, Fier, Gjirokaster, Korçë, Lezhë, Shkodër, ca (Ahti & Stenroos 2013). It is very common Tirana and Vlorë counties. It grows on slopes and and frequent with several chemotypes. bare soils of Carpinus orientalis formations and also Quercus coccifera, on sandstone and lime- 4. Cladonia carneola (Fr.) Fr., Lichenogr. Eur. stones substrates, from 40-1000 m altitude. Tra- Reform.: 233 (1831) ditionally considered a chemotype of C. humilis This species has only been reported from with fumarprotocetraric and bourgeanic acids Shkodër County by Zahlbruckner (1889: 7) nevertheless, should be considered a different (Hafellner 2007). Species with bipolar distri- species in base of the results of Pino-Bodas et al. bution (Ahti & Stenroos 2013). (2012).

*5. Cladonia cervicornis (Ach.) Flot., Jahres- *10. Cladonia cryptochlorophaea Asahina, J. ber., Schles. Ges. Vaterl. Cult. 27: 105 (1849) Jap. Bot. 16: 711 (1940) This species is not frequent, only encoun- This is a very rare species only collected tered in Durrës, Shkodër and Vlorë counties. in Vlorë County at 186 m altitude growing on It grows from sea level to 865 m altitude on limestones with Erica arborea and deciduous limestone and sandy and acid soils. It has a Quercus. Is a terricolous species with arctic/ Holarctic distribution with uncertain records antarctic to northern temperate distribution in South America (Ahti & Stenroos 2013). (Ahti & Stenroos 2013).

6. Cladonia chlorophaea (Flörke ex Som- *11. Cladonia cyathomorpha Stirt. ex Walt. merf.) Spreng., Syst. Veg., Edn 16, 4(1): 273 Watson, J. Bot. 73: 156 (1935) (1827) It was collected from Elbasan, Kukës, Shkod- A rare species collected in Gjirokastër and ër and Tiranë counties growing at 120-540 m Vlorë counties. But it was early reported from altitude on limestone and serpentine substrates Kukës and Shkodër districts (Hafellner & Kas- with Carpinus orientalis, deciduous Quercus Burgaz, A. R. et al. Bot. complut. 43, 2019: 15-40 23 and Arbutus unedo. It has a western European distribution with some outpost in southern South 15. Cladonia furcata (Huds.) Schrad., Spicil. America (Ahti & Stenroos 2013). Fl. Germ. 1: 107 (1794) incl. C. subrangiformis Sandst. 12. Cladonia digitata (L.) Hoffm., Deutschl. This common species was collected in Durrës, Fl., Zweiter Theil (Erlangen): 124 (1796) Elbasan, Gjirokastër, Korçë and Vlorë counties This species was only reported by Svobo- from sea level to 1700 m altitude. Most of the da et al. (2012) from Dibër district growing samples contain fumarprotocetraric acid and on rotten wood at 1700-1730 m altitude. It is occasionally physodalic acid. The morphotype mainly a Holarctic species with some outposts with atranorin and fumarprotocetraric acid or in East Africa (Ahti & Stenroos 2013). atranorin and bourgeanic acid, traditionally identified as C. subrangiformis, was collected 13. Cladonia fimbriata (L.) Fr., Lich. Eur. Re- from Korçë, Shkodër and Vlorë counties. This form. (Lund): 222 (1831) taxon is considered to fall into the chemical syn. C. major (K. G. Hagen) Sandst., C. minor variability of C. furcata supported by the DNA (K. G. Hagen) Szat. studies of Pino-Bodas et al. (2015). A very common species encountered in C. furcata was reported from Dibër, Dibër, Durrës, Elbasan, Gjirokastër, Korçë, Përmet, Shkodër and Tiranë districts by Mark- Kukës, Lezhe, Shkodër, Tiranë and Vlorë graf (1931, 1932), Hafellner & Kashta (2003) counties. It grows from sea level to 1750 m and Svoboda et al. (2012) without chemical in- altitude on rotten wood, tree bases and hu- dications. It grows on humose, shaded or open mose soil over limestone, sandstone and sandy soil over limestone, serpentine, and conglom- soil. The species was reported by Hafellner & erate substrates in a wide varirty of forests. It Kashta (2003) and Svoboda et al. (2012) from has a subcosmopolitan distribution in boreal to Dibër, Has, Kukës and Shkodër districts. It is a tropical areas (Ahti & Stenroos 2013). cosmopolitan species (Ahti & Stenroos 2013). *16. Cladonia humilis (With.) J.R. Laundon, 14. Cladonia foliacea (Huds.) Willd., Fl. Berol. Lichenologist 16: 220 (1984) Prodr.: 363 (1787) A rare species only collected in Elbasan incl. C. convoluta (Lam.) Anders and Vlorë counties from 50 to 540 m alti- This is a very common species found in tude. It grows on limestone substrate with Berat, Dibër, Durrës, Elbasan, Gjirokastër, Ko- Carpinus orientalis, Quercus spp. and Buxus rçë, Kukës, Lezhe, Shkodër, Tiranë and Vlorë sempervirens. The samples analysed contain counties from 78 to 1550 m altitude. It is a terri- atranorin and fumarprotocetraric acid. It has colous species growing on grasslands and open a Holarctic distribution, with several out- soil, preferably calcareous. It is a Western Eur- posts in the Southern Hemisphere, occurring asian, Macaronesian and North African species from boreal to arctic areas (Ahti & Stenroos which extends in Asia as far as Mongolia and it 2013), although its distribution is not well grows in temperate to subtropical areas (Ahti & known. Stenroos 2013). The specimens containing fumarprotoce- 17. Cladonia macilenta Hoffm., Deutschl. Fl., traric and usnic acids, additionally physodal- Zweiter Theil (Erlangen): 126 (1796) ic acid are the commonest, but samples with This is a rare species only collected in fumarprotocetraric, usnic and psoromic acids Lezhë County growing on bark of Pinus nigra. were found in Korçë, Gjirokastër and Vlorë It was reported from Vlorë district (Hafellner counties on sandstone at 211-953 m altitude. & Kashta 2003). This species has a subcosmo- Most of the early reports (Markgraf 1927, politan distribution (Ahti & Stenroos 2013). 1931, 1932; Hafellner & Kashta 2003; Svo- boda et al. 2012) were published as C. convo- *18. Cladonia merochlorophaea Asahina, J. luta but we include it in the concept of C. fo- Jap. Bot. 6: 713 (1940) liacea because there are no molecular differ- A very rare species only collected in Korçë ences between these taxa (Pino-Bodas et al. County, on calcareous soil, at 1625 m altitude. 2010a, 2018), although specimens called C. This is a subcosmopolitan species but its taxo- convoluta are of larger size due to its adapta- nomic status is not fully clear (Ahti & Stenroos tion to calcareous soil and dry situations. 2013). 24 Burgaz, A. R. et al. Bot. complut. 43, 2019: 15-40

*19. Cladonia parasitica (Hoffm.) Hoffm., 24. Cladonia rangiformis Hoffm., Deutschl. Deutschl. Fl. 2: 127 (1796) Fl., Zweiter Theil (Erlangen): 114 (1796) This is a very rare species only collected A very common species collected in Berat, in Vlorë County at 828 m altitude growing on Dibër, Durrës, Elbasan, Fier, Gjirokastër, Ko- Castanea sativa stumps. This is a Holarctic rçë, Kukës, Lezhë, Shkodër, Tiranë and Vlorë species with continental tendencies (Ahti & counties. It grows on grasslands and open soil Stenroos 2013). of sandy substrate, serpentines, limestones and slates from sea level to 1770 m altitude. Its dis- *20. Cladonia peziziformis (With.) J.R. Laun- tribution is mainly restricted to western Eurasia don, Lichenologist 16: 223 (1984) and North Africa with one outpost in eastern A very rare species only collected in Canada (Ahti & Stenroos 2013). The common- Shkodër County, on serpentine soil at 120 m est chemotype contains atranorin, fumarproto- altitude. This is a subcosmopolitan species cetraric and rangiformic acids sometimes with easily overlooked due to its small size (Ahti & additional physodalic acid, but the chemotype Stenroos 2013). with atranorin and rangiformic acid (common elsewhere) is very rare. One specimen of a sore- *21. Cladonia polycarpoides Nyl. in Zwackh, diate morph (‘var. sorediophora’) was found in Lich. Exs. Nos. 626, 626bis (1892) Shkodër. syn. C. subcariosa auct. It was reported from Dibër, Gjirokastër, It is a rare species collected in Kukës and Kukës, Shkodër, Tiranë and Vlorë counties Shkodër counties growing on serpentine soil by Szatala & Timkó (1926), Markgraf (1927, from sea level to 600 m altitude. It is a sub- 1932) and Svoboda et al. (2012), but without cosmopolitan species especially abundant un- indication of their chemistry. der suboceanic to oceanic conditions (Ahti & Stenroos 2013). *25. Cladonia rei Schaer. Lich. Helv. Spicil. 1(1): 34 (1823) It is an uncommon species collected in 22. Cladonia pyxidata (L.) Hoffm., Deutschl. Durrës, Kukës and Vlorë counties. It grows on Fl., Zweiter Theil (Erlangen): 121 (1796) limestone and sandstone substrates at 90-330 incl. C. pocillum (Ach.) Grognot; C. monomorpha m altitude on limestone and sandstones sub- Aptroot, Sipman & van Herk strates. It has a Holarctic distribution with con- A common species collected in Dibër, Durrës, tinental preferences (Ahti & Stenroos 2013). Elbasan, Fier, Gjirokastër, Korçë, Shkodër, One sample analysed contains homosekikaic Tiranë and Vlorë counties growing on bare soils, acid only, but other specimens contain homo- grasslands and rarely on wood. Most of the samples sekikaic and fumarprotocetraric acids. Some analysed contain fumarprotocetraric acid and time ago C. rei was considered a homosekika- often additional physodalic acid. A chemotype ic chemotype of C. subulata (Spier & Aptroot with fumarprotocetraric and psoromic acids was 2007) but molecular studies maintained them collected at 540 m altitude from . as different species (Pino-Bodas et al. 2010b). Cladonia pyxidata was early reported from Kukës, Lezhë, Shkodër and Vlorë districts (Zahlbruckner *26. Cladonia subulata (L.) F.H. Wigg., Prim. 1889, Markgraf 1931, Szatala & Timkó 1926, Fl. Holsat.: 90 (1780) Murati 1979, Hafellner & Kashta 2003, Svoboda This is a rare species only collected in et al. 2012). Some of these reports refer to C. Kukës and Lezhë counties on limestone susb- pocillum but we do not distinguish it as a different strate at 40-330 m altitude. It is a subcosmo- entity because molecular studies do not support politan species (Ahti & Stenroos 2013). it (Kotelko & Piercey-Normore 2010). It has a cosmopolitan distribution (Ahti & Stenroos 2013). 27. Cladonia symphycarpa (Flörke) Fr., Sched. Crit. Lich. Suec. 8-9. 20 (1826) *23. Cladonia ramulosa (With.) J.R. Laun- incl. C. dahliana Kristinsson don, Lichenologist 16: 225 (1984) A very common species collected in Elbasan, It was collected in Durrës and Lezhë coun- Korçë and Shkodër counties growing on lime- ties on sandy soil and limestone from sea lev- stone substrate at 400 to 1800 m altitude. It has el to 40 m altitude. This is a subcosmopolitan been reported only from Shkodër County by Svo- species (Ahti & Stenroos 2013). boda et al. (2012). The most frequent chemotype Burgaz, A. R. et al. Bot. complut. 43, 2019: 15-40 25 has atranorin and norstictic acid, additionally fatty studies are necessary to clarify the identity of acid; the other chemotype with atranorin, norstic- these specimens. tic and fumarprotocetraric acids is very rare. It has a Holarctic distribution with some outliers in South America (Ahti & Stenroos 2013). Acknowledgements

Cladonia sp. A. R. Burgaz received financial support from Some specimens collected in Shkodër and the project CGL2013-41839-P, Ministry of Vlorë were difficult to identify when the pode- Economy and Competitiveness, Spain. Chro- tia are not well developed and do not bear split matographic studies (HPTLC) carried in Hun- proliferations in the margin of the scyphi. These gary were supported by the Hungarian Na- specimens belong to the Cladonia humilis tional Research Development and Innovation group (Pino-Bodas et al. 2013) but molecular Fund [NKFI K 124341].

References

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Annex 1. Selected specimens examined oxycedrus, pastureland, sandstone, 1200 m, 23- IV-2017, A.R. Burgaz (MACB 111453). District Cladonia borealis S. Stenroos of Pogradec, Udënisht, SH3 Piskutat-Pogradec, Barbatic acid: KORÇË: District of Ko- shore Ohrid lake, 40º58’55”N 20º38’21”E, rçë, Dardhë, SH90 Boboshticë-Dardhë, Pinus halepensis, Juniperus oxycedrus, Buxus 40º31’26”N 20º48’06”E, Fagus sylvatica, Ju- sempervirens, sandstone, 710 m, 22-IV-2017, niperus communis, Abies sp., sandstone, 1550 A.R. Burgaz (MACB 111452). KUKËS: m, 23-IV-2017, A.R. Burgaz (MACB 111457). District of Kukës, Kolsh, SH5 close Drin river, 42º04’18”N 20º20’32”E, deciduous Quercus, Cladonia caespiticia (Pers.) Flörke Juniperus oxycedrus, Buxus sempervirens, Fumarprotocetraric acid: LEZHË: Dis- serpentine soil, 510 m, 21-IV-2017, A.R. Burgaz trict of Kurbin, Milot, left side Mat river, SH25 (MACB 111433). District of Kukës, Rruga Milot-Zogu Bridge, 41º41’54”N 19º45’10”E, Tirana, 42º04’46”N 20º24’45”E, Quercus cerris, Carpinus orientalis, Erica arborea, lime- deciduous Quercus, limestone, 330 m, 21-IV- stone, 40 m, 21-IV-2017, A.R. Burgaz (MACB 2017, A.R. Burgaz (MACB 111435, 111434). 111430, GenBank accession nº: MK543953). Atranorin and rangiformic acid chemotype: ELBASAN: , Urakë, SH9 Qafë Thanë, Cladonia cariosa (Ach.) Spreng. 41º04’24”N 20º36’18”E, Quercus macrolepis, Atranorin chemotype: KUKËS: District deciduous Quercus, Juniperus oxycedrus, of Kukës, Kukës, Rruga Tirana, 42º04’46”N limestone, 1000 m, 22-IV-2017, A.R. Burgaz 20º24’45”E, Quercus cerris, deciduous Quercus, (MACB 111447). KORÇË: District of Kolonjë limestone, 330 m, 21-IV-2017, A.R. Burgaz (Rrethi i Kolonjës), c. 1.9 km W of village (MACB 111436). SHKODËR: Vau i Dejës, SH25 Pepellash, 40°28’56.18”N 20°40’21.27”E, on Vau i Dejës-Koman, 42º00’55”N 19º39’37”E, acid soil in moist road slope, 549 m, 18-X-2011, Pinus pinaster, Carpinus orientalis, serpentine Z. Barina, L. Lőkös, D. Pifkó (BP 96585). District soil, 120 m, 20-IV-2017, A.R. Burgaz (MACB of Korçë, Dardhë, SH90 Boboshticë-Dardhë, 111420). Vau i Dejës, Karmë, SH25 Vau i Dejës- 40º31’18”N 20º49’22”E, Fagus sylvatica, Pinus Koman, 42º04’02”N 19º40’59”E, Quercus nigra, Juniperus communis, Betula pendula, macrolepis, Q. pubescens, Juniperus oxycedrus, Abies sp., limestone, 1390 m, 23-IV-2017, A.R. Erica arborea, slates, 130 m, 20-IV-2017, A.R. Burgaz (MACB 111460). District of Korçë, Burgaz (MACB 111429). District of Malësi e Dardhë, SH90 Boboshticë-Dardhë, 40º31’26”N Madhe, Bogë, SH21 -Bogë, 42º21’46”N 20º48’06”E, Fagus sylvatica, Juniperus 19º37’56”E, Pinus nigra, Juniperus oxycedrus, communis, Abies sp., sandstone, 155 m, 23-IV- limestone, 865 m, 20-IV-2017, AR. Burgaz 2017, A.R. Burgaz (MACB 111456). District (MACB 111298). Atranorin and norstictic of Korçë (Rrethi i Korçës), on the ridge of Mts acid chemotype: ELBASAN: Prrenjas, Urakë, Mali i Moravës above village Dardhë, near peak SH9 Qafë Thanë, 41º04’24”N 20º36’18”E, Buzeja e Boboshticës (1701 m), 40°32’23.97”N Quercus macrolepis, deciduous Quercus, 20°49’14.20”E, on acid soil in beech forest on Juniperus oxycedrus, limestone, 1000 m, 22-IV- gravel conglomerate, 1550 m, 10-VIII-2010, 2017, A.R. Burgaz (MACB 111446). District of L. Lőkös (BP 96563). KUKËS: District of Elbasan, , SH3 Bradashesh-Graze, Kukës, Kolsh, SH5 close Drin river, 42º04’18”N 41º06’33”N 19º59’49”E, Carpinus betulus 20º20’32”E, deciduous Quercus, Juniperus Juniperus oxycedrus, limestone karst, 530 m, 15- oxycedrus, Buxus sempervirens, serpentine IV-2017, A.R. Burgaz (MACB 111402). KORÇË: soil, 510 m, 21-IV-2017, A.R. Burgaz (MACB District of Korçë, Dardhë, SH90 Boboshticë- 111462). SHKODËR: Vau i Dejës, SH25 Vau Dardhë, 40º31’18”N 20º49’22”E, Fagus i Dejës-Koman, 42º03’01”N 19º39’45”E, sylvatica, Pinus nigra, Juniperus communis, Pinus pinaster, Juniperus oxycedrus, Quercus Betula pendula, Abies sp., limestone, 1390 m, 23- cerris, Erica arborea, Cotinus coggygria, IV-2017, A.R. Burgaz (MACB 111459). District serpentine soil, 130 m, 20-IV-2017, A.R. of Korçë, Dardhë, SH90 Boboshticë-Dardhë, Burgaz (MACB 111426). TIRANË: District 40º31’26”N 20º48’06”E, Fagus sylvatica, of Tiranë, Mushqeta, SH3 Mushqeta-, Juniperus communis, Abies sp., sandstone, 1550 41º12’38”N 19º57’09”E, Arbutus unedo, Erica m, 23-IV-2017, A.R. Burgaz (MACB 111455). arborea, Platanus orientalis, boulders, 480 m, District of Korçë, Boboshticë, SH90 Boboshticë- 22-IV-2017, A.R. Burgaz (MACB 111440). Dardhë, 40º32’37”N 20º47’11”E, Juniperus Atranorin, rangiformic acid and zeorin 28 Burgaz, A. R. et al. Bot. complut. 43, 2019: 15-40 chemotype: SHKODËR: Vau i Dejës, SH25 Vau 39º55’08”N 20º08’48”E, Quercus coccifera, i Dejës-Koman, 42º03’01”N 19º39’45”E, Pinus Carpinus betulus, Pinus halepensis, Cercis pinaster, Juniperus oxycedrus, Quercus cerris, siliquastrum, limestone slopes, 125 m, 17-IV- Erica arborea, Cotinus coggygria, serpentine 2017, A.R. Burgaz (MACB 111193). soil, 130 m, 20-IV-2017, A.R. Burgaz (MACB 111425). Vau i Dejës, Karmë, SH25 Vau i Dejës- Cladonia coccifera (L.) Willd. Koman, 42º04’02”N 19º40’59”E, Quercus Usnic acid and zeorin: SHKODËR: Pukë macrolepis, Q. pubescens, Juniperus oxycedrus, District (Rrethi i Pukës), Mount Rras’ e Qer- Erica arborea, slates, 130 m, 20-IV-2017, A.R. retit above village Qerret, 42°03’36.98”N Burgaz (MACB 111428). 19°49’53.45”E, in serpentine rocky area, on acid soil in pine woodland, 735 m, 6-V-2015, Cladonia cervicornis (Ach.) Flot. L. Lőkös (BP 96621). Fumarprotocetraric acid: DURRËS: District of Durrës, Fushë-Draçi, San Pietro Cladonia coniocraea (Flörke) Spreng. beach track, 41º31’01”N 19º31’12”E, Pinus Fumarprotocetraric acid: DURRËS: pinea, sandy soil, 1 m, 19-IV-2017, A.R. Burgaz District of Durrës, Fushë-Draçi, San Pietro (MACB 111225). SHKODËR: District of beach track, 41º31’01”N 19º31’12”E, Pi- Malësi e Madhe, Poica, SH21 Koplik-Bogë, nus pinea, sandy soil, 1 m, 19-IV-2017, A.R. 42º15’53”N 19º29’47”E, Juniperus oxycedrus, Burgaz (MACB 111224). LEZHË: District of Cupressus sempervirens, boulders, 320 m, 20- Kurbin, Milot, left side Mat river, SH25 Mi- IV-2017, A.R. Burgaz (MACB 111292). District lot-Zogu Bridge, 41º41’54”N 19º45’10”E, of Malësi e Madhe, , SH21 Koplik-Bogë, Carpinus orientalis, Erica arborea, lime- 42º18’04”N 19º32’32”E, Carpinus betulus, stone, 40 m, 21-IV-2017, A.R. Burgaz (MACB Juniperus oxycedrus, limestone rocks, 495 m, 111305). VLORË: District of Delvinë (Rre- 20-IV-2017, A.R. Burgaz (MACB 111299). thi i Delvinës), Mali i Gjerë, on the southern District of Malësi e Madhe, Bogë, SH21 slope of 1331 m high mount above Delvinë, Koplik-Bogë, 42º21’46”N 19º37’56”E, Pinus 39°57’42.12”N 20°07’59.66”E, in chestnut nigra, Juniperus oxycedrus, limestone, 865 m, woodland, on bark (Castanea sativa), 828 m, 20-IV-2017, A.R. Burgaz (MACB 111303). 1-V-2009, Z. Barina & D. Pifkó (BP 96542). Pukë District (Rrethi i Pukës), Mount Rras’ e Qerretit above village Qerret, 42°03’36.98”N Cladonia conista (Nyl.) Robbins 19°49’53.45”E, in serpentine rocky area, on Fumarprotocetraric and bourgeanic acids: acid soil in pine woodland, 735 m, 6-V-2015, L. DURRËS: District of Durrës, Fushë-Draçi, Lőkös (BP 96620). District of Shkodër (Rrethi i road cape of Rodon, 41º32’50”N 19º31’08”E, Shkodrës), Qafa e Çelës above village Mirdicë, Carpinus betulus, Arbutus unedo, Pistacia 42°0’11.98”N 19°43’33.16”E, on soil in roadside lentiscus, Cistus salviifolius, sandstone, 90 m, dry rocky grassland, 470 m, 12-X-2011, L. Lőkös 19-IV-2017, A.R. Burgaz (MACB 111412). (BP 96756). VLORË: , Muzinë, Muzinë, District of Durrës (Rrethi i Durrësit), west of 39º54’48”N 20º11’20”E, deciduous Quercus, village Shënpjetër, at the southern foot of hill Erica arborea, limestone, 186 m, 17-IV-2017, M. e Multit (159.9 m), near the coast of the A.R. Burgaz (MACB 111199). Adriatic Sea, 41°32’41.0”N 19°29’16.0”E, on soil in grasslands, 0 m, 26-IV-2009, L. Lőkös Cladonia chlorophaea (Flörke ex Sommerf.) (BP 96551). ELBASAN: Prrenjas, Urakë, SH9 Spreng. Qafë Thanë, 41º04’24”N 20º36’18”E, Quercus Fumarprotocetraric and physodalic acids: macrolepis, deciduous Quercus, Juniperus GJIROKASTËR: District of Tepelenë, Këlcyrë, oxycedrus, limestone, 1000 m, 22-IV-2017, A.R. Këlcyrë gorge, left side Vjosa river, 40º17’52»N Burgaz (MACB 111449). District of Librazhd 20º10’05»E, Carpinus betulus, Pinus halepensis, (Rrethi i Librazhdit), Polis Mts, above village sandstone, 180 m, 18-IV-2017, A.R. Burgaz, Polis, on the ridge between the valley of brook (MACB 111411). LEZHË: District of Mirditë “Sut Kukit” and river Shkumbin, 41°09’16.50”N (Rrethi i Mirditës), in the valley of stream Fani 20°15’08.79”E, in bushland, on conglomeratic i Vogël above village Thirrë, in serpentine rocky flysh sediments, 555 m, 12-X-2010, L. area, on bark (Pinus nigra), 42°0’20.98”N Lőkös (BP 96571). Elbasan county, in open 20°14’33.25”E, 905 m, 6-V-2015, L. Lőkös boxwood scrubland above village Kutërman, (BP 96627). VLORË: Finiq, Bistricë, Muzinë, 41°13’40.99”N 20°14’41.04”E, on acid soil in Burgaz, A. R. et al. Bot. complut. 43, 2019: 15-40 29 rocky serpentine area, 510 m, 16-V-2016, L. D. Pifkó (BP 96543). District of Vlorë (Rrethi i Lőkös (BP 96660). FIER: , Pylli i Vlorës), between village Ilias and manastir Shën Levanit, above village Levan, 40°40’48.99”N Theodor, 40°08’07.8”N 19°40’26.4”E, on soil in 19°30’11.70”E, on calcareous soil in oak olive plantation, 211 m, 29-IV-2009, L. Lőkös woodland, 95 m, 5-V-2017, L. Lőkös (BP 96678). (BP 96557). GJIROKASTËR: Gjirokastër District (Rrethi i Gjirokastrës), Mali i Nikollaqit above village Cladonia cryptochlorophaea Asahina Sofratikë, 39°59’41.22”N 20°12’40.05”E, on Cryptochlorophaeic acid: VLORË: Finiq, calcareous soil in rocky grasslands, 260 m, Muzinë, Muzinë, 39º54’48”N 20º11’20”E, de- 8-V-2015, L. Lőkös (BP 96631). District of ciduous Quercus, Erica arborea, limestone, Gjirokastër (Rrethi i Gjirokastrës), Kurvelesh 186 m, 17-IV-2017, A.R. Burgaz (MACB i Sipërm uplands above village Picar, 111410). 40°11’0.39”N 20°02’31.51”E, on calcareous soil in rocky grasslands, 750 m, 29-III-2015, L. Cladonia cyathomorpha Stirt. ex Walt. Watson Lőkös (BP 96600). District of Përmet (Rrethi i Fumarprotocetraric acid: ELBASAN: Përmetit); left part of river Vjosë, between town Prrenjas, Prrenjas, SH3 Qukës-Prrenjas, Këlcyrë and village Mbrezhdan, 40°19’11.24”N 41º04’24”N 20º30’15”E, Carpinus orientalis, 20°05’43.54”E, in scrubland, on flysh soil, 205 Quercus macrolepis, deciduous Quercus, m, 9-V-2015, L. Lőkös (BP 96633). District of Buxus sempervirens, limestone slopes, 540 m, Tepelenë (Rrrethi i Tepelenës), c. 500 m N of 22-IV-2017, A.R. Burgaz (MACB 111443). village Mirnë, 40°21’53.91”N 19°58’15.44”E, in KUKËS: District of Kukës, Kukës, Rruga dry grassland, on flysh soil, 160 m, 14-X-2012, Tirana, 42º04’46”N 20º24’45”E, Quercus L. Lőkös (BP 96595). KORÇË: Kolonjë District cerris, deciduous Quercus, limestone, 330 m, (Rrethi i Kolonjës), between town Leskovik and 21-IV-2017, A.R. Burgaz (MACB 111438). village Vrëpckë, 40°10’30.62”N 20°38’22.95”E, SHKODËR: Vau i Dejës, SH25 Vau i Dejës- on acid soil in rocky serpentine grasslands, 1030 Koman, 42º00’55”N 19º39’37”E, Pinus pinaster, m, 10-V-2015, Z. Barina, L. Lőkös & D. Pifkó Carpinus orientalis, serpentine soil, 120 m, (BP 96637). District of Korçë (Rrethi i Korçës), 20-IV-2017, A.R. Burgaz (MACB 111421). above the water reservoir c 2.5 km WSW of village TIRANË: District of Tiranë, Mushqeta, SH3 Ujëbardhë, 40°31’17.31”N 20°39’50.62”E, on Mushqeta-Gracen, 41º12’38”N 19º57’09”E, acid soil in dry rocky grassland, 1105 m, 18-X- Arbutus unedo, Erica arborea, Platanus 2011, L. Lőkös (BP 96586). LEZHË: Kurbin, orientalis, boulders, 480 m, 22-IV-2017, A.R. Milot, left side Mat river, SH25 Milot-Zogu Burgaz (MACB 111442). Bridge, 41º41’54”N 19º45’10”E, Carpinus orientalis, Erica arborea, limestone, 40 m, Cladonia fimbriata (L.) Fr. 21-IV-2017, A.R. Burgaz (MACB 111431). Fumarprotocetraric acid: DIBËR: District SHKODËR: Pukë District (Rrethi i Pukës), of Mat (Rrethi i Matit), Mali i Skënderbeut, on Mount Rras’ e Qerretit above village Qerret, the northern forehills of Mt. “maja e 42°03’36.98”N 19°49’53.45”E, in serpentine Mëllezit”, near village Shkopet, 41°41’38.1”N rocky area, on bark (Juniperus), 735 m, 6-V- 19°49’16.4”E, on soil in Ostrya carpinifolia 2015, L. Lőkös (BP 96622). TIRANË: District scrubland, 75 m, 24-IV-2009, L. Lőkös (BP of Kavajë (Rrethi i Kavajës), by the coast of the 96549). DURRËS: District of Durrës, Fushë- Adriatic Sea between villages Spille and Rreth Draçi, San Pietro beach track, 41º31’01”N (Rreth-Greth), 41°05’07.6”N 19°27’31.5”E, 19º31’12”E, Pinus pinea, sandy soil, 1 m, on sandy soil in sand dune area, 10 m, 27-IV- 19-IV-2017, A.R. Burgaz (MACB 111413). 2009, L. Lőkös (BP 96552). VLORË: Finiq, District of Krujë, Krujë, SH38 to Qafë-Shtamë Bistricë, Muzinë, 39º55’08”N 20º08’48”E, N. P., 41º31’03”N 19º49’27”E, Carpinus Quercus coccifera, Carpinus betulus, Pinus betulus, pasturelands, limestone, 720 m, 19- halepensis, Cercis siliquastrum, limestone IV-2017, A.R. Burgaz (MACB 111414). slopes, 125 m, 17-IV-2017, A.R. Burgaz (MACB ELBASAN: Prrenjas, SH3 Qukës-Prrenjas, 111409). District of Delvinë (Rrethi i Delvinës), 41º04’24”N 20º30’15”E, Carpinus orientalis, Mali i Gjerë, on the southern slope of 1331 m Quercus macrolepis, deciduous Quercus, high mount above Delvinë, 39°57’42.12”N Buxus sempervirens, limestone slopes, 540 m, 20°07’59.66”E, in chestnut woodland, on bark 22-IV-2017, A.R. Burgaz (MACB 111445). (Castanea sativa), 828 m, 1-V-2009, Z. Barina, Prrenjas, Urakë, SH9 Qafë Thanë, 41º04’24”N 30 Burgaz, A. R. et al. Bot. complut. 43, 2019: 15-40

20º36’18”E, Quercus macrolepis, deciduous boulders, 480 m, 22-IV-2017, A.R. Burgaz Quercus, Juniperus oxycedrus, limestone, 1000 (MACB 111441). VLORË: District of Sarandë, m, 22-IV-2017, A.R. Burgaz (MACB 111448). Pasqirave, Butrint N. P., SH81, Shengjergj GJIROKASTËR: Gjirokastër County, Bredhi monastery, 39º48’42”N 20º00’43”E, Quercus i Sotirës above village Sotirë, 39°48’53.21”N coccifera, limestone slopes, 50 m, 17-IV-2017, 20°21’41.98”E, in dry scrubland area, on A.R. Burgaz (MACB 111347). Finiq, Bistricë, bark (Platanus orientalis), 550 m, 6-V-2017, Muzinë, 39º55’08”N 20º08’48”E, Quercus L. Lőkös (BP 96680). District of Tepelenë, coccifera, Carpinus betulus, Pinus halepensis, Këlcyrë, Këlcyrë gorge, left side Vjosa river, Cercis siliquastrum, limestone slopes, 125 m, 40º17’52”N 20º10’05”E, Carpinus betulus, 17-IV-2017, A.R. Burgaz (MACB 111348). Pinus halepensis, sandstone, 180 m, 18-IV- Finiq, Muzinë, 39º54’48”N 20º11’20”E, 2017, A.R. Burgaz (MACB 111350). KORÇË: deciduous Quercus, Erica arborea, limestone, District of Korçë, Dardhë, SH90 Boboshticë- 186 m, 17-IV-2017, A.R. Burgaz (MACB Dardhë, 40º31’18”N 20º49’22”E, Fagus 111349). District of Delvinë (Rrethi i Delvinës), sylvatica, Pinus nigra, Juniperus communis, Mali i Gjerë, on the southern slope of 1331 m Betula pendula, Abies sp., limestone, 1390 m, 23- high mount above Delvinë, 39°57’42.12”N IV-2017, A.R. Burgaz (MACB 111458). District 20°07’59.66”E, in chestnut woodland, on bark of Korçë, Dardhë, SH90 Boboshticë-Dardhë, (Castanea sativa), 828 m, 1-V-2009, Z. Barina 40º31’26”N 20º48’06”E, Fagus sylvatica, & D. Pifkó (BP 96544). Juniperus communis, Abies sp., sandstone, 1550 m, 23-IV-2017, A.R. Burgaz (MACB Cladonia foliacea (Huds.) Willd. 111454). District of Korçë (Rrethi i Korçës), Fumarprotocetraric and usnic acids Mali i Strugarit above village Belorta, chemotype: BERAT: District of Berat (Rrethi i 40°39’56.56”N 20°50’9.79”E, open grasslands Beratit), Tomor Mountains (Mali i Tomorrit), c. on conglomerate rocks, 1010 m, 11-V-2015, Z. 700 m north of village Karkanjoz, 40°41’55.61”N Barina, L. Lőkös, D. Pifkó (BP 96642). District 20°03’28.69”E, in Mediterranean scrubland, on of Pogradec, Udënisht, SH3 Piskutat-Pogradec, calcareous soil in rocky grassland, 490 m, 25-V- shore Ohrid lake, 40º58’55”N 20º38’21”E, 2004, Z. Barina & Cs. Németh (BP 96540). Pinus halepensis, Juniperus oxycedrus, Buxus District of Skrapar (Rrethi i Skraparit), above the sempervirens, sandstone, 710 m, 22-IV-2017, gorge of river Osum next to village Dhorës, A.R. Burgaz (MACB 111451). KUKËS: District 40°28’27.79”N 20°15’16.11”E, in open of Kukës, Kukës, Rruga Tirana, 42º04’46”N bushland, on calcareous soil, 420 m, 13-X-2010, 20º24’45”E, Quercus cerris, deciduous L. Lőkös (BP 96575). DIBËR: Dibër county, on Quercus, limestone, 330 m, 21-IV-2017, A.R. the mount between villages Fushë Studë and Burgaz (MACB 111437). LEZHË: District of Zabzun, 41°19’54.43”N 20°24’38.25”E, on Lezhë (Rrethi i Lezhës), on the slope of Mt Guri calcareous soil in rocky grasslands, 1295 m, 18- i Kuq above village Fregën, 41°51’41.58”N V-2016, L. Lőkös (BP 96668). DURRËS: District 19°44’11.22”E, on acid soil in open serpentine of Durrës, Fushë-Draçi, San Pietro beach track, grasslands, 575 m, 30-III-2015, L. Lőkös (BP 41º31’01”N 19º31’12”E, Pinus pinea, sandy 96606). SHKODËR: Vau i Dejës, SH25 Vau i soil, 1 m, 19-IV-2017, A.R. Burgaz (MACB Dejës-Koman, 42º03’01”N 19º39’45”E, Pinus 111288). Prrenjas, Urakë, SH9 Qafë Thanë, pinaster, Juniperus oxycedrus, Quercus cerris, 41º04’24”N 20º36’18”E, Quercus macrolepis, Erica arborea, Cotinus coggygria, serpentine deciduous Quercus, Juniperus oxycedrus, soil, 130 m, 20-IV-2017, A.R. Burgaz (MACB limestone, 1000 m, 22-IV-2017, A.R. Burgaz 111424). District of Malësi e Madhe, Bogë, (MACB 111313). ELBASAN: District of SH21 Koplik-Bogë, 42º21’46”N 19º37’56”E, Elbasan, Bradashesh, SH3 Bradashesh-Graze, Pinus nigra, Juniperus oxycedrus, limestone, 41º06’05”N 20º00’26”E, Olea europea, 865 m, 20-IV-2017, A.R. Burgaz (MACB limestone karst, 390 m, 15-IV-2017, A.R. Burgaz 111417). TIRANË: Rrogozhinë, Grethi i (MACB 111278). District of Elbasan, Mesem, 41º05’00”N 19º27’35”E, Pinus Bradashesh, SH3 Bradashesh-Graze, 41º06’33”N pinaster, Juniperus oxycedrus, sandy soil, 0 19º59’49”E, Carpinus betulus, Juniperus m, 15-IV-2017, A.R. Burgaz (MACB 111346). oxycedrus, limestone karst, 530 m, 15-IV-2017, District of Tiranë, Mushqeta, SH3 Mushqeta- A.R. Burgaz (MACB 111279). Elbasan County Gracen, 41º12’38”N 19º57’09”E, Arbutus (Qarku i Elbasanit), on the mount above bridge unedo, Erica arborea, Platanus orientalis, Ura e Murrashit, between Mirakë and Librazhd, Burgaz, A. R. et al. Bot. complut. 43, 2019: 15-40 31

41°11’1.64”N 20°16’33.36”E, on acid soil in 20°49’14.20”E, on acid soil in beech forest on serpentine scrublands, 290 m, 3-IV-2016, L. gravel conglomerate, 1550 m, 10-VIII-2010, L. Lőkös (BP 96657). Elbasan county, in village Lőkös (BP 96561). Korçë, Boboshticë, SH90 Floq, 41°16’1.19”N 20°14’19.17”E, on soil by Boboshticë-Dardhë, 40º32’37”N 20º47’11”E, the roadside, 650 m, 17-V-2016, L. Lőkös (BP Juniperus oxycedrus, pastureland, sandstone, 96664). GJIROKASTËR: , Grapsh, 1200 m, 23-IV-2017, A.R. Burgaz (MACB SH78, 39º56’50”N 20º14’31”E, deciduous 111317). District of Korçë (Rrethi i Korçës), Quercus, limestone slope rocks, 440 m, 17-IV- Mali i Strugarit above village Belorta, 40° 39’ 2017, A.R. Burgaz (MACB 111285). Dropull, 56.56”N 20°50’9.79”E, open grasslands on Goranxi, to Kishae Ravenes, 40º00’56”N conglomerate rocks, 1010 m, 11-V-2015, Z. 20º11’15”E, limestone rocks, 268 m, 17-IV- Barina, L. Lőkös, D. Pifkó (BP 96643). District 2017, A.R. Burgaz (MACB 111286). District of of Devoll (Rrethi i Devollit), on the mount Gjirokastër (Rrethi i Gjirokastrës), Kurvelesh i between village Tren and lake Liqeni i Prespës së Sipërm uplands above village Picar, Vogël, 40°40’10.55”N 20°59’12.62”E, on 40°10’21.66”N 20°02’40.64”E, on calcareous calcareous soil in rocky grasslands, 910 m, 12-V- soil in rocky grasslands, 390 m, 29-III-2015, L. 2015, Z. Barina, L. Lőkös & D. Pifkó (BP 96648). Lőkös (BP 96602). District of Përmet (Rrethi i Korçë County, Mali i Lenies at village Gopesh, Përmetit), c. 1.3 km SE of village Kosinë, along 40°43’37.56”N 20°23’56.77”E, on calcareous river Vjosë, 40°15’40.11”N 20°19’08.84”E, on soil in rocky pastures, 1005 m, 3-V-2017, L. calcareous conglomerate rocks, 215 m, 17-X- Lőkös (BP 96676). Korçë County, Mt “Golines” 2011, Z. Barina, L. Lőkös & D. Pifkó (BP 96580). at village Rakickë, 40°43’8.88”N 20°59’0.97”E, District of Përmet (Rrethi i Përmetit), right part on calcareous soil in rocky grasslands, 1105 m, of river Vjosë between villages Golemas and 1-V-2017, L. Lőkös (BP 96672). District of Kutal, 40°15’22.93”N 20°19’27.80”E, on Pogradec, Udënisht, SH3 Piskutat-Pogradec, conglomerate rocks, 220 m, 10-V-2015, Z. shore Ohrid lake, 40º58’55”N 20º38’21”E, Pinus Barina, L. Lőkös & D. Pifkó (BP 96641). District halepensis, Juniperus oxycedrus, Buxus of Tepelenë, Dukaj, SH76 Tepelenë-Sinanaj, sempervirens, sandstone, 710 m, 22-IV-2017, 40º18’45”N 19º57’46”E, Quercus coccifera, A.R. Burgaz (MACB 111316). KUKËS: District limestone, 280 m, 18-IV-2017, A.R. Burgaz of Kukës, Kolsh, SH5 close Drin river, 42º04’18”N (MACB 111287). District of Tepelenë (Rrrethi i 20º20’32”E, deciduous Quercus, Juniperus Tepelenës), c. 1.4 km S of village Vasjar, oxycedrus, Buxus sempervirens, serpentine soil, 40°21’27.84”N 19°55’57.13”E, on calcareous 510 m, 21-IV-2017, A.R. Burgaz (MACB 111308). soil in exposed rocky grasslands, 155 m, 14-X- LEZHË: District of Lezhë (Rrethi i Lezhës), near 2012, L. Lőkös (BP 96596). KORÇË: Kolonjë the pass between Mt. “Sheshi i Velës” (1049.2 m) District (Rrethi i Kolonjës), between town and Mt. “Velës” (1170.1 m), above village Leskovik and village Vrëpckë, 40°10’30.62”N Kallmet, 41°50’28.1”N 19°42’54.5”E, on soil in 20°38’22.95”E, on acid soil in rocky serpentine rocky forest, 961 m, 23-IV-2009, L. Lőkös (BP grasslands, 1030 m, 10-V-2015, Z. Barina, L. 96548). SHKODËR: Shkodër District (Rrethi i Lőkös, D. Pifkó (BP 96638). District of Korçë Shkodrës), c. 1.5 km NE of village Gomsiqe e (Rrethi i Korçës), above the water reservoir c 2.5 Epërme, near road SH5, 41°59’31.61”N km WSW of village Ujëbardhë, 40°31’17.31”N 19°48’12.99”E, on acid soil in dry serpentine 20°39’50.62”E, on acid soil in dry rocky grasslands, 533 m, 6-V-2015, L. Lőkös (BP grassland, 1105 m, 18-X-2011, L. Lőkös (BP 96618). District of Shkodër (Rrethi i Shkodrës), 96587). District of Devoll (Rrethi i Devollit), on Qafa e Çelës above village Mirdicë, 42°0’11.98”N the hills above village Ponçarë, 40°31’17.63”N 19°43’33.16”E, on soil in roadside dry rocky 20°57’54.23”E, in dry grasslands, on flysh soil, grassland, 470 m, 12-X-2011, L. Lőkös (BP 1035 m, 11-V-2015, Z. Barina, L. Lőkös & D. Pifkó 96578). Pukë District (Rrethi i Pukës), Mount (BP 96646). District of Korçë, Dardhë, SH90 Rras’ e Qerretit above village Qerret, Boboshticë-Dardhë, 40º31’26”N 20º48’06”E, 42°03’36.98”N 19°49 53.45”E, in serpentine Fagus sylvatica, Juniperus communis, Abies sp., rocky area, on acid soil in pine woodland, 735 m, sandstone, 550 m, 23-IV-2017, A.R. Burgaz 6-V-2015, L. Lőkös (BP 96623). Shkodër District (MACB 111319). District of Korçë (Rrethi i (Rrethi i Shkodrës), in the valley of stream Përroi Korçës), on the ridge of Mts Mali i Moravës i Çibunit E of village Prekal, 42°10’44.69”N above village Dardhë, near peak Buzeja e 19°43’18.46”E, on calcareous soil in scrubland, Boboshticës (1701 m), 40°32’23.97”N 220 m, 5-V-2015, L. Lőkös (BP 96617). Shkodër 32 Burgaz, A. R. et al. Bot. complut. 43, 2019: 15-40

District (Rrethi i Shkodrës), between villages Kukës, Rruga Tirana, 42º04’46”N 20º24’45”E, -Egç-Lepurosh and Kurtaj, 42°11’10.22”N Quercus cerris, deciduous Quercus, limestone, 19°33’19.74”E, on calcareous soil in dry 330 m, 21-IV-2017, A.R. Burgaz (MACB scrubland, 260 m, 4-V-2015, Z. Barina, L. Lőkös 111310). TIRANË: District of Tiranë, Mushqeta, & D. Pifkó (BP 96613). District of Shkodër SH3 Mushqeta-Gracen, 41º12’38”N 19º57’09”E, (Rrethi i Shkodrës), in the valley of river Përroi i Arbutus unedo, Erica arborea, Platanus Thatë at village Lashaj, along the road nr. E 762, orientalis, boulders, 480 m, 22-IV-2017, A.R. 42°14’19.70”N 19°25’24.82”E, on calcareous Burgaz (MACB 111311). Fumarprotocetraric, soil in open dry rocky grasslands, 78 m, 30-III- usnic and psoromic acids chemotype: KORÇË: 2015, L. Lőkös (BP 96603). District of Malësi e District of Kolonjë (Rrethi i Kolonjës), Mt. Madhe, Poica, SH21 Koplik-Bogë, 42º15›53»N Melesin (1410.8 m), 40°08’50.1”N 20°35’37.0”E, 19º29›47»E, Juniperus oxycedrus, Cupressus on limestone rocks, 953 m, 3-V-2009, L. Lőkös sempervirens, boulders, 320 m, 20-IV-2017, A.R. (BP 96559). VLORË: District of Vlorë (Rrethi i Burgaz (MACB 111293). District of Malësi e Vlorës), between village Ilias and monastir Shën Madhe, Dedaj, SH21 Koplik-Bogë, 42º18’04”N Theodor, 40°08’07.8”N 19°40’26.4”E, on soil in 19º32’32”E, Carpinus betulus, Juniperus olive plantation, 211 m, 29-IV-2009, L. Lőkös oxycedrus, limestone rocks, 495 m, 20-IV-2017, (BP 96558). A.R. Burgaz (MACB 111294). District of Malësi e Madhe, Bogë, SH21 Koplik-Bogë, 42º21’46”N Cladonia furcata (Huds.) Schrad. 19º37’56”E, Pinus nigra, Juniperus oxycedrus, Fumarprotocetraric acid chemotype: limestone, 865 m, 20-IV-2017, A.R. Burgaz DIBËR: Dibër county, on the mount between (MACB 111302). Malësi e Madhe District villages Fushë Studë and Zabzun, 41°19’54.43”N (Rrethi i Malësisë së Madhe), on Mount Bukovik 20°24’38.25”E, on calcareous soil in rocky above village Rrapshë-Starë, 42°23’6.12”N grasslands, 1295 m, 18-V-2016, L. Lőkös (BP 19°29’25.56”E, on calcareous soil in rocky 96669). DURRËS: District of Durrës, Fushë-Draçi, grasslands, 715 m, 3-V-2015, L. Lőkös (BP road cape of Rodon, 41º32’50”N 19º31’08”E, 96608). TIRANË: District of Kavajë (Rrethi i Carpinus betulus, Arbutus unedo, Pistacia lentiscus, Kavajës), on hills east of village Stërbeg-Bragash, Cistus salviifolius, sandstone, 90 m, 19-IV-2017, 41°06’36.2”N 19°32’07.0”E, on soil in poplar A.R. Burgaz (MACB 111220). ELBASAN: woodland, 107 m, 28-IV-2009, L. Lőkös (BP District of Librazhd (Rrethi i Librazhdit): Polis Mts, 96554). VLORË: Himarë, Ilias, Himarë, between villages Gurrëshpat and Vilan, 40º08’29”N 19º40’11”E, Quercus coccifera, 41°05’38.45”N 20°16’15.72”E, on acid soil among Cupressus sempervirens, Cistaceae shrubs, serpentine rocks, 1270 m, 12-X-2010, L. Lőkös limestone, 300 m, 16-IV-2017, A.R. Burgaz (BP 96574). District of Elbasan, Bradashesh, SH3 (MACB 111283). District of Vlorë (Rrrethi i Bradashesh-Graze, 41º06’33”N 19º59’49”E, Vlorës), on the E ridge of Mt maja e Thanasit Carpinus betulus Juniperus oxycedrus, limestone (1352.1 m) above Llogora pass, 40°11’51.54”N karst, 530 m, 15-IV-2017, A.R. Burgaz (MACB 19°35’27.82”E, on calcareous soil in mixed 111403). District of Librazhd (Rrethi i Librazhdit): forest, 1075 m, 12-VIII-2010, L. Lőkös (BP Polis Mts; above village Polis, on the ridge between 96568). District of Vlorë, Dukat i Ri, SH3 the valley of brook “Sut Kukit” and river Shkumbin, Orikum-Dhërmi, Llogora N. P., 40º12’00”N in bushland, 41°09’16.50”N 20°15’08.79”E, on 19º35’26”E, Pinus nigra, Juniperus oxycedrus, conglomeratic flysh sediments, 555 m, 12-X-2010, Buxus sempervirens, limestone, 1030 m, 16-IV- L. Lőkös (BP 96572). Elbasan County (Qarku i 2017, A.R. Burgaz (MACB 111282). District of Elbasanit), on the mount above bridge Ura e Vlorë, Dukat i Ri, SH3 Orikum-Dhërmi, Llogora Murrashit, between Mirakë and Librazhd, N. P., 40º15’11”N 19º32’14”E, Quercus 41°11’1.64”N 20°16’33.36”E, on acid soil in coccifera, Pinus halepensis, limestone, 320 m, serpentine scrublands, 290 m, 3-IV-2016, L. Lőkös 16-IV-2017, A.R. Burgaz (MACB 111280). (BP 96658). Elbasan county, in village Floq, Fumarprotocetraric, usnic and physodalic 41°16’1.19”N 20°14’19.17”E, on soil by the acids chemotype: KORÇË: District of Korçë, roadside, 650 m, 17-V-2016, L. Lőkös (BP 96665). Dardhë, SH90 Boboshticë-Dardhë, 40º31’18”N GJIROKASTËR: District of Gjirokastër (Rrethi i 20º49’22”E, Fagus sylvatica, Pinus nigra, Gjirokastrës), between settlements Picar and Andon Juniperus communis, Betula pendula, Abies sp., Poçi in the valley of river Vjosë near road SH4, limestone, 1390 m, 23-IV-2017, A.R. Burgaz 40°10’41.12”N 20°04’45.55”E, in grazed (MACB 111320). KUKËS: District of Kukës, grassland, on flysh soil, 170 m, 28-III-2015, L. Burgaz, A. R. et al. Bot. complut. 43, 2019: 15-40 33

Lőkös (BP 96598). District of Tepelenë, Këlcyrë, coccifera, Carpinus betulus, Pinus halepensis, Këlcyrë gorge, left side Vjosa river, 40º17’52”N Cercis siliquastrum, limestone slopes, 125 m, 17- 20º10’05”E, Carpinus betulus, Pinus halepensis, IV-2017, A.R. Burgaz (MACB 111197). District of sandstone, 180 m, 18-IV-2017, A.R. Burgaz Vlorë (Rrethi i Vlorës), Mts. Mali i Çikës, in the (MACB 111215). KORÇË: Kolonjë District valley below Mt. “Maja e Pandelejmonit” (805.8 (Rrethi i Kolonjës), between town Leskovik and m), north of village Dhërmi, 40°09’32.2”N village Vrëpckë, 40°10’30.62”N 20°38’22.95”E, 19°38’49.4”E, in dry rocky grassland, on calcareous on acid soil in rocky serpentine grasslands, 1030 m, soil, 260 m, 29-IV-2009, L. Lőkös (BP 96555). 10-V-2015, Z. Barina, L. Lőkös & D. Pifkó (BP Fumarprotocetraric and physodalic acids 96639). District of Kolonjë (Rrethi i Kolonjës), c. chemotype: DURRËS: District of Durrës, Fushë- 1.9 km W of village Pepellash, 40°28’56.18”N Draçi, San Pietro beach track, 41º31’01”N 20°40’21.27”E, on acid soil in moist road slope, 19º31’12”E, Pinus pinea, sandy soil, 1 m, 19-IV- 549 m, 18-X-2011, Z. Barina, L. Lőkös & D. Pifkó 2017, A.R. Burgaz (MACB 111228). ELBASAN: (BP 96582). District of Korçë (Rrethi i Korçës), on Prrenjas, Urakë, SH9 Qafë Thanë, 41º04’24”N the ridge of Mts Mali i Moravës above village 20º36’18”E, Quercus macrolepis, deciduous Dardhë, near peak Buzeja e Boboshticës, Quercus, Juniperus oxycedrus, limestone, 1000 m, 40°31’52.6”N 20°48’36.1”E, on acid soil in beech 22-IV-2017, A.R. Burgaz (MACB 111314). forest on gravel conglomerate, 1625 m, 10-VIII- GJIROKASTËR: Dropull, Goranxi, to Kishae 2010, L. Lőkös (BP 96565). District of Korçë, Ravenes, 40º00’56”N 20º11’15”E, limestone rocks, Dardhë, SH90 Boboshticë-Dardhë, 40º31’18”N 268 m, 17-IV-2017, A.R. Burgaz (MACB 111206). 20º49’22”E, Fagus sylvatica, Pinus nigra, GJIROKASTËR: District of Përmet, Piskovë, left Juniperus communis, Betula pendula, Abies sp., side Lumica river, K75 to Frashër, 40º16’44”N limestone, 1390 m, 23-IV-2017, A.R. Burgaz 20º17’07”E, Juniperus oxycedrus, Paliurus spina- (MACB 111344). District of Devoll (Rrethi i christi, Arbutus unedo, Pistacia terebinthus, Devollit), on the mount between village Tren and sandstone, 230 m, 18-IV-2017, A.R. Burgaz lake Liqeni i Prespës së Vogël, 40°40’10.55”N (MACB 111218). District of Tepelenë, Këlcyrë, 20°59’12.62”E, on calcareous soil in rocky Këlcyrë gorge, left side Vjosa river, 40º17’52”N grasslands, 910 m, 12-V-2015, Z. Barina, L. Lőkös 20º10’05”E, Carpinus betulus, Pinus halepensis, & D. Pifkó (BP 96649). Korçë County, Mali i sandstone, 180 m, 18-IV-2017, A.R. Burgaz Lenies at village Gopesh, 40°43’37.56”N (MACB 111213). District of Tepelenë, Dukaj, 20°23’56.77”E, on calcareous soil in rocky pastures, SH76 Tepelenë-Sinanaj, 40º18’45”N 19º57’46”E, 1005 m, 3-V-2017, L. Lőkös (BP 96677). LEZHË: Quercus coccifera, limestone, 280 m, 18-IV-2017, Pukë District (Rrethi i Pukës); Mount Rras’ e A.R. Burgaz (MACB 111221). KORÇË: District of Qerretit above village Qerret, 42°03’36.98”N Korçë, Dardhë, SH90 Boboshticë-Dardhë, 19°49’53.45”E, in serpentine rocky area, on acid 40º31’18”N 20º49’22”E, Fagus sylvatica, Pinus soil in pine woodland, 735 m, 6-V-2015, L. Lőkös nigra, Juniperus communis, Betula pendula, Abies (BP 96624). SHKODËR: District of Shkodër sp., limestone, 1390 m, 23-IV-2017, A.R. Burgaz (Rrethi i Shkodrës), Qafa e Çelës above village (MACB 111321). District of Pogradec, Udënisht, Mirdicë, 42°0’11.98”N 19°43’33.16”E, on soil in SH3 Piskutat-Pogradec, shore Ohrid lake, roadside dry rocky grassland, 470 m, 12-X-2011, L. 40º58’55”N 20º38’21”E, Pinus halepensis, Lőkös (BP 96579). District of Shkodër (Rrethi i Juniperus oxycedrus, Buxus sempervirens, Shkodrës), in the valley of river Përroi i Thatë at sandstone, 710 m, 22-IV-2017, A.R. Burgaz village Lashaj, along the road nr. E 762, (MACB 111315). KUKËS: District of Kukës, 42°14’19.70”N 19°25’24.82”E, on calcareous soil Kolsh, SH5 close Drin river, 42º04’18”N in open dry rocky grasslands, 78 m, 30-III-2015, L. 20º20’32”E, deciduous Quercus, Juniperus Lőkös (BP 96604). TIRANË: oxycedrus, Buxus sempervirens, serpentine soil, (Qarku i Tiranës); at the foot of Mt Kalaja e Dorëzit, 510 m, 21-IV-2017, A.R. Burgaz (MACB 111307). between villages Pezë and Dorëz, 41°13’30.08”N District of Kukës, Kukës, Rruga Tirana, 42º04’46”N 19°43’53.26”E, in grasslands, on flysh soil, 180 m, 20º24’45”E, Quercus cerris, deciduous Quercus, 2-IV-2016, L. Lőkös (BP 96652). VLORË: District limestone, 330 m, 21-IV-2017, A.R. Burgaz of Sarandë (Rrrethi i Sarandës), on a hill between (MACB 111309). LEZHË: District of Kurbin, villages Kulluricë and Kodër, 39°48’43.85”E Milot, left side Mat river, SH25 Milot-Zogu Bridge, 20°06’34.02”E, in dry grassland, on flysh soil, 55 41º41’54”N 19º45’10”E, Carpinus orientalis, m, 13-X-2012, L. Lőkös (BP 96591). Finiq, Bistricë, Erica arborea, limestone, 40 m, 21-IV-2017, A.R. Muzinë, 39º55’08”N 20º08’48”E, Quercus Burgaz (MACB 111306). SHKODËR: District of 34 Burgaz, A. R. et al. Bot. complut. 43, 2019: 15-40

Malësi e Madhe, Poica, SH21 Koplik-Bogë, calcareous soil in open, rocky scrublands, 1295 m, 42º15’53”N 19º29’47”E, Juniperus oxycedrus, 1-V-2017, L. Lőkös (BP 96674). Cupressus sempervirens, boulders, 320 m, 20-IV- 2017, A.R. Burgaz (MACB 111291). District of Cladonia humilis (With.) J.R. Laundon Malësi e Madhe, Dedaj, SH21 Koplik-Bogë, Atranorin and fumarprotocetraric acid: 42º18’04”N 19º32’32”E, Carpinus betulus, ELBASAN: Prrenjas, Prrenjas, SH3 Qukës- Juniperus oxycedrus, limestone rocks, 495 m, 20- Prrenjas, 41º04’24”N 20º30’15”E, Carpinus IV-2017, A.R. Burgaz (MACB 111297). District of orientalis, Quercus macrolepis, deciduous Malësi e Madhe, Bogë, SH21 Koplik-Bogë, Quercus, Buxus sempervirens, limestone slopes, 42º21’46”N 19º37’56”E, Pinus nigra, Juniperus 540 m, 22-IV-2017, A.R. Burgaz (MACB oxycedrus, limestone, 865 m, 20-IV-2017, A.R. 111444). VLORË: District of Sarandë, Pasqirave, Burgaz (MACB 111301). VLORË: Finiq, Muzinë, Butrint N. P., SH81, Shengjergj monastery, Muzinë, 39º54’48”N 20º11’20”E, deciduous 39º48’42”N 20º00’43”E, Quercus coccifera, Quercus, Erica arborea, limestone, 186 m, 17-IV- limestone slopes, 50 m, 17-IV-2017, A.R. Burgaz 2017, A.R. Burgaz (MACB 111200). Finiq, (MACB 111407). Bistricë, Muzinë, 39º55’08”N 20º08’48”E, Quercus coccifera, Carpinus betulus, Pinus Cladonia macilenta Hoffm. halepensis, Cercis siliquastrum, limestone slopes, Barbatic and thamnolic acids: LEZHË: 125 m, 17-IV-2017, A.R. Burgaz (MACB 111195). District of Mirditë (Rrethi i Mirditës); in the Atranorin and fumarprotocetraric acid: valley of stream Fani i Vogël above village KORÇË: District of Korçë (Rrethi i Korçës), on the Thirrë; in serpentine rocky area, on bark (Pinus ridge of Mts Mali i Moravës above village Dardhë, nigra). 42° 0’ 20.98”N 20°14’33.25”E, 905 m, near peak Buzeja e Boboshticës (1701 m), 6-V-2015, L. Lőkös (BP 96626). 40°32’23.97”E 20°49’14.20”E, on acid soil in beech forest on gravel conglomerate, 1550 m, 10- Cladonia merochlorophaea Asahina Fumarprotocetraric and merochlorophaeic VIII-2010, L. Lőkös (BP 96562). Atranorin, acids: KORÇË: District of Korçë (Rrethi i fumarprotocetraric and physodalic acids Korçës), on the ridge of Mts Mali i Moravës chemotype: KORÇË: District of Korçë, Dardhë, above village Dardhë, near peak Buzeja e SH90 Boboshticë-Dardhë, 40º31’26”N Boboshticës, on calcareous soil, 40°31’52.6”N 20º48’06”E, Fagus sylvatica, Juniperus communis, 20°48’36.1”E, 1625 m, 10-VIII-2010, L. Lőkös Abies sp., sandstone, 1550 m, 23-IV-2017, A.R. (BP 96566). Burgaz (MACB 111318). SHKODËR: District of Malësi e Madhe, Poica, SH21 Koplik-Bogë, Cladonia parasitica (Hoffm.) Hoffm. 42º15’53”N 19º29’47”E, Juniperus oxycedrus, Barbatic and thamnolic acids: VLORË: Cupressus sempervirens, boulders, 320 m, 20-IV- District of Delvinë (Rrethi i Delvinës), Mali i 2017, A.R. Burgaz (MACB 111290). District of Gjerë, on the southern slope of 1331 m high Malësi e Madhe, Dedaj, SH21 Koplik-Bogë, mount above Delvinë; in chestnut woodland, 42º18’04”N 19º32’32”E, Carpinus betulus, on bark (Castanea sativa), 39°57’42.12”N 20° Juniperus oxycedrus, limestone rocks, 495 m, 20- 07’59.66”E, 828 m, 1-V-2009, Z. Barina & D. IV-2017, A.R. Burgaz (MACB 111295, 111296). Pifkó (BP 96545). Malësi e Madhe, Bogë, SH21 Koplik-Bogë, 42º21’46”N 19º37’56”E, Pinus nigra, Juniperus Cladonia peziziformis (With.) J.R. Laundon oxycedrus, limestone, 865 m, 20-IV-2017, A.R. Fumarprotocetraric acid: SHKODËR: Vau Burgaz (MACB 111300). VLORË: Finiq, Muzinë, i Dejës, SH25 Vau i Dejës-Koman, 42º00’55”N Muzinë, 39º54’48”N 20º11’20”E, deciduous 19º39’37”E, Pinus pinaster, Carpinus orientalis, Quercus, Erica arborea, limestone, 186 m, 17-IV- serpentine soil, 120 m, 20-IV-2017, A.R. Burgaz 2017, A.R. Burgaz (MACB 111284). District of (MACB 111418). Vlorë, Dukat i Ri, SH3 Orikum-Dhërmi, Llogora N. P., 40º15’11”N 19º32’14”E, Quercus coccifera, Cladonia polycarpoides Nyl. (= C. subcariosa Pinus halepensis, limestone, 320 m, 16-IV-2017, auct.) A.R. Burgaz (MACB 111281). Atranorin, Norstictic acid chemotype: SHKODËR: Vau bourgeanic and fumarprotocetraric acids: i Dejës, SH25 Vau i Dejës-Koman, 42º03’01”N KORÇË: Korçë County, Mt Golines at village 19º39’45”E, Pinus pinaster, Juniperus oxycedrus, Rakickë, 40°43’42.80”N 20°59’28.06”E, on Quercus cerris, Erica arborea, Cotinus coggygria, Burgaz, A. R. et al. Bot. complut. 43, 2019: 15-40 35 serpentine soil, 130 m, 20-IV-2017, A.R. Burgaz (MACB 111341). District of Korçë (Rrethi i (MACB 111427). Atranorin and norstictic acid Korçës); Mali i Strugarit above village Belorta, chemotype: KUKËS: District of Kukës, Kolsh, 40°39’56.56”N 20° 50’ 9.79”E, open grasslands SH5 close Drin river, 42º04’18”N 20º20’32”E, on conglomerate rocks, 1010 m, 11-V-2015, Z. deciduous Quercus, Juniperus oxycedrus, Buxus Barina, L. Lőkös, D. Pifkó (BP 96644). District sempervirens, serpentine soil, 510 m, 21-IV-2017, of Devoll (Rrethi i Devollit), on the mount A.R. Burgaz (MACB 111432). between village Tren and lake Liqeni i Prespës së Vogël, 40°40’10.55”N 20°59’12.62”E, on Cladonia pyxidata (L.) Hoffm. calcareous soil in rocky grasslands, 910 m, 12-V- Fumarprotocetraric acid chemotype: DIBËR: 2015, Z. Barina, L. Lőkös, D. Pifkó (BP 96650). Dibër county, Golloborde, in the dry valley Gropa Korçë County, Mt Golines at village Rakickë, Gjugarica above village Sebisht, 41°23’34.22”N 40°43’8.88”N 20°59’0.97”E, on calcareous soil 20°25’16.33”E, on calcareous soil in rocky in rocky grasslands, 1105 m, 1-V-2017, L. Lőkös grasslands, 1015 m, 19-V-2016, L. Lőkös (BP (BP 96673). District of Korçë (Rrethi i Korçës), 96670). Dibër county, Mt Serakoli above villae Thatë Mountains (Mali i Thatë), c. 3.0 km north of Ploshtan, 41°50’44.99”N 20°26’43.85”E, on village Zvezdë, c. 1.85 km south-southeast of the calcareous soil in rocky pastures, 1200 m, 21-V- peak of Mount “maja e Shëngjergjit” (1921 m), 2016, L. Lőkös (BP 96671). DURRËS: District 40°44’37.16”N 20°52’35.53”E, on calcareous soil of Krujë, Krujë, SH38 to Qafë-Shtamë N. P., in rocky grasslands, 1165 m, 18-X-2011, L. Lőkös 41º31’03”N 19º49’27”E, Carpinus betulus, (BP 96589). SHKODËR: Vau i Dejës, SH25 Vau pasturelands, limestone, 720 m, 19-IV-2017, A.R. i Dejës-Koman, 42º03’01”N 19º39’45”E, Pinus Burgaz (MACB 111229). ELBASAN: Elbasan pinaster, Juniperus oxycedrus, Quercus cerris, county, in open boxwood scrubland above village Erica arborea, Cotinus coggygria, serpentine soil, Kutërman, 41°13’40.99”N 20°14’41.04”E, on 130 m, 20-IV-2017, A.R. Burgaz (MACB 111423). acid soil in rocky serpentine area, 510 m, 16-V- Shkodër District (Rrethi i Shkodrës), in the valley 2016, L. Lőkös (BP 96661). FIER: of stream Përroi i Çibunit E of village Prekal, (Rrethi i Fierit), Mali i Kalivaçit, S of village 42°10’50.54”N 19°43’42.57”E, on calcareous Kalivaç, above river Vjosë, 40°24’14.86”N soil in scrubland, 250 m, 5-V-2015, L. Lőkös (BP 19°48’4.08”E, on calcareous soil in Quercus 96615). District of Malësi e Madhe, Dedaj, SH21 coccifera scrubland, 100 m, 8-V-2015, L. Lőkös Koplik-Bogë, 42º18’04”N 19º32’32”E, Carpinus (BP 96629). GJIROKASTËR: Gjirokastër betulus, Juniperus oxycedrus, limestone rocks, County, Bredhi i Sotirës above village Sotirë, 495 m, 20-IV-2017, A.R. Burgaz (MACB 39°48’53.21”N 20°21’41.98”E, on mossy rocks 111326). District of Malësi e Madhe, Bogë, SH21 in dry scrublands, 550 m, 6-V-2017, L. Lőkös (BP Koplik-Bogë, 42º21’46”N 19º37’56”E, Pinus 96681). Gjirokastër County, at village Derviçian, nigra, Juniperus oxycedrus, limestone, 865 m, 20- 40°01’24.45”N 20°10’39.21”E, on calcareous IV-2017, A.R. Burgaz (MACB 111327). Malësi soil in rocky grasslands, 260 m, 7-V-2017, L. e Madhe District (Rrethi i Malësisë së Madhe), Lőkös (BP 96684). District of Përmet (Rrethi i on Mount Bukovik above village Rrapshë-Starë, Përmetit), c. 1.3 km SE of village Kosinë, along 42°23’6.12”N 19°29’25.56”E, on calcareous river Vjosë, 40°15’40.11”N 20°19’8.84”E, on soil in rocky grasslands, 715 m, 3-V-2015, L. calcareous conglomerate rocks, 215 m, 17-X- Lőkös (BP 96609). TIRANË: District of Tiranë, 2011, Z. Barina, L. Lőkös, D. Pifkó (BP 96581). Mushqeta, SH3 Mushqeta-Gracen, 41º12’38”N District of Përmet (Rrethi i Përmetit), left part of 19º57’09”E, Arbutus unedo, Erica arborea, river Vjosë, between town Këlcyrë and village Platanus orientalis, boulders, 480 m, 22-IV-2017, Mbrezhdan, 40°18’17.42”N 20°12’17.52”E, A.R. Burgaz (MACB 111337). Tirana County in scrubland, on flysh soil, 205 m, 9-V-2015, (Qarku i Tiranës), between village Pëllumbas L. Lőkös (BP 96686). Tepelenë District and cave Shpella e Pëllumbasit, 41°15’4.10”E (Rrethi i Tepelenës), in village Mezhgoran, 19°57’34.16”E, on calcareous soil in Carpinus 40°19’11.24”N 20°05’43.54”E, on calcareous orientalis woodland, 445 m, 3-IV-2016, L. Lőkös soil, 660 m, 9-V-2015, Z. Barina, L. Lőkös, D. (BP 96654). Tirana County (Qarku i Tiranës), Pifkó (BP 96634). KORÇË: District of Pogradec, near cave Shpella e Pëllumbasit above village Udënisht, SH3 Piskutat-Pogradec, shore Ohrid Pëllumbas, 41°15’21.11”N 19°57’50.31”E, on lake, 40º58’55”N 20º38’21”E, Pinus halepensis, calcareous soil in Carpinus orientalis woodland, Juniperus oxycedrus, Buxus sempervirens, 450 m, 3-IV-2016, L. Lőkös (BP 96656). sandstone, 710 m, 22-IV-2017, A.R. Burgaz VLORË: District of Sarandë (Rrrethi i Sarandës), 36 Burgaz, A. R. et al. Bot. complut. 43, 2019: 15-40 on the eastern slope of Mt Maja e Milletit, above Quercus, Buxus sempervirens, limestone slopes, village Sopik, 39°46’12.49”N 20°06’20.58”E, on 540 m, 22-IV-2017, A.R. Burgaz (MACB calcareous soil in rocky grasslands, 160 m, 13-X- 111312). 2012, L. Lőkös (BP 96594). District of Sarandë (Rrrethi i Sarandës), on a hill between villages Cladonia ramulosa (With.) J.R. Laundon Kulluricë and Kodër, 39°48’43.85”N 20° 06’ Fumarprotocetraric acid: DURRËS: 34.02”E, in dry grassland, on flysh soil, 55 m, District of Durrës, Fushë-Draçi, San Pietro 13-X-2012, L. Lőkös (BP 96592). Ilias, Himarë, beach track, 41º31’01”N 19º31’12”E, Pinus 40º08’29”N 19º40’11”E, Quercus coccifera, pinea, sandy soil, 1 m, 19-IV-2017, A.R. Bur- Cupressus sempervirens, Cistaceae shrubs, gaz (MACB 111226). LEZHË: Kurbin, Milot, limestone, 300 m, 16-IV-2017, A.R. Burgaz left side Mat river, SH25 Milot-Zogu Bridge, (MACB 110961). District of Vlorë (Rrrethi i 41º41’54”N 19º45’10”E, Carpinus orientalis, Vlorës); on the E ridge of Mt maja e Thanasit Erica arborea, limestone, 40 m, 21-IV-2017, (1352.1 m) above Llogora pass, 40°11’51.54”N A.R. Burgaz (MACB 111332, GenBank acces- 19°35’27.82”E, on calcareous soil in mixed sion nº: MK543954). forest, 1075 m, 12-VIII-2010, L. Lőkös (BP 96569). Fumarprotocetraric and physodalic Cladonia rangiformis Hoffm. acids chemotype: GJIROKASTËR: Dropull, Atranorin and rangiformic acid chemotype: Grapsh, SH78, 39º56’50”N 20º14’31”E, BERAT: District of Berat (Rrethi i Beratit), Tomor deciduous Quercus, limestone slope rocks, 440 Mountains (Mali i Tomorrit), c. 700 m north of m, 17-IV-2017, A.R. Burgaz (MACB 111201, village Karkanjoz, 40°41’55.61”N 20°03’28.69”E, 111203). Dropull, Goranxi, to Kishae Ravenes, in Mediterranean scrubland, on calcareous soil in 40º00’56”N 20º11’15”E, limestone rocks, 268 rocky grassland, 490 m, 25-V-2004, Z. Barina, Cs. m, 17-IV-2017, A.R. Burgaz (MACB 111205). Németh (BP 96541). ELBASAN: Elbasan county, District of Tepelenë, Kodër, left Vjosa river bank, in open boxwood scrubland above village 40º17’20”N 20º04’25”E, Paliurus spina-christi, Kutërman, 41°13’40.99”N 20°14’41.04”E, on Spartium junceum, Phlomis shrubs, limestone acid soil in rocky serpentine area, 510 m, 16-V- slopes, 160 m, 18-IV-2017, A.R. Burgaz (MACB 2016, L. Lőkös (BP 96662). GJIROKASTËR: 111208, 111209). District of Tepelenë, Këlcyrë, District of Përmet (Rrethi i Përmetit), left part of Këlcyrë gorge, left side Vjosa river, 40º17’52”N river Vjosë, between town Këlcyrë and village 20º10’05”E, Carpinus betulus, Pinus halepensis, Mbrezhdan, 40°18’17.42”N 20°12’17.52”E, in sandstone, 180 m, 18-IV-2017, A.R. Burgaz scrubland, on flysh soil, 205 m, 9-V-2015, L. (MACB 111212, 111214). VLORË: Finiq, Bistricë, Lőkös (BP 96636). KORÇË: Kolonjë District Muzinë, 39º55’08”N 20º08’48”E, Quercus (Rrethi i Kolonjës); between town Leskovik and coccifera, Carpinus betulus, Pinus halepensis, village Vrëpckë, 40°10’30.62”N 20°38’22.95”E, Cercis siliquastrum, limestone slopes, 125 m, 17- on acid soil in rocky serpentine grasslands, 1030 IV-2017, A.R. Burgaz (MACB 111192). District of m, 10-V-2015, Z. Barina, L. Lőkös & D. Pifkó (BP Vlorë, Dukat i Ri, SH3 Orikum-Dhërmi, Llogora N. 96640). District of Devoll (Rrethi i Devollit), on P., 40º12’00”N 19º35’26”E, Pinus nigra, Juniperus the hills above village Ponçarë, 40°31’17.63”N oxycedrus, Buxus sempervirens, limestone, 1030 20°57’54.23”E, in dry grasslands, on flysh soil, m, 16-IV-2017, A.R. Burgaz (MACB 110957, 1035 m, 11-V-2015, Z. Barina, L. Lőkös & D. 110958). District of Vlorë, Dukat i Ri, SH3 Orikum- Pifkó (BP 96647). Korçë County, Mal Moravë Dhërmi, Llogora N. P., 40º15’11”N 19º32’14”E, above village Hoçisht, 40°36’12.59”N Quercus coccifera, Pinus halepensis, limestone, 20°54’33.39”E, on calcareous soil in oak 320 m, 16-IV-2017, A.R. Burgaz (MACB woodland, 1030 m, 2-V-2017, L. Lőkös (BP 110954, 110955). Fumarprotocetraric and 96675). District of Korçë (Rrethi i Korçës), Mali i psoromic acids chemotype: BERAT: District of Strugarit above village Belorta, 40°39’56.56”N Berat (Rrethi i Beratit), Tomor Mountains (Mali 20°50’9.79”E, open grasslands on conglomerate i Tomorrit), c. 1800 m SSW of village Tomor; in rocks, 1010 m, 11-V-2015, Z. Barina, L. Lőkös & Mediterranean scrubland, on calcareous soil in D. Pifkó (BP 96645). District of Devoll (Rrethi i rocky grassland, 40°40’53.98”N 20° 07’06.64”E, Devollit), on the mount between village Tren and 1080 m, 25-V-2004, Z. Barina & Cs. Németh lake Liqeni i Prespës së Vogël, 40°40’10.55”N (BP 96539). ELBASAN, Prrenjas, Prrenjas, SH3 20°59’12.62”E, on calcareous soil in rocky Qukës-Prrenjas, 41º04’24”N 20º30’15”, Carpinus grasslands, 910 m, 12-V-2015, Z. Barina, L. Lőkös orientalis, Quercus macrolepis, deciduous & D. Pifkó (BP 96651). District of Pogradec, Burgaz, A. R. et al. Bot. complut. 43, 2019: 15-40 37

Udënisht, SH3 Piskutat-Pogradec, shore Ohrid Bradashesh-Graze, 41º06’33”N 19º59’49”E, lake, 40º58’55”N 20º38’21”E, Pinus halepensis, Carpinus betulus, Juniperus oxycedrus, limestone Juniperus oxycedrus, Buxus sempervirens, karst, 530 m, 15-IV-2017, A.R. Burgaz (MACB sandstone, 710 m, 22-IV-2017, A.R. Burgaz 111404). District of Librazhd (Rrethi i Librazhdit): (MACB 111342). KUKËS: District of Kukës, Polis Mts, above village Polis, on the ridge Kolsh, SH5 close Drin river, 42º04’18”N between the valley of brook “Sut Kukit” and river 20º20’32”E, deciduous Quercus, Juniperus Shkumbin, 41°09’16.50”N 20°15’08.79”E, in oxycedrus, Buxus sempervirens, serpentine soil, bushland, on conglomeratic flysh sediments, 555 510 m, 21-IV-2017, A.R. Burgaz (MACB 111334). m, 12-X-2010, L. Lőkös (BP 96573). Elbasan LEZHË: District of Lezhë (Rrethi i Lezhës), on County (Qarku i Elbasanit), on the mount above the slope of Mt Guri i Kuq above village Fregën, bridge Ura e Murrashit, between Mirakë and 41°51’41.58”N 19°44’11.22”E, on acid soil in Librazhd, 41°11’1.64”N 20°16’33.36”E, on acid open serpentine grasslands, 575 m, 30-III-2015, L. soil in serpentine scrublands, 290 m, 3-IV-2016, L. Lőkös (BP 96607). SHKODËR: Shkodër District Lőkös (BP 96659). Elbasan county, in village (Rrethi i Shkodrës), c. 1.5 km NE of village Floq, 41°16’1.19”N 20°14’19.17”E, on soil by the Gomsiqe e Epërme, near road SH5, 41°59’31.61”N roadside, 650 m, 17-V-2016, L. Lőkös (BP 96666). 19°48’12.99”E, on acid soil in dry serpentine FIER: District of Fier (Rrrethi i Fierit), Pishë-Poro grasslands, 533 m, 6-V-2015, L. Lőkös (BP Nature Reserve, between village Pishë and the 96619). Pukë District (Rrethi i Pukës); Mount Adriatik Sea, 40°40’3.67”N 19°22’17.76”E, on Rras’ e Qerretit above village Qerret, soil in stone pine forest, 5 m, 12-VIII-2010, L. 42°03’36.98”N 19°49’53.45”E, in serpentine Lőkös (BP 96567). Lushnjë District (Rrethi i rocky area, on acid soil in pine woodland, 735 m, Lushnjës), Golda e Kullarit W of village Kular, ca 6-V-2015, L. Lőkös (BP 96625). District of 4 km NW of Divjakë, 41°01’6.54”N Shkodër (Rrethi i Shkodrës), in the valley of river 19°29’24.54”E, in pine woodland on sandy Përroi i Thatë at village Lashaj, along the road nr. E seacoast area, 5 m, 7-V-2015, L. Lőkös (BP 762, 42°14’19.70”N 19°25’24.82”E, on calcareous 96628). Fier District (Rrethi i Fierit), Mali i soil in open dry rocky grasslands, 78 m, 30-III- Kalivaçit, S of village Kalivaç, above river Vjosë, 2015, L. Lőkös (BP 96605). VLORË: District of 40°24’14.86”N 19°48’4.08”E, on calcareous soil Vlorë (Rrrethi i Vlorës), at seashore rrugët e Bardha in Quercus coccifera scrubland, 100 m, 8-V-2015, west of village Palasë, 40°10’30.46”N L. Lőkös (BP 96630). Fier County, Pylli i Levanit, 19°36’9.32”E, on calcareous soil in dry roadside above village Levan, 40°40’48.99”N scrubland, 225 m, 12-X-2012, L. Lőkös (BP 19°30’11.70”E, on calcareous soil in oak 96590). Atranorin, fumarprotocetraric and woodland, 95 m, 5-V-2017, L. Lőkös (BP 96679). rangiformic acids chemotype: BERAT: District GJIROKASTËR: Gjirokastër County, Bredhi i of Skrapar (Rrethi i Skraparit), above the gorge of Sotirës above village Sotirë, 39°48’53.21”N river Osum next to village Dhorës, 40°28’27.79”N 20°21’41.98”E, on mossy rocks in dry scrublands, 20°15’16.11”E, in open bushland, on calcareous 550 m, 6-V-2017, L. Lőkös (BP 96682). Gjirokastër soil, 420 m, 13-X-2010, L. Lőkös (BP 96576). County, N of village Koshovicë, in Platanus DIBËR: District of Mat (Rrethi i Matit), Mali i woodland, 39°50’11.52”N 20°23’51.90”E, on soil Skënderbeut, on the northern forehills of Mt. “maja in rocky grasslands, 430 m, 6-V-2017, L. Lőkös e Mëllezit”, near village Shkopet, 41°41’28.5”N (BP 96683). Dropull, Grapsh, SH78, 39º56’50”N 19°49’14.8”E, on soil, 222 m, 24-IV-2009, L. 20º14’31”E, deciduous Quercus, limestone slope Lőkös (BP 96550). ELBASAN: Prrenjas, SH3 rocks, 440 m, 17-IV-2017, A.R. Burgaz (MACB Qukës-Prrenjas, 41º04’24»N 20º30’15»E, 111204). Dropull, Goranxi, to Kishae Ravenes, Carpinus orientalis, Quercus macrolepis, deciduous 40º00’56”N 20º11’15”E, limestone rocks, 268 m, Quercus, Buxus sempervirens, limestone slopes, 17-IV-2017, A.R. Burgaz (MACB 111207). 540 m, 22-IV-2017, A.R. Burgaz (MACB 111339). Gjirokastër District (Rrethi i Gjirokastrës), Mount Prrenjas, Urakë, SH9 Qafë Thanë, 41º04’24”N Maja e Lalucit above village Saraqinishtë, 20º36’18”E, Quercus macrolepis, deciduous 40°06’57.10”N 20°14’20.54”E, on calcareous soil Quercus, Juniperus oxycedrus, limestone, 1000 m, in rocky grasslands, 1055 m, 9-V-2015, L. Lőkös 22-IV-2017, A.R. Burgaz (MACB 111340). District (BP 96632). District of Gjirokastër (Rrethi i of Elbasan, Bradashesh, SH3 Bradashesh-Graze, Gjirokastrës), between settlements Picar and 41º06’05”N 20º00’26”E, Olea europea, limestone Andon Poçi in the valley of river Vjosë near road karst, 390 m, 15-IV-2017, A.R. Burgaz (MACB SH4, 40°10’41.12”N 20°04’45.55”E, in grazed 111401). District of Elbasan, Bradashesh, SH3 grassland, on flysh soil, 170 m, 28-III-2015, L. 38 Burgaz, A. R. et al. Bot. complut. 43, 2019: 15-40

Lőkös (BP 96599). District of Gjirokastër (Rrethi Pinus pinaster, Juniperus oxycedrus, Quercus i Gjirokastrës); Kurvelesh i Sipërm uplands above cerris, Erica arborea, Cotinus coggygria, village Picar, 40°11’0.39”N 20°02’31.51”E, on serpentine soil, 130 m, 20-IV-2017, A.R. Burgaz calcareous soil in rocky grasslands, 750 m, 29-III- (MACB 111329). Vau i Dejës, Karmë, SH25 Vau 2015, L. Lőkös (BP 96601). District of Tepelenë, i Dejës-Koman, 42º04’02”N 19º40’59”E, Quercus Kodër, left Vjosa river bank, 40º17’20”N macrolepis, Q. pubescens, Juniperus oxycedrus, 20º04’25”E, Paliurus spina-christi, Spartium Erica arborea, slates, 130 m, 20-IV-2017, A.R. junceum, Phlomis shrubs, limestone slopes, 160 Burgaz (MACB 111331). District of Shkodër m, 18-IV-2017, A.R. Burgaz (MACB 111210). (Rrethi i Shkodrës), in the valley of river Përroi i Tepelenë District (Rrethi i Tepelenës), in village Thatë at village Lashaj, along the road nr. E 762, Mezhgoran, 40°19’11.24”N 20°05’43.54”E, on 42°14’19.70”N 19°25’24.82”E, on calcareous soil calcareous soil, 660 m, 9-V-2015, Z. Barina, L. in open dry rocky grasslands, 78 m, 30-III-2015, Lőkös, D. Pifkó (BP 96635). District of Tepelenë L. Lőkös (BP 96605). LEZHË: District of Lezhë (Rrrethi i Tepelenës), c. 1.4 km S of village Vasjar, (Rrethi i Lezhës), Mali i Rrencit, near the peak of 40°21’27.84”N 19°55’57.13”E, on calcareous soil Mt Burga (262 m), above village Shëngjin, in exposed rocky grasslands, 155 m, 14-X-2012, 41°49’28.2”N 19°35’03.9”E, in open oak L. Lőkös (BP 96597). KORÇË: Kolonjë District woodland, on limestone together with bryophytes, (Rrethi i Kolonjës); between town Leskovik and 255 m, 22-IV-2009, L. Lőkös (BP 96547). village Vrëpckë, 40°10’30.62”N 20°38’22.95”E, SHKODËR: Shkodër District (Rrethi i Shkodrës), on acid soil in rocky serpentine grasslands, 1030 in the valley of stream Përroi i Çibunit E of village m, 10-V-2015, Z. Barina, L. Lőkös, D. Pifkó (BP Prekal, 42°10’50.54”N 19°43’42.57”E, on 96640). District of Kolonjë (Rrethi i Kolonjës), c. calcareous soil in scrubland, 250 m, 5-V-2015, L. 1.9 km W of village Pepellash, 40°28’56.18”E Lőkös (BP 96616). Shkodër District (Rrethi i 20°40’21.27”N, on acid soil in moist road slope, Shkodrës), between villages Linaj-Egç-Lepurosh 549 m, 18-X-2011, Z. Barina, L. Lőkös, D. Pifkó and Kurtaj, 42°11’10.22”N 19°33’19.74”E, on (BP 96583). District of Devoll (Rrethi i Devollit), calcareous soil in dry scrubland, 260 m, 4-V-2015, on the hills above village Ponçarë, 40°31’17.63”N Z. Barina, L. Lőkös, D. Pifkó (BP 96614). Malësi 20°57’54.23”E, in dry grasslands, on flysh soil, e Madhe District (Rrethi i Malësisë së Madhe), on 1035 m, 11-V-2015, Z. Barina, L. Lőkös, D. Pifkó Mount Bukovik above village Rrapshë-Starë, (BP 96647). District of Korçë (Rrethi i Korçës), 42°23’6.12”N 19°29’25.56”E, on calcareous soil above the water reservoir c 2.5 km WSW of village in rocky grasslands, 715 m, 3-V-2015, L. Lőkös Ujëbardhë, 40°31’17.31”N 20°39’50.62”E, on (BP 96610). TIRANË: District of Kavajë (Rrethi i acid soil in dry rocky grassland, 1105 m, 18-X- Kavajës), by the coast of the Adriatic Sea between 2011, L. Lőkös (BP 96588). District of Korçë, villages Spille and Rreth (Rreth-Greth), Boboshticë, SH90 Boboshticë-Dardhë, 41°05’07.6”N 19°27’31.5”E, on sandy soil in 40º32’37”N 20º47’11”E, Juniperus oxycedrus, sand dune area, 10 m, 27-IV-2009, L. Lőkös (BP pastureland, sandstone, 1200 m, 23-IV-2017, A.R. 96553). Tirana County (Qarku i Tiranës), at the Burgaz (MACB 111343). District of Korçë (Rrethi foot of Mt Kalaja e Dorëzit, between villages Pezë i Korçës); Mts Mali i Moravës, near the peak of and Dorëz, 41°13’30.08”N 19°43’53.26”E, in Mt Maja e Shëndëllisë (1365 m) above village grasslands, on flysh soil, 180 m, 2-IV-2016, L. Mborje, 40°36’39.81”E 20° 48’ 55.00”N, on acid Lőkös (BP 96653). Tirana County (Qarku i soil in serpentine rocky area, 1330 m, 9-VIII-2010, Tiranës), between village Pëllumbas and cave L. Lőkös (BP 96560). KUKËS: District of Kukës, Shpella e Pëllumbasit, 41°15’4.10”N Kolsh, SH5 close Drin river, 42º04’18”N 20º20’32”E, 19°57’34.16”E, on calcareous soil in Carpinus deciduous Quercus, Juniperus oxycedrus, Buxus orientalis woodland, 445 m, 3-IV-2016, L. Lőkös sempervirens, serpentine soil, 510 m, 21-IV-2017, (BP 96655). VLORË: Sarandë, Pasqirave, Butrint A.R. Burgaz (MACB 111333). District of Kukës, N. P., SH81, Shengjergj monastery, 39º44’51”N Rruga Tirana, 42º04’46”N 20º24’45”E, Quercus 20º00’24”E, Quercus coccifera, Olea europea, cerris, deciduous Quercus, limestone, 330 m, 21-IV- Phlomis sp., limestone, 60 m, 17-IV-2017, A.R. 2017, A.R. Burgaz (MACB 111335). SHKODËR: Burgaz (MACB 111408). District of Sarandë Vau i Dejës, SH25 Vau i Dejës-Koman, (Rrethi i Sarandës), Mt maja e Dhema, between 42º00’55”N 19º39’37”E, Pinus pinaster, Carpinus village Ksamil and Pasqyrë, 39°47’28.03”N orientalis, serpentine soil, 120 m, 20-IV-2017, 20°0’31.43”E, on calcareous soil, 60 m, 17-X-2010, A.R. Burgaz (MACB 111422). Vau i Dejës, SH25 L. Lőkös (BP 96577). District of Sarandë (Rrrethi i Vau i Dejës-Koman, 42º03’01”N 19º39’45”E, Sarandës), on a hill between villages Kulluricë and Burgaz, A. R. et al. Bot. complut. 43, 2019: 15-40 39

Kodër, 39°48’43.85”N 20°06’34.02”E, in dry 20º10’05”E, Carpinus betulus, Pinus halepensis, grassland, on flysh soil, 55 m, 13-X-2012, L. Lőkös sandstone, 180 m, 18-IV-2017, A.R. Burgaz (BP 96593). District of Sarandë (Rrethi i Sarandës), (MACB 111216). District of Tepelenë, Dukaj, north of village Gjashtë, towards Delvinë, SH76 Tepelenë-Sinanaj, 40º18’45”N 19º57’46”E, 39°53’34.58”N 20°01’58.26”E, in open rocky Quercus coccifera, limestone, 280 m, 18-IV-2017, grassland, on limestone together with bryophytes, A.R. Burgaz (MACB 111219). District of Përmet, 45 m, 1-V-2009, Z. Barina, L. Lőkös & D. Pifkó Piskovë, left side Lumica river, K75 to Frashër, (BP 96546). Finiq, Muzinë, Muzinë, 39º54’48”N 40º16’44”N 20º17’07”E, Juniperus oxycedrus, 20º11’20”E, deciduous Quercus, Erica arborea, Paliurus spina-christi, Arbutus unedo, Pistacia limestone, 186 m, 17-IV-2017, A.R. Burgaz terebinthus, sandstone, 230 m, 18-IV-2017, A.R. (MACB 111202). Finiq, Bistricë, Muzinë, Burgaz (MACB 111217). SHKODËR: Vau i 39º55’08”N 20º08’48”E, Quercus coccifera, Dejës, SH25 Vau i Dejës-Koman, 42º00’55”N Carpinus betulus, Pinus halepensis, Cercis 19º39’37”E, Pinus pinaster, Carpinus orientalis, siliquastrum, limestone slopes, 125 m, 17-IV-2017, serpentine soil, 120 m, 20-IV-2017, A.R. Burgaz A.R. Burgaz (MACB 111194). Himarë, Ilias, (MACB 111304). Karmë, SH25 Vau i Dejës- Himarë, 40º08’29”N 19º40’11”E, Quercus coccifera, Koman, 42º04’02”N 19º40’59”E, Quercus Cupressus sempervirens, Cistaceae shrubs, macrolepis, Q. pubescens, Juniperus oxycedrus, limestone, 300 m, 16-IV-2017, A.R. Burgaz (MACB Erica arborea, slates, 130 m, 20-IV-2017, A.R. 111406). District of Vlorë (Rrethi i Vlorës), Mts. Burgaz (MACB 111330). District of Malësi e Mali i Çikës, in the valley below Mt. “Maja e Madhe, Poica, SH21 Koplik-Bogë, 42º15’53”N Pandelejmonit” (805.8 m), north of village 19º29’47”E, Juniperus oxycedrus, Cupressus Dhërmi, 40°09’32.2”N 19°38’49.4”E, in dry sempervirens, boulders, 320 m, 20-IV-2017, A.R. rocky grassland, on calcareous soil, 260 m, 29-IV- Burgaz (MACB 111324). District of Malësi e 2009, L. Lőkös (BP 96556). District of Vlorë Madhe, Dedaj, SH21 Koplik-Bogë, 42º18’04”N (Rrrethi i Vlorës), on the E ridge of Mt maja e 19º32’32”E, Carpinus betulus, Juniperus Thanasit (1352.1 m) above Llogora pass, oxycedrus, limestone rocks, 495 m, 20-IV-2017, 40°11’51.54”N 19°35’27.82”E, on calcareous soil A.R. Burgaz (MACB 111325). District of Malësi in mixed forest, 1075 m, 12-VIII-2010, L. Lőkös e Madhe, Bogë, SH21 Koplik-Bogë, 42º21’46”N (BP 96570). Atranorin, fumarprotocetraric, 19º37’56”E, Pinus nigra, Juniperus oxycedrus, rangiformic and physodalic acids chemotype: limestone, 865 m, 20-IV-2017 m, A.R. Burgaz DURRËS: District of Durrës, Fushë-Draçi, San (MACB 111328). TIRANË: Rrogozhinë, Grethi i Pietro beach track, 41º31’01”N 19º31’12”E, Pinus Mesem, 41º05’00”N 19º27’35”E, Pinus pinaster, pinea, sandy soil, 1 m, 19-IV-2017, A.R. Burgaz Juniperus oxycedrus, sandy soil, 1 m, 15-IV-2017, (MACB 111227). Krujë, Krujë, SH38 to Qafë- A.R. Burgaz (MACB 110951). VLORË: District Shtamë N. P., 41º31’03”N 19º49’27”E, Carpinus of Sarandë, Pasqirave, Butrint N. P., SH81, betulus, pasturelands, limestone, 720 m, 19-IV-2017, Shengjergj monastery, 39º44’51”N 20º00’24”E, A.R. Burgaz (MACB 111289). District of Durrës, Quercus coccifera, Olea europaea, Phlomis sp., Fushë-Draçi, road cape of Rodon, 41º32’50”N limestone, 60 m, 17-IV-2017, A.R. Burgaz 19º31’08”E, Carpinus betulus, Arbutus unedo, (MACB 110964). District of Sarandë, Pasqirave, Pistacia lentiscus, Cistus salviifolius, sandstone, 90 Butrint N. P., SH81, Shengjergj monastery, m, 19-IV-2017, A.R. Burgaz (MACB 111223). 39º48’42”N 20º00’43”E, Quercus coccifera, ELBASAN: District of Elbasan, Bradashesh, SH3 limestone slopes, 50 m, 17-IV-2017, A.R. Burgaz Bradashesh-Graze, 41º06’05”N 20º00’26”E, Olea (MACB 110963). Himarë, Ilias, Himarë, europaea, limestone karst, 390 m, 15-IV-2017, A.R. 40º08’29”N 19º40’11”E, Quercus coccifera, Burgaz (MACB 110952). District of Elbasan, Cupressus sempervirens, Cistaceae shrubs, Bradashesh, SH3 Bradashesh-Graze, 41º06’33”N limestone, 300 m, 16-IV-2017, A.R. Burgaz 19º59’49”E, Carpinus betulus Juniperus (MACB 110962). Himarë, Gjilek, Rruget te oxycedrus, limestone karst, 530 m, 15-IV-2017, bardha, 40º10’29”N 19º36’09”E, Quercus A.R. Burgaz (MACB 110953). GJIROKASTËR: coccifera, Pistacia lentiscus, limestone, 225 m, District of Tepelenë, Kodër, left Vjosa river bank, 16-IV-2017, A.R. Burgaz (MACB 110960). 40º17’20”N 20º04’25”E, Paliurus spina-christi, District of Vlorë, Dukat i Ri, SH3 Orikum-Dhërmi, Spartium junceum, Phlomis shrubs, limestone Llogora N. P., 40º12’00”N 19º35’26”E, Pinus slopes, 160 m, 18-IV-2017, A.R. Burgaz (MACB nigra, Juniperus oxycedrus, Buxus sempervirens, 111211). District of Tepelenë, Këlcyrë, Këlcyrë limestone, 1030 m, 16-IV-2017, A.R. Burgaz gorge, left side Vjosa river, 40º17’52”N (MACB 110959). Vlorë, Dukat i Ri, SH3 Orikum- 40 Burgaz, A. R. et al. Bot. complut. 43, 2019: 15-40

Dhërmi, Llogora N. P., 40º15’11”N 19º32’14”E, of Malësi e Madhe, Dedaj, SH21 Koplik-Bogë, Quercus coccifera, Pinus halepensis, limestone, 42º18’04”N 19º32’32”E, Carpinus betulus, 320 m, 16-IV-2017, A.R. Burgaz (MACB 110956). Juniperus oxycedrus, limestone rocks, 495 m, 20-IV-2017, A.R. Burgaz (MACB 111415). Cladonia rei Schaer. District of Malësi e Madhe, Bogë, SH21 Kopik- Homosekikaic acid chemotype: VLORË: Bogë, 42º21’46”N 19º37’56”E, Pinus nigra, Finiq, Muzinë, Muzinë, 39º54’48”N 20º11’20”E, Juniperus oxycedrus, limestone, 865 m, 20-IV- deciduous Quercus, Erica arborea, limestone, 186 2017, A.R. Burgaz (MACB 111416). Malësi e m, 17-IV-2017, A.R. Burgaz (MACB 111198). Madhe District (Rrethi i Malësisë së Madhe), on Homosekikaic and fumarprotocetraric acids Mount Bukovik above village Rrapshë-Starë, chemotype: DURRËS: District of Durrës, 42°23’6.12”N 19°29’25.56”E, on calcareous Fushë-Draçi, road cape of Rodon, 41º32’50”N soil in rocky grasslands, 715 m, 3-V-2015, L. 19º31’08”E, Carpinus betulus, Arbutus unedo, Lőkös (BP 96611). Malësi e Madhe District Pistacia lentiscus, Cistus salviifolius, sandstone, (Rrethi i Malësisë së Madhe); Rapshë Gorge 90 m, 19-IV-2017, A.R. Burgaz (MACB between villages and Lagjja e Re, 111222). KUKËS: District of Kukës, Kukës, 42°25’12.46”N 19°30’30.02”E, on calcareous Rruga Tirana, 42º04’46”N 20º24’45”E, Quercus soil in rocky grasslands, 225 m, 3-V-2015, L. cerris, deciduous Quercus, limestone, 330 m, 21- Lőkös (BP 96612). Shkodër County, at the foot IV-2017, A.R. Burgaz (MACB 111336). of Mt Maja e Sokolit between villages Tamarë Cladonia subulata (L.) F.H. Wigg. and Selcë, 42°29’0.61”N 19°35’0.98”E, on Fumarprotocetraric acid chemotype: calcareous soil in rocky grasslands, 385 m, 8-V- KUKËS: District of Kukës, Rruga Tirana, 2017, L. Lőkös (BP 96685). VLORË: District of 42º04’46”N 20º24’45”E, Quercus cerris, deci- Vlorë, Dukat i Ri, SH3 Orikum-Dhërmi, Llogora duous Quercus, limestone, 30 m, 21-IV-2017, N. P., 40º12’00”N 19º35’26”E, Pinus nigra, A.R. Burgaz (MACB 111439). Juniperus oxycedrus, Buxus sempervirens, limestone, 1030 m, 16-IV-2017, A.R. Burgaz Cladonia symphycarpa (Flörke) Fr. (MACB 111405). Atranorin, norstictic acid Atranorin and norstictic acid chemotype: and fatty acid chemotype: ELBASAN: KORÇË: District of Kolonjë (Rrethi i Elbasan county, in village Floq, 41°16’1.19”E Kolonjës), c. 1.9 km W of village Pepellash, 20°14’19.17”E, on soil by the roadside, 650 m, 40°28’56.18”N 20°40’21.27”E, on acid soil 17-V-2016, L. Lőkös (BP 96667). Atranorin, in moist road slope, 549 m, 18-X-2011, Z. norstictic and fumarprotocetraric acids Barina, L. Lőkös, D. Pifkó (BP 96584). District chemotype: ELBASAN: Prrenjas, Urakë, SH9 of Korçë, Dardhë, SH90 Boboshticë-Dardhë, Qafë Thanë, 41º04’24”N 20º36’18”E, Quercus 40º31’18”N 20º49’22”E, Fagus sylvatica, macrolepis, deciduous Quercus, Juniperus Pinus nigra, Juniperus communis, Betula oxycedrus, limestone, 1000 m, 22-IV-2017, pendula, Abies sp., limestone, 1390 m, 23-IV- A.R. Burgaz (MACB 111450). 2017, A.R. Burgaz (MACB 111461). District of Korçë (Rrethi i Korçës), on the ridge of Mts Cladonia sp. Mali i Moravës above village Dardhë, near peak Fumarprotocetraric acid chemotype. Buzeja e Boboshticës (1701 m), 40°32’23.97”N SHKODËR: Vau i Dejës, SH25 Vau i Dejës- 20°49’14.20”E, on acid soil in beech forest on Koman, 42º00’55”N 19º39’37”E, Pinus gravel conglomerate, 1550 m, 10-VIII-2010, L. pinaster, Carpinus orientalis, serpentine soil, Lőkös (BP 96564). ELBASAN: Elbasan county, 120 m, 20-IV-2017, A.R. Burgaz. VLORË: Finiq, in open boxwood scrubland above village Bistricë, Muzinë, 39º55’08”N 20º08’48”E, Kutërman, 41°13’40.99”N 20°14’41.04”E, on Quercus coccifera, Carpinus orientalis, Pinus acid soil in rocky serpentine area, 510 m, 16-V- halepensis, Cercis siliquastrum, limestone 2016, L. Lőkös (BP 96663). SHKODËR: District slopes, 125 m, 17-IV-2017, A.R. Burgaz.