Journal of Micropalaeontology, 16: 73-84. 0262-821X/97 $10.00 0 1997 British Micropalaeontological Society.
Revision of the Gault foraminifera from the Hollis and Neaverson Collection (1921)
c. HERRERO' & J. R. HAYNES~ ' Departamento de Paleontologia, Facultad de Ciencias Geol6gicas, Universidad Complutense de Madrid, Ciudad Universitaria, 28040 Madrid, Spain. Geology Department, Institute of Earth Studies, University of Wales, Aberystwyth, Dyfed SY23 3DB, UK.
ABSTRACT - In 1921, Hollis and Neaverson listed 135 species and varieties of foraminifera from the Gault Phosphatic Nodule Bed at Ford, Buckinghamshire, England. The material has been reexamined, the list is brought up to date and the species are figured. The Collection has yielded a diverse and well preserved foraminifera1 fauna of 90 species. Nodosariacea are the dominant group, though Ataxophragmiacea, Hedbergellacea and Mitiolacea are abundant. The recorded assemblages allow us to date the Phosphatic Nodule Bed at Ford as belonging to the uppermost Euhoplires luutus Zone to lower part of Morfoniceras inflaturn Zone within the Albian standard ammonite zonation. J. Micropalaeontol. 16(1): 73-84, May 1997.
INTRODUCTION Ford for phosphatic nodules is not known exactly. In their Hollis and Neaverson (1921) published a short paper on the paper, Hollis and Neaverson (1921) commented that the Gault foraminifera from the workings for phosphatic nodules at workings from which Hollis had collected his material in 1919 Ford, four miles southwest of Aylesbury, Buckinghamshire were abandoned and nothing remained but the spoil-heap which (England). They listed 135 species and varieties of foraminifera was by then (1921) completely grassed over. The available but neither descriptions nor illustrations were given by the information, about the fields where the workings are said to authors. After their paper no other workers have given have been (Fig. 1), was provided by the locals or elderly relatives additional infomiation about the foraminifera of the Gault at of the locals to the Buckinghamshire County Museum (pers. Ford. In fact, no references to the paper of Hollis and comm. Ms K. M. Hawkins, Keeper of Biology and Geology of Neaverson (1921) have been found in the Early Cretaceous that Museum). literature of Great Britain except for one by Crittenden (1988: p. The only description of the nodules beds is that of Keeping, 19, Unpublished Ph.D. Thesis, Plymouth Polytechnic). who saw earlier workings near Ford in 1876. According to The location of the site in which the Gault was worked at Jukes-Brown, who incorporated his observations into his
09
Fig. 1. Cretaceoiis outcrops in southern Britain and location of the site of phosphatic nodules ('coprolites') workings, modified from sheet SP 70 NE (by courtesy of Ms K. M. Hawkins, Buckinghamshire County Museum, Aylesbury).
73 Herrero & Haynes memoir (1900), the nodules were worked on this occasion Gault, foraminiferal ranges now being regarded as somewhat between the years 1875 and 1884. Keeping’s remarks include the more ‘critical’ than in the time of Hollis and Neaverson who following: were under the heavy influence of Chapman and the ‘English School’. The coprolite seam is 3 to 4 inches thick and is constant throughout the pit, though the bed is irregular in position. The MATERIAL irregularity is of two kinds, the commonest being slips of a few The state and characteristics of the collections are described inches (4 to lo), cutting off the seam with a clear face marked by below. slickensides; in other cases the bed is bent downwards to a similar extent, and this bending is due, I am inclined to think, to Buckinghamshire County Museum (Aylesbury) subsequent folding and not to irregularity of deposition. The set consists of slides made of cedar-wood, originally without The matrix of the seam is a stiff calcareous clay crowded with cover-glasses (covered in the course of this study). The accession phosphatised shells and lumps of ‘coprolite’which in my cursory register, number 38, dates from 29 January 1920, and to quote search, I could not prove to have suffered from erosion previous from it, describes the collection as consisting of ‘108 microscope to being embedded in the Gault, while some of them, such as slides, foraminifera from Gault, Ford, Bucks., collected by Hamites were in such a condition that they could not have Hollis 1919’. They are well set out and labelled but the sustained much knocking about on a shore. information on them is limited. The labels are arranged in horizontal orientation and comprise on the left side the generic Above the [nodule] bed comes a hard clay with iron-stained joint and/or specific identification and the locality and unit (Gault, planes, which is covered by a true clay, somewhat lighter in Ford, Bucks.) and on the right side the specific classification, or colour than ordinary Gault, and containing a second irregular nothing. On the back of the slides is written ‘E. Hollis 1919’ and coprolite zone in a series of lenticular patches. Its nodules are the slide number (see Tables 1 & 2). As it now stands, the smaller than in the regular seam below. collection includes 124 slides (Trays 38:20). Of them, 17 slides It should be noted that the emphasis on structural relationships contain specimens as yet unidentified (perhaps not originally in this account may be explained by the fact that Keeping was registered) and 107 slides with identified specimens. Of the 107, teaching geology at Aberystwyth at this time and attempting to four have specimens with only generic classification [Nodosaria work out the structure of Plynlimmon! (two slides), Pleurostornella (one slide) and Pulvinulina (one Jukes-Brown visited the workings in 1885 after they had been slide)], two have specimens left in open nomenclature (Nodosaria abandoned but was able to collect fossils. Collections were also sp.) and 101 have specimens with specific identification. Of all made by a local schoolmaster (Mr Hayter of Monks Risbor- the species and varieties that are identified on the slides, 106 are ough) which put together with those of Keeping presented a included in Hollis and Neaverson’s list published in 1921 and 28 ‘curious assemblage for the Lower Gault’, including ‘Ammo- are not included. In addition, there are 29 species on the list nites’ auritus, cristatus and lautus. which were not found in the slides (see Tables 1 & 2). Hollis and Neaverson in their remarks on the ammonites said, Natural History Museum (London) Judging by specimens of ammonites in the Bucks. County As in the Aylesbury collection the slides are made of cedar-wood Museum, Aylesbury, the deposit worked belonged to the zones and without cover-glasses. The labels are arranged in the same of Hoplites auritus and H. luutus. At the typical locality of way. The only difference is that there are no labels on the back Folkestone, these zones occupy 17 feet out of a total of 99 feet of the slides. The accession register indicates that the material for the whole of the Gault; but at Ford only three or four feet was deposited by E. Hollis in 1926 and the register numbers are seem to have been worked. Critical zonal study is, of course, from P23018 to P23232 (Tray 494). All the slides have specific impossible; . . . identification and of those, 56 are included in the published list We were pleasantly surprised to discover that there were three of Hollis and Neaverson, 10 are not in it and 79 species and different sets of slides of the Hollis and Neaverson Collection of varieties of the list were not found in the slides (see Tables 1 & the Gault at Ford. One set is located in the Buckinghamshire 2). County Museum (Aylesbury, England), the other one in the Natural History Museum (London, England) and the last one in Micropalaeontological Museum, University of Wales (Aberyst- the Micropalaeontological Museum of the University of Wales WYth) (Aberystwyth, Wales). The first two sets in the Museums were This last set consists of 47 slides, also made of cedar-wood, donated by E. Hollis while the third one was eventually originally without cover-glasses and now with them, attached in deposited at Aberystwyth when acquired by Prof. A. Wood the course of this work. The labels on the slides are arranged in from Neaverson who had been his research supervisor at vertical orientation. The label includes at the top the following Liverpool University. items: ‘Foraminifera, Gault, Ford, Bucks., Aug. 1919, Eng.’. The aim of this work is to bring up to date the list of The generic and specific identification is written down in ink or foraminifera of the Hollis and Neaverson paper, document the in pencil at the bottom of the label. The accession number is three scattered collections of slides and their contents and JH.389 and now all the slides have been numbered (see Tables 1 illustrate all the species, which are mostly represented by very & 2). Two of the slides contain specimens as yet unidentified, well preserved specimens. On this basis we will attempt to relate another two with only generic classification (Bulimina and the fauna to the modern, foraminiferal biostratigraphy of the Nodosaria) and 43 with specific identification. Of all the species
74
SL
@, @, Reuss Reuss slide. slide. the written (Reuss) book; register and/or slide on written is on on is is macrodisca Crisiellaria
@, @, @, @, is is on variety variety the book; register the sub-genus the with written name specific the book; register and/or slide in in ‘acutungulum’ ‘acutungulum’ (Dentalinu)
a, a, &, &,
0, 0, sub-genus sub-genus the without written is name specific the pencil; in written identification specific the slide; empty (Aberystwyth). (Aberystwyth). is is (Dentalinu) (Dentalinu)
Wales Wales University Museum, Micropalaeontological MM MM (London); Museum History Natural The NHM: (Aylesbury); Museum of of UWA): UWA):
of of County County Buckinghamshire BCM: museums. three the in material the location and Neaverson list published Original Table (1921) (1921) Hollis of I. I. & &
6R 6R Reuss Reuss Nodmorru Nodmorru orrhoihm orrhoihm
Reuss Reuss 61 61 - prismuricd Nrnloruriu
l 135 135
(Reuss) (Reuss)
.vpinuiifrru .vpinuiifrru
Puivinulinu Puivinulinu XIEZ-OE EZd 9E
66 66 Reuss Reuss ID ID fcnuiciirru fcnuiciirru Nodonrurio
- - - )
pol I34
fR6mer) fR6mer) currrcollu currrcollu Pulvinulinrr (D (D Reuss Reuss 65 65 Nodosuriu Nodosuriu ohscum ohscum ) )
I 133 ~iegunr ~iegunr
Puivinuiinu Puivinuiinu 90 (dOrbtgny) Q9Z-PZIEZd a SE
64 64
(Berthelin) (Berthelin) (D.)fonrunnu.rr (D.)fonrunnu.rr
- 'OP Nodowriu
I32 I32
(Reuss) (Reuss) urnmonoides urnmonoides Anomulinu Anomulinu ,EIEZ-LEIEZd IS
63 63 Reuss Reuss
(D (D Nodouriu Nodouriu
1 v )puuperculu 13 13 I I I I
€0I (dOrbigny) Discorbinu rugoru rugoru Discorbinu Z EZ-6 EZd LE 62 62 Reuss Reuss wuprrum wuprrum Nodmunu
101 130
dOrbigny dOrbigny
crmzceu crmzceu Gbbigrrino Gbbigrrino IOlEZd 3OP-6L
61 61 Brady Brady D.) D.) inrurrrllulurrr inrurrrllulurrr Nodorurru
- 1
Wright Wright 129 129 uruleuru uruleuru
I Rumulinu EP
MI MI Chapman Chapman Nodmuriu Nodmuriu
- bomhu.ru
128 128
Brady Brady OP globuliferu globuliferu
86 Rumulinu a 59 59 dOrbigny dOrbigny Nodr,.sunu Nodr,.sunu
- - hivyidu
I27 I27 (Berthelin) (Berthelin) Sugrinu Sugrinu
86 alcumru 58 58 Reuss Reuss Nodoruriu Nodoruriu
- ,.olomo~~hu
Chapman Chapman Sugrinu usperulu usperulu Sugrinu
za 126 ZP 57 57
Chapman Chapman
rurisrriuru rurisrriuru 'D. Nidmuriu
8 8C )
125 125 EOlCZd
Reuss Reuss var horridu horridu 28 P~,i,[ymo~hinu~rrfori.,
D D 56 56 (Neugeboren) (Neugeboren) mucronuru mucronuru Nndowrru
- - - )
124 124
(Romer) (Romer)
Pol~morphinufusifnfomtis Pol~morphinufusifnfomtis 55 55
d'Orbigny d'Orbigny
(D.) (D.) Nodmurio Nodmurio
18 0 cr)mmunis
I23 I23
(I. (I. 1 1 iucrru iucrru var. var. LE & & C C ucuplucrnru ucuplucrnru
'6L Polymorphrnu LZlEZd a 54 54 (Neugeboren) (Neugeboren) (D. (D. rocrneri rocrneri Nohsuriu
- - - J
122 122
Berthelin Berthelin drudemuru drudemuru
Crisrrlluriu Crisrrlluriu
53 53 Reuss Reuss (D.) (D.) Nndmurio Nndmurio Iegumm Iegumm
I21 I21 9P
Berthelin Berthelin
68 srrrnulis Crrsrrlloriu WlEZd
52 52 Reuss Reuss riphioidus riphioidus (D. Nodosariu Nodosariu
- J
120 120 6P Berthelin Berthelin
guulrinu guulrinu Crisrelluriu Crisrelluriu :ZlEZ-ZZIEZd
51 51
Reuss Reuss huniulifrru huniulifrru (D. Nodo,wrio
)I J
96-56 119 IEZ-601EZd Reuss Reuss var. roruluru roruluru Cricrelluriu mucrrxliscus mucrrxliscus zs
50 50 (Reurs) (Reurs) c!lindroider c!lindroider (D Niidorurro
- - J
(Lamarck) (Lamarck) I8 I8
roruluru roruluru Cristrllarkr
1 1 LP
(dorbigny) (dorbigny) 49 consohrinu consohrinu (D.)
58 - - Nodosiiriei I I I7 I7
Bornemann Bornemann
convrrgenr convrrgenr Crisrrliurhr (D (D
48 48 (dOrbigny) (dOrbigny) )puuprruru )puuprruru
I 0s Nodowmu gibbu gibbu dOrbigny dOrbigny 116 116
98 Crisrelluriu 9E EZd (D.) (D.) (dOrbigny) (dOrbigny) 47 47 lornt'iunu lornt'iunu Nod~i.~urra
16 I5
I I K) I (Costa) (Costa) Crirtrllorio Crirtrllorio
ioburu ioburu EZ-SO1EZd zs
(dOrblgnY) (dOrblgnY) 46 46 (D.) (D.) NcJdo.roriii
- - Kmcr1ir
14 14 1 1 Berthelin Berthelin 8P circumcidunru circumcidunru Crisrelloriu
45 45
Chapman Chapman var.
soluro soluro pulchrllu pulchrllu (D.)
- - 8P N'odo.ruria I3 I3 1 1
Reuss Reuss
rurgidulu rurgidulu Crisrelluriu
(Reuss) (Reuss) 44 44 (D (D Nodosuri[, snluru snluru
- J
112 112
Chapman Chapman brodyona brodyona
L8 Crixrellurio IP
13 13
a Reuss jfurcimen jfurcimen ID. Nodoerrii~
- (Soldmi)
I I I I 1 1 t
Reuss Reuss
Crisrelluriu Crisrelluriu
complunuru complunuru IOICZ-LO1 Zd IP
42 42 a Reuss (L (L joneri joneri var var rudiculu Nodowrin
- )
110 110
Berthelin Berthelin 8P vrsriru vrsriru L6 Crirrrllurru
41 41 Reuss Reuss (G. (G. Nodorono cylidruceu cylidruceu
- - j
109 109
(Defrance) (Defrance) irulica irulica
L6 Cri.srelluriu
40 40 (Reuss) (Reuss) (C.) (C.) muruhilir muruhilir
16 - PP Nodo.rrrriii 108 108
Berthelin Berthelin bononienJis bononienJis Crisrrlluriu
39 39 Rorner Rorner (G.) humilis humilis
- ;I 1 I I tP Nodosurrgi 107 107
Berthelin Berthelin
Crirrrllorio Crirrrllorio .sciralo .sciralo EZ-Z EZd
38 38 Reuss Reuss h;.spidu h;.spidu
- Ligenu
106 106 $1 Berthelin Berthelin IEZ-91 rrunculuru rrunculuru
Cristelloriu Cristelloriu lEZd 5-9
(Seguenza) (Seguenza) 31 31
gwcillimu gwcillimu
- Ligrnu
I05 I05 dOrbigny dOrbigny rriungulurir rriungulurir
Crrs~elluriu Crrs~elluriu PIEZ-OPIEZd ZE
36 36 Reuss Reuss
irpiiuluru irpiiuluru var. var.
rmuciuiu rmuciuiu
- - Lrgmir
104 104 Reuss Reuss rulrifrru rulrifrru Crisrrlluriu Crisrrlluriu EE
Reuss Reuss 35 Lrgrnir Lrgrnir
- crniculuru
dOrbigny dOrbigny 103 103 nuwicuiu nuwicuiu
88 Crisrrlluriu PE
34 34 (Montague) (Montague) glohosu glohosu
- krgcnu
7PI 102 102 (Ficchel (Ficchel Moll) Moll) EZ-ZPI IE & & crepidulo crepidulo
Crisrrlluriu Crisrrlluriu EZd
33 33
Schwager Schwager
- - ulrernuns Plruro.rrrrmrllu
101 101 Reuss Reuss humilis humilis L6 Crisrelluriu
32 32 ohrum ohrum Berthelin Berthelin
E6 Plrur~i.srornullo
Reus? Reus? purullelu purullelu Crirrrlluriu
I(w) I(w) ,PI EZ-8PIEZd 62
I I 3 3
d'Orbigny d'Orbigny
uJj7nr.s uJj7nr.s Bulimmii
Reuss Reuss 99 99 exi1i.r exi1i.r
P6 Crrsrdluriu .PI EZ-SPI tZd 62
30 30 d'Orbigny d'Orbigny
- hruvis Buliminu
Chapman Chapman 98 98
26 priceunu Vuginulinu LZ
29 29
il'Orbigny il'Orbigny ohrusu ohrusu
- Bulirninb
97 97 Berthelin Berthelin biochri biochri
SL Vuginulinu 8Z
28 28
d'Orbigny d'Orbigny murchisonrunu murchisonrunu
9L - Bulimino Berthelin Berthelin 96 96 85 Vuginulina Vuginulina Ruulrinu Ruulrinu
Reuna Reuna 27 27 orhign>i orhign>i
- Buliminu
LL 95 Reuss Reuss Vuginulinu Vuginulinu rruncuru rruncuru LZ
ZL (Reuss) (Reuss) 94 94 ,srrrgilluro ,srrrgilluro Vuginulinu Vuginulinu &lEZ-&SltZd 82-95
8L - - Reuss Reuss 93 93 rrcru rrcru Vuginulina
24 24
(Reuss) (Reuss)
PL - complunuru Sprroplecru
Reuss Reuss 92 92 jonrri jonrri Murginulinu Murginulinu PZ
23 23
Jones) Jones) (Parker & & unnucrrns unnucrrns
IL Spiroplucru I I Reuss Reuss 9 9 ,rtrrurocosruru ,rtrrurocosruru
Murginulinu Murginulinu I81EZ-LLIEZd zz-oz
Reuss Reuss 22
pvrumidoru pvrumidoru - - Triruxro
89 90 Reuss Reuss
urquivocu urquivocu Morgtnulinu
2 2 I I
Reuss Reuss rricorinuru rricorinuru Triruritr
lL 89 Berthelin Berthelin I91 l debiir debiir
MurRinulinu MurRinulinu EZ-8S EZd sz
(Munster) (Munster) 20 20
L9 - - rriycwrru Vurnrurlrnu Reuss Reuss 88 88
lineurrs lineurrs Murginulino
(Reuss) (Reuss) I9 I9 Turruluriu Turruluriu
OL complunuru
Reuss Reuss 87 87 inurquu1i.r inurquu1i.r Murginulinu Murginulinu :9l EZ-191EZd sz
Reuss Reuss Tuxruluw Tuxruluw prurlongu prurlongu - I8
dOrbigny dOrbigny
gluhru gluhru Murglnulinu
- 86 ,LIEZ-ELIEM OZ
I7 I7 dOrbtgny dOrbtgny ugg1urinun.r ugg1urinun.r - Tuxruluriu
85 85 Reuss Reuss
Rhubdogonium Rhubdogonium
excuvucum excuvucum zz
dOrbigny dOrbigny mnicu mnicu Turrulurio
99 - 16
(dOrbigny) (dOrbigny) 84 84 rricurinurum rricurinurum
Rhubdogonium Rhubdogonium PZ
I5 I5
dOrblgny dOrblgny
Tanrloriu Tanrloriu
0 - iurri.!
99 Reuss
83 83 Frondrtulurru Frondrtulurru
rordui rordui EZ
dOrbigny dOrbigny I4 I4 frochu.\ frochu.\
E9 Tuxruluriu Chapman Chapman prmvuru prmvuru Frondiculuriu Frondiculuriu
- 82 rYlfZ-P9IEZd EZ
Berthelin Berthelin TuxruL,riu minuru minuru TuxruL,riu
- I3
XI XI Reuss Reuss microdiircu microdiircu
- Frondrculurru EZ
Chapman Chapman
millurrrunus millurrrunus Ammodireus Ammodireus
- I2
Reuss Reuss guesrphulicu guesrphulicu Frondiruluriu
80 80 zz
(dOrbigny) (dOrbigny) urms urms in( Ammodr.rcu.v Ammodr.rcu.v
c9 - - I I Chapman Chapman Frondicuiuriu plunrfolru plunrfolru Frondicuiuriu 79 79
Brady Brady u1bIcern.s u1bIcern.s Thurunminu
19 10
Reuss Reuss purkrri purkrri Frondiculuriu Frondiculuriu
78 78 L60EZd 9 '5
Chapman Chapman .rhrrborni .rhrrborni rich? Hciplo~
t9-59 9
Reuss Reuss Frondrculunu Frondrculunu ungrrr ungrrr
77 77 :61EZ-68lEZd 91
(P. (P.
& & J J ~lobigrrinifimnu ~lobigrrinifimnu Hrrplophrugmium
) )
19 R Reuss Reuss cf cf .srrigillutu .srrigillutu Frondiculuriu
- 76 WEZ-PO5 EZd SI
Hanken Hanken HupIoj*t:rogmrum ucuirdor.wium ucuirdor.wium HupIoj*t:rogmrum
- 7 Perner Perner
frir,rchi frir,rchi Frondicuiuriu
75 75 0 El
Montfon Montfon .\corpiuru.\ .\corpiuru.\
Rriiphu~ Rriiphu~
65 6
Reuss Reuss gUuhuI gUuhuI Frondrculurru
74 74 'OZEZ-OOZtZd ZI
(dOrbigny) (dOrbigny) Mrlrolww/urui Mrlrolww/urui
iucir iucir I9 5 Chapman Chapman denrrculricurinuru denrrculricurinuru Frondicuiurru
73 73 OZEZ-LOZEZd II
(dOrbigny (dOrbigny
rr~co~~in~ircr rr~co~~in~ircr Miliolincr ) )
09 4 Berthelin Berthelin lopi lopi Frondiculuriu
72 72 'ZZEZ-PZZEZd 0 I
(Karrerl (Karrerl vrnr~.~rer vrnr~.~rer
09 Mrliolmo 3 3 Reuss Reuss Lingulrna zrmiornuru zrmiornuru Lingulrna
71 71 'ZOEZ-ZZOEZd 6
dOrbigny dOrbigny nirrdu nirrdu Slurdoculinrr Slurdoculinrr
Lb 2 Reuss Reuss
nodmunu nodmunu Lingulm
70 70 'COEZ-5ZOEZd 01 8
Chapman Chapman n,idulrrsu n,idulrrsu 85 Niibr' ul.iriu ul.iriu
I I
Reusb Reusb rerrugonu rerrugonu Nr,do.curiu Nr,do.curiu
69 69 SOEZ-EPOtZd L -OP 'EOEZ-SEOEZd z - ~~~~ aqmnu iaqumu aqmnu- aPllS iaisrSaa aPllS
w38 WHN
The The foraminifera foraminifera Giiult Giiult Explanation of Plate I. Hollis and Neaverson Collection of the Gaul1 of Ford (Buchnghamshire, England). Unless specified. all the specimens are from the Micropalaeontological Museum, University of Wales (Aberystwyth). Scale bars 100pm. Fig. 1. Thurmmiw sp.. AYBCM.1920.38.1.6, Buckinghamshire County Museum (Aylesbury). Fig. 2. Ammodisms crefaceus (Reuss, 1845), JH.389.7.1. Fig. 3. Haplophrugmoides nonioninoides (Reuss, 1863), JH.389.5.3. Fig. 4. Haplosfiche?sherborni Chapman, 1892, JH.389.6.14. Fig. 5. Spiropleclinafa onnectem (Parker & Jones, 1863), JH.389.14.12. Fig. 6. Dorothiu turris (d’ Orbigny, 1840), JH.389.9.8. Pip. 7. Texfularia minuta Berthelin, 1880, JH.389.8.6. Fig. 8. Texruloria chapmani Lalicker, 1935, JH.389.8.5. Fig. 9. Bigerina? asperula (Chapman, 1896), JH.389.38.2. Fig. 10. Nodobomlaria nodulosa (Chapman, l891), JH.389.1.16. Fig. 11. Gaudryina dirpnmo Chapman, 1892, JH.389.14.10. Fig. 12. Gaudryina gradnta Berthelin, 1880, JH.389.10.11. Fig. 13. Tritaxiapyrmidnta Reuss, 1863, JH.389.13.13. Fig. 14. Quinqueloculina antiqua (Franke, 1928), JH.389.3.17. Fig. 15. Nubeculinella? sp., AYBCM.1920.38.3.1, Buckinghamshire County Museum (Aylesbury). Fig. 16. Eggerellim mriae Ten Dam, 1950, JH.389.9.9. Fig. 17. Arenobulimiw chapmoni Cushman, 1936, JH.389.15.7. Fig. 18. Reophax sp., JH.389.4.4. Fig. 19. Spiroloculina guultina (Ten Dam, 1950), JH.389.2.19. Fig. 20. Spiroloculiw cretaceo Reuss, 1854, JH.389.2.18. Fig. 21. Nubecularin? depressa Chapman, 1891, AYBCM.1920.38.4.1, Buckinghamshire County Museum (Aylesbury). Fig. 22. Cornuspira crefacea (Reuss, 1845), JH.389.7.15. Fig. U. Trochammiw concava Chapman, 1892, AYBCM.1920.38.18.1, Buckinghamshire, County Museum (Aylesbury), (a) dorsal view, (h) ventral new, (c) lateral view.
76 The Gaul1 foraminifera
- In Table 3, the updated names used in this work are in the left BCM column and are arranged following the foraminifera1 classifica- Slide tion of Haynes (1981). The right column includes the - I Anomulinu rudis (Reuss) 102 equivalencies of the list of Hollis and Neaverson (1921). The 2 Cluvulina unguluris dOrbigny 19 species with an asterisk are those that are not included in the 3 Crisrelluriu lurlfrons Brady 97 published list but were found in the slides. The ‘p.p.’ for some 4 Crisrelluriu 1ineurt.r Reuss 84 5 Cri.rre1lurru mumilligeru Karrer 9a species is explained at the end of the table. 6 Crisrelluriu rripleuru Reuss 97 The agglutinated foraminifera are represented by two species 7 Flubellim~didymu (Beflhelin) 65 8 Frondiculuriu didjmu Berthelin - of the Order Astrorhizida and 14 of the Order Lituolida. Of 9 Globigrriiau ueyuilurerulis Brady 99 them, two species, Thurammina sp. and Reophax sp., have been 10 Huplopbuqmium nonioninoides Reuss 14 1 I Murginulinufolken~loniensis Chapman 69 left in open nomenclature and another, Bigerina? asperula 12 Nodorurio inflaru Reuss 40 (Chapman), with doubtful generic identification due to the bad I3 Nodosurio (G.J lueuiguru dOrbigny 38 14 Nubeculuriu depressu Chapman 4 preservation of the initial portion of the test in all the examined 15 Nub~ruluriuribiu (Jones &Parker) 3 specimens. In addition, Sagrina calcarata (Berthelin) although 16 Polymorpliinu ungusm Egger 79 on the list could not be identified because it is represented in the 17 Polymorphinu criminunis dOrbigny 80 I8 Polymorpbinu rompressu dOrbigny 79 slide by two indeterminate fragments of agglutinated foramini- I9 Polymorphinu gibbu dOrbigny 81 fera only. The species Ammodiscus millettianus Chapman was 20 Polymorphinu lucreu (Walker) 81 21 Polvmorphinu sosoriu Reuss 81 misindentified by Hollis and Neaverson (1921) as in the slide 22 Polymorphi~sosoI~iu var. cuspidoru Brady 80 there are only four fragments of organic carbonates. 23 Pulvinulin~rrericuluru Reuss 105 24 Spiroloculinu usperulu Karrer - The porcelaneous foraminifera, Order Miliolida, are repre- 25 Textuluriu grumen d‘Orbigny 24 sented by a significant number of specimens belonging to seven 26 Thummminu pupilloru Brady 17 species. One of them, Nubecularia? depressa Chapman has been 27 Truchumminu COIIC~IVUChapman 18 28 Vuginulr~urguru Neuss 77 left with doubtful generic identification as it is not possible to see 29 Vuginulinu rerru var. renuisrriuru Chapman 77 the initial portion of the test in the specimen of the collection. 30 VURinU/I?Iurruncuru var. robusru B. & C. 73 - The specimen that Hollis and Neaverson named Nubeculariu Table 2. List of species found in the slides and not included in the tibia (Parker & Jones) has been left in open nomenclature as published list of Hollis and Neaverson (1921). a,the specific only the last two free chambers are preserved. identification is written in pencil. The Order Nodosariida is the best, well-represented group. Sixty species have been identified (50 of the Superfamily and varieties, 103 (75 written in ink and 28 in pencil) are Nodosariacea, three of the Superfamily Polymorphinacea and included in the list of Hollis and Neaverson, three are not in it seven of the genus Ramulina). The specimens of Lingulina and 32 specific names on the list were not found in the slides (see nodosaria Reuss, Nodosaria (Dentalina) mucronata (Neugebo- Tables 1 & 2). ren) and Nodosaria prismatica Reuss of the list are completely Combining the three sets of slides, 1311 specimens of broken and badly preserved so they have been left as foraminifera of the Gault at Ford have been examined (423 indeterminate fragments of Lingulina, Dentalina and Nodosaria, from the Buckinghamshire County Museum, 214 from the respectively. Six other species have been left in open nomen- Natural History Museum and 674 from the Micropalaeontolo- clature (Dentalina sp., Frondicularia sp., Marginulina sp., gical Museum of the University of Wales). Specimens of two Nodosaria sp., Planularia sp. and Ramulina sp.). species and two varieties of the original published list of Hollis The other groups represented are the Order Buliminida, and Neaverson [Haplophragmium globigeriniforme (Parker & Superfamily Cassidulinacea, the Order Robertinida, Super- Jones), Gaudryina oxycona Reuss, Polymorphina fusiformis family Ceratobuliminacea, the Order Rotaliida, Superfamily Roemer var horrida Reuss and Polymorphina lactea (Walker Discorbacea, and the Order Globigerinida, Superfamily Hed- & Jacobs) var. acuplacenta (J. & C.)] are not in either of the three bergellacea. Although only a few species have been identified in sets (see Table l), s80unfortunately they could not be included in all these groups there are a significant number of specimens, this work. especially in the case of the Hedbergellacea [Hedbergella infracretacea (Glaessner)] and Discorbacea [Gavelinella inter- RESULTS media (Berthelin)]. In bringing up-to-date the list of species names used by Hollis The material under the names Clavulina angularis d’orbigny, and Neaverson for the Foraminifera. of the Gault (see Table l), Polymorphina communis d’orbigny, Polymorphina sosoria var. it has been necessary to change most of the generic names and a cuspidata Brady and the specimens of Reophax scorpiurus number of specific names as well. In part this simply reflects Montfort of the Buckinghamshire County Museum, are here progress in taxonomic discrimination but also the tendency of considered as incertae sedis. the members of the ‘English School’, such as Chapman, to apply A brief and quick analysis of the composition of the Recent names to fossil species. However, in his work on the assemblages in the Gault at Ford shows that in number of Gault, Chapman (I 891-1898) recognized many new species and specimens, the dominant species is Hedbergella infracretacea accepted many set up by Continental authors. Interestingly, this (Glaessner). Gavelinella intermedia (Berthelin) is also abundant. preceded the disastrous ‘discovery’ of the supposed Cambrian Among the Nodosariida, Vaginulina recta Reuss and Lenticulina foraminifera of the Malverns in 19010 which seemed to confirm gaultina (Berthelin) are the commonest species. Arenobulimina all the prejudices of the ‘English School’. chapmani Cushman, Spiroplectinata annectens (Parker & Jones)
77 Herrero & Haynes
Order Astrorhizida ~ehili\(Berthelin. 1880) Murginulinu dehih Berthelin Suwrfamily Ammodiscacea 'I 2. fig 3 Ammodi$c 11, c wru( YU\ (Reu\\. 1845) p p Amrnodi.scus incerru,\ (d'Orbignyj. 1 lenrulinu disrincru Reugs. I860 Murginulinu 1ineuri.s Reus\ PI I. hg z 'I 2. fig 4 Nodownu 1G ) cylindruceu Reuss Thuriiniminu \p 11p Thurumminu ulhicuns Brddy. 2 Nodo.,unu (D) cjlindrorde.\ Reus\ PI I. fig I Thurumminu pupilluru Brady* Nodo\uriu (D I humuliferu Reu\\ Nodowriu (D ) lorneiunu (d'Orbigny) Nodo.suriu iD I rnluru (Reucs) t Nodo.\uriu (D I xiphwide\ Reuh\ I Biyerinu " uipericlu (Chapman. 1896) .%Igrinu u\peru/u Chapman PI I. fig 9 pp N fDj communi.\ dorbigny, 9 Huplophruyni,ii,lr\ niinioninoidev (Reur\. Di.scorhinu rugosu (d'Orbignyi p p Nodo.\uriu (D) furcirnen Reusa, 10 1863) PI I. fig 3 Huplophrugmiurn ucuridorsurum Hantk Denrulinu gruci1i.s (d'Orbigny. 1840) Nodmariu (D i grucili.\ (d'Orbigny) Rriiphu r \p p p Reophux .scorpiuru.s Montfort, 3 PI 2, fig I1 p p N (D) consohrinu (dOrbigny), I I PI I.fig 18 p p Nodosuriu (D) consohrinu (Orb.), 4 Denfullnu lrfiurnen (Reus\. 1845) Murginulinu folke\ronien.riv Chapman* Texruluriu c hapmuni Lalicker. 1935 11 p Texruluriu conicu dOrbigny, 5 PI 2, fig. 8. Niidosuriu (D.) le,qumen Reuu PI I. fig 8 p p Texruluriu pruelongu Reuss, 6 Denrulinu pseudonunu Ten Dam. 1950 Nodmunu fD1 roemeri (Neugcboreni Trrrulurio minuru Berthelin. 1880 Texruluriu rninuru Berthelin PI. 2, fig. 10 PI 1. fig 7 Denrulinu pulchellu (Chapman. 1893) Nodo.suriu (D ) .\oluru (Reus\) var Huplow: he" iherhirni Chapman. I892 Huplmiche sherhnmi Chapman PI 2, fig 7. pulchellu Chapman PI I, flg 3 Denrulinu sp aff D. inrerce1luluri.s Nodosuriu (D ) inrercelluluri\ Brady (Brady. 1881) PI 2,fig 13 Nodmuriu (D) rurisrriufu Chapman Dentulinu \p. Nodo.\uriu (D J puuperuru dOrbigny PI 2,fig 21 11 p Nodosuriu (Dlfurcirnen Reuss. 10 Frondiculuriu denriculncurinutu F'rondiculuriu denric ulo( urrnufu Chapman, 1894 PI. 2, fig 14. Chapman Frondiculuriu fiuulrinu Reuss, I860 Frondrculuriu guulrinu Reus\ PI 2, fig 9 Frondiculuriu pinnuejorrnis Frondic uluriu frit.\<.hiReus\ Chapman, 1894 PI. 2, F-. 6 Frnndiculuriu pliinifihi Chapman, 1894 ~Frondi Frondiculuriu SD PI 2, fig I? Lugenu upiculuru (Reuss. I85 I) Lugenu ~pic~l~r~Reuss PI 2. fig 16 Lugenu upiculuru var emuciuru Reu\\ pp Lugenu hi3pidu Reuat. 12 Lugenu aff hispidu Reu\s. I863 Lugenu grucillimu (Seguenza) PI 2. fig. 15 [I.[). Lugenu hispidu Reuas, 13 Lenric ulinu ciri.umciduneu (Berthelin, Crisrellurh (iniimcidunru Berthelin 1880) PI 2,fig 17. Lenriculmu diudrrnuru (Berthelm, 1880) Cri.srelluriu diudrrnuru Berthelin PI 2,fig. IS. 1,enriculinu guulfinu (Berthelin. 1880) Cri.\relluriu guulrinu Berthelin PI ?.fig 20. Cn.\relluriu Rihhu dOrbigny Nodohuculuriu nodulo su Nuhrculuriu nodulo.\u Chapman Cri\rrlluriu mumrlligeru Karrer* (Chapman. 1891) PI I.fig 10. Crr\relluriu rofuluru (Lamdrck) C. roruloru (L ) var mucroditcut Reus\ Nuhei uluriu " depre.\\u Chapman. 1891 Nuheculuriu depressu Chapman* Cri srelluriu .\ternuli.\ Berthelin Cri.\relluriu rurgidulu Reus\ Lenriculinu rurgidulu (Reuss. 1863) Cri.srelluriri conwr~enrBornemann PI. 2. fir 19 Cri.srelluriu lohuru (Co\tai SiInPrhmilv Milinlaced lCrirtrlluriu sukrfrru Reuss Lindinu loni Berthelin. 1880 I Frondiculuriu lomi Berthelin PI 2. fig 22. Lingulinu wmiornum Reus\. 1863 Linjiulinu .\cmiornuru Reusa PI 2, fig 21 Murginulinu inuequuli.\ Reuss, 1860 Cri.\rellurru humills Reu\s PI 2. fig. 24 Murginulinu gluhru dOrbigny Murginulinu inuequuli.! Reuss Murginulinu jone.si (Reuss. 1863) Murginulinu ueyuivo~uReu\\ PI. 3. fig 1 Murginulinu jime~iReuss Crisrelluriu lorifrim Brady A,roc,,lu\ rriplruru Reu\\. I860 * Murginulinu lineuris (Reuss. 1863) Crisrelluriu Iineuri! Reu\s* Cri.srelluriu rripleuru Reuss* PI. 2. fig. I PI 3, fig. 2. Cirhurinellu drrlvmu (Berthelin. 1880) Fluhrllinu didymu (Berhelin)* Murginulinu striutoco.sruru (Reus. 1863) Murginulinu \friufocosruru Reur\ PI 2. fig 2 Frondiculariu didvmu Berthelin' Friindiculuriu perowfu Chapman Table 3. Left column: updated names used in this work. Right column: list of Hollis and Neaverson. Footnote explanations at end of table. 78 The Gault foraminifera Table 3 continued -CrisrelluriuCrrstelluriu erilisexilis Reuss Reuss wlap.- Rumulinu aff gloho-ruhulo.\~i p p Rumulinu ,y/~~hu/iferuBrad). 19, 21 Cushmdn, 1938 PI 4, fig. 7 p p Rumulinu uculeuru Wrr&ht Niidiisuricr cf. ujjinr.! Reuss. 184q p p Nodiisuriu orth~~pleuruReuss. 14 p p Nodosuriu orlhopleuru Reuss. 14 Rumulinu aff 1uevi.r Jones. I875 p.p Rumulinu gliihulitrru Bradv. 19. 21 PI 4. fig I0 pp Rumulrnu urulrutu Wnght Nodmrru cf humhuru Chapman, 1893 Nodosurru humbu,\u Chapman Rumulrnu muriculinu luebhch & Tdppdn pp Rumulinu glohuliferu Brady 1Y 23 PI 3. fi .8. 1949 PI 4,figII pp Rumulinu u 79 Herrero & Haynes However, looking at the stratigraphical distribution of foraminifera as seen in several key sections of southern and eastern England, Northern Ireland, northwestern Scotland and the North Sea Basin (Carter & Hart, 1977; Hart, 1973, 1990; LJ Hart et al., 1981, 1989, 1990) it is apparent that some of the P species recorded in the Gault at Ford, such as Haplostiche? ~~ P 5 sherborni Chapman, Eggerellina mariae Ten Dam, Tritaxiu E varicosum R pyramidata Reuss, Textularia chapmani Lalicker, Quinquelocu- lina antiqua (Franke), Frondicularia pinnaeformis Chapman and Vuginulina mediocarinata Ten Dam, are indicative of the Upper G A M. inflaturn Albian. I: orbignyi The presence of Frondicularia pinnaeformis Chapman, an L important zonal indicator in the Upper Albian (Carter & Hart, A T 4a N 1977), in association with the typical Upper Albian Arenobuli- - mina chapmani Cushman, Nodobacularia nodulosa (Chapman) M. L cristutum and poorly developed specimens of Epistomina spinulifera (Reuss), characteristic of the lower Gault Clay (Hart & Carter, A c 1975; Carter & Hart, 1977; Price, 1977), points to the possibility L t! E. luutus duviesi B t 4 of the Phosphatic Nodule Bed being in the Epistomina I L spinuliYeralFrondicularia pinnaeforrnis Concurrent Range Zone A 7 (Zone 4a) of the benthonic zonal scheme of Carter & Hart N nitidus (1977). However, the presence of Eggerellina mariae Ten Dam -_ --- could indicate at least the Frondicularia pinnaeformis Assem- blage Zone (Zone 5) of the same authors. Fig. 2. Owen’s (1971, 1973,1975) Standard ammonite zonation (columns So, according to the benthonic zonal scheme proposed by 1 and 2) and Hart’s (1973) foraminifera1 zonation (column 3) at the Middle-Upper Albian boundary. Note that Hart includes the crisfuturn Hart (1973) and Carter & Hart (1977) it is most probable that Subzone in the luutus rather than the injratum Zone. the foraminifera of Ford were recovered from the uppermost lower Gault Clay-lower part of the upper Gault Clay, Zones 4a- and Tritaxia; pyramidata Reuss are the best represented among 5 (from the uppermost Middle Albian to the lower part of the the agglutinated foraminifera and Quinqueloculinu antiqua Upper Albian), equivalent to the uppermost Euhoplites luutus (Franke) and Spiroloculina cretacea Reuss are the commonest Zone (cristatum Subzone) to lower part of Mortoniceras inflaturn species of the porcelaneous group. Zone (orbignyi-varicosum-lowermost auritus? Subzones) of the Albian ammonite zonation (see Fig. 2). DISCUSSION: AGE OF THE ASSEMBLAGES It must be noted that Hart’s treatment of the ammonite zones Many of the species recorded in the Phosphatic Nodule Bed of differs slightly from that of Owen (1971, 1973, 1975) in that the the Gault at Ford are also present in the Chapman Collection of cristatum Subzone is placed in the lautus Zone, i.e. the Middle the Gault of Folkestone (England) which has been examined in Albian rather than the Upper Albian. Hart (1973, p. 272) the Natural History Museum during the course of the present comments, in relation to the Copt Point section at Folkstone, work (see Chapman, 1891-1898). Hollis and Neaverson (1921) ‘The cristatum Subzone contains a distintive bed of rounded pointed out that the relative abundances of the different groups nodules . . . which represents a non-sequence marking the were not the same for these two geographical areas. However, Lower-Upper Gault boundary (although the top of the Walters (1958, Unpublished Ph.D. Thesis, U.C.W. Aberyst- cristatum Subzone is some 30 cms above this level)’. The Ford wyth) studying the foraminifera from several outcrops in section is therefore quite similar both lithologically and faunally Southeast England, including Folkstone, shows similar relative to the Folkstone section. abundances to those at Ford. These apparently contradictory Although Hart (1973) states that his faunal scheme, ‘can, results could be due to different approaches in the procedures unfortunately, only be described as being of local application,’ used to pick and count the specimens. the type Albian at L‘Aube, again shows a similar, major faunal The recorded assemblages at Ford, which are dominated by change over the Middle-Upper Albian boundary (lower-upper the superfamily Nodosariacea, do not seem to be typical of Gault boundary). Here, as at Folkstone, the passage to the the Middle Albian because in that interval the dominant group Upper Albian is marked by the appearance of Arenobulimina is the Superfamily Robertinacea (e.g. Hart & Carter, 1975; Hart chapmani Cushman and Frondicularia pinnaeformis Chapman et al., 1981, 1989). At the same time they are not quite the and their overlap with Epistomina spinulifera (Reuss) (see table 3 same as the typical assemblages of the Upper Albian because of Magniez-Jannin, 1975). in the upper part of the Gault Clay Formation, the fauna totally changes and becomes dominated by agglutinated taxa ACKNOWLEDGEMENTS although nodosarids and gavelinellids are quite numerous (Hart, The help of Ms K. M. Hawkins of the Buckinghamshire County 1990; Talwar, 1990, Unpublished M.Sc. Thesis, U.C.W. Museum (Aylesbury), who arranged the loan of the Hollis and Aberystwyth; Toogood, 1973, Unpublished M.Sc. Thesis, Neaverson’s set of specimens from that Museum and also U.C.W. Aberystwyth). offered helpful information on the site of the phosphatic nodules 80 Explanation of Plate 2. Hollis and Neaverson Collection of the Gault of Ford (Buckinghamshire. England). Unless specified, all the specimens are from the Micropalaeontological Museum, University of Wales (Aberysrtwyth). Scale bars 100pm. Fig. 1. AsfucoluF tripleura Reuss, 1860, AYBCM. 1920.38.97.4, Buckinghamshire County Museum (Aylesbury). Fig. 2. Cifharinella didyma (Berthelin, 1880), JH.389.26.20. Fig. 3. Dentalha debilis (Berthelin, 1880), JH.389.28.21. Fig. 4. Dentalino distincfa Reuss, 1860, JH.389.20.23. Fig. 5. Frondiculariaplanifolia Chapman, 1894, JH.389.24.31. Fig. 6. Friondiculariupinnneformis Chapman, 1894, JH.389.26.30. Fig. 7. Dentalino pulchella (Chapman, 1893), AYBCM.1920.38.41.2, Buckinghamshire County Museum (Aylesbury). Fig. 8. Dentalino legumen (Reuss, 1845), .JH.389.19.24. Fig. 9. Frondiculnrio goultina Reuss, 1860, AYBCM.1920.38.61.4, Buckinghamshire County Museum (Aylesbury). Fig. 10. Dentalinapseudononw Ten Dam, 1950, JH.389.20.25. Fig. 11. Denfdinogracilis (d.Orhigny, 1840), AYBCM.1920.38.50.2, Buckinghamshire County Museum (Aylesbury. Fig. 12. Frondicularia sp., JH. 389.26.32. Fig. 13. Dentalino sp. aff. D. infercellularir (Brady, 1881), JH. 389.19.26. Fig. 14. Frondiculurio denriculocarinata Chapman, 1894, JH.389.25.28. Fig. 15. Lageno afY. kispida Reuss, 1863, AYBCM.1920.:18.33.1, Buckinghamshire County Museum (Ayleshury). Fig. 16. Lageno apiculata (Reuss, 1851), P23140, Natural History Musewn (London). Fig. 17. Lenriculina circumcidanea (Berthelin, IBXO), JH.389.35.34. Fig. 18. Lmriculina diademafa (Berthelin, 1880), JH.389.34.35. Fig. 19. Lenticulina turgiddo (Reuss, 1863). JH.389.32.38. Fig. 20. Lenficulino gaulrina (Berthelin, I 880), JH.389.35.37. Fig. 21. Dentalina sp., JH.389.19.27. Fig. 22. Lingulinu loryi Berthelin, 1880, JH.389.24.29. Fig. 23. Lingulina semiornufa Reuss, 1863, JH.389.23.39. Fig. 24. ,Marginulinn inarqualis Rizuss, 1860, JH.389.28.40. 81 Explanation of Plate 3. Hollis and Neaverson Collection of the Gault of Ford (Buckinghamshire, England). Unless speciEed, all the specimens are from the Micropalaeontological Museum, University of Wales (Aberystwyth). Scale bars 1OOpm. Fig. 1. Marginulina jonesi (Reuss, 1863), JH.389.28.41. Fig. 2. Morginulina linearis (Reuss, 1863), AYBCM.1920.38.38.84.1, Buckinghamshire County Museum (Aylesbury). Fig. 3. Marginulim strialocostafa (Reuss, 1863). JH.389.28.42. Fig. 4. Marginulina sp., JH.389.31.43. Fig. 5. Nodosoria cf. afinis Reuss, 1845, JH.389.21.44 Fig. 6. Nodosaria/onfannesi(Berthelin, 1880), JH.389.21.46. Fig.7. Nodosaria orthopleura Reuss, 1863, JH.389.21.48. Fig. 8. Nodosoria cf. bambura Chapman, 1893, JH.389.21.45. Fig. 9. Nodosoria sp., AYBCM.1920.38.35.3, Buckinghamshire County Museum (Aylesbury). Fig. 10. Nodosaria obscura Reuss, 1845, JH.389.22.47. Fig. 11. Planularia bradyam (Chapman, 1894), JH.389.32.51.Fig. 12. Pseudonodosaria mulabilis (Reuss, 1863), P23103, Natural History Museum (London) Fig. 13. Nodosariapauperculu Reuss, 1845, JH.389.22.49. Fig. 14. Planularia cf. priceana (Chapman, 1894), JH.389.30.52. Fig. 15. Planularia vesrita (Berthelin, 1880), JH.389.33.53. Fig. 16. Planularia sp., JH.389.32.54. Fig. 17. Saracenaria navicula (d’orbigny, 1840), JH.389.31.56 Fig. 18. Soracenoria triangulark (d’orbigny, 1840), JH.389.31.57. Fig. 19. Nodosaria sceptrum Reuss, 1863, JH.389.22.50. Fig. 20. Vaginulinn recto Reuss, 1863, JH.389.29.61. Fig. 21. Vaginulina gaulfina Berthelin, 1880, JH.389.30.60. Fig. 22. Vaginulinn mediocarinnfa Ten Dam, 1950, JH.389.30.62. Fig. 23. Vaginulina humilis (Reuss, 1863), JH.389.32.61. Fig. 24. TrLtix excavofum (Reuss, 1863), JH.389.27.58. Fig. 25. Vaginulinn parallelu (Reuss, 1863), JH.389.32.63. Fig. 26. Tristix gaulrina Khan, 1950. JH.389.28.59. 82 Explanation of Plate 4. Hollis and Neaverson Collection of the Gault of Ford (Buckinghamshire, England). Unless specified, all the specimens are from the Micropalaeontological Museum, University of Wales (/\beryslwyth). Scale bars 100pm. Fig. 1. Clobufinafacrima(Rcuss, 1845), fistulose form, JH 389.37.65. Fig. 2. Ramufinaapliemis Banenstein & Brand, 1951, JH.389.39.69. Fig. 3. Pyrulina alT. bucculenta (Berthelin, l880), fistulose form, JH.389.37.66. Fig. 4. Pyrulina obfusa (Reuss, 1863), JH.389.37.67. Fig. 5. Ramulinafusiformis Khan, 1950, JH.389.39.70. Fig. 6. Pleurosfomella barroisi Berthelin, 1880, JH.389.16.75. Fig. 7. Ramuiina aff. globo-fubulosncushman, 1938, JH.389.39.71.Fig. 8. Ramulim sp., P23189, Natural History Museum (London). Fig. 9. Ramulina aculeara (d’Orbigny, 1840), JH.389.21.68. Fig. 10. Rumulino aff. laevir Jones, 1875, JH.389.39.72. Fig. 11. Ramulina muricarino Loehlich & Tappan, 1949, JH.389.39.73. Fig. 12. Pleurosrornella reussi Berthelin, 1880, JH.389.16.76. Fig. 13. Hoeglundina chapmani (Ten Dam, 1948). JH.389.42.79, (a) dorsal view, (b) venlral new, (c) lateral view. Fig. 14. Epirtomina spinulifea (Reuss, 1863), JH.389.43.78, (a) dorsal view, (b)ventral view, (c)lateral view. Fig. 15. Episromina aff. ornafa (Roemer, 1841). JH.389.42.77, (a) dorsal view, @) ventral view, (c) laleral view. Fig. 16. Cadinella intermedia (Berthelin, 1880), JM.389.41.80, (a) dorsal view, (b) ventral view, (c) lateral view. Fig. 17. Hedbergelia infracrefacea (Glaessner, 1937). JH.389.40.81, (a) dorsal view, (b) ventral view, (c) lateral view. 83 Herrero & Haynes Liverpool Geological Society and Manchester Geological Association, workings, is gratefully acknowledged. The first author would Special issue, 5 267-288. also like to express her gratitude to Dr John Whittaker, Mr Greg Hart, M. B. 1990. Cretaceous agglutinated foraminifera of the UK: A Welsh and Mr Richard Hodgkinson for their assistance and review. In Hemleben, C., Kaminski, M. A,, Kuhnt, W. & Scott, D. B. permission to examine the Gault foraminifera1 collections of (Eds) Faleoecology, Biostratigraphy, Paleoceanography and Taxonomy Hollis and Neaverson and Chapman in the Natural History of Agglutinated Foraminifera, 929-943. Kluwer, The Netherlands. Hart, M. B. & Carter, D. J. 1975. Some observations on the Cretaceous Museum (London). The photographic work of Mr D. D. Foraminiferida of SE England. Journal of Foraminifera1 Research, 5 Griffiths (UWA) and the photographs of the specimens of the 114-126. NHM provided by Dr John Whittaker are also gratefully Hart, M. B., Bailey, H. W., Fletcher, B. N., Price, R. J. & Swiecicki, A. acknowledged. This research has been partially supported by the 1981. Cretaceous. In Jenkins, D. G. & Murray, J. W. (Eds) Direccibn General de Investigacibn Cientifica, Ministerio de Stratigraphical Atlas of Fossil Foraminifera, 149-221. 1st Edition, British Micropalaeontological Society Series. Ellis Honvood, Chiche- Educacibn y Ciencia (Spain). ster. pp. 149-227. We are also indebted to Ms K. M. Hawkins for biographical Hart, M. B., Bailey, H. W., Crittenden, S., Fletcher, B. N., Price, R. J. & details of Hollis (Curator at the Aylesbury Museum from 1908 Swiecicki, A. 1989. Cretaceous. In Jenkins, D. G. & Murray, J. W. to 1941) and to Mr I. P. Roe, Headmaster at Aylesbury (Eds) Stratigraphical Atlas of Fossil Foraminifera, 213-311, 2nd. Grammar School, and Prof. C. Paul of the Department of Earth Edition, British Micropalaeontological Society Series. Ellis Horwood, Chichester. Sciences at Liverpool University, for their efforts in attempting Hart, M. B., Carter, D. J., Leary, P. N. & Talwar, A. D. 1990. to trace the movements of Neaverson. Agglutinated foraminiferida from the Albian/Cenomanian boundary NB. Unfortunately, although it is clear that Neaverson was in SE England. In Hemleben, C., Kaminski, M. A,, Kuhnt, W. & Science Master in Aylesbury Grammar School in the early 1920s Scott, D. B. (Eds) Paleoecology, Biostratigraphy, Paleoceanography and Taxonomy of Agglutinated Foraminifera, 945-960. Kluwer, The and went on to become Lecturer and Senior Lecturer at Netherlands. Liverpool, the dates when he entered and left these institutions Haynes, J. R. 1981. Foraminfera. Macmillan, London, 433 pp. are as yet unknown. Hollis, E. & Neaverson, E. 1921. Gault foraminifera from Ford, Bucks. Please note that the Aberystwyth Theses mentioned in the text Naturalist, Hull, 1 July: 235-240. can be obtained on loan from Hugh Owen Library (University Jukes-Brown, A. J. 1900. Gault and Upper Greensand (Selbomian) in Oxfordshire and Buckinghamshire. In: The Cretaceous Rocks of of Wales, Aberystwyth). Also photocopies from the National Britain, 1, The Gault and Upper Greensand (Selbornian) of England. Library of Wales (Aberystwyth). Memoir of the Geological Survey, London, 1: 275-283. 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