New Species and New Faunistic Data of West Palearctic Bisnius STEPHENS, 1829 (Coleoptera: Staphylinidae: Staphylininae)

106 Koleopt. Rdsch. 89 (2019) Koleopterologische Rundschau 89 107–114 Wien, September 2019

PACE, R. 1992: Aleocharinae nepalesi del Museo di Ginevra Parte VII (conclusione): Oxypodini e Aleocharini (Coleoptera, Staphylinidae) (115° Contributo alla conoscenza delle Aleochrinae). – Revue Suisse de Zoologie 99 (2): 263–342. New species and new faunistic data of PACE, R. 1999a: Aleocharinae dell Cina: Parte V (conclusione) (Coleoptera, Staphylinidae). – Revue West Palearctic Bisnius STEPHENS, 1829 Suisse de Zoologie 106 (1): 107–164. (Coleoptera: Staphylinidae: Staphylininae) PACE, R. 1999b: Aleocharinae del Cile (Coleoptera, Staphylinidae). – Bollettino del Museo Civico di Storia Naturale di Verona 23: 119–210. H. SCHILLHAMMER PACE, R. 2005: Aleocharinae of the Australian Region collected by Dr. P. M. Giachino and Dr. M. Daccordi (Coleoptera, Staphylinidae). – Monografie di Museo Regionale di Scienze naturali, Torino 42: 371–434. Abstract PACE, R. 2012: Biodiversità delle Aleocharinae della Cina: Hygronomini e Oxypodini (Coleoptera, Two new species of West Palearctic Bisnius STEPHENS, 1829 (Coleoptera: Staphylinidae: Staphylin- Staphylinidae). – Beiträge zur Entomologie, Keltern 62 (1): 125–163. inae, Philonthina) are described and illustrated: B. karkarensis (Armenia) and B. iranicus (Iran). Additional faunistic data are provided for Bisnius microphthalmus SCHILLHAMMER, 2011, B. schill- PAŚNIK, G. 2001: The North Korean Aleocharinae (Coleoptera, Staphylinidae): diversity and biogeo- hammeri HROMADKA, 2001 and B. zhuk (GUSAROV, 1995). Bisnius microphthalmus SCHILLHAMMER, graphy. – Acta Zoologica Cracoviensia 44 (3): 185–234. 2011 is recorded from Kazakhstan for the first time. QUEDENFELDT, M. 1881: Diagnose einer neuen europäischen Art der Staphylinen-Gattung Echidnoglossa Wollast. – Berliner Entomologische Zeitschrift 25 (1–2): 293. Key words: Coleoptera, Staphylinidae, Staphylininae, Philonthina, Bisnius, new species, new data, West Palearctic. SAINTE-CLAIRE DEVILLE, J. 1913: Nouvelle capture du Blepharrhynemus [sic] mirandus Fauv. [Col. Staphylindae]. – Bulletin de la Société Entomologique de France 1913 (1): 48–49. Introduction SCHEERPELTZ, O. 1954: Die paläarktischen Arten der Gattung Blepharrhymenus Solier. (Col. Staphyl.). – Koleopterologische Rundschau 32 [1951–1954]: 132–143. The genus Bisnius STEPHENS, 1829 is a moderately speciose (current knowledge) genus in the subtribe Philonthina with an almost strictly Holarctic distribution. SCHÜLKE, M. & SMETANA, A. 2015: Staphylinidae, pp. 304–1134. – In Löbl, I. & Löbl, D. (eds): Catalogue of Palaearctic Coleoptera. Volume 2. Hydrophiloidea – Staphylinoidea. Revised and There is quite a number of potential Bisnius species still residing in the genus Philonthus which updated edition. – Leiden: Brill, xxvi + 1702 pp. have not yet been studied for correct generic assignment. In addition, the monophyly of the SKALITZKY, C. 1884: Zwei neue europäische Staphylinenarten aus Portugal. – Wiener Entomologische genus is doubtful and the systematic position of the fimetarius group, outside of Bisnius s.str. in Zeitung 3 (4): 97–99. CHANI-POSSE et al. (2018), should be assessed in detail. SOLIER, A.J.J. 1849: Orden III. Coleopteros; pp. 105–380, 414–511. – In Gay, C. (ed.): Historia fisica y In this paper, two new species of Bisnius are described: B. karkarensis (Armenia) and B. iranicus politica de Chile segun documentos adquiridos en esta republica durante doce años de residencia (Iran), and new or additional faunistic data are provided for Bisnius microphthalmus SCHILL- en ella y publicada bajo los auspicios del supremo gobierno. Zoologia. Tomo Cuarto. – Paris: C. HAMMER, 2011, B. schillhammeri HROMADKA, 2001 and B. zhuk (GUSAROV, 1995). Gay, 511 pp. SONG, J.-H. & AHN, K.-J. 2013: Blepharhymenus koreanus Paśnik and Tachyusa wei Pace new to South Korea (Coleoptera: Staphylinidae: Aleocharinae). – Korean Journal of Applied Entomology 52 Abbreviations and acknowledgements (4): 311–314. The material treated in this paper is deposited in the following institutional and private SUN, X. (ed.) 1989: Atlas of the People’s Republic of China. – Beijing: Foreign Languages Press, China collections: Cartographic Publishing House, 113 pp., 51 pls. CAH coll. Volker Assing, Hannover, Germany VÍT, S. & HOZMAN, P. 1980: Coléoptères intéressants et nouvaux pour la faune suisse. – Mitteilungen der CGO coll. Vladimir Gusarov, Oslo, Norway Schweizerischen Entomologischen Gesellschaft 53: 285–295. CSB coll. Michael Schülke, Berlin, Germany WOLLASTON, T.V. 1864: Catalogue of the coleopterous insects of the Canaries in the collection of the HUB Museum für Naturkunde – Leibniz Institute for Evolution and Biodiversity Science, Berlin, Germany British Museum. – London: Order of the Trustees, 648 pp. (J. Frisch) NMP Národní muzeum (National Museum), Praha, Czechia (J. Hájek) ZANETTI, A., SETTE, A., POGGI, R. & TAGLIAPIETRA, A. 2016: Biodiversity of Staphylinidae (Coleoptera) NMW Naturhistorisches Museum Wien, Austria in the Province of Verona (Veneto, Northern Italy). – Memorie della Società Entomologica Italiana 93 (1–2): 3–237. The curators mentioned above are thanked for providing specimens and help in various ways. In addition, I am grateful to Adam J. Brunke (Ottawa, Canada) for valuable comments to im- Dr. Volker ASSING prove the manuscript. Gabelsbergerstr. 2, D – 30163 Hannover, Germany ([email protected])

108 Koleopt. Rdsch. 89 (2019)

Bisnius zhuk (GUSAROV, 1995) Philonthus zhuk GUSAROV 1995: 93. TYPE MATERIAL EXAMINED: Holotype : “Krim, nad Massandroj, Gusarov, 31.V.1990 [handwritten, in Russian] \ na excrementakh korovy” (CGO). REDESCRIPTION: 6.2–7.3 mm long (3.1–3.5 mm, abdomen excluded). Black, moderately shining; elytra brown to dark reddish brown, distinctly darkened in basal depression, deflexed anterior portion between shoulders and neck sometimes reddish; abdominal tergites with posterior margins obscurely reddish, tergite X pale brown; legs reddish, medial faces of tibiae, especially metatibiae, infuscate. Head rounded quadrangular, about as long as wide, eyes moderately large, tempora 1.35–1.60 times as long as eyes; medial interocular punctures widely separated, as large as lateral interocular punctures, situated in front of level of lateral interocular punctures, vertex impunctate; antennae with segments 4–5 inconspicuously oblong, segments 6–10 about as long as wide; pronotum subparallel-sided, 1.10–1.15 times as long as wide, dorsal rows each with four more or less equidistant punctures; head and pronotum with very short-meshed microsculpture; elytra rather densely and coarsely punctate, distance between individual punctures variable, generally about a puncture diameter, but also in places slightly less or slightly more; scutellum very finely, sparsely punctate, surface with very fine transverse microsculpture; abdominal tergites III–VI with two basal lines, elevated area between basal lines impunctate on tergites III–V, that on tergite VI with a sparse row of fine setiferous punctures; remaining surface of tergites finely, moderately densely punctate, punctures almost forming indistinct transverse rows. Aedeagus (Figs. 1–2) short, broad, with rounded apex; in lateral view broad, but apical portion rather flat; paramere (Figs. 1c, 2c) broad, slightly broader than median lobe (the holotype is slightly teneral with a partly collapsed median lobe, therefore the paramere appears much broader than the median lobe), apical margin deeply emarginate with additional small notch medially, with dense row of peg setae but without peg setae at medial notch. Generally, the aedeagus is similar to that of Bisnius fimetarius GRAVENHORST, 1802, but in the paramere of B. fimetarius the apical margin is only weakly emarginate, distinctly sinuate and with a much deeper medial notch. DIAGNOSIS: Externally, the species is very similar to B. fimetarius except for the smaller eyes (tempora less than 1.2 times as long as eyes in B. fimetarius). From the former as well as from B. schillhammeri HROMADKA, 2001 and B. karkarensis it is distinguished mainly by the brownish elytra. ADDITIONAL MATERIAL: T U R K M E N I S T A N: 2 : “USSR – Turkmenia, goldener Bach, westl. Aschchabad, 7.V.89, Wrase” (CSB, NMW); 1 : “Turkm., Aschchabad, 7.V.[19]89, Wrase” (CSB). DISTRIBUTION: The species has been recorded from southern Russia, Azerbaijan and Turk- menistan. BIONOMICS: The specimens of the type series have been collected from excrements of various animals, but also from decaying mushrooms.

108 Koleopt. Rdsch. 89 (2019) SCHILLHAMMER: New species and new faunistic data of West Palearctic Bisnius (STAPHYLINIDAE) 109

Bisnius schillhammeri HROMADKA, 2001 Bisnius schillhammeri HROMADKA 2001: 139. TYPE MATERIAL: Holotype  (not studied for this paper): “S Iran, 29 km Yasui, 2300 m, 16.-17.6.1973, Loc. No. 245, Exp. Mus. Nat. Praha” (NMP), cited from the original description. Figs. 1–2: Aedeagus of Bisnius zhuk: 1) holotype, 2) specimen from Turkmenistan; ventral view (a), lateral view (b), paramere (c). – Scale bars: 0.50 mm (a, b), 0.25 mm (c). 110 Koleopt. Rdsch. 89 (2019)

Figs. 3–4: Aedeagus of 3) Bisnius schillhammeri; 4) B. karkarensis; ventral view (a), lateral view (b), paramere (c). – Scale bars: 0.50 mm (a, b), 0.25 mm (c). 110 Koleopt. Rdsch. 89 (2019) SCHILLHAMMER: New species and new faunistic data of West Palearctic Bisnius (STAPHYLINIDAE) 111

Figs. 3–4: Aedeagus of 3) Bisnius schillhammeri; 4) B. karkarensis; ventral view (a), lateral view (b), Figs. 5–7: 5) Aedeagus of Bisnius iranicus; ventral view (a), lateral view (b), paramere (c); 6–7) head of paramere (c). – Scale bars: 0.50 mm (a, b), 0.25 mm (c). B. iranicus (6) and B. microphthalmus (7). – Scale bars: 0.50 mm (a, b), 0.25 mm (c). 112 Koleopt. Rdsch. 89 (2019)

The redescription is based on a single, newly collected male specimen because some parts of the original description are doubtful (especially the oblong head and the body size). The holotype has to be re-studied at a given time. REDESCRIPTION: 6.8 mm long (3.3 mm, abdomen excluded). Black, shining; elytra dark brown to black brown, posterior margins of abdominal tergites broadly but obscurely reddish, palpi dark brown, last segment of maxillary palpi paler, legs dark brown, medial faces of meso- and metatibiae infuscate, tibiae narrowly reddish proximally, tarsi reddish with first segment somewhat darker. Head rounded quadrangular, about as long as wide, eyes rather small, tempora subparallel, 1.6 times as long as eyes; medial interocular punctures widely separated, slightly shifted anteriad, as large as lateral interocular punctures, distance between medial interocular punctures about four times distance between medial and lateral interocular puncture, vertex medially impunctate, with two pairs of larger punctures laterally behind level of posterior margin of eye and a pair of smaller punctures posteriorly; antennae with segment 4 inconspicuously oblong, segments 5–7 about as long as wide, 8–10 slightly transverse; pronotum subparallel-sided, 1.15 times as long as wide, dorsal rows each with four punctures, anterior three almost equidistant, distance between 3 and 4 slightly wider; head and pronotum with short-meshed, but rather obsolete microsculpture, especially on medial portion of pronotum barely visible; elytra rather densely and coarsely punctate, distance between punctures about a puncture diameter or slightly more; scutellum moderately coarsely and moderately densely punctate; abdominal tergites III–VI with two basal lines, elevated area between basal lines impunctate; remaining surface of tergites finely, moderately densely punctate. Aedeagus (Fig. 3) with median lobe short and broad in ventral view, but apical portion slightly less broad than in B. zhuk; paramere (Fig. 3c) with much flatter apical emargination than in B. zhuk, medially, without notch and even slightly produced anteriad; apical margin with dense uninterrupted row of peg setae. DIAGNOSIS: The species differs from Bisnius zhuk in the darker elytra, from B. fimetarius, B. zhuk and B. karkarensis in the very weak microsculpture on head and pronotum, and from B. karkarensis in the more robust body and longer antennae. ADDITIONAL MATERIAL: I R A N: Isfahan Province: 1 : S of Fereydun Shahr, Gukan, 2260 m, 32°42'36"N 50°05'16"E, 2.VII.2004, leg. J. Frisch (HUB). DISTRIBUTION: The species is at present known only from Iran (provinces of Isfahan and Kohgīlūyeh va Būyer Aḩmad). BIONOMICS: Nothing is known about the habitat requirements of the species.

Bisnius karkarensis sp.n. Holotype : “ARMENIA [46] – 30 km NW Sisian, Mt. Karkar, 39°46'51"N 45°56'30"E, 2990 m, manure sifted, 11.VII.2017, V. Assing” (CAH). Paratypes (10 exs.): 5 : same label data as holotype (4 CAH, 1 NMW); 1 , 2 : “ARMENIA [AR17-46] 30 km NW Sisian, Mt. Karkar, 39°46'51"N 45°56'30"E, 2990 m, straw manure sifted, 11.VII.2017, leg. M. Schülke” (2 CSB, 1 NMW); 1 : “ARMENIA [AR17-27] SW Gavar, 40°14'31"N 45°01'41"E, 2570 m, stream valley, moist litter and roots near stream sifted, 4.VII.2017, leg. M. Schülke” (CSB); 1 : “ARMENIA [AR17-42] 40 km NW Sisian, Vorotan Pass, 39°40'33"N 45°45'07"E, 2140 m, stream valley with Salix, litter and roots, sifted, 10.VII.2017, M. Schülke” (CSB). DESCRIPTION: 5.2–6.5 mm long (2.8–3.0 mm, abdomen excluded). Black, moderately shining; tips of mandibles, outer faces of tibiae, tarsi and tergite X reddish brown to pale brown, reddish 112 Koleopt. Rdsch. 89 (2019) SCHILLHAMMER: New species and new faunistic data of West Palearctic Bisnius (STAPHYLINIDAE) 113

The redescription is based on a single, newly collected male specimen because some parts of the stripe on tibiae often reduced to short and narrow portion proximally, basal one or two tarso- original description are doubtful (especially the oblong head and the body size). The holotype meres often darkened to almost black. has to be re-studied at a given time. Head rounded quadrangular, as long as wide or inconspicuously wider than long, eyes REDESCRIPTION: 6.8 mm long (3.3 mm, abdomen excluded). Black, shining; elytra dark moderately large, tempora 1.36–1.50 times as long as eyes; medial interocular punctures widely brown to black brown, posterior margins of abdominal tergites broadly but obscurely reddish, separated, smaller than lateral interocular punctures, situated in front of level of lateral inter- palpi dark brown, last segment of maxillary palpi paler, legs dark brown, medial faces of meso- ocular punctures, vertex impunctate; antennae with segment 4 inconspicuously oblong, segments and metatibiae infuscate, tibiae narrowly reddish proximally, tarsi reddish with first segment 5 and 6 about as long as wide, 8–10 weakly transverse; pronotum subparallel-sided, 1.10–1.15 somewhat darker. times as long as wide, dorsal rows each with four more or less equidistant punctures; head and Head rounded quadrangular, about as long as wide, eyes rather small, tempora subparallel, 1.6 pronotum with very short-meshed microsculpture; elytra rather densely and coarsely punctate, times as long as eyes; medial interocular punctures widely separated, slightly shifted anteriad, as punctures separated by a puncture diameter or less, slightly less dense basally; abdominal large as lateral interocular punctures, distance between medial interocular punctures about four tergites III–VI with two basal lines, elevated area between basal lines impunctate on tergites III– times distance between medial and lateral interocular puncture, vertex medially impunctate, with V, that on tergite VI with a sparse row of fine setiferous punctures; remaining surface of tergites two pairs of larger punctures laterally behind level of posterior margin of eye and a pair of finely, moderately densely punctate, punctures almost forming indistinct transverse rows. smaller punctures posteriorly; antennae with segment 4 inconspicuously oblong, segments 5–7 Aedeagus (Fig. 4) smaller than in the previous two species, median lobe more slender with about as long as wide, 8–10 slightly transverse; pronotum subparallel-sided, 1.15 times as long shorter and less flat (lateral view) apical portion; paramere (Fig. 4c) less broad, with shallow as wide, dorsal rows each with four punctures, anterior three almost equidistant, distance apical emargination, apical margin with dense uninterrupted row of peg setae. between 3 and 4 slightly wider; head and pronotum with short-meshed, but rather obsolete DIAGNOSIS: Bisnius karkarensis differs from both previous species in the smaller body size, microsculpture, especially on medial portion of pronotum barely visible; elytra rather densely generally more slender build, shorter antennae and the completely black abdomen. and coarsely punctate, distance between punctures about a puncture diameter or slightly more; scutellum moderately coarsely and moderately densely punctate; abdominal tergites III–VI with DISTRIBUTION: The species is at present known only from the type locality. two basal lines, elevated area between basal lines impunctate; remaining surface of tergites BIONOMICS: The specimens have been sifted from different kinds of plant material, mostly finely, moderately densely punctate. from leaf litter along streams. All specimens are from elevations above 2000 m. Aedeagus (Fig. 3) with median lobe short and broad in ventral view, but apical portion slightly DERIVATIO NOMINIS: The species is named after the type locality. less broad than in B. zhuk; paramere (Fig. 3c) with much flatter apical emargination than in B. zhuk, medially, without notch and even slightly produced anteriad; apical margin with dense uninterrupted row of peg setae. Bisnius microphthalmus SCHILLHAMMER, 2011 DIAGNOSIS: The species differs from Bisnius zhuk in the darker elytra, from B. fimetarius, Bisnius microphthalmus SCHILLHAMMER 2011: 136. B. zhuk and B. karkarensis in the very weak microsculpture on head and pronotum, and from ADDITIONAL MATERIAL: B. karkarensis in the more robust body and longer antennae. K A Z A K H S T A N: Turkistan Region: 1 : “Kazakhstan: Aksu-Djabagly, Djabagy riv., Kshi-Kaindy, 19.06.2001, V. Kastcheev” (NMW). ADDITIONAL MATERIAL: I R A N: Isfahan Province: 1 : S of Fereydun Shahr, Gukan, 2260 m, 32°42'36"N 50°05'16"E, 2.VII.2004, leg. First record for Kazakhstan! J. Frisch (HUB).

DISTRIBUTION: The species is at present known only from Iran (provinces of Isfahan and Bisnius iranicus sp.n. Kohgīlūyeh va Būyer Aḩmad). Holotype : “IRAN (Kohkīlyeh va Būyer Ahmadī [Kohgīlūyeh va Būyer Aḩmad]) Zagros Mts., Dena Mts., BIONOMICS: Nothing is known about the habitat requirements of the species. Sisakht, Kugol vall., 2750 m, 30°49'49"N 51°32'50"E, (subalpine pasture with tall herbaceous vegetation/under stones), 25.IV.2018, Wrase & Laser [18]” (CSB). Paratype , same data as holotype (NMW). Bisnius karkarensis sp.n. DESCRIPTION: 8.0–9.5 mm long (4.5–4.9 mm, abdomen excluded). Black, matt, elytra reddish, abdominal tergites with posterior margins obscurely reddish, palpi pale reddish brown, legs dark Holotype : “ARMENIA [46] – 30 km NW Sisian, Mt. Karkar, 39°46'51"N 45°56'30"E, 2990 m, manure sifted, reddish, medial faces of metatibiae infuscate. 11.VII.2017, V. Assing” (CAH). Paratypes (10 exs.): 5 : same label data as holotype (4 CAH, 1 NMW); 1 , 2 : “ARMENIA [AR17-46] 30 km NW Sisian, Mt. Karkar, 39°46'51"N 45°56'30"E, 2990 m, straw manure Head rounded subquadrangular, 1.11–1.16 times as wide as long, widest at about level of half- sifted, 11.VII.2017, leg. M. Schülke” (2 CSB, 1 NMW); 1 : “ARMENIA [AR17-27] SW Gavar, 40°14'31"N length of tempora, eyes very small, tempora 2.7 (female paratype) to 3.2 (male holotype) times 45°01'41"E, 2570 m, stream valley, moist litter and roots near stream sifted, 4.VII.2017, leg. M. Schülke” (CSB); as long as eyes; with four interocular punctures, medial interocular punctures separated by about 1 : “ARMENIA [AR17-42] 40 km NW Sisian, Vorotan Pass, 39°40'33"N 45°45'07"E, 2140 m, stream valley with Salix, litter and roots, sifted, 10.VII.2017, M. Schülke” (CSB). twice the distance between medial and lateral interocular puncture, all interocular punctures arranged in approximately one line at about level of half-length of eye; for chaetotaxy of head DESCRIPTION: 5.2–6.5 mm long (2.8–3.0 mm, abdomen excluded). Black, moderately shining; see Fig. 6; neck very broad, broader than half-width of head; antennae slightly sexually di- tips of mandibles, outer faces of tibiae, tarsi and tergite X reddish brown to pale brown, reddish morphic – male: segment 4 slightly oblong, segments 5–6 about as long as wide, 7–10 transverse; female: segment 4 as long as wide, 5 slightly transverse, segments 6–10 more distinctly 114 Koleopt. Rdsch. 89 (2019) transverse; pronotum about as long as wide, sides very weakly rounded; dorsal rows each with four punctures, anterior three punctures equidistant, distance between punctures 3 and 4 slightly larger; both head and pronotum with dense and distinct microsculpture of transverse waves, which are curved anteriad at base of head; elytra along shoulders much longer than pronotum along midline, finely punctate, punctures separated by about a puncture diameter or slightly more, due to small size of punctures punctation appearing rather dense; scutellum moderately densely, finely punctate, with exceedingly fine, short-meshed microsculpture; abdominal tergites III–VI with two basal lines, elevated area between basal lines densely, moderately strongly punctate and with dense isodiametrical microsculpture, in addition, with shallow depression immediately posterior of second basal line, with isodiametrical microsculpture in depression and very fine wavy microsculpture on remaining surface, tergites VII–VIII with a similar pattern of microsculpture but without depression at base. Aedeagus (Fig. 5) with moderately broad median lobe, lateral outline of apical portion slightly sinuately narrowed toward subtruncate apex; paramere (Fig. 5c) moderately long, subparallel- sided, with rounded apex; with a subapical cluster of 12 peg setae arranged in two almost regular rows. Median lobe of B. microphthalmus with more slender apical portion of median lobe and broader paramere with subapical cluster of irregularly arranged peg setae (see SCHILLHAMMER 2011: Fig. 3). DIAGNOSIS: The species is very similar to Bisnius microphthalmus, but immediately differs in the reddish elytra. In addition, the medial interocular punctures are more widely separated (Fig. 6) (more or less equidistant in B. microphthalmus – see Fig. 7). DISTRIBUTION: The species is at present known only from the type locality. BIONOMICS: The type specimens were collected from under stones at an elevation of 2750 m. DERIVATIO NOMINIS: The species is named after the country of its origin.

Zusammenfassung Zwei neue westpaläarktische Arten der Gattung Bisnius STEPHENS, 1829 werden beschrieben und illustriert: Bisnius karkarensis (Armenien) und B. iranicus (Iran). Zusätzliche Funddaten für Bisnius microphthalmus SCHILLHAMMER, 2011, B. schillhammeri HROMADKA, 2001 and B. zhuk (GUSAROV, 1995) werden präsentiert.

References CHANI-POSSE, M.R., BRUNKE, A.J., CHATZIMANOLIS, S., SCHILLHAMMER, H., SOLODOVNIKOV, A.Yu. 2018: Phylogeny of the hyper-diverse rove beetle subtribe Philonthina with implications for classification of the tribe Staphylinini (Coleoptera: Staphylinidae). – Cladistics 34: 1–40. GUSAROV, V. 1995: New and little known Palearctic staphylinids (Coleoptera, Staphylinidae). – Entomo- logicheskoe Obozrenie 74 (1): 81–96 [in Russian]. HROMADKA, L. 2001: Zwei neue Staphyliniden-Arten aus Iran. – Reichenbachia 34 (16): 137–141. SCHILLHAMMER, H. 2011: Old and new Staphylinini from the Palearctic and Oriental regions. – Koleo- pterologische Rundschau 81: 133–163.

Dr. Harald SCHILLHAMMER Naturhistorisches Museum Wien, Burgring 7, A – 1010 Wien, Austria ([email protected])