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Generic Classification of the Beetle Family Tetratomidae (Coleoptera, Tenebrionoidea) of the World, with Description of New Taxa

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Generic Classification of the Beetle Family Tetratomidae (Coleoptera, Tenebrionoidea) of the World, with Description of New Taxa Nikolai B. Nikitsky Generic classification of the beetle family Tetratomidae (Coleoptera, Tenebrionoidea) of the world, with description of new taxa Pensott Series Faunistica No. 9 Generic classification of the beetle family Tetratomidae (Coleoptera, Tenebrionoidea} of the world, with description of new taxa by Nikolai B. Nikitsky Sofia - Moscow 1998 ...PEJtSV E..I@Publishers An international publishing house in allfields of environmental & life sciences Acad. G. Bonchev Str., Bl. 6 1113 Sofia, Bulgaria Tel./Fax+359-2-7133460 or-7133406 E-mail: [email protected] © PENSOFfPublishers ISBN 954-642-039-5 Firstpublished 1998 All rights reseIVed Editor: Dr. S. Golovatch Graphic and coverdesign: Dr.V. Milgram Onthe cover: Pisenuschujoi Miyatake, 1960 (original) Printed by Biolnform SCIVices, Ltd., Moscow, Russia, January, 1998 CONTENTS Ab stract .................................................................................................................. 5 Introduction.. .. ... .. ... ... ... ... .... .. .. .. ..... .. .. .. .. .. .. .. .. .. ... .. .. .. ... .. 6 Mozphologicalc cs .................................................................................. 6 haracteristi gnosIS. .............................................................................................................. 10 Dia Key tosub families,tribes and genera of Tetratomidaebas edon imago ................ 10 Key tosub familiesand genera ofTetratomidae bas edon larvae ........................... 14 SubfamilyTetratominae Bill berg,18 20 ................................................................. 16 Genus TetratomaFabricius, 1790 .......................................................................... 16 Key tosubgenera of Tetratomabas edon imago.................................................... 19 Tetratoma(Tetratoma ) tiiscoguttat.asp. nov . ....... ........ ............. ........ .. ... .... .. ..... .. ... 21 Te tratoma(T.) nepalensissp.nov . ......................................................................... 23 Te tratoma (T.) wittrJJ.erisp. nov . ............................................................................ 25 Key to speciesof Tetratomabas edon larvae................ ......................................... 26 SubfamilyPiseninae Miyatake, 19 60 ····························································�······· 27 Genus Pis enllSCasey, 1900 .................................................................................. 27 Key tospecies of Pis enllS basedon larvae.................... ........................................ 27 Genus No topisenllSNikitsky et Lawrence, 1992 .................................................. 29 GenusTri phylliaReitter, 1898 .............................................................................. 29 SubfamilyPentbinae Lacordaire, 1859 ................................................................. 29 � Genus Cyanopenth c gen. nov . ........... ..... .... ............. ..... ........ .. ......... ............ ... ... 29 Cyanopenthc sp. nov . ......................................................................... 30 thailandica Genus Pcnth cNewman, 1838 ............................................................................... 32 Pcnth c reitterisp. nov ............................................................................................ 32 SubfamilyHall omeninaeMulsant, 1856 .............................................................. 33 Genus HallomcnllS Panzer, 1794 .......................................................................... 33 Ha llomcnllS (HallomcnllS} chinensissp . nov .................................. ; ...................... 35 HallomenllS (H.) orient.alissp.nov . ........................... ........................................... 36 Key tospecies of Hall omenllS basedon larvae .................................................... 39 Genus MycetomaDejean, 1834 ............................................................................ 39 SubfamilyEustrop hinaeGistel, 1856 ························'.···················· ...... ....... ...... ... 39 TribeHolostrophini, tribus nov . ............................................................................. 39 Genus PseudoholostrophllSNikitsky, 1983 ........................................................... 40 Key tospecies of Pseudoholostroph11Sbased on imago ........................................ 40 PseudoholostrophllS (Holostrophinussub gen. nov. ) discolor (Hom, 1888), comb. nov .............................................................................................................. 43 Pseudoholostroph11S (Pseudoholostrophus) chinensissp. nov . ..... ......................... 44 PseudoholostrophllS{P.)klapperichi(Pic, 1954) ................................................... 45 Pseudoholostrophus(P.) impressicollis(LeConte, 1874), comb. nov . .......... ..... .... 47 GenusHo lostroph usHom, 1888 .......................................................................... 48 4 Holostrophusminimus sp.nov . ............................................................................. 48 Holostrophussimilis sp.nov . ................................................................................. 51 Holostroph usorien. talis Lewis,1895 ..................................................... ...... .......... 52 ?Holostrophusaureofasciatus(Pic, 1954) ............................................................. 52 Key tospecies of Holostroph usb�ed on larvae ................................................... 53 TribeEustrophini Gistel, 1856 .............................................................................. 53 GenusEus trophusIlli ger,1802 ............................................................................. 54 Eustroph us)'UI1I111J1 en.sissp. nov. ........................................................................... 54 Genus SynstrophusSeidlitz,1898 ......................................................................... 58 Genus Eustrophopsis Champion, 1889 ................................................................ 58 Eustrophopsis similis sp.nov.. ............................................................................... 59 Eustrophopsis sexm.aculatasp. nov .. ..................................................................... 60 Eustrophopsis ochracea(Motschoulsky, 1872) ..................................................... 63 Genus CurteustrvphinusPie, 1952 ........................................................................ 63 Acknowledgm.ents ................................................................................................ 63 References. ................................................ .. .......................................................... 64 Index ................................................................................................................ 79 ABSTRACT The beetlefamily Tetratomidae is diagnosed, withdetailed keys to subfamilies,genera and some species on TetratomaF. consideredas thesole constituent genus of based adults and laivae. is Tetratominaeincluding the subgenera Tetraromas. str.,AbstruliaCasey, Falsoxantbalia Pie, stat nov., Incolia Casey, nov. (bothpreviously regardedas genera), andParatetrafo 1111l stat full-rank subgen.nov. (type-species: Tetrafo1111lsabgutiiNakane,1955). new species described Three are genus: (T.) tiiscoguttBta nov. and (T.) nepalensis nov. fromNepal, and in this T. sp. T. sp. T.(T.) wittmeri sp. nov. from Bhutan. A new genus, Cyan.openth.e gen. nov., is established in the subfamilyPenthinae, with Penth.e me on, 1916 asits type-species and Cyan.openth.e taJlicaChampi nov., a,.q a second congener.Penth.e r:eitteri nov. descn"bed tha.il;mdica sp. fromThailand, sp. is from Sechuan, In thesubfamily Hallomeninae,which onlyincludes the genera HaJlomenus China. andMyceto1111l Dej.,the subgenus Parahaflomenus Nomuraet 1958, genusHall omenus, Pz. KatO, isconsidered as a new subjectivejunior synonym (syn. nov.) of thesubgenus XeuxesChampion, 1889, stat nov., thelatter tax.on previously regardedas agenus of full Basedrank. on a restudyof Pie, 1930, Hallomenus 1817, reticul1Jtus type specimens,XeuxeY diversicomis paJlensGyll.enbal, H Motschoulsky, 1872, H fuscosutumlis Blatchley, 1913 and H irmatusKangas, 1959 actually belongto neitherTetratomidae nor Melandryidae.Hall omenus variegatus Motschoulsky, 1872 and H 8D11Spioides Motschoulsky,1845 obviously do not belongin Tetratomidaeeither, but this could not reconfirmedby pertinent Twonew species described Hallomenus: be material. are in H chinensis nov. from Sechuan, China and orient.a1is nov. from India and sp. H sp. Kashmir, Pakistan.In the subfamily Eustrophinae,which includesthe genera Pseudoholostrophus Hazara, Nikitsky,1983, Holostrophus Horn,1888, Eustrophus Illiger, 1802, Eustrophopsis Champion, 1889 and S)'IlStrophus Seidlitz, 1898, a new tribe is established,Holostropbini tribus nov., to incorporateHolostrophusaruiPseJXkJholostrophU£ Holostrophusis typified forthe first time (type­ species: Eustrophus bifasciatus Say, 1824), with H minimus sp. nov. describedfrom Luzon, Philippinesand H similis sp.nov. from Sarawak,Malaysia. Based on a restudyof thetype of H multinotatus Pie, 1911, fromTaiwan, this taxon is considered as aninfraspecific color variation (syn. nov., stat nov.) of H orietJ.t.a1is Lewis, 1895. Eustrophinus aureofascliitus Pie, 1954, described from China,is transferredinto Holostrophus: ?H aureofasciatus(Pie, 1954), comb. nov. Pseudoholostrophusisconsidered to encompass two subgenera: Pseudoholostrophus
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