Contr. Tert. Quatern. Geol. 29(1-2) 27-37 1 fig., 3 pis. Leiden, June 1992

New Pliocene and Oligocene Olividae

(Mollusca, Gastropoda) from France and

the Mediterranean area

Pierre Lozouet

Museum National d'Histoire Naturclle

Paris, France

— Lozouet, Pierre. New Pliocene and Oligocene Olividae (Mollusca, Gastropoda) from France and the Mediterranean area. Contr.

Tert. Quatern. Geol., 29(1-2): 27-37, 1 fig., 3 pis. Leiden, June 1992.

Two olivid Olivella and Amalda cf. obsoleta recorded for the first time from the Mediterranean species, clanzigi n. sp. (Brocchi, 1814), are

Pliocene of Tunisia and Spain, respectively. It has generally been assumed that the Olividae had disappeared from the Mediterranean

area during the Late Miocene, having fallen victim to the Messinian salinity crisis. The climatic deterioration during the Pliocene would

of not have permitted their re-introduction into the area. In addition, a new species of Amalda,A. abessensis, from the Late Oligocene

France is described and illustrated. This have match in Cainozoic faunas and is close Amalda species appears to no European to papillata

the of the from the Miocene ofAustralia. In view of the Tate, 1885, type species subgenus Alocospira Cossmann, 1899, Early this, most

Gracilancilla G. important Late Oligocene-Miocene Ancillinae of Europe are here illustrated; it is shown that Thiele, 1925 (type species:

ofAncillina 1882 Ancilla Fuchs in sumatrana Thiele, 1925, western Indo-Pacific) is a junior synonym Bellardi, (type species: pusilla Karrer,

1877, European Miocene).

Key words Mollusca, Gastropoda, Olividae, Oligocene-Pliocene, Mediterranean area, France, new taxa.

des Invertebres Marins de P. Lozouet, Museum National d'Histoire Naturelle, Laboratoire de Biologie et Malacologie, 55 rue Buffon,

75005 Paris, France.

Contents to describe more exhaustively the Pliocene popula-

tions of the Mediterranean The of area. discovery Introduction p. 27 two olivid from and species Spain Tunisia, may Systematic descriptions p. 27 illustration in this serve as an respect. Conclusions 32 p. Field work carried fifteen out during nearly years Acknowledgements p. 32 in the Oligocene ofAquitaine (France) has led to the References p. 32 of which discovery numerous new species, amongst

of Amalda described herein. In is the new species Introduction view of this, most of the Oligocene and Neogene

Field work carried out in co-operation with Mr Ancillinae of Europe are here illustrated.

Sylvain Clanzig in Spain, in the counties of The material described is housed in the following

Maghreb (Morocco, Algeria and Tunisia) and in institutions: MNHN (Museum National d'Histoire

northern Italy from March to July of 1983 has Naturelle de Paris, Laboratoire de Biologie des

yielded diverse molluscan faunas from the Oli- Invertebres Marins et Malacologie) and SMF

gocene (Italy) and the Neogene (Spain and (Senckenberg Naturmuseum, Frankfurt am Main,

Maghreb). This material includes an important col- Germany).

lection from the Pliocene of northern Tunisia. This

SYSTEMATIC particularly fossiliferous Pliocene, described by DESCRIPTIONS

Fekih (1975), complements our views of the Medi- Genus Amalda H. & A. Adams, 1853 terranean Pliocene gleaned mainly from Italian

Material from — Ancillaria tankervillii strata. Malaga (Spain) now available Type species Swainson, 1825.

shows the of in detail the — necessity studying greater Discussion The European Oligocene and Miocene

strata in other countries as well in order to be able Ancillinae have traditionally been assigned to Ancilla 28

location of localities in the Fig. 1. Map showing fossil referred to text.

Lamarck, 1799 (type species: Ancilla cinnamonea A number of species have also been assigned to

1810 buc- Lamarck, 1801). This classification was adopted by Ancillus de Montfort, (type species: Ancilla

who another be cinoides which is Davoli (1989), considered genus to Lamarck, 1805), occasionally con- valid, viz. Ancillarina Bellardi, 1882. However, the sidered to be a subgenus of Ancilla. In fact, as presence in the two principal species, A. obsoleta Kilburn (1977) has already noted, Ancillus is appar-

and A. related Amalda. The (Brocchi, 1814) glandiformis (Lamarck, 1811), ently most closely to genus of a divided basal fasciole, a deep siphonal sinus and Ancillus is the oldest known representative of the

callus indicates that should Ancillinae with acutulata a strong secondary these the species Eoancilla Step- in fact be referred to Amalda. This interpretation is henson, 1941 from the Maastrichtian (Late Cre- in with accordance the typical features of the genera taceous) of the United States (see Sohl, 1964, pi. 36,

and Ancilla worked Chavan Amalda as out by (1965) figs 1-7). and Kilburn (1977, 1981). Finally, only Ancillarina (type species: Ancilla cana- 29

lifera Lamarck, 1802) is clearly a representative of specimens are housed in MNHN collections, and

Ancilla of which it is the Amalda probably precursor this study has shown that the Late Oligocene

Ancillarina but few anomala from Wilhelmshohe differs from Late Oli- (Kilburn, 1977). comprises spe-

and have been in A. from the in cies appears to quasi-stasis during gocene glandiformis Aquitaine only

the Eocene to Late Miocene, when it became having a less well-developed spire callus. In fact,

extinct. these two species cannot be separated. In the Early

The which A. Ancillinae, are so poorly represented Miocene of Aquitaine, glandiformis frequently

in the and Miocene have shows the which Oligocene of Europe, spiral ridges on parietal region, are

apparently had the following distribution pattern: unknown in Late Oligocene specimens, but a few

- Six species of Ancillinae are known with certainty individuals (e.g. from Saucats, Moulin de Lagus)

from the Late show columellawith small folds. Amalda European Oligocene: a numerous

1 - Northern area: glandiformis from the Middle and Late Miocene dif-

Ancillus indivisus (Koch & Wiechmann, 1872) (PI. 2, fers in generally having only two high columellar

Figs 8, 9) folds and in lacking ridges on the parietal region.

Ancillus karsteni The above leads conclude that the (Beyrich, 1853) (PI. 2, Figs 10, 11). me to presence

Amalda obsoleta (Brocchi, 1814) (PI. 1, Figs 9-12) of numerous columellar folds is an ancestral feature

Amalda anomala which A. from the glandiformis (Schlotheim, 1820) (PI. 1, permits to separate glandiformis

Figs 5-8) Late Oligocene, in addition to the absence or pres-

2 - of the Mesogean area: ence ridges on parietal region. Following

Ancillus raulini(Peyrot, 1936) (PI. 2, Figs 5-7) Glibert (1960) I therefore considerAmalda anomala to

Ancillarina subcanalifera (d'Orbigny, 1852) (PI. 2, Figs be an Oligocene subspecies widely distributed in

12, 13) Europe.

Amalda abessensis n. sp. (PI. 1, Figs 1-4).

3 - Mesogean and Northern areas: From the European Miocene (when distinctions

Amalda glandiformis anomala (Schlotheim, 1820) between the Northern and Mesogean areas tend to

blur) I have been able to recognise five species, viz.

Remark — Ancillus raulini differs principally from Amaldaglandiformis (Lamarck, 1811) (PI. 1, Figs 5-8)

Amalda glandiformis in the weakness of the ancillid Amalda obsoleta (Brocchi, 1814) (PI. 1, Figs 9-12)

band and the absence of secondary spire callus. A. Ancillarina subcanalifera (d'Orbigny, 1852) (PI. 2, Figs glandiformis is also a much larger-sized species. Sau- 12, 13)

bade al. did Ancillus raulini Ancillus sismondanus in et (1978) not accept (d'Orbigny Bellardi, 1882) (PI.

(Peyrot) and united this taxon with a population of 2, Figs 2-4)

Amaldaglandiformis comprising specimens of reduced Ancillina pusilla (Fuchs in Karrer, 1877) (PI. 3, Figs 8,

size. 11).

According to Janssen (1979) Amalda anomala from

the North Sea Basin Oligocene differs from Remark — Davoli (1989) recorded from the Euro-

Miocene A. in Miocene two additional of glandiformis having a poorly developed pean species Ancillinae,

columellawith small folds Amalda in spire callus, a numerous viz. patula (Doberlein Bellardi, 1882) and

and in the side. Amalda lacking spiral ridges on parietal I sowerbyi (Michelotti, 1847). Judging from

examined of A. from Davoli's the first be but forma of specimens glandiformis a illustrations, may a

number of Miocene localities Miocene A. whereas the second be (Early [Bur- glandiformis, appears to

digalian] of the Aquitaine [Saucats, Meilhan, Mim- conspecific with A. obsoleta.

Gracilancilla baste], Middle Miocene of the Aquitaine [Sau- Thiele, 1925 was erected for Ancilla

brigues, St-Jean-de-Marsacq], Middle Miocene of sumatrana Thiele, 1925 of the western Indo-Pacific;

the Loire Basin [Le Louroux], Middle Miocene of it is a junior synonym of Ancillina Bellardi, 1882

Austria [Soos, Steinabrunn], Middle Miocene of (type species: A. pusilla Fuchs, 1877). The differences

and Late Miocene of and A. Italy [Turin hills] Italy between A. pusilla sumatrana are less impor-

In I of A. and the I have been able [Stazzano], addition, studied specimens tant are at species level. to

from several Late in convince of glandiformis Oligocene localities myself this by an examination of speci-

the Aquitaine (St-Paul-les-Dax [Abesse], St- mens of A. pusilla (SMF collections) and of A. su-

Etienne-d'Orthe, Peyrehorade [Peyrere] and from matrana (MNHN, Bouchet & Triclot Collection,

the Late Oligocene of Germany (Wilhelmshohe). All Musorstom II, station 106, 13°47 N - 120°30 E, 30

640-668 m; PI. 3, Figs 9, 10). So far as differences in preservation: from the Early Miocene of Aquitaine

the the protoconchs are concerned, in A. sumatrana it is (St-Paul-les-Dax [Cabanes]), from Middle

much more acuminate, of brownish colour and Miocene of Aquitaine (Saubrigues, St-Jean-de-Mar-

clearly separated from the teleoconch. Chavan sacq), from the Middle Miocene of Belgium

(1965) suggested that Gracilancilla be classed as a (Edegem), from the Middle Miocene of Germany

subgenus of Ancillina. (Dingden), from the Middle Miocene of Austria

(Soos, Voslau), from the Late Miocene of the Loire

Amalda abessensis Basin and from the Late (Alocospira) n. sp. (St-Clement-de-la-Place)

PI. 1, Figs 1-4 Miocene of Italy (Montegibbio, Tortonese and

Santa Agata). In a number of specimens I noted

Stratum — Late faluns bleu typicum Oligocene, a only very shallow and inconspicuous spiral grooves,

Miogypsinoides. not comparable with the spiral ornament seen in A.

Locus — Chateau olivid in typicus d'Abesse, St-Paul-lès-Dax abessensis. There is no other species of the

(Landes, France). European Cainozoic that is likely to be confused

Derivatio nominis — After the with the type locality. present new taxon. — Types Holotype (MNHN), seven paratypes (four Amalda abessensis is referred to the subgenus

specimens, MNHN, Lozouet et al. Collection; three Alocospira Cossmann, 1899 (type species: Amalda

specimens, Lesport Collection). Additional material papillata Tate, 1885 from the Early Miocene of Aus- from the nine Extra- same locality comprises fragmentary tralia, see Ninomiya, 1990, pi. 2, fig. 8).

shells (MNHN, Lozouet et al. Collection). Australian records of this subgenus include the

— The 12.2 Dimensions holotype measures mm Early Miocene of New Zealand (A. hebera Hutton,

and 5 (height) mm (maximum diameter). 1873), and the Late Miocene ofJava (A. nitida Wan-

— The medium At Description shell is of size, ovate, ner & Hahn, 1935). present, Alocospira contains with thick test. The number of whorls cannot be fewer than ten species distributed in the Pacific, determined. The is Australian but cyrtoconoid spire relatively mainly on coasts (Ninomiya, 1990)

covered in callus which masks the also south in the short, a strong present ofJapan (Ninomiya, 1988),

The takes the form of suture completely. apex a south China Sea and near New Caledonia (Kilburn mamillate button. The callus extends over part of & Bouchet, 1990). Amalda abessensis is closely related the last whorl and is continuous with A. but is nevertheless slender and the callus over to papillata, more the of the that of the is much parietal part aperture; spire possesses a more clearly cyrtoconoid spire. peculiar in being sectioned by 7-8 effacing spiral Michaux (1991) elevated the taxon Alocospira to lirae. The last whorl than the rank of and the represents more two genus considered spiral grooves thirds of the total shell The be feature'in it. height. aperture (dam- to an important species referred to aged in the holotype) would have been lanceolate, He also recognised three subgenera of Amalda, viz.

in its narrower adapical part and forming a large Gracilispira, Baryspira and Spinaspira. The other sub-

notch. The columella is of siphonal convex clearly genera Amalda, especially those recently erected twisted the columella which and at pillar, on some ten by Ninomiya (1988, 1990) are poorly defined

folds be counted. At its base, a included in Michaux's oblique may fairly were not paper. Michaux large band (the ancillid band) envelops the neck, remarked that A. montrouzieri of Bouchet & Kilburn

delimited and being anteriorly posteriorly by two (1988) from New Caledonia was probably a mixed

The neck is covered in callus grooves. entirely a lot, their figures 1-6 representing Alocospira mon-

of basal fasciolar which is consisting a large band, trouzieri (Souverbie, 1860) and figures 7, 8 appearing divided into and of the dorsal columella. Amalda of two zones, to represent an (Baryspira) the mucronata

Discussion — Amalda abessensis bears some resem- examination of Bouchet & group. However, an

A. in but blance to obsoleta general shape, differs in Kilburn's material from New Caledonia does not

the bulbous and the The - spire apex spiral groove. permit me to in fact distinguish two species and a

ofA. abessensis before well fortiori - is specimens me are not very two genera. Michaux's (1991) statement preserved and the spiral ornament may have been confusing and in need of clarification, which is why accentuated erosion/abrasion. For this I I here the Amalda for the by reason, prefer to accept only genus examined specimens of A. obsoleta from various time being. localities (MNHN collections) with various states of 31

Amalda cf. obsoleta Olivella (Brocchi, 1814) clanzigi n. sp.

PI. 2, Fig. 1 PI. 3, Figs 1-4

and — Pliocene Stratum — Falun Age locality Early (F. Serrano, pers. typicum with Potamides (Early

comm.), Colonia Santa Ines (Province of Malaga, Pliocene).

— Spain). Locus typicus Oued el Galaa, southeast of Lac dc — Discussion Amalda obsoleta is known with certainty Bizerte, Tunisia.

from from — Europe the Early Miocene onwards (Bur- Derivatio nominis Dedicated to Mr Sylvain

digalian of St-Paul-les-Dax, Moulin de Cabanes). Clanzig.

Rossi-Ronchetti indicated that Broc- — (1955, p. 238) Types Holotype (MNHN), a single paratype chi's type specimen originates from the Piemont (MNHN, Clanzig and Lozouet Collection).

— Pliocene. Davoli contested this Dimensions The 14 (1989) provenance holotype measures mm

and considered it have to come from the Italian (height) and 6 mm (maximum diameter).

Late Miocene. This species differs from Amalda glan- Description — Shell of moderate size, with 3 Vz

with which it has often been in elon- diformis, confused, cylindroconical whorls, thick test, aperture much having a larger basal band (ancillid band). gate, wide, conical spire short. Only 2 '/a whorls of

Glibert (1952, calculated ratios of the the these p. 353) (size spire are preserved; are separated by a

band total shell for vs. height) populations of A. deep suture. The last whorl is ovate and occupies

obsoleta and A. which make the three glandiformis, differ- quarters of the total shell height. Its elongate

between ences these clear far is but in its species appear cut as aperture wide, narrower adapical part this is as feature concerned. Glibert noted for A. where it terminates in a wide anal notch extended

obsoleta value of 6.4 of variation 5.64 the The of a mean (range by suture. abapical part the aperture is

to and for A. forma widerand notch in 6.57) glandiformis elongata displays a large siphonal a highly

(Deshayes, a meanof 4.65 from 4.35 to U-form. The columella is 1832) (range asymmetrical very

The two from the Pliocene is near-rectilinear 4.95). specimens Spanish oblique, and terminates in a

before with values of6.4 and 6.66 should thus twisted columella me be pillar, covered in a callus forming

referred A. obsoleta. to a fairly large bulge adapically and showing some ten

Saubade al. did consider the size of et (1978) not subparallel ridges on the columella pillar; these

' ’ the basal band in their biometric of Ancilla study ridges are slightly more pronounced in the abapical

from the Miocene of The result of where into Early Aquitaine. part they converge a strong basal limbus.

this was that certain of A. The is populations glandiformis lip slightly convex, with bevelled margin, and

and A. obsoleta were mixed In this A. obsoleta is smooth up. way, interiorly. Protoconch unknown.

from the ofSt-Paul-les-Dax Remarks — The last whorl Burdigalian (Moulin de of the paratype specimen

Cabanes, see PI. 1, 11, was to be is better and Figs 12) interpreted preserved permits to enlarge upon the

an elongate form of A. glandiformis. typical features of this species. The callus of the

Chavan used the Amalda obsoleta (1951) name var. columella forms a fairly strong bulge in the parietal

marcaisi Chavan for much form The columella shows a more elongate part. pillar some ten oblique

from the Pliocene of Morocco el folds and terminates in limbus (Oued Arjat, near a strong accentuated

Amalda marinhensis from Rabat). (Cox, 1941) the by a groove, below which extend two folds. The

Pliocene of has similar that callus of the base Portugal a profile to of extends over c 'A of the total

the Malaga specimens. However, the Portuguese height of the last whorl. This callus subdivides into

broad ancillid specimen possesses a band, which two fasciolar bands of equal magnitude. The first

makes it attributable to the A. fasciolar glandiformis group. (posterior band) differs by a narrower There thus additional are two species of Amalda in inductura. The second (anterior fasciolar band)

the Pliocene of the western of the Mediterra- attaches to the notch part siphonal and growth lines may

nean area. One of these be A. be It is delimited small may assigned to the seen. adapically by a bourre- obsoleta lineage, while the other is referred to the A. let and abapically by the protuberance of the col- glandiformis lineage. umella pillar.

Discussion — Olivella grateloupi (d'Orbigny, 1852) (PI. Genus Olivella Swainson, 1831 3, Fig. 7), the sole species of Olivella known to date Oliva Type species dama (Mawe, 1828). from the European Cainozoic (Late Oligocene- 32

A. which Early Miocene), is smaller and possesses but 5 to 6 (Alocospira) papillata group, at present

folds which in Australian waters. Its in columellar are, however, more pro- occurs mainly presence

nounced. This species is known from the Aquitaine European waters is fairly easily explained, as it

from to the closure of the Basin (Late Oligocene-Early Miocene) and appears prior principal Tethys

Italy (Early Miocene of the Turin hills). which, judging from the faunas, occurred at the

also be with O. of the Miocene. different Olivella clanzigi may compared start Early Quite a story

the which in pulchella (Duclos, 1836) (PI. 3, Figs 5, 6) from holds for Ancillina pusilla, occurs commonly

Recent West African of which it be the the Middle Miocene of This or a coast, may Europe. species,

The differs from related is known the Mid- precursor. new species specimens closely form, not prior to of O. pulchella from Senegal and Angola (MNHN, dle Miocene and is close to the Recent A. sumatrana

in from leg. Pin, leg. Gofas) having an anterior fasciole the west Indo-Pacific. Consequently, it with much wider inductura and columellatermi- be another that was intro- a appears to yet species

nating in a much stronger fold. duced from the west Indo-Pacific at the Early/Mid-

dle Miocene boundary or during the Middle CONCLUSIONS Miocene. This is in of the that there support view

Davoli (1989) emphasised the absence of Olividae have in fact been intermittent communications

from the Mediterranean Pliocene which he thought between the Paratethys and the Indian Ocean at

The time. was attributable to climatic factors. present-day that

Olividae ofthe North Atlantic are restricted to trop- ACKNOWLEDGEMENTS and held have ical subtropical regions. They are to

I wish to thank Mr C. Salas and Mr S. Gofas for disappeared from the Mediterranean at the end of Pliocene material from Malaga, Dr R. the Miocene, having fallen victim to the Messinian supplying for the loan of material from the Sencken- salinity crisis. The climatic conditions prevailing in Janssen Naturmuseum collections, Mr D. the Mediterranean the Pliocene did not berg Gourgues during per- and Mr for the loan of material from mit their re-introduction. J.F. Lesport

the of the Adour Basin and Mr F. Despite this, the Pliocene fauna of the Mediterra- Upper Oligocene

Serrano for data. Messrs S. nean comprises numerous tropical elements biostratigraphical

faunas: Gofas, P. Maestrati, Drs M.C. Cadee and R.Jans- amongst gastropod Ficidae, Cancellariidae, read the for which I Mitridae, Conidae and Terebridae (Marasti & sen critically manuscript, am grateful. Raffi, 1979). In addition, Olividae have been

recorded from the Portuguese Pliocene (Cox, 1941) REFERENCES and from Morocco (Tetouan) (Gentil & Boistel, Chavan, A., 1951. Deux interessants Gasteropodes du Pliocene This that the climatic conditions 1905). means marocain. Notes et Memoires du Service geologique du

does not suffice to the absence of hypothesis explain Maroc, 83: 135-139, 2 figs.

Olividae from the area; the discovery of the new Chavan, A., 1965. Essai de reclassification des Olividae Ancil-

linae — Bulletin de la Societe de species described herein is in fact not surprising. (Gastropodes). geologique France, 102-109, 1 fig., 1 pi. in the Pliocene of the Mediter- (7)7: Appear as they may Cox, L.R., 1941. New species of Gastropoda from the Pliocene Olividae remain ranean area, rare. of Portugal. Publica<;oes do Museu Mineralogico e Geo- Amalda abessensis of the Late of n. sp. Oligocene Noticias, logico da Universidade de Coimbra, Memorias e

Aquitaine is the sole European representative of the 10: 7-12, 1 pi.

PLATE 1

1-4. Amalda abessensis Late Château d’Abesse Figs (Alocospira) n. sp., Oligocene, St-Paul-lès-Dax, (Landes, France). 1, 2 - holotype, height

4 & 12.2 - 14 Lozouet Maestrati Coll. mm; 3, paratype, height mm. (MNHN).

Figs 5-8. Amalda glandiformis anomala (Schlotheim, 1820). Late Oligocene. 5-7 from St-Paul-lès-Dax, ‘Estoti’ (Landes, France), height 32

mm (5, 6), height 36 mm (7); 8 from Peyrehorade, ‘Peyrère’ (Landes, France), height 15 mm. Lozouet & Maestrati Coll.

(MNHN).

Figs 9-12. Amalda obsoleta (Brocchi, 1814). Early and Middle Miocene. 9, 10 from Saubrigues (Landes), Middle Miocene (Langhian), height

21 mm, Staadt Coll. (MNHN); 11, 12 from St-Paul-lès-Dax, ‘Cabanes’ (Landes), Early Miocene (Burdigalian), height 29 mm,

Lozouet, Maestrati & Senut Coll. (MNHN).

in Specimens Figs 1-10 were coated with ammonium chloride prior to photography. 33\1

PLATE 1 34

Davoli, F., 1989. Olividae (Gastropoda) miocenici: ultima testi- molluscs. State of the problem. In: Vllth International Con-

Mediterranean Athens Parma monziana nell'area mediterranea di un clima intertropicale. gress on Neogene, 1979, (Insti-

— Bollettino della Societa Paleontologica Italiana, 28(1): tute of Geology and Paleontology, University of Parma), 8

2 2 6 101-132, figs, tabs, pis. pp.

marin Nord de la Michaux, B., 1991. The evolution of the Ancillinae with special Fekih, M., 1975. Paleoecologie du Pliocene au — Annales des Mines de la 27: 193 reference New Zealand and Recent of Tunisie. et Geologie, pp., to Tertiary species

8 figs, 14 tabs, pis 1-4 + 42 pis. Amalda H. & A. Adams (Gastropoda: Olividae: Ancillinae). - Gentil, L., & A. Boistel, 1905. Sur l'existence d'un remarquable Venus, 50(2): 130-149, 9 figs, 2 tabs.

— five ofthe gisement pliocene a Tetouan (Maroc). Comptes rendus de Ninomiya, T., 1988. A new subgenus and new species

1'Academie des Sciences, Paris, 140: 1725-1727. Ancillinae (Gastropoda: Olividae) from southwestern Austra-

1952. du Miocene du Bassin de lia, and Taiwan. — Venus, 1 1 Glibert, M., Gastropodes moyen Japan 47(3): 141-153, fig., pi.

— ofthe la Loire. Deuxieme partie. Memoires de l'Institut royal des Ninomiya, T., 1990. A new subgenus and five new species

Sciences naturelles de Belgique, (2)46: 243-449, 15 pis. Ancillinae (Gastropoda: Olividae) from Australia. — Venus,

Glibert, M., 1960. Les Volutacea fossiles du Cenozoique 49(2): 69-82, 8 figs, 2 pis.

des collections de l'lnstitut des Sciences 1955. I della "Conchiologia fossile etranger royal Rossi-Ronchetti, C., tipi

naturelles de Belgique. — Memoires de l'lnstitut royal des subapennina" di Brocchi. Parte II: Gastropoda, Scafopodi. — Sciences naturelles de Belgique, (2)61: 1-109. Rivista Italiana di Paleontologia e Stratigrafia, Memoria,

Janssen, R., 1979. Die Mollusken des Oberoligozans (Chattium) 5: 91-343, figs 38-185.

im Nordseebecken. 2. Neogastropoda, Euthyneura, Ceph- Saubade, A.-M., L. Rousselle & S. Chavanon, 1978. Essai

alopoda. Archiv fiir Molluskenkunde, 109(4-6): 277-376, d'interpretation d'Ancilla glandiformis (Lk) (Gasteropode) et

obsoleta A. raulini Cossm. pis 15-18, 18a. des formes voisines (A. (Broc.) et et — ofAmalda and dans le Tertiaire Bulletin de l'lnstitut Kilburn, R.N., 1977. Descriptions ofnew species Peyr.) aquitain.

Chilotygma (Gastropoda: Olividae: Ancillinae) with a note geologique du Bassin d'Aquitaine, 24: 85-105, 14 figs, 1 pi. — the of Ancillus and Ancillista. 1964. and Basom- on systematics Amalda, Sohl, N.F., Neogastropoda, Opisthobranchia

Annals of the Natal Museum, 23(1): 13-21, 7 figs. matophora from the Ripley, Owl Creek, and Prairie Bluff

1981. Revision of the Ancilla Lamarck, Formations Cretaceous in Tennessee and Kilburn, R.N., genus [Late gastropods

— — States Professio- 1799 (Mollusca: Olividae: Ancillinae). Annals ofthe Natal Mississippi]. United Geological Survey,

Museum, 24(2): 349-463, 258 figs. nal Paper, 331-B: iv + 153-344, figs 12-18, 2 tabs, pis 19-52.

& P. 1988. The Amalda in New Kilburn, R.N., Bouchet, genus

Caledonia (Mollusca, Gastropoda, Olividae, Ancillinae). —

Bulletin du Museum national d'Histoire naturelle de Paris,

(4)10(A2): 277-300, 58 figs.

Marasti, R., S. RafTi, 1979. Observations on the & paleoclimatic Manuscript received 6 November 1991; revised version accepted

and biogeographic meaning of the Mediterranean Pliocene 3 May 1992.

PLATE 2

1. obsoleta MNHN A. Fig. Amalda cf. (Brocchi, 1814). Early Pliocene. Colonia Santa Ines (Malaga, Spain), height 12 mm, (leg.

Barrajon).

Figs 2-4. Ancillus sismondanus (d’Orbigny in Bellardi, 1882). Middle Miocene (Langhian), Turin hills (Italy), height 18.4 mm (2), height 20

mm (3, 4), Staadt Coll. (MNHN).

Figs 5-7. Ancillus raulini (Peyrot, 1936). Late Oligocene, St-Etienne-d’Orthe (Landes, France), height 16 mm (5, 6), height 15 mm (7),

Lozouet & Maestrati Coll. (MNHN).

Figs 8, 9. Ancillus indivisus (Koch & Wiechmann, 1872). Late Oligocene (Chattian), Rumeln (Germany), height 12 mm. Gorges Coll.

(SMF no. 250936).

11. karsteni (Germany), SMF Coll. 251543a). Figs 10, Ancillus (Beyrich, 1853). Late Oligocene (Chattian), Rumeln height 9.8 mm, (no.

& Figs 12, 13. Ancillarina subcanalifera (d’Orbigny, 1852). Late Oligocene, St-Etienne-d’Orthe (Landes, France), height 25 mm, Lozouet

Maestrati Coll. (MNHN).

All specimens were coated with ammonium chloride prior to photography. 35\2

PLATE 2 36

PLATE 3

1-4. Olivella southeast of Lac de Oued et Galaa 14 Figs clanzigi n. sp. Early Pliocene, Bizerte, (Tunisia), 1, 4 - holotype, height mm; 2,

3 - paratype, height 15.6 mm, Clanzig & Lozouet Coll. (MNHN).

MNHN Figs 5, 6. Olivella pulchella (Duclos, 1836). Joal, at depth 10-20 m (Senegal), height 16 mm, (leg. Pin).

Fig. 7. Olivella grateloupi (d’Orbigny, 1852). Early Miocene (Burdigalian), St-Paul-lès-Dax, ‘Cabanes’ (Landes, France), height 11

mm, Lozouet & Maestrati Coll. (MNHN).

11. 6 SMF Figs 8, Ancillina pusilla (Fuchs in Karrer, 1877). Middle Miocene, Kostej (Rumania), height mm (8), height 6.5 mm (11),

(Boettger Coll.).

Figs 9, 10. Ancillina sumatrana (Thiele, 1925). Campagne Musorstom 3, station 106, 13°47 N - 120°"30 E (at depth 640-668 m), Philip-

MNHN pine Islands, height 5.6 mm, (Bouchet & Triclot Coll.).

Specimens in Figs 1-4 and 8 were coated with ammonium chloride prior to photography. 37\3

PLATE 3 38

Maria Tembrock's scientific work, produced under the List of Dr M.L. Tembrock's papers

list includes those that have been above-mentioned circumstances, is impressive and has [This only papers actually publis- hed in scientific journals] justly found internationalacclaim. The main objectives

of her studies were the gastropods of the Cainozoic Tembrock, M.L., 1958. Einige Zahlenverhaltnisse bei den haufig-

North Sea Basin. The basic that she consis- des Freienwalde und concepts sten Molluskenarten Rupeltones von Joach-

3: tently adhered to are the following: imstal. Ber. geol. Gesellsch. DDR, 136-138.

- Determination Tembrock, M.L., 1960. Zum Problem der von the gastropod shell is to be considered an entirety Gastropodenschalen. Ber. geol.Gesellsch. DDR, 5: 365-381, and yields a lot of information on ontogenetic and 2 pis.

phylogenetic developments. Molluskenfauna der Tembrock, M.L., 1962. Bemerkungen zur — of taxonomic are the especial importance embryonic Bohrungen GolBen 2, Drehna 5 (Lausitz) und des Fundpunktes

whorls. These were relatively conservative during evo- Sollingen (Braunschweig). — Geologie, 11:118-123, 1 tab.

Muriciden Mittel- und Tembrock, M.L., 1963. aus dem Oberoligo- lution and thus often preserve features which may indi- Palaont. zan und den Vierlandschichten des Nordseebeckens. cate relationships. Abhandl., (A) 1(4): 299-351, 3 tabs, 10 pis. — the of and of in development genera groups genera a Tembrock, M.L., 1964a. Taxionomie des Formcnkreises "Fusus becomes conceivable and regional setting tangible by — multisulcatus" Nyst, 1843 (Gastropoda). Ber. geol. Gesellsch. in time of faunas and proper arrangement geological DDR, 9: 304-310, 3 pis.

1964b. von Faziesabhangigkeit detailed studies of features in relation to chronostrati- Tembrock, M.L., Einige Beispiele — bei tertiaren Gastropoden. Ber. geol. Gesellsch. DDR, 9: Maria Tembrock's muri- graphical position. papers on 311-337, 6 pis. cids and the from the North Sea on genus Scalaspira Tembrock, M.L., 1965a. Erlauterungen zur Gattung Streptochctus Basin this well. exemplify very Cossmann (Gastropoda). — Senckenb. leth. (Weiler-Festschrift), in addition in to morphological studies, keeping 46a: 427-439, 4 figs, 1 pi.

mind the in ofconditions Tembrock, M.L., 1965b. Zur Systematik problematischer development time, an analysis einiger

Caeciden-Gattungen (Gastropoda). — Abhandl. Zentr. Geol. of life should be considered, if one is to avoid serious Inst. Berlin, 1: 81-93, 1 pi. conditions misinterpretations. Ecological may cause Tembrock, M.L., 1965c. Zum Artproblem bei Gastropoden, erlau- far-reaching but reversible changes, oflimi- phenotypic tert an den Scalaspira (al. "Aquilofusus")-Arten des norddeut-

ted taxonomic and phylogenetic value. At the other schen Oligozans und Miozans. - Ber. geol. Gesellsch. DDR, 10:

shell be sufficient 429-438, 1 pi. extreme, morphology may not to 1967. Book review Miozan-Mediterranen forms in different environments Tembrock, M.L., [Die distinguish living very Gastropoden Ungarns, by L. Strausz]. Ber. deutsch. without overlap, so that these should be considered to Gesellsch. geol. Wiss., A. Geol. Palaont., 12: 741-743. species. represent separate Tembrock, M.L., 1968. Taxionomisch-stratigraphische Studie zur

Maria Tembrock started her in — When career the Scalaspira-Gruppc (Gastropoda, Tertiar). Palaont. Abhandl.,

these of (A)3(2): 193-366, 2 figs, 2 tabs, 18 pis. 1960s, principles were hardly matter course Tembrock, M.L., 1982. Eine Sammlung obereozaner, oligozaner but became and amongst malacologists, accepted more

— und miozaner Mollusken aus der La-Tene-Zeit. Z. geol. this Maria Tembrock's work more. In respect, was Wiss., 10: 1387-1391, 1 pi. Several extensive the pioneering. papers (e.g. one on Tembrock, M.L., 1989. Neue Spiratella-Arten (Gastropoda, Opis-

Dentaliidae from the North Sea had been Z. 35: 1 Basin) com- thobranchier, Pteropoda). angew. Geol., 242-244, pi.

for but Strauch, F., & M.L. Tembrock, 1978. Marine Molluskenreste. In: K. pleted over twenty years, were not published Brunnacker (ed.). Geowissenschaftliche Untersuchungen in Gon- because of a misplaced state security policy. As soon as nersdorf: 231-233. Wiesbaden. these restrictions dropped, Maria Tembrock updated

she could these papers, but unfortunately not complete

these studies.

Maria Tembrock her was always prepared to pass on

others. author of the own experience to The present

obituary notice will always remember with gratitude

the discussions he had with her scientific many on as

unselfish well as personal matters, and her helpfulness.

I)r Joachim Griindel

Franz-Dahlem-StraBe 14

D(O)-1055 Berlin, Germany