On the of the genus Rosacris 43

Entomologie heute 28 (2016): 43-52

On the Taxonomy of the Genus Rosacris Bolívar, 1931 (: )

Zur Taxonomie der Gattung Rosacris Bolívar, 1931 (Orthoptera: Tetrigidae)

JOSIP SKEJO

Summary: The genus Rosacris Bolívar, 1931 is a pygmy (Orthoptera: Tetrigidae) ge- nus hitherto known only from a holotype male of its type species Rosacris antennata Bolívar, 1931 from Makiling Mt. (Luzon island, the Philippines) and originally assigned to the Discotettiginae. Redescription of the genus and its type species are based on the holotype and a new photographic record of the species (Isarog Mt.) found in Flickr (www.fl ickr.com). The holotype is found to be male nymph rather than an adult . The species is morphologically similar to members of the genus Metamazarredia Günther, 1939 from the Philippines. Therefore the genus is transferred from the subfamily Discotettiginae to Metrodorinae. Assumption exists that the specimens hitherto identifi ed as R. antennata represent nymphs of Metamazarredia fuscipes (Stål, 1877).

Key words: Discotettiginae, Metrodorinae, the Philippines, Flickr, nature macro-photography

Zusammenfassung: Die Gattung Rosacris Bolívar, 1931, ein Zwerg-Grashüpfer (Orthoptera: Tetrigidae), ist bislang nur durch den Gattungsholotypen Rosacris antennata Bolívar, 1931 vom Mount Makiling (Insel Luzon, Philippinen) bekannt. Sie wurde ursprünglich der Unterfamilie Discotetti- ginae zugeordnet. Die Wiederbeschreibung des Gattungstypus‘ basiert auf dem Holotypus und einem neuen Foto der Art vom Mount Isarog auf dem Fotoportal Flickr (www.fl ickr.com). Der Holotypus scheint eher eine männliche Nymphe als ein adultes Tier zu sein. Die Art ist morpho- logisch den Vertretern der Gattung Metamazarredia Günther, 1939, ebenfalls von den Philippinen, ähnlich. Deshalb wurde die Gattung aus der Unterfamilie Discotettiginae in die der Metrodorinae überführt. Es besteht die Vermutung, dass die bisherigen Exemplare von R. antennata Nymphen von Metamazarredia fuscipes (Stål, 1877) sind.

Schlüsselwörter: Discotettiginae, Metrodorinae, die Philippinen, Flickr, Natur-Makrofotografi e

1. Introduction Discotettigiae in BOLÍVAR 1931). The original description, which includes a drawing of The genus Rosacris was established by I. the species’ habitus and brief comments BOLÍVAR (1931) for a single species named on its relationship with Hirrius Bolívar, 1887 R. antennata Bolívar, 1931, based on a sin- (another member of the Discotettiginae), gle male specimen from Mt. Makiling, a was never cited again during later revisions dormant volcano in the Laguna province of the family Tetrigidae. Even GÜNTHER (Luzon Is., the Philippines) with an elevation (1938), who reviewed the Discotettiginae up to 1,090 m a.s.l. Based on the antennal seven years later, overlooked the publication. morphology, i.e. the presence of fl attened OTTE (1997), in the fi rst printed version antennal segments, the species was originally of the Orthoptera Species File, listed this placed in the subfamily Discotettiginae (= species as of uncertain placement within

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Tetrigidae. The type specimen was listed in other Caeliferan families by (I) the long the catalogue of Bolívar’s collection (PARÍS pronotum, usually covering the abdomen, 1994). Two additional records of the genus (II) tarsal formula 2-2-3, (III) lack of an were recently found (1) on April 15th 2015 arolium between the claws and (IV) presence on Flickr (photos by P. BERTNER, dorsal and of a sternomentum (modifi ed fi rst sternum). lateral habitus of a nymph, from Isarog Mt.) Composition: Nine subfamilies (Batrachide- and (2) on April 14th 2016 on eBay (female inae, Cladonotinae, Cleostratinae, Discotet- specimen for sale, from Luzon, without tiginae, Lophotettiginae, Metrodorinae, specifi ed locality). , Tetriginae, Tripetalocerinae) The aims of this article are (1) to present a and a few tribes (Clinophaestini, Coptotet- new record of Rosacris antennata from Mt. tigini, Criotettigini, Dinotettigini, Miriatrini, Isarog on Luzon Is. (the Philippines), (2) Xerophyllini) with 270-290 genera and 1800- to redescribe the genus and its type species 2000 currently described species (EADES et and (3) to discuss the position of the genus al. 2016). within the current Tetrigidae taxonomy and Distribution: Globally except for the Ant- its relationship to other genera. arctic continent, the majority of the species (about 75%) occur in the tropical region, 2. Material and methods with the highest known biodiversity of the family in the Indomalayan and Wallacean For this study, the curator of the Orthoptera regions (EADES et al. 2016). collection in the Museo Nacional de Ciencias Since taxonomy follows Orthoptera Species Naturales (MNCN, Madrid, Spain) MERCEDES File (EADES et al. 2016) where the genus is PARÍS sent me the photos of the lectotype denoted as of uncertain placement within (= holotype) of Rosacris antennata Bolívar Tetrigidae and since in the original descrip- and the original description (Bolívar 1931) tion the genus was assigned to the subfamily of the genus and the species. Morphological Discotettiginae, here I present diagnosis and terminology and measurements follow TUM- composition of Discotettiginae, as well as BRINCK (2014) and SKEJO & CABALLERO (2016). that of Metrodorinae, the subfamily I assign Measurements of the holotype were obtained the genus to after morphological compari- using the ImageJ software (ABRAMOFF et al. son with Metamazarredia spp. 2004) after cali bration with a scale added by the curator after processing the photographs. 3.2. Subfamily Discotettiginae Han- A photographic record was found (1) on cock, 1907 Flickr (a social network to share photos) [https://www.flickr.com/photos/rainfo- Diagnosis: The paranota not square shaped, rests/13064225934/in/photolist-kUrzdJ] antennae 11-14 segmented, with all or only a and later more photos were obtained directly few subapical segments widened. The only from the photographer, P. BERTNER, and (2) true diagnostic character of this subfamily on eBay (from user Philinsectbugs). is the morphology of antennae (see discus- sion). The group is polyphyletic. It is prob- 3. Results ably synonymous with true Scelimeninae based upon the morphology of the genus 3.1. Family Tetrigidae Rambur, 1838 (su- Discotettix Costa, 1864 (SKEJO et al. in press). perfamily Tetrigoidea Rambur, 1838) Composition and distribution. One tribe composed of a single genus (Discotettigini: Diagnosis: Family of short-horned grass- Discotettix) and fi ve more genera (excluding hoppers () easily separated from Rosacris) without tribal placement. Distri- On the taxonomy of the genus Rosacris 45 buted in SE Asia (ancient Sundaland and names), tropical Asia including Malesia, surroundings). Melanesia and Papuasia (42 genera, 163 spp), 1) Stål, 1877: Mindanao (the Phil- Australia (2 genera, 3 spp.). ippines, 2 spp.), 2) Discotettix Costa, 1864 (type genus of the 3.4. Genus Rosacris Bolívar, 1931 subfamily): peninsular Malaya (1 sp.), Suma- Diagnosis: Small body size (~ 10 mm); U- tra (2 spp.), Borneo (2 spp.), Mindanao (the shaped carinae of the vertex; 14-segmented Philippines, 1 sp.), antennae with widened subapical segments 3) Flatocerus Liang & Zheng, 1984: China (10 (10th-12th); pronotum (and median carina) spp., probably including a few synonymous strongly elevated in frontal 2/3 of its length; names) [the genus is likely synonymous with organs of fl ight not visible (= nanopronotal, Phaesticus (SKEJO & STOROZHENKO, unpub- apterous); tibiae of fore and mid legs robust lished data)], and rectangular in section. Very similar 4) Hirrius Bolívar, 1887: Mindanao (the Phi- to members of the genus Metamazarredia lippines, 2 spp.), Sulawesi (3 spp.), Günther, 1939, more specifi cally M. fuscipes 5) Kraengia Bolívar, 1909: Sulawesi (1 sp.), (Stål, 1877) (Fig. 3) the only difference being 6) Phaesticus Uvarov, 1940: China (3 spp.), lack of the organs of fl ight in R. antennata. India: Assam (1, not identifi ed sp.), Thailand Original etymology and vernacular name: (1 sp.), peninsular Malaya, Sumatra, Java Rosa from the family name of professor DA- (1 sp. in the whole region), Borneo (1 sp.). NIELLE ROSA, and (a)cris from ancient Greek akrís (άκρίς), meaning locust. Here I propose 3.3. Subfamily Metrodorinae Bolívar, an English common name for this genus 1887 and species – Rosa’s turtlehopper, because the morphology of the pronotum of the Diagnosis (after PAVÓN-GONZALO et al. species resembles a pygmy jumping turtle. 2012): Mainly characterized by having the Type species: Rosacris antennata Bolívar, 1931 median ocellus and the antenna placed by original monotypy. below the eyes, a relatively small divergence Composition: Monotypic – Rosacis antennata of the rami of the frontal costa not forming Bolívar. wide scutellum, and a similar length of the Distribution: Luzon Island (the Philippines): fi rst and third segments of the hind tarsus. Mt. Makiling [N14.129735, E121.199807] Many species of Metrodorinae also exhibit (type locality of R. antennata) and Mt. Isarog the posterior angles of the lateral lobes of [N13.658922, E123.372851 ] (locality of the the pronotum produced outwards (the main new record) – volcanic mountains. character used when the subfamily was esta- blished) often becoming acutely spinose. All 3.5. Species Rosacris antennata Bolívar, these characters together separate the sub- 1931 (Figs 1, 2) family from the other eight subfamilies of Tetrigidae, although none of them is enough Literature records: Rosacris antennata: BOLÍVAR to characterize Metrodorinae by itself. 1931, PARÍS 1994, YIN, SHI & YIN 1996 Composition and distribution: 85 genera and Locus typicus: the Philippines: Luzon Isl.: 528 species in all the continents excluding Makiling Mt. Europe and Antarctica. Africa including Material examined: Holotypus*: 1♂ (Figure Madagascar (22 genera, 85 spp.), N America 1) [nymph!] Monte Makiling, Luzón [the (1 sp. in Mexico), S America (15 genera, 78 Philippines] Baker [leg.] (fi rst label, print- spp.), temperate Asia (20 genera, 216 spp., ed); Orctacris* [?] gen. n. antennatus (second probably including numerous synonymous label, handwritten by I. Bolívar); Inventory

Entomologie heute 28 (2016) 46 JOSIP SKEJO

Fig. 1: Holotype (male) of Rosacris antennata from MNCN, Madrid, Spain: dorsal habitus, lateral habitus, head and labels. Photo: MERCEDES PARÍS. Abb. 1: Holotyp (Männchen) von Rosacris antennata aus MNCN, Madrid, Spanien: Dorsal- und Lateralansicht, Kopf und Beschriftungen. Foto: MERCEDES PARÍS. number Cat. Tipos No 250, MNCN_Ent 14.IV.2015. (eBay) seller: Philinsectbugs, 138792 (MNCN). det. J. SKEJO. Other material studied: 1♂ (Figure 2) [* PARÍS (1994) designated the only specimen (nymph) the Philippines: Luzon Isl.: Isarog of the species as lectotype. However, it is Mt. 3.III.2014. (Flickr) photo: P. BERTNER, clear from the original description that the det. J. SKEJO; 1♀ (nymph) Luzon Isl. [with- generic and the specifi c name were based on out specifi ed locality, photos of bad quality] a single male specimen from Mt. Makiling. On the taxonomy of the genus Rosacris 47

Fig. 2: Rosacris antennata specimen from Isarog Mt. Photo: PAUL BERTNER. Abb. 2: Exemplare von Rosacris antennata vom Mount Isarog. Foto: PAUL BERTNER.

Thus, the specimen PARÍS designated lecto- Distribution: the Philippines: Luzon Isl.: type is in fact the holotype (a single specimen Makiling Mt. and Isarog Mt. in the type series) and thus I correct the de- signation here and cite it as holotypus. PARÍS Holotype (male) re-description (Fig. 1) (1994) also stated that BOLÍVAR changed the generic name from the originally proposed General characters:. Small species (body (on the label: Orctacris) to Rosacris in the ori- length about 10 mm). Body smooth, fi nely ginal description, the name derived after the granulated with very small and smooth family name of Professor DANIELLE ROSA.] tubercles that are not placed on the carinae.

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Nanopronotal, apterous species. General basal segments, each distal one longer than body color brown. Antennae black with the preceding proximal, 8th central segment white-yellowish apical segments. Head about 8 time longer than wide, 8th-9th distal has the same color as the rest of the body. cylindrical segments, >5 times longer than Compound eyes black. Pronotum with pale wide, 10th-12th subapical segments, 10th and and dark colored stripes in the infrascapular 11th large and widened, 12th reduced in com- area. Fore and mid femora brown with weak parison to previous two, but folliaceous as pale bands, fore and mid tibiae paler than well, apical 13th-14th reduced in size. femora. Fore tibiae have distal third of dark Pronotum: Pronotum elevated in frontal two brown color. Hind femora brown with two thirds, strongly descending in the last third; pale colored stripes from the dorsal carina covering 4/5 of the abdomen, reaching towards the external area. Hind tibiae brown. about half the length of the hind femora. Proximal tarsal segments of fore and mid Anterior margin of pronotum truncated, legs dark, distal segments with proximal half slightly angular. Median carina continuous, yellowish, distal half dark. First and second running from the anterior margin to the segment of hind tarsi dark colored, third apex. Pronotal carinae present, well develop- pale colored, except a dark part in the apex ed. Extralateral carinae present, but weak. (before the claws). Interscapular area very wide at the level Head: Head exerted slightly above the level between coxae of mid and hind legs, then of the pronotum. Fastigium of vertex not narrowing towards the apex of pronotum, produced in front of the eyes in dorsal view, sinuate. Lateral area narrow in anterior part concave in frontal view. Anterior margin of and widened towards the apex. Humero- fastigium of vertex slightly convex. Fossulae apical carinae weak and straight. Lateral area present and deep. Median carina of vertex wide. Pronotal apex blunt. A single paranotal present in the distal half of the vertex lobe present, directed downwards, slightly length. Lateral carinae of vertex present, outwards. Apex truncated. Pronotal sulci elevated, U-shaped. Supraocular lobes pres- strong and deep. Lateral pronotal projection ent. Vertex narrower than compound eye. directed ventrally, slightly laterally, with trun- Median ocellus situated in the distal margin cated apex, ventral sinus present, tegminal of scutellum, between facial carinae. Paired sinus absent. ocelli situated between the compound eyes, Wings: Fore and hind wings absent. slightly below the middle of the compound Legs: Dorsal margin of fore and mid femo- eye height. Frontal costa in lateral view ra straight. Ventral margin of fore femora visible, very prominent. Frontal costa bifur- straight, ventral margin of mid femora undu- cation between the compound eyes, slightly late. Fore femora almost circular in section. above double ocelli. Scutellum narrow, facial Fore and mid tibiae widened, rectangular in carinae almost parallel, slightly divergent. section, with sulcate outer margin. Distal Antennal grooves and scapus considerably tarsal segments of fore and mid legs con- wider than scutellum, fl agellum as wide as siderably longer than proximal one. Hind scutellum. Maxillar palpi fl attened, brown. femora slender (ratio length: maximum Eyes in dorsal view elliptic, in lateral view width 3.15). External median area with globular with truncated lower margin, in seven transverse ridges. Dorsal margin of frontal view irregularly globular. Antennal the hind femora concave in distal half. Geni- grooves situated at the level of the lower cular and antigenicular teeth low and sharp, margins of the compound eyes, large, situated on the elevated dorsal carina. Hind almost touching scutellum. Antenna with tibiae robust, with a few spines that are not 14 segments: 1st scapus, 2nd pedicell, 3rd-7th prominent. First and third tarsal segments On the taxonomy of the genus Rosacris 49 almost equal in length. Pulvilli angular, but that may contribute to a better camoufl age not acute spinose. of the animal within algae and mosses in its Measurements: Body length (from fastigium natural habitat. This symbiosis is not unusual to the end of the abdomen) 10.11 mm, in Tetrigidae (it was observed also in e.g. pronotum length 8.11 mm, pronotum width Discotettix spp. specimens from Sumatra, (between lateral pronotal projections) 2.59 Borneo and Mindanao). mm, pronotum height (from the lowest part of lateral projection to the highest part of 4. Discussion and conclusions median carina) 3.21 mm, fore femur length 2.65 mm, fore femur width 0.83 mm, mid According to the current definition of femur length 2.45 mm, mid femur width Tetrigidae subfamilies, the genus Rosacris is 0.69 mm, hind femur length 5.21 mm, hind a member of the Metrodorinae subfamily, femur width 1.69 mm. Vertex width 0.48 based on extreme morphological similarity mm, compound eye width 0.65 mm. to Metamazarredia spp. Rosacris antennata is quite different in morphology from repre- On the specimen from the Isarog Mt. sentatives of the subfamily Discotettiginae. (Fig. 2) Among Discotettiginae, the antennae pos- sess three slightly widened segments similar The male nymph photographed on the Isa- to R. antennata, which can be found in mem- rog Mt. (the Philippines: Luzon Isl.) shows bers of the genus Flatocerus, but are usually considerable similarity with the holotype of followed by a few more segments that are Rosacris antennata from MNCN and I refer to slightly widened. Also, the projected frontal it as to the same species as the holotype, de- costa and vertex in Flatocerus are similar to spite of possibility that it represents nymph Phaesticus, but strongly projected supraocular of Metamazarredia fuscipes (see discussion). lobes are not present in the latter genus. Fur- However, a few differences are to be noted. thermore, the pronotal morphology differs (I) The pronotum of the specimen from from Phaesticus and Flatocerus, but is almost Isarog is slightly more robust than that of the same as that of certain Metamazarredia the holotype. (II) The color of the Isarog species (genus within Metrodorinae). All the specimen is paler (light and with clear dark members of the subfamily Discotettiginae bands on fore and mid femora, as well as exhibit similar morphology of widened an- in infrascapular area) than in holotype. It tennal segments. However, similar pattern is important to note that nymphs of most of compressed antennal segments can be Orthoptera are paler than adults and that observed in members of at least 11 genera dried museum specimens may darken com- assigned to subfamilies other than Dis- pared to living specimen, so I think there cotettiginae: (I) Tripetalocera, Tripetalocerodes, is no taxonomic value in these differences. Clinophaestus and Birmana, (Tripetalocerinae: (III) The dorsal carina of the hind femora is SE Asia), (II) Ophiotettix (Metrodorinae: New more compressed and higher in Mt. Isarog Guinea), (III) Metamazarredia (Metrodorinae: specimen, which is a typical juvenile charac- Borneo and the Philippines), (IV) Andriana ter and based upon this feature it is possible and Hybotettix (Metrodorinae: Madagascar), to conclude that it is a male nymph and not (V) Hyperyboella (Metrodorinae: New Cale- an adult specimen. An interesting observa- donia), (VI) Chiriquia Morse, 1900 (Metro- tion that can be noted after examination of dorinae: N South America), (VII) Lophotettix the photographs are epizoic interactions (Lophotettiginae: S and C America). with algae (which are visible on the legs, The holotype of R. antennata is probably not pronotum, vertex, especially right fossula), an adult animal, but a last instar nymph, not

Entomologie heute 28 (2016) 50 JOSIP SKEJO

Fig. 3: Holotype (female) of Metamazarredia fuscipes from Naturhistoriska Riksmuseet, Stockholm, Sweden: dorsal habitus, lateral habitus, head and labels. Photo: JOSEF TUMBRINCK. Abb. 3: Holotyp (Weibchen) von Metamazarredia fuscipes aus dem Naturhistorischem Reichsmuseum, Stockholm, Schweden: Dorsal-, und Lateralansicht. Kopf und Beschriftungen. Foto: JOSEF TUMBRINCK. having genicular and antigenicular teeth, but teeth, but projections of the elevated dorsal possessing high dorsal carina of hind femora margin. running towards the knee and forming two There are many morphological characters teeth-like projections. The larval stages in shared between Metamazarredia spp. and R. Tetrigidae can be clearly recognized by the antennata. In conclusion, the only character missing incision between the antegenicular separating Rosacris and Metamazarredia is the teeth and the knee of the hind femur (TUM- absence of fl ight organs in Rosacris. BRINCK 2014). These teeth were drawn in the Taxonomic position of the genus Metamazar- original descriptive paper as fully incised redia was discussed within the comprehen- teeth of an adult animal, but after holotype sive revision of the Metrodorinae (GÜNTHER examination, I conclude that the drawing 1939). Morphology of Metamazarredia spp. is wrong and that these teeth are not true was discussed and the author did not assign On the taxonomy of the genus Rosacris 51 the genus to Discotettiginae, a conclusion students association – BIUS, Zagreb, Cro- with which I completely agree. GÜNTHER atia) for checking the manuscript and for (1939) made a comparison of the genus their comments, to PAUL BERTNER (Canada) with Mazarredia spp. The specimens I as- and JOSEF TUMBRINCK (NABU, Germany) signed to R. antennata could be nymphs of for photos of alive Rosacris antennata and Metamazzaredia fuscipes (Fig. 3). The species holotype of Metamazarredia fuscipes. Thanks is already known from Isarog Mt. (male to MICHAEL JARED THOMAS (Illinois Natural from Museum für Naturkunde, Berlin), so History Survey, Champaign, Illinois, USA) the specimen from Isarog here identifi ed as for comments and for improving the English R. antennata is likely to represent a nymph of the manuscript. Thanks go to reviewers, of M. fuscipes. For a fi nal conclusion on the whose comments improved the quality of relation of R. antennata and M. fuscipes, a the study. series of specimens from Makiling Mt. and other similar mountains are needed, as well Literature as research on the nymphal morphology of Metamazzaredia spp. ABRAMOFF, M.D., MAGALHAES, P.J., & RAM, S.J. R. antennata is one of the species that seem (2004): Image Processing with ImageJ. Bio- to have a mutualistic relationship with algae photonics International 11(7): 36-42. (and possibly also mosses), which are found BOLÍVAR, I. (1931): Nuevo acridino del grupo growing on the fi ne tubercula of the inte- Discotettigiae. Archivio Zoologico Italiano gument of the vertex, pronotum and legs. 15: 29-33. The species lives in the volcanic mountains EADES, D.C., OTTE, D., CIGLIANO, M.M., & Makiling Mt. (a dormant volcano) with an BRAUN, H. (2016): Orthoptera Species File. elevation up to 1090 m a.s.l. and Isarog Mt. Version 5.0/5.0 [01.05.2016.] http://ortho- ptera.speciesfi le.org (a potentially active stratovolcano) with an GÜNTHER, K. (1938): Revision der Acrydiinae, elevation up to 2000 m a.s.l. These moun- I. Sectiones Tripetalocerae, Discotettigiae, tains display a variety of habitats (warm Lophotettigiae, Cleostrateae, Bufonidae, grasslands, wet grasslands, lowland forests, Cladonotae, Scelimenae verae. Mitteilungen wet and cool mountainous forests) and are aus dem Zoologischen Museum in Berlin rich in biodiversity. There are many similar 23: 299-437. volcanic mountains on Luzon Isl. that have GÜNTHER, K. (1939): Revision der Acrydiinae not yet been investigated in terms of the (Orthoptera), III. Sectio Amorphopi (Me- Tetrigidae fauna. I assume that similar spe- trodorae Bol. 1887, aut.). Abhandlungen und Berichte aus den Staatlichen Museen für cimens will be found in many other localities Tierkunde und Völkerkunde in Dresden (Ser. after comprehensive systematic research and A: Zool.) (N.F.). 20(1):16-335. comparison with nymphs of Metamazarredia OTTE, D. (1997): Tetrigoidea and fuscipes, thus giving the fi nal conclusion on (Orthoptera: Caelifera) and Addenda to OSF identity of specimens hitherto identifi ed as Vols. 1-5. Orthoptera Species File 6: 1-261. R. antennata. PARÍS, M. (1994): Catálogo de tipos de ortop- teroides (Insecta) de Ignacio Bolivar, I: Acknowledgments Blattaria, Mantodea, Phasmoptera y Orthop- tera (, Rhaphidophoroidea, Tettigonioidea, , Tetrigoidea). Eos, I am very grateful to MERCEDES PARÍS Revista española de Entomología 69: 143-264. (MNCN, Madrid, Spain) for sending me PAVÓN-GONZALO, P., MANZANILLA, J., & GARCÍA- photos of the holotype and the original PARÍS, M. (2012): Taxonomy and morpho- description of the genus and the species; to logical characterization of Allotettix simoni DORA PAPKOVIĆ and FRAN REBRINA (Biology (Bolívar, 1890) and implications for the sys-

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tematics of Metrodorinae (Orthoptera: Te- descriptions of new genera and species from trigidae). Zoological Journal of the Linnean New Guinea and New Caledonia. Biodiversity, Society 164(1): 52-70. Biogeography and Nature Conservation in SKEJO, J., & CABALLERO, J.H.S. (2016): A hidden Wallacea and New Guinea 2: 345-396 (plates pygmy devil from the Philippines: Arulenus 64-91). miae sp. nov. – a new species serendipitously YIN, X.C., SHI, J.P., & YIN, Z. (1996): A synonymic discovered in an amateur Facebook post catalogue of and their allies of (Tetrigidae: Discotettiginae). Zootaxa 4067(3): the world (Orthoptera: Caelifera). China Fore- 383-393. stry Publishing House, Beijing, China 1266 pp. SKEJO, J., TUMBRINCK, J., & PUSKHAR, T.I. (in press): Taxononomic revision of the genus Bsc Josip Skejo, graduate student Discotettix Costa, 1864 and comments on University of Zagreb, Faculty of Science, the related genus Kraengia Bolívar, 1909 Department of Biology (Tetrigidae: Discotettiginae) Zootaxa. Rooseveltov trg 6, TUMBRINCK, J. (2014): Taxonomic revision of the Cladonotinae (Orthoptera: Tetrigidae) from HR-10000 Zagreb the islands of South-East Asia and from Croatia Australia, with general remarks to the classifi - SIGTET – Special Interest Group Tetrigidae cation and morphology of the Tetrigidae and E-Mail: [email protected]