Arch. Biol. Sci., Belgrade, 62 (3), 755-765, 2010 DOI:10.2298/ABS1003755G

DIVERSITY OF SPECIES OF THE FAMILY (COLEOPTERA) IN

B. GAVRILOVIĆ1 and S. B. ĆURČIĆ2

1Institute of Forestry, 11030 Belgrade, Serbia 2Institute of Zoology, Faculty of Biology, University of Belgrade, 11000 Belgrade, Serbia

Abstract - On the basis of literature data and personally collected entomological material, we made a faunistic and zoogeographical analysis of the family Scarabaeidae in Serbia. A total of 178 species, 83 genera, 15 tribes, and 7 subfamilies of the family Scarabaeidae are recorded from Serbia. The majority of findings are from the territory of northern Serbia, while western Serbia is the least explored region. Most of the species have a wide distribution range; therefore on the basis of the zoogeographical classification, 65.17% of the species belong to some of the Holarctic chorotypes, 34.27% of the species are included in some of the European chorotypes, while 0.56% are subcosmopolitan.

Keywords: Scarabaeidae, Coleoptera, diversity, distribution, Serbia

UDC 595.764(497.11):574/575

INTRODUCTION by Mikšić (1958, 1962, 1965). The majority of other authors focused only on certain localities in The family Scarabaeidae is a large and very diverse Serbia. One of the goals of this work is to provide a group of Coleoptera. According to some estimates, close estimate of the composition and number of more than 28,000 species are known worldwide species that are distributed in Serbia. In this paper (Grimaldi and Engel, 2005). The monophyly of the the analysis of species from the Scarabaeidae family is well established and is based on the family has been undertaken. Using data from the following characteristics: (1) adult antennae are works of numerous authors and from the first with a club or lamellate apex; (2) the fore legs are author collection, a table overview of all registered adapted for burrowing, with large fore coxae and species and localities where they were collected is tibiae with a series of teeth; (3) the hind wings have given (Table 1). reduced venation and a characteristic folding mechanism; (4) tergite VIII forms a true pygidium and is not concealed by tergite VII; (5) there are MATERIAL AND METHODS four Malpighian tubules as part of the excretory This paper is based on the works of numerous system; and (6) larvae are of scarabaeiform type. authors published from the end of the XIX century Phylogenetic research indicates, however, that till today. Also, the entomological material from the the family Scarabaeidae is not a monophyletic collection of the first author was analyzed. All group. Today, everything from 13 (American species and genera names are compatible with those authors) to as many as 25 (European authors) that are in use today, while various synonyms are families are recognized (Kohlmann, 2006). omitted. For practical reasons, the taxa and authors are listed alphabetically. Localities were given The most comprehensive research on according to the geographical regionalization of Scarabaeidae from the region that included Serbia Serbia. The zoogeographical distribution is defined (as well as other ex-Yugoslav countries) was made according to Vigna Taglianti et al. (1999).

755 756 B. GAVRILOVIĆ ET AL.

Table 1. Overview of all analyzed species and subspecies with their distribution in Serbia, including the authors that collected them. Abbreviations: N - North Serbia, S - South Serbia, W - West Serbia, SW - Southwest Serbia, E - East Serbia, C - Central Serbia, ? - unknown distribution in Serbia. Taxon Authors Distribution Family SCARABAEIDAE Subfamily APHODIINAE Acanthobodilus immundus (Creutzer, 1799) Mikšić (1962); Petrik (1958); Wohlberedt-Triebes (1909) N, E, SW Acrossus bimaculatus (Laxman, 1770) Mikšić (1962); Petrik (1958) N Acrossus depressus (Kugelann, 1792) Janković (1972); Mikšić (1962); Živojinović (1950) N, C, E Acrossus luridus (Fabricius, 1775) Gradojević (1963); Janković (1972); Košanin (1904); Mamontov (1937); Mikšić (1962); Teleki and Csiki (1922- N, C, E, SW, S 1940); Wohlberedt-Triebes (1909); Živojinović (1950) Acrossus rufipes (Linnaeus, 1758) Gavrilović (2006); Gavrilović and Stojanović (2008); Janković N, C, E (1972); Mikšić (1962); Živojinović (1950) Agoliinus piceus (Gyllenhal, 1808) Teleki and Csiki (1922-1940) S Agolius abdominalis abdominalis (Bonelli, 1812) Košanin (1904) C Agrilinus ater (De Geer, 1774) Mikšić (1962); Wohlberedt-Triebes (1909) N, SW Agrilinus rufus (Moll, 1782) Košanin (1904); Mamontov (1937); Mikšić (1962) N, C, S Agrilinus sordidus (Fabricius, 1775) Gavrilović (2006); Gavrilović and Stojanović (2008); Mikšić N (1962) Alocoderus hydrochaeris (Fabricius, 1798) Mikšić (1962) N Amidorus obscurus obscurus (Fabricius, 1792) Janković (1972); Košanin (1904); Mikšić (1962); Teleki and C, E, S Csiki (1922-1940) Amidorus thermicola (Sturm, 1800) Košanin (1904); Mikšić (1962); Teleki and Csiki (1922-1940) N, E, S Ammoecius brevis Erichson, 1848 Mikšić (1962) N Aphodius coniugatus (Panzer, 1795) Mikšić (1962); Živojinović (1950) N, E Aphodius fimetarius (Linnaeus, 1758) Jakšić et al., (2006); Janković (1972); Košanin (1904); Mikšić (1962); Teleki and Csiki (1922-1940); Wohlberedt-Triebes C, W, E, SW, S (1909); Živojinović (1950) Aphodius foetens (Fabricius, 1787) Mamontov (1937); Mikšić (1962) N, C Aphodius foetidus (Herbst, 1783) Mikšić (1962) N Biralus satellitius (Herbst, 1789) Mikšić (1962) N Bodilus circumcinctus (Schmidt, 1840) Mikšić (1962) N Bodilus ictericus ictericus (Laicharting, 1781) Gradojević (1963); Košanin (1904); Mikšić (1962); N, C, SW Wohlberedt-Triebes (1909) Bodilus lugens (Creutzer, 1799) Gavrilović (2006); Gavrilović and Stojanović (2008); N, C, E Mamontov (1937); Mikšić (1962) Bodilus punctipennis (Erichson, 1848) Mikšić (1962) N Calamosternus granarius (Linnaeus, 1767) Gavrilović (2006); Košanin (1904); Mamontov (1937); Mikšić N, C, E (1962); Petrik (1958); Živojinović (1950) Chilothorax conspurcatus (Linnaeus, 1758) Mikšić (1962); Živojinović (1950) N, E Chilothorax distinctus (Müller, 1776) Gavrilović (2006); Gavrilović and Stojanović (2008); Janković N, C, E (1972); Mikšić (1962); Živojinović (1950) Chilothorax lineolatus (Illiger, 1803) Mikšić (1962) E Chilothorax melanostictus (Schmidt, 1840) Mikšić (1962) N, C, E Chilothorax pictus (Sturm, 1805) Mikšić (1962) N Colobopterus erraticus (Linnaeus, 1758) Janković (1972); Košanin (1904); Mamontov (1937); Mikšić (1962); Teleki and Csiki (1922-1940); Wohlberedt- C, E, SW, S Triebes (1909); Živojinović (1950) Coprimorphus scrutator (Herbst, 1789) Janković (1972); Košanin (1904); Mikšić (1962); N, W, C, E, S Živojinović (1950) Diastictus vulneratus (Sturm, 1805) Gradojević (1963); Mikšić (1962); Mitić and Gradojević N, E (1983); Petrik (1958); Živojinović (1950) Esymus merdarius (Fabricius, 1775) Janković (1972); Mikšić (1962); Živojinović (1950) N, C, E Esymus pusillus (Herbst, 1789) Mikšić (1962); Živojinović (1950) C, E Eudolus quadriguttatus (Herbst, 1783) Mikšić (1962); Živojinović (1950) E Euheptaulacus carinatus carinatus (Germar, 1824) Mikšić (1962); Teleki and Csiki (1922-1940) E, S

DIVERSITY OF SPECIES OF THE FAMILY SCARABAEIDAE (COLEOPTERA) IN SERBIA 757

Table 1. Ctd. Taxon Authors Distribution Euheptaulacus villosus (Gyllenhal, 1806) Hristovski et al., (1996) S Euorodalus coenosus (Panzer, 1798) Gavrilović (2006); Gavrilović and Stojanović (2008); N Mikšić (1962) Eupleurus subterraneus (Linnaeus, 1758) Mikšić (1962) N, C Heptaulacus testudinarius (Fabricius, 1775) Mikšić (1962) N Labarrus lividus (Olivier, 1789) Mikšić (1962); Petrik (1958) N, C Limarus maculatus (Sturm, 1800) Mikšić (1962) N, C Liothorax kraatzi (Harold, 1868) Mikšić (1962) N Liothorax plagiatus (Linnaeus, 1767) Mikšić (1962) N Loraphodius suarius (Faldermann, 1835) Mikšić (1962) N Melinopterus consputus (Creutzer, 1799) Mikšić (1962); Teleki and Csiki (1922-1940) S Melinopterus prodromus (Brahm, 1790) Gradojević (1963); Janković (1972); Košanin (1904); Mamontov (1937); Mikšić (1962); Wohlberedt-Triebes N, C, E, SW (1909); Živojinović (1950) Melinopterus sphacelatus (Panzer, 1798) Mikšić (1962) N Mendidius ivanovi (Lebedev, 1912) Gradojević (1963) N Neagolius limbolarius limbolarius (Reitter, 1892) Janković (1972); Mikšić (1962) C, S Nialus varians (Duftschmidt, 1805) Gavrilović and Stojanović (2008); Gradojević (1963); Košanin (1904); Mamontov (1937); Mikšić (1962); Teleki N, C, E, SW, S and Csiki (1922-1940); Wohlberedt-Triebes (1909); Živojinović (1950) Nimbus contaminatus (Herbst, 1783) Janković (1972); Mikšić (1962) C Nimbus obliteratus (Panzer, 1823) Mikšić (1962); Teleki and Csiki (1922-1940); Živojinović N, C, E, S (1950) Otophorus haemorrhoidalis (Linnaeus, 1758) Janković (1972); Košanin (1904); Mamontov (1937); Mikšić (1962); Wohlberedt-Triebes (1909), Živojinović C, E, SW (1950) Oxyomus sylvestris (Scopoli, 1763) Košanin (1904); Mikšić (1962) N, C, E Phalacronothus biguttatus (Germar, 1824) Gradojević (1963); Mikšić (1962); Živojinović (1950) N, C, E Phalacronothus quadrimaculatus (Linnaeus, Janković (1972); Mikšić (1962); Živojinović (1950) N, C, E 1761) Plagiogonus putridus (Fourcroy, 1785) Mikšić (1962) N, C Pleurophorus caesus (Creutzer, 1796) Gradojević (1963); Košanin (1904); Mikšić (1962); Petrik N, E (1958); Živojinović (1950) Psammodius asper (Fabricius, 1775) Gradojević (1963, 1981); Košanin (1904); Mikšić (1962); N, C, S Mitić and Gradojević (1983); Petrik (1958) Psammodius laevipennis Costa, 1844 Mikšić (1962); Živojinović (1950) N, E Psammoporus sabuleti (Panzer, 1797) Mamontov (1937) C Rhyssemus germanus (Linnaeus, 1767) Mikšić (1962); Petrik (1958); Teleki and Csiki (1922-1940) N, C, S Sigorus porcus (Fabricius, 1792) Mikšić (1962) N Subrinus sturmi (Harold, 1780) Mikšić (1962) ? Teuchestes fossor (Linnaeus, 1758) Janković (1972); Mikšić (1962); Živojinović (1950) N, C, E Trichonotulus scrofa (Fabricius, 1787) Mikšić (1962); Živojinović (1950) N, E Volinus sticticus (Panzer, 1798) Mikšić (1962); Živojinović (1950); Wohlberedt-Triebes N, C, E, SW (1909) Subfamily CETONIINAE Tribe CETONIINI Cetonia aurata aurata (Linnaeus, 1761) Bobić (1891); Gavrilović (2006); Gavrilović and Stojanović (2008); Gradojević (1963); Grozdanić (1956); Hristovski et al., (1996); Ivanović (1962); Jakšić et al., (2006); Janković N, C, W, E, (1972); Košanin (1904); Mamontov (1937); Mikšić (1965); SW, S Petrik (1958); Teleki and Csiki (1922-1940); Tripković- Čubrilović (1960); Wohlberedt-Triebes (1909); Živojinović (1950) 758 B. GAVRILOVIĆ ET AL.

Table 1. Ctd. Taxon Authors Distribution Oxythyrea cinctella (Schaum, 1841) Mikšić (1965) E Oxythyrea funesta (Poda, 1761) Bobić (1891); Gavrilović (2006); Gavrilović and Stojanović (2008); Ivanović (1962); Jakšić et al., (2006); Janković (1972); Jovanić (1962); Košanin (1904); Mamontov (1937); N, C, E, SW, S Mikšić (1965); Petrik (1958); Teleki and Csiki (1922-1940); Wohlberedt-Triebes (1909); Živojinović (1950) Protaetia (Cetonischema) aeruginosa (Linnaeus, Bobić (1891); Ivanović (1962); Jakšić et al., (2006); Janković N, C 1767) (1972); Mamontov (1937); Mikšić (1965) Protaetia (Eupotosia) affinis affinis (Andersch, Košanin (1904); Mikšić (1965); Wohlberedt-Triebes N, C, E, SW 1797) (1909); Živojinović (1950) Protaetia (Netocia) cuprea (Fabricius, 1775) Gavrilović (2006); Gavrilović and Stojanović (2008); Gradojević (1963); Grozdanić (1956); Janković (1972); N, C, E Živojinović (1950) Protaetia (Netocia) cuprea metallica (Herbst, Košanin (1904); Mamontov (1937); Mikšić (1965) N, C, E, S 1782) Protaetia (Netocia) cuprea obscura (Andersch, Mikšić (1965); Petrik (1958) N 1797) Protaetia (Netocia) fieberi (Kraatz, 1880) Mikšić (1965) N, C Protaetia (Liocola) lugubris (Herbst, 1786) Ivanović (1962); Košanin (1904); Mikšić (1965) N, C Protaetia (Netocia) ungarica ungarica (Herbst, Gradojević (1963); Grozdanić (1956); Mikšić (1965); Petrik N 1790) (1958) Tropinota (Epicometis) hirta (Poda, 1761) Čamprag et al., (1975); Djurkić et al., (1973-1974); Gavrilović (2006); Gavrilović and Stojanović (2008); Gradojević (1963); Ivanović (1962); Jakšić et al., (2006); Janković (1972); Jovanić (1962); Košanin (1904); N, C, E, SW, S Mamontov (1937); Mikšić (1965); Petrik (1958); Teleki and Csiki (1922-1940); Tripković-Čubrilović (1960); Vukasović et al., (1962, 1964); Wohlberedt-Triebes (1909); Živojinović (1950) Tropinota (Tropinota) squalida squalida Mikšić (1965) C (Scopoli, 1783) Tribe TRICHIINI Gnorimus nobilis (Linnaeus, 1758) Jakšić et al., (2006); Košanin (1904); Mikšić (1965); C, E, S Živojinović (1950) Gnorimus variabilis (Linnaeus, 1758) Gavrilović (2006); Gavrilović and Stojanović (2008); Ivanović (1962); Mamontov (1937); Mikšić (1965); N, C, E Živojinović (1950) Osmoderma eremita (Scopoli, 1763) Ivanović (1962); Janković (1972); Košanin (1904); Mikšić (1965); Radović et al., (1995); Tripković-Čubrilović (1960); N, C, E Živojinović (1950) Trichius fasciatus (Linnaeus, 1758) Ivanović (1962); Janković (1972); Košanin (1904); Mikšić N, C, S (1965) Trichius sexualis Bedel, 1906 Bobić (1891); Gavrilović (2006); Gavrilović and Stojanović (2008); Jakšić et al., (2006); Janković (1972); Košanin (1904); N, C, E, S Mikšić (1965); Petrik (1958); Teleki and Csiki (1922-1940); Živojinović (1950) Tribe VALGINI Valgus hemipterus (Linnaeus, 1758) Bobić (1891); Gavrilović (2006); Gavrilović and Stojanović (2008); Gradojević (1963); Ivanović (1962); Jakšić et al., (2006); Janković (1972); Košanin (1904); Mamontov (1937); N, C, E, SW, S Mikšić (1965); Petrik (1958); Teleki and Csiki (1922-1940); Tripković-Čubrilović (1960); Wohlberedt-Triebes (1909); Živojinović (1950)

DIVERSITY OF SPECIES OF THE FAMILY SCARABAEIDAE (COLEOPTERA) IN SERBIA 759

Table 1. Ctd. Taxon Authors Distribution Subfamily DYNASTINAE Tribe ORYCTINI Oryctes (Oryctes) nasicornis (Linnaeus, 1758) Bobić (1891); Ivanović (1962); Janković (1972); Košanin (1904); Mamontov (1937); Petrik (1958); Tripković-Čubrilović N, C, E, SW, S (1960); Wohlberedt-Triebes (1909); Živojinović (1950) Oryctes nasicornis holdhausi (Linnaeus, 1758) Mikšić (1965) N Oryctes nasicornis kuntzeni (Linnaeus, 1758) Mikšić (1965) ? Tribe PENTODONTINI Pentodon idiota idiota (Herbst, 1789) Gavrilović (2006); Gavrilović and Stojanović (2008); Čamprag et al., (1968, 1973); Košanin (1904); Mamontov (1937); Mikšić N, C, E (1965); Petrik (1958); Vukasović et al., (1962, 1964); Živojinović (1950) Subfamily Tribe HOPLIINI Hoplia argentea (Poda, 1761) Gavrilović (2006); Gavrilović and Stojanović (2008); Jakšić et al., (2006); Janković (1972); Jovanić (1962); Mikšić (1965); N, C, E, SW, S Teleki and Csiki (1922-1940); Wohlberedt-Triebes (1909); Živojinović (1950) Hoplia auriventris Apfelbeck, 1912 Mikšić (1965); Winkler (1927-1932) N, C Hoplia brunnipes Bonelli, 1812 Mikšić (1965) N Hoplia dilutipes Reitter, 1890 Mikšić (1965); Winkler (1927-1932) N, E Hoplia graminicola (Fabricius, 1792) Košanin (1904); Petrik (1958) N, C Hoplia praticola Duftschmidt, 1805 Jovanić (1962); Mikšić (1965), Petrik (1958) N, C Hoplia subnuda Reitter, 1903 Mikšić (1965) N Tribe MELOLONTHINI Anoxia (Protanoxia) orientalis (Krynicky, 1832) Čamprag et al., (1973, 1987); Gavrilović (2006); Gavrilović and Stojanović (2008); Gradojević (1963); Grozdanić (1956); N, C, E Košanin (1904); Mikšić (1965); Petrik (1958); Vukasović et al., (1964); Živojinović (1963) Anoxia (Anoxia) pilosa (Fabricius, 1792) Čamprag et al., (1973, 1974, 1987); Gradojević (1963); Grozdanić (1956); Mikšić (1965); Petrik (1958); Vukasović et N, E al., (1964); Živojinović (1950); Živojinović (1963); Živojinović and Tomić (1956) Melolontha hippocastani Fabricius, 1801 Mikšić (1965); Živojinović (1950) N, E Melolontha melolontha (Linnaeus, 1758) Bobić (1891); Čamprag et al., (1987); Gavrilović (2006); Gavrilović and Stojanović (2008); Gradojević (1963); Grozdanić (1956); Janković (1972); Jovanić (1962); Košanin (1904); Mamontov (1937); Mikšić (1965); Petrik (1958); N, C, E, SW, S Vukasović et al., (1964); Wohlberedt-Triebes (1909); Živojinović (1950); Živojinović (1963); Živojinović and Petrović (1955) Melolontha pectoralis Megerle, 1812 Janković (1972); Mikšić (1965); Teleki and Csiki (1922-1940); N, C, S Živojinović (1963) Polyphylla fullo (Linnaeus, 1758) Bobić (1891); Čamprag et al., (1973, 1987); Gradojević (1963, 1981); Grozdanić (1956); Hristovski et al., (1996); Janković (1972); Košanin (1904); Mikšić (1965); Petrik (1958); N, C, SW, S Vukasović et al., (1964); Wohlberedt-Triebes (1909); Živojinović and Tomić (1956); Živojinović (1963) Tribe RHIZOTROGINI Amphimallon assimile (Herbst, 1790) Gavrilović (2006); Gavrilović and Stojanović (2008); Hristovski et al., (1996); Janković (1972); Košanin (1904); N, C, E, S Mikšić (1965) Amphimallon atrum (Herbst, 1790) Mikšić (1965) N Amphimallon burmeisteri Brenske, 1886 Mikšić (1965) C Amphimallon caucasicum (Gyllenhal, 1817) Mikšić (1965) E Amphimallon ochraceus (Knoch, 1801) Mikšić (1965) N Amphimallon ruficorne (Fabricius, 1775) Košanin (1904) S 760 B. GAVRILOVIĆ ET AL.

Table 1. Ctd. Taxon Authors Distribution Amphimallon solstitiale (Linnaeus, 1758) Čamprag et al., (1968, 1973, 1974, 1987); Djurkić et al., (1973- 1974); Gavrilović (2006); Gavrilović and Stojanović (2008); Hristovski et al., (1996); Jakšić et al., (2006); Janković (1972); N, C, W, E, SW, Jovanić (1962); Košanin (1904); Mamontov (1937); Teleki and S Csiki (1922-1940); Vukasović et al., (1962, 1964); Wohlberedt- Triebes (1909); Živojinović (1950); Živojinović (1963); Živojinović and Petrović (1955) Amphimallon solstitiale matutinale Nonveiller, 1963 Mikšić (1965) ? Haplidia transversa transversa (Fabricius, 1801) Mikšić (1965) N, C Holochelus (Miltotrogus) aequinoctialis (Herbst, Bobić (1891); Čamprag et al., (1968, 1974, 1975); Djurkić et al., 1790) (1973-1974); Grozdanić (1956); Jovanić (1962); Košanin (1904); Mamontov (1937); Mikšić (1965); Petrik (1958); N, C, S Tripković-Čubrilović (1960); Vukasović et al., (1962, 1964); Živojinović and Petrović (1955); Živojinović and Tomić (1956); Živojinović (1963) Holochelus (Miltotrogus) fallax (Marseul, 1879) Mikšić (1965) S Holochelus (Miltotrogus) nocturnus (Nonveiller, Mikšić (1965) C 1958) Holochelus (Miltotrogus) pilicollis (Gyllenhal, 1817) Mikšić (1965) N Holochelus (Miltotrogus) vernus (Germar, 1823) Čamprag et al., (1968); Djurkić et al., (1973-1974); Mikšić N (1965); Vukasović et al., (1962, 1964); Živojinović (1963) Pseudotrematodes frivaldszkyi (Ménétriès, 1836) Mikšić (1965) S aestivus (Olivier, 1789) Jakšić et al., (2006); Janković (1972); Košanin (1904); N, C, S Mamontov (1937); Mikšić (1965); Petrik (1958) Rhizotrogus marginipes Mulsant, 1842 Živojinović (1950) E Tribe SERICINI Maladera (Maladera) holosericea (Scopoli, 1772) Čamprag et al., (1968, 1973, 1974, 1975, 1987); Djurkić et al., (1973-1974); Gradojević (1963); Košanin (1904); Mikšić N, C, S (1965); Mitić and Gradojević (1983); Petrik (1958); Vukasović et al., (1964) Omaloplia (Omaloplia) alternata Küster, 1849 Janković (1972) C Omaloplia (Omaloplia) erythroptera Frivaldszky, Mikšić (1965); Petrik (1958) N, E 1835 Omaloplia (Omaloplia) marginata (Fuessly, 1775) Jakšić et al., (2006); Janković (1972); Mikšić (1965); Teleki and N, C, E, S Csiki (1922-1940); Živojinović (1950) Omaloplia (Omaloplia) ruricola (Fabricius, 1775) Gradojević (1963); Hristovski et al., (1996); Mikšić (1965); N, E, S Živojinović (1950) Omaloplia (Acarina) spireae (Pallas, 1776) Mikšić (1965); Petrik (1958); Živojinović (1950) N, E Serica brunnea (Linnaeus, 1758) Janković (1972); Mikšić (1965) N, C Triodonta aquila (Laporte de Castelnau, 1840) Gradojević (1963); Janković (1972); Mikšić (1965); N, C, SW, S Wohlberedt-Triebes (1909) Subfamily ORPHNINAE Chaetonyx robustus robustus Schaum, 1862 Gradojević (1963); Mikšić (1962) N Subfamily RUTELINAE Tribe ANOMALINI Anisoplia (Anisoplia) agricola (Poda, 1761) Bobić (1891); Gavrilović (2006); Gavrilović and Stojanović (2008); Čamprag et al., (1968); Janković (1972); Jovanić N, C, W, S (1962); Košanin (1904); Mikšić (1965); Teleki and Csiki (1922- 1940) Anisoplia (Autanisoplia) austriaca austriaca Čamprag et al., (1968, 1973, 1974, 1975, 1987); Djurkić et al., (Herbst, 1783) (1973-1974); Gradojević (1963); Jovanić (1962); Mamontov N, C, E (1937); Mikšić (1965); Petrik (1958); Vukasović et al., (1962, 1964); Živojinović (1950) Anisoplia (Anisoplia) bromicola (Germar, 1817) Bobić (1891); Mikšić (1965) N, C Anisoplia (Anisoplia) bulgarica Apfelbeck, 1909 Mikšić (1965) E Anisoplia (Anisoplia) bureschi Zacharieva-Stoilova, Mikšić (1965) C 1958 DIVERSITY OF SPECIES OF THE FAMILY SCARABAEIDAE (COLEOPTERA) IN SERBIA 761

Table 1. Ctd. Taxon Authors Distribution Anisoplia (Anisoplia) deserticola Fischer von Djurkić et al., (1973-1974); Gradojević (1963); Mikšić (1965); N Waldheim, 1824 Petrik (1958); Živojinović (1963) Anisoplia (Anisoplia) erichsoni Reitter, 1883 Janković (1972); Mikšić (1965) W, C, S Anisoplia (Anisoplia) flavipennis Brullé, 1832 Gavrilović and Stojanović (2008); Janković (1972); Živojinović N, C (1963) Anisoplia (Anisoplia) lata lata Erichson, 1847 Čamprag et al., (1968, 1973, 1974); Gradojević (1963, 1981); Jovanić (1962); Košanin (1904); Mamontov (1937); Mikšić N, C (1965); Petrik (1958); Vukasović et al., (1962, 1964) Anisoplia (Anisoplia) monticola Erichson, 1847 Živojinović (1950) E Anisoplia (Anisoplia) tempestiva Erichson, 1847 Janković (1972); Košanin (1904); Mikšić (1965) C, E Anomala dubia (Scopoli, 1763) Gradojević (1963); Košanin (1904) Mikšić (1965); Petrik N, C, E (1958); Živojinović (1950); Živojinović (1963) Anomala errans errans (Fabricius, 1775) Gradojević (1963, 1981); Mikšić (1965); Živojinović (1963) N Anomala solida Erichson, 1847 Gradojević (1981); Grozdanić (1956); Mikšić (1965); Petrik (1958); Vukasović et al., (1964); Winkler (1927-1932); N, C Živojinović (1963) Anomala vitis (Fabricius, 1775) Bobić (1891); Gavrilović (2006); Gavrilović and Stojanović (2008); Košanin (1904); Mikšić (1965); Živojinović and Tomić N, C, E (1956) Blitopertha arenaria (Brullé, 1832) Gradojević (1963, 1981); Mikšić (1965) N, S Blitopertha lineata (Fabricius, 1798) Janković (1972); Košanin (1904); Mikšić (1965) C, E, S Blitopertha lineolata (Fischer von Waldheim, 1824) Teleki and Csiki (1922-1940) S Chaetopteroplia segetum segetum (Herbst, 1783) Čamprag et al., (1968, 1973, 1974, 1987); Gradojević (1963); Jovanić (1962); Košanin (1904); Mikšić (1965); Petrik (1958); N, C, E, S Vukasović et al., (1964); Živojinović (1950); Živojinović (1963) Exomala (Trichopertha) hirtella (Brullé, 1832) Janković (1972); Mikšić (1965); Živojinović (1950) C, E Phyllopertha horticola (Linnaeus, 1758) Čamprag et al., (1973); Janković (1972); Košanin (1904); N, C Mamontov (1937) Subfamily SCARABAEINAE Tribe SCARABAEINI Scarabaeus (Scarabaeus) typhon (Fischer, 1824) Grozdanić (1956); Mikšić (1958); Petrik (1958) N, E, S Scarabaeus (Scarabaeus) pius (Illiger, 1803) Gradojević (1963); Grozdanić (1956); Mikšić (1958); Petrik N (1958) Scarabaeus (Scarabaeus) sacer Linnaeus, 1758 Bobić (1891); Gradojević (1963), Košanin (1904) N, C, E Gymnopleurus mopsus (Pallas, 1781) Gradojević (1963, 1981); Grozdanić (1956); Janković (1972); N, C, S Mikšić (1958); Petrik (1958) Gymnopleurus sturmii MacLeay, 1821 Mikšić (1958); Košanin (1904) N, E, S Gymnopleurus geoffroyi (Fuessly, 1775) Mikšić (1958); Košanin (1904); Teleki and Csiki (1922-1940); N, C, E, SW, S Wohlberedt-Triebes (1909) Sisyphus schaefferi (Linnaeus, 1758) Hristovski and Ilioska (1996); Košanin (1904); Mamontov (1937); Mikšić (1958); Teleki and Csiki (1922-1940); N, C, SW, S Wohlberedt-Triebes (1909) Tribe COPRINI Copris lunaris (Linnaeus, 1758) Bobić (1891); Gavrilović (2006); Gavrilović and Stojanović (2008); Gradojević (1963); Jakšić et al., (2006); Janković N, C, E, SW, S (1972); Košanin (1904); Mamontov (1937); Mikšić (1958); Petrik (1958); Wohlberedt-Triebes (1909); Živojinović (1950) Tribe ONITINI Cheironitis ungaricus (Herbst, 1789) Mikšić (1958) N Tribe ONITICELLINI Euoniticellus fulvus (Goeze, 1777) Jakšić et al., (2006); Košanin (1904); Mamontov (1937); Mikšić (1958); Petrik (1958); Teleki and Csiki (1922-1940); N, C, E, SW, S Wohlberedt-Triebes (1909); Živojinović (1950) Euoniticellus pallipes (Fabricius, 1798) Mikšić (1958) N Tribe ONTHOPHAGINI Caccobius histeroides (Ménétriès, 1832) Gradojević (1963) N

762 B. GAVRILOVIĆ ET AL.

Table 1. Ctd. Taxon Authors Distribution Caccobius schreberi (Linnaeus, 1758) Gradojević (1963); Jakšić et al., (2006); Košanin (1904); Mamontov (1937); Mikšić (1958); Petrik (1958); Teleki and N, C, E, SW, S Csiki (1922-1940); Wohlberedt-Triebes (1909); Živojinović (1950) Euonthophagus amyntas (Olivier, 1789) Gradojević (1963); Janković (1972); Košanin (1904); Mikšić N, C, E, S (1958); Petrik (1958) Euonthophagus gibbosus (Scriba, 1790) Janković (1972); Mikšić (1958); Teleki and Csiki (1922-1940) C, S Onthophagus (Onthophagus) illyricus (Scopoli, 1763) Mikšić (1958) N, C Onthophagus (Onthophagus) taurus (Schreber, Csiki (1904); Hristovski and Ilioska (1996); Košanin (1904); 1759) Mamontov (1937); Mikšić (1958); Petrik (1958); Teleki and N, C, W, E, SW, Csiki (1922-1940); Wohlberedt-Triebes (1909); Živojinović S (1950) Onthophagus (Palaeonthophagus) coenobita Gradojević (1963); Košanin (1904); Mamontov (1937); Mikšić (Herbst, 1783) (1958); Teleki and Csiki (1922-1940); Wohlberedt-Triebes N, C, SW, S (1909) Onthophagus (Palaeonthophagus) fissicornis Steven, Mikšić (1958) N, E 1809 Onthophagus (Palaeonthophagus) fracticornis Gavrilović (2006); Gavrilović and Stojanović (2008); (Preyssler, 1790) Gradojević (1963); Janković (1972); Košanin (1904); N, C, E Mamontov (1937); Mikšić (1958); Petrik (1958); Živojinović (1950) Onthophagus (Palaeonthophagus) gibbulus (Pallas, Košanin (1904); Mamontov (1937); Mikšić (1958) N, C, S 1781) Onthophagus (Palaeonthophagus) grossepunctatus Mikšić (1958) N Reitter, 1905 Onthophagus (Palaeonthophagus) lemur (Fabricius, Gradojević (1963, 1981); Košanin (1904); Mikšić (1958); Mitić 1781) and Gradojević (1983); Teleki and Csiki (1922-1940); N, C, E, SW, S Wohlberedt-Triebes (1909); Živojinović (1950) Onthophagus (Palaeonthophagus) lucidus (Sturm, Mikšić (1958) N 1800) Onthophagus (Palaeonthophagus) nuchicornis Hristovski et al., (1996); Košanin (1904); Mamontov (1937); (Linnaeus, 1758) Mikšić (1958); Petrik (1958); Wohlberedt-Triebes (1909); N, C, E, SW Živojinović (1950) Onthophagus (Palaeonthophagus) ovatus (Linnaeus, Košanin (1904); Mamontov (1937); Mikšić (1958); Petrik 1767) (1958); Teleki and Csiki (1922-1940); Wohlberedt-Triebes N, C, E, SW, S (1909); Živojinović (1950) Onthophagus (Palaeonthophagus) ruficapillus Mikšić (1958) N, C Brullé, 1832 Onthophagus (Palaeonthophagus) semicornis Mikšić (1958) N (Panzer, 1798) Onthophagus (Palaeonthophagus) vacca (Linnaeus, Gavrilović (2006); Gavrilović and Stojanović (2008); Janković 1767) (1972); Košanin (1904); Mamontov (1937); Mikšić (1958); N, C, E, SW, S Teleki and Csiki (1922-1940); Wohlberedt-Triebes (1909), Živojinović (1950) Onthophagus (Palaeonthophagus) verticicornis Janković (1972); Košanin (1904); Mikšić (1958); Živojinović N, C, W, E, S (Laicharting, 1781) (1950) Onthophagus (Palaeonthophagus) vitulus (Fabricius, Mikšić (1958); Wohlberedt-Triebes (1909) N, SW 1776) Onthophagus (Furconthophagus) furcatus Gradojević (1963, 1981); Jakšić et al., (2006); Košanin (1904); (Fabricius, 1781) Mamontov (1937); Mikšić (1958); Mitić and Gradojević N, C, E, S (1983); Petrik (1958); Radović et al., (1995); Teleki and Csiki (1922-1940); Živojinović (1950)

DIVERSITY OF SPECIES OF THE FAMILY SCARABAEIDAE (COLEOPTERA) IN SERBIA 763

RESULTS AND DISCUSSION species from central Serbia, two species from east Serbia, one species from south-west Serbia, and With regard to the results of this study, presented in one species from south Serbia. From the subfamily Table 1, the existence of 178 species, 83 genera, 15 Melolonthinae, 29 species were recorded from tribes, and 7 subfamilies from the family Scara- north Serbia, 20 species from central Serbia, one baeidae have been confirmed. Today, with over species from west Serbia, 14 species from east 28,000 known species, Scarabaeidae is the largest fa- Serbia, five species from south-west Serbia, and 15 mily of the superfamily , comprising species from south Serbia. The subfamily Orphni- 91% of the total number of species of the superfami- nae is represented with one species from north ly. Within Scarabaeidae, the subfamilies Aphodiinae Serbia. For the subfamily Rutelinae, 13 species and Scarabaeinae, with approximately 6,850 species have been noted to be from north Serbia, 15 spe- worldwide, represent 22% of the total species of cies from central Serbia, two species from west Scarabaeoidea, or 25% of the known species of Serbia, nine species from east Serbia, and six spe- Scarabaeidae. The subfamilies Orphninae, Melolon- cies from south Serbia. The subfamily Scarabaei- thinae, Dynastinae, Rutelinae, and Cetoniinae in- nae is represented with 31 species from north Ser- clude about 20,950 species, or 69% of all known bia, 21 species from central Serbia, two species Scarabaeoidea worldwide and 75% of the known from west Serbia, 17 species from east Serbia, 12 Scarabaeidae (Ratcliffe and Jameson, 2004). species from south-west Serbia, and 18 species from south Serbia. The genera Onthophagus Latreille, 1802 (Scarabaeinae), with 17 noted species (9.55% of A total of 142 species have been recorded from the total analyzed species), Anisoplia Schoenherr, north Serbia, 107 species from central Serbia, eight 1817 (Rutelinae), with 11 species (6.18%), and species from west Serbia, 86 species from east Hoplia Illiger, 1803 and Amphimallon Berthold, Serbia, 33 species from south-west Serbia, and 62 1827 (Melolonthinae), each with seven species species from south Serbia. For five of the species the (3.93% each), stand out as the most numerous in exact locality of their collection is not known. Con- Serbia. The tribes Anomalini (Rutelinae) and cerning Scarabaeidae, it follows that north Serbia Onthophagini (Scarabaeinae) each have 21 species (Vojvodina Province) is the best characterized area, (11.80% each), while the tribe Rhizotrogini (Melo- while west Serbia is the least explored region. lonthinae) is represented with 16 species (9.00%). The largest subfamily Aphodiinae has 68 species On the basis of the general geographical distri- (38.20%), and following after it are the subfamilies bution of each species, the zoogeographical distri- Melolonthinae, with 37 species (20.78%), and bution was determined using the zoogeographical Scarabaeinae, with 32 species (17.98%). classification suggested by Vigna Taglianti et al., (1999). All analyzed species fall under two major Within the family Scarabaeidae and subfamily groups of chorotypes – chorotypes of the species Aphodiinae, 52 species have been recorded from widely distributed in the Holarctic region and north Serbia, 34 species from central Serbia, two chorotypes of those that are more or less widely species from west Serbia, 33 species from east spread in Europe. From the total number of Serbia, 10 species from south-west Serbia, and 15 species, 116 species (65.17%) belong to some of the species from south Serbia. The subfamily Ceto- chorotypes of Holarctic, 61 species (34.27%) are niinae is represented with 14 species from north related to some of the European chorotypes, while Serbia, 15 species from Central Serbia, one species one species (0.56%) is subcosmopolitan. Within from West Serbia, 11 species from East Serbia, five the chorotypes of the Holarctic region, two species species from south-west Serbia, and seven species (1.12%) are part of the Holarctic chorotype, 27 from south Serbia. The subfamily Dynastinae was species (15.17%) belong to the Palaearctic found to have two species from north Serbia, two chorotype, 32 species (17.98%) are included within 764 B. GAVRILOVIĆ ET AL.

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ДИВЕРЗИТЕТ ВРСТА ФАМИЛИЈЕ SCARABAEIDAE (COLEOPTERA) У СРБИЈИ

Б. ГАВРИЛОВИЋ1 и С. Б. ЋУРЧИЋ2

1Институт за шумарство, 11030 Београд, Србија 2Институт за зоологију, Биолошки факултет, Универзитет у Београду, 11000 Београд, Србија

У студији је утврђено oквирно стање фауне фа- Србије до сада. Такође, приказано је и њихово милије Scarabaeidае у Србији. Коришћени су ли- регионално распрострањење у Србији. На тературни подаци разних аутора од краја XIX основу зоогеографске поделе урађена је зоогео- века до данас, а анализирана је и збирка ентомо- графска анализа. Северна Србија се издваја као лошког материјала првог аутора. Укупно 178 област у оквиру које су највише вршена истра- врста, 83 рода, 15 трибуса и седам подфамилија живања, док је западна Србија слабо истражена. Scarabaeidae је до сада евидентирано из Србије. Готово све анализиране врсте су са широким Табеларно је дат приказ свих врста фамилије распрострањењем у области Холарктика и Scarabaeidae које су прикупљене са територије Европе.