The Bee Flies (Diptera: Bombyliidae) of Jordan

Home , Anastoechus, Andrena, Anthracinae, Anthrax (fly), Apolysis (fly), Bombyliidae

The bee flies (Diptera: Bombyliidae) of Jordan

A. KATBEH-BADER & S. ARABYAT

Abstract: Weekly field trips to collect bee flies were conducted from October 2001 to November 2002. More than 870 specimens from 72 localities were collected. Specimens of Bombyliidae collected previously from Jordan and preserved in the University of Jordan Insects Museum as well as other Jordanian collections were also studied. The specimens were found to belong to 132 species in 41 genera and eight subfamilies. Of these, 124 species are recorded for the first time. The number of specimens examined and collecting sites and dates in Jordan is provided for each species followed by remarks.

Key words: bee flies, Bombyliidae, Jordan, ">

Introduction are important as natural enemies of major pests including locust, grasshoppers, army- Bombyliidae or bee flies are world wide worms, nettle caterpillars and tsetse flies. in distribution. Nearly 4500 species are Others develop in nests of solitary wasps and known, distributed among 16 subfamilies bees, and they are occasionally considered (EVENHUIS &. GREATHEAD 1999). to be pests when they kill the larvae of bees Many bee flies can be recognised by being bred as pollinators for crops such as al- their woolly covering of hairs and their long falfa in United States (EVENHUIS & GREAT- snout up to 10 mm (BODENHEIMER 1935). HEAD 1999). They have attractively patterned wings and There are no known publications on the brightly coloured bodies. They are more Bombyliidae of Jordan except that of (EVEN- abundant in arid and semi arid regions HUIS & GREATHEAD 1999) who listed the (HULL 1973). They are mostly medium to following 14 species; Anaswechus exafoidus large insects (1-20 mm). The adults are (WIEDEMANN), A. nivifrons WALKER, A. mostly strong fliers, except for small or vnsignatus (PORTSCHINSKY), Bombylisoma minute species and Usiinae, which are en- flavibarbum LOEW, Bombylius aregentifacies countered resting inside flowers (GREAT- AUSTEN, Chakochitm argyrocephalus MAC- HEAD & EVENHUIS 1997). QUART, Cytherea aureus FABRICIUS, C. albo- Adult bee flies are often the major polli- Uneata BEZZI, C. barbara SACK, C. delicata nators of many flowering plants, especially (BECKER), C. dispar (LOEW), C. nucleorum those occurring in the more desertic regions (BECKER), Spogostylum ocyak (WIEDEMANN), of our planet. Recent studies have shown Caecanthrax arabicus (MACQUART). Howev- that some plants depend on bee flies for pol- er, several studies were conducted on the lination so that the survival of some endan- Bombyliidae of the Middle East. From Pales- gered plant species may depend on preserv- tine, BODENHEIMER (1937) listed 72 species ing their bee flies pollinators (EVENHUIS & in 4 subfamilies. AUSTEN (1937) recorded GREATHEAD 1999). Immatures are para- 128 species in 31 genera and provided keys sitoids or hyperparasitoids on several orders to subfamilies, genera and species in addi- of insects such as Lepidoptera, Orthoptera, tion to the description of some new species. Hymenoptera, Diptera, Coleoptera, Neu- Many new records, including new species, roptera and Blattaria (DU MERLE 1975, were published by ZAITZEV (1995, 1996, Denisia 14, zugleich Kataloge der OÖ. Landesmuseen YEATES & GREATHEAD 1997). Some species 1997, 1998, 1999). From Arabia, GREAT- Neue Serie 2 (2004), 353-384

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Table 1: Collecting sites and their coordinates (1989) listed 13 species from Al-Kuwait.

Coordinate Collecting sites Coordinate Collecting sites From Egypt, EL-HAWAGRY (1998) described 32°14'N-35°481E Dibbin 32°03'N-35°52'E Abu Nusayr two new species of Anthrax, in (2001) he 32°08'N-35°45'E Eastern Bayudah 29°37'N-35°33'E Ad Disah recorded four species of Xeramoeba, one 31°23'N-35°40'E Faqu'a 32°18'N-35°43'E Ajlun species of them was new to science (EL- 31°02'N-35°28'E Ghawr As Safi 32°40'N-35°37'E Al Adasiyah HAWAGRY 2001) and in 2002 he described 32°05'N-35°39'E Hummrit As Sahin 32°07'N-35°39'E Al Aridah three new species (EL-HAWAGRY 2002). EL- 31°59'N-35°4VE Ira 31°52'N-36°50'E Al Azraq HAWAGRY et al. (2000) recorded 31 species 32O33'N-35°5VE Irbid 31°26'N-35°41'E Al Badhiyah of Anthracinae; two species of them were 32°17'N-35O54'E Jarash 31°53'N-35°46'E Al Bahhath described as new species to science. From 32°36'N-35°43'E Kaf r Asad 32°04'N-35°5VE Al Baqa'a Oman, GREATHEAD (1980a) recorded 38 o o 32 04'N-35 42'E Kafr Huda 32°38'N-35°43'E Al Baqurah species including a new genus and 4 new 32°22'N-35°55'E Kafr Khal 32°03'N-37°08'E Al Buqei'wiya species. 32°12'N-35°36'E King Talal Dam 32°01'N-35°46'E Al Fuhays 32°18'N-35°42'E Kufrinjah 30°59'N-35°24'E Al Ghawr Since there are no known studies about 32°16'N-35°36'E Kurayyimah 32°29'N-35°52'E Al Husson the Bombyliidae of Jordan, this study was 31°41'N-35°42'E Ma'ain 32°01'N-35°52'E Al Jubayhah conducted to identify species collected pre- 30°12'N-35°44'E Ma'an 31°52'N-35°38'E Al Kafrayn viously from Jordan and kept in the Univer- 31°43'N-35°48'E Madaba 32°12'N-35°41'E Al Karak sity of Jordan Insect Museum (UJIM) and 31°59'N-35°45'E Mahis 32-21'N-36°12'E Al Mafraq other available specimens in Jordanian col- 31°59'N-35°59'E Marka 32°42'N-35°41'E Al Mukhaybah al Fawqa lections. Additional specimens were collect- 32°39'N-35°48'E Muzyrib 32°39'N-35°40'E Al Mukhaybah at Tahta ed for one year from different parts in Jordan 32°17'N-35°50'E Nahlah 31°48'N-35°47'E Al Mushaqqar by conducting weekly or biweekly field trips. 31°52'N-35°50'E Na'or 31°46'N-36°13'E Al Muwaqqar This paper is based on an article which is go- 30°19'N-35°29'E Petra 31°39'N-35°43'E Al Wala ing to be published in the journal Zootaxa. 32°14'N-35°42'E Rajib 31°57'N-35°56'E Amman 30°57'N-35°37'E Rihab 31°16'N-35°40'E Ar Rabbah 29°35'N-35°26'E Rum 32°30'N-36°04'E Ar Ramtha Materials and Methods 32O11'N-35°37'E Ruwayhat al Ghawr 32°06'N-35°47'E Ar Rumaymin Weekly field trips were conducted to dif- 32°27'N-36°14'E Sama As Sarhan 32°09'N-35°49'E Ar Rumman ferent locations in Jordan from October 31°59'N-35°59'E Sayl Az Zarqa 30-41'N-35°34'E Ar Rummanah 2001 to November 2002. Adult bee flies 32°O2'N-35°5O'E Suwaylih 32°11'N-37°06'E As Safawi were collected by sweeping nets. In addition 31°46'N-35°36'E Suwaymah 32°03'N-35°44'E As Salt to the collected specimens, additional mate- 32°27'N-35°37'E Tabaqt fahl 32°35'N-35°59'E Ash Shajarah 32°39'N-35°40'E Um Qays 30°30'N-35°3VE Ash Shawbak rials of Bombyliidae were examined from 32°30'N-37°17'E Wadi al Hashad 31°50'N-36°49'E Ash Shumary Reserve three Jordanian collections: Al Al-Bayt Uni- 31°51'N-36°48'E Wadi ar Ratam 31°54'N-35°38'E Ash Shunah versity Collection, Al Balqa'a Applied Uni- 32°23'N-35°35'E Wadi al Yabis 32°08'N-36°04'E As Sukhnah versity Collection, and Natural History Mu- 31°57'N-35°49'E Wadi as Sir 30°50'N-35°36'E At Tafilah seum at Al Yarmouk University. The first 31°10'N-35°32'E Wadi Marsad 32°36'N-35°56'E Ayn Almuallaqah two museums contained specimens collected 32°01'N-35°35'E Wadi Shu'ayb 32°43'N-35°47'E Ayn Aqraba previously by Prof. Ahmad Katbeh and his 31°24'N-35°43'E Wadi Shuqayq 32°35'N-35°57'E Ayn esh Shallaleh team, while the last collection was collected 32o10'N-35°43'E Western Bayudah 32°05'N-36°06'E Az Zarqa by Mr. Suhail Ismael, the museum curator. 32°01'N-36°00'E Yajuz 31°12'N-35°40'E Badhan O All the collected specimens were kept in 31 59'N-35°42'E Yarqa 32°26'N-35°44'E Birqish 32o31'N-35o50'E Zabdah 30-41'N-35°37'E Dana the UJIM. Some specimens were kept in Dr. 32°33'N-35°48'E Zahar 32°15'N-36°49'E Dayr al Kahf Greathead's collection, Imperial College, 32°06'N-35°45'E Zai 32°05'N-35°36'E Dayr Alia London, who helped in the identification or 32°26'N-35°46'E Zubiya 31°36'N-35°33'E Dead Sea confirmation of identified species. The distribution of species in Jordan is mentioned HEAD (1980b) recorded 118 species, 13 of in the material examined section for each them were new to science. In 1988, he species. Localities are presented in alphabet- added 29 species and 2 subspecies; two of ical order. Names of sites were according to them were described as new species. In addi- Gazetteer of Jordan (ANONYMUS 1990). The tion, he provided identification keys to sub- sites and their coordinates are given in families, genera and species. AL-HOUTY Table 1. Dates within each locality are

354 © Biologiezentrum Linz/Austria; download unter www.biologiezentrum.at arranged chronologically. The number of Bowbyleüa simulans (AUSTEN 1937) specimens examined or collected is given Bombomyia discoidea (FABRICIUS 1794) between brackets after the dates. Unless Bombomyia suctica (FABRICIUS 1794) otherwise mentioned, the specimens are de- Bombylius fuscus FABRICIUS 1781 posited at the UJ1M. Bombylius medius LINNAEUS 1758 Bombylius trichurus PALLAS 1818 Results Bombylius fimbriatus MEIGEN 1820 Bombylius major LINNAEUS 1758 Bombylius modestus LOEW 1873 Species List Bombylius pumilus MEIGEN 1820 The following 132 species of Bombyli- Bombylius fulvescens WlEDEMANN 1820 idae are recorded in this study. These belong Bombylius canescens MlKAN 1796 to 41 genera in 8 subfamilies. Bombylius mendax AUSTEN 1937 Bombylius cinerascens MlKAN 1796 Subfamily Usiinae BECKER Bombylius posacus FABRICIUS 1805. Bombylius (Zephyrectes) quadrifarius Tribe Apolysini EVENHUIS LOEW 1855 Apolysis sp. 1 Dischistus syriacus (ViLLENEUVE 1912) Neobombybdes giganteus (ViLLENEUVE 1920) Tribe Usiini BECKER Anastoechus trisignatus Usia ignorata BECKER 1906 (PORTSCHINSKY 1881) Usia aenea (ROSSI 1794) Anastoechus bahirae BECKER 1915 Usia bicobr MACQUART 1855 Anastoechus nitidulus (FABRICIUS 1794) Parageron grams (LOEW 1856) Anastoechus exalbidus (WlEDEMANN 1820) Parageron sp. 1

Parageron sp. 2 Subfamily Ecliminae HALL Eclimus gracilis LOEW 1844 Subfamily Toxophorinae SCHINER

Subfamily Cythereinae BECKER Tribe Toxophorini SCHINER Calbstoma fasdpenne MACQUART 1840 Toxophora fasciculata (VILLERSI 1789) Cytherea obscura FABRICIUS 1794

Tribe Gerontini HESSE Cytherea aureus FABRICIUS 1794 Cytherea fenestrata (LOEW 1873) Geron mystacinus BEZZI 1924 Cytherea nucleorum (BECKER 1902) Geron olivieri MACQUART 1840 Cytherea delicata (BECKER 1906) Geron erythropus BEZZ! 1925 Cytherea dispar (LOEW 1873) Geron smimovi ZAITZEV 1978 Chalcochiton spedosus (LOEW 1844) Geron krymensis PARAMONOV 1929 Chalcoc/iiton pallasi (LOEW 1856) C/talcoc/iiton syriacus (LOEW 1869) Subfamily Phthiriinae BECKER Amictus obliquenotatus AUSTEN 1937 Tribe Phthiriini BECKER Amictus validus LOEW 1869 Amictus sp. near tigrinus AUSTEN 1937 Phthiria sp. 1 Amictus sp. near zinamominus BECKER 1906 Amictus virgatus AUSTEN 1937 Subfamily Bombyliinae LATREILLE Amictus sp. near pictus LOEW 1869 Tribe Conophorini BECKER

Conophorus giaucescens (LOEW 1863) Subfamily Lomatiinae SCHINER Conophorus nobiUs (LOEW 1873) Tribe lomatiini SCHINER Legnotomyia trichorhoea (LOEW 1855) Legnotomyia tineracea AUSTEN 1937 Lomatia abbreviata ViLLENEUVE 1911 Lomatia polyzona LOEW 1869 Tribe Bombyliini LATREILLE Lomatia vysiphone LOEW 1860 Systoechus bngirostris BECKER 1916 Bombylelia atra (SCOPOLI 1763)

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Subfamily Anthracinae UTREILLE Thyridanthrax perspirillaris ssp. perspicillaris (lX)EW 1869) Tribe Aphoebantini BECKER Thyridanthrax perspiciüaris ssp. idolus Cononedys stenura (LoEW 1871) HESSE 1956 Cononedys inomata (GREATHEAD 1967) Thyridanthrax polyphemus (WiEDEMANN 1819) Tribe Prorostomatini HULL Thyridanthrax sp. near griseolus Stomylomyia europaea (LOEW 1869) (KLUG 1832) Plesiocera algira MACQUART 1840 Thyridanthrax incanus (KLUG 1832) Thyridanthrax elegans (WiEDEMANN 1818) Tribe Xeramoebini HULL Pachyanthrax fulvifacies (AUSTEN 1937) Desmawneura sp. 1 Pachyanthrax telamon (LOEW 1869) Xeramoeba semirufa (SACK 1909) ? Pachyanthrax nimrodkus ZAITZEV 1998 Xeramoeba sabulonis (BECKER 1906) Exhyalanthrax afer (FABRICIUS 1794) Xeramoeba salwae EL HAWAGRY 2001 Exhyalanthrax melanchlaenus (LOEW 1869) Petrorossia hespera (Rossi 1790) Exhyalanthrax contrarius (BECKER 1916) Pecrorossia letho (WiEDEMANN 1828) Exhyalanthrax muscarius (PALLAS 1818) Petrorossia albula ZAITZEV 1962 Veribuboanus (WiEDEMANN 1828) Pipuncubpsis stackelbergi ZAITZEV 2000 Veribubo saffra GREATHEAD 1981 Pipuncubpsis sp. 1 Veribubo misellus (LOEW 1869)

Tribe Anthracini LATREILLE Tribe Exoprosopini BECKER Spogostylum sordidum SACK 1909 Micomitra iris (LOEW 1869) Spogostylum ocyale (WiEDEMANN 1828) Micomitra sp. near chrystallina (BEZZI 1924) Spogostylum griseipenne (MACQUART 1850) Heterabnia (Zygodipla) singularis Spogostylum sp. 1 (MACQUART 1840) Spogostylum nitidum AUSTEN 1937 Heterabnia (Zygodipla) bagdadensis Spogostylum perpusittum AUSTEN 1937 (MACQUART 1840) Spogostylum candidum (SACK 1909) Heterabnia (Zygodipla) hermani Spogostylum isis (ME1GEN 1820) (FRANCOIS 1967) Anthrax virgo EGGER 1859 Heterabnia (Zygodipla) mucorea Anthrax dentata (BECKER 1906) (KLUG 1832) Anthrax aethiops (FABRICIUS 1781) Heterabnia (Acrodisca) suffusa (KLUG 1832) Anthrax greatheadi EL HAWAGRY 1998 Heteralonia (Homobnia) megerlei Anthrax sticticus KLUG 1832 (MEIGEN 1820) Anthrax niger (AUSTEN 1937) Exoprosopa pandora (FABRICIUS 1805) Anthrax candidapex (AUSTEN 1937) Exoprosopa minos (MEIGEN 1804) Exoprosopa rivularis (MEIGEN 1818) Tribe Villini HULL Exoprosopa sp. near pusitia MACQUART 1840 Oestranthrax brunnescens (LOEW 1857) Oestranthrax sp. near alfierii Subfamily Usiinae PARAMONOV 1931 Oestranthrax sp. near pallifrons BEZZI 1926 Tribe Apolysini EVENHUIS Villa ixion (FABRICIUS 1794) Apolysis sp. 1 (Fig. 1a) Villa sp. near laevis BECKER 1915 Material: One specimen: Ayn Aqraba 25.111.02 Villa insignis AUSTEN 1937 (1 9 ). Sweeping vegetation. Villa atricauda AUSTEN 1937 Af>o!ysis is poorly known genus and more Villa sp. near stenozona (LOEW 1869) material of both sexes is required for identi- Villa niphobleta (LOEW 1869) fication. Our specimen is black. Villa bivirgata AUSTEN 1937 Villa sp. 1 Tribe Usiini BECKER Villa fasciculata BECKER 1916 Hemipenthes velutina (MEIGEN 1820) Usia ignorata BECKER 1906 (Fig. 1b) Caecanthrax arabicus (MACQUART 1840) Material: 34 specimens: AI Aridah 4III.02 Thyridanthrax btus (LOEW 1869) (6CO-+699 ) from Pink Flax flowers; Dayr Alia

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18.11.02 (5 99). 21.11.90 (19), 27.111.02 species among Jordanian Bombyliidae (1,7- (799+7ccr); Sayl AI Karak 1.1V.02 (I9); 3,5 mm). Many specimens were picked by Tabaqt fahl 7.111.84 (lcr). hand from the flowers of Pink Flax, Linum Distribution: Afrotropical: Sudan. pubescens BANKS & SOL. at sunset in As Salt Palaearctic: Algeria, Egypt, Palestine, Mo- and from Blue flax Linum peyronii POST. rocco, Spain, Syria, and Tunisia. Parageron gratus (LOEW 1856) (Fig. 1e) Usia ignorata is small black shiny bee fly. It appears to be common in Jordan Valley. Material: 42 specimens: Al Jubayhah 7.111.81 (2O-O-+3 99); Al Mafraq 19.111.01 (19); Dayr Many specimens were collected from the Alia 18.11.02 (499), 4.111.02 (1799), 27.111.02 flowers of Pink Flax, Linum pubescens BANKS (15 99)- & SOL (Linaceae). Distribution: Palaearctic: Canary Is, Egypt, Italy, Morocco, Spain. Usia aenea (Rossi 1794) (Fig. 1c) Material: 140 specimens: AI Aridah 4.II1.02 It appears to be a common beefly. It was (10 99 +2O-O-), from Pink Flax flowers; Al Jubay- collected from the flowers of Palestine hah 26.111.94 (1 9); 3.1V.88 (1 9); Al Mukhybah chamomile, Anthem« pakstina REUT. (Com- Al Fawqa 9.111.02 (37 99 +48o-cr), 11.IH.02 positae) and Poppy Papaver syriacum BoiSS. (299), from Pink and Blue Flax flowers; Al & BLANCHE (Papaveraceae). Colour of Mukhybah At Tahta 11.111.02 (2 99+30-c), mesonotum of available specimens is usually from Pink Flax flowers; Aqraba 25.111.02 with 2 black stripes or may be entirely black. (499+40-cr), Ar Rumman 11.1V.88 (I9), 6.V.O2 (lcr); As Salt 13.1V.02 (19+120-CT), Parageron sp. 1 (Fig. 1f) 16.IV.O2 (4 99 +5aa), from Pink Flax flowers; Irbid 25.111.02 (19); Wadi Shu'ayb 28.1.95 Material: One specimen: Al Jubayhah 7.111.81 (19)- (lcr). Distribution: Palaearctic: Algeria, Aus- Our 4 mm long specimen differs from P. tria, Bulgaria, Croatia, Czech Republic, gratus by having the eyes in contact for the France, Germany, Greece, Hungary, Iran, length of ocellar tubercle (less than the Italy, Morocco, Portugal, Slovakia, Slove- length in P. gratus), black hair fringe behind nia, Spain, Switzerland, Turkey. eyes (white in P. gratus), broad black stripes on mesonotum (narrow in P. gratus). Other It is common in the highlands and Jor- characters are similat to P. gratus. dan Valley. Many specimens were picked by hand from the flowers of Pink Flax, Linum Parageron sp. 2 (Fig. 1g) pubescens BANKS & SOL. at sunset in As Material: One specimen: Al Aridah 4.111.02 (lcr). Salt. It was also collected from Blue Flax, Linum peyronii POST (Linaceae). This specimen is similar to the above species but its eyes are in contact for twice the length of the ocellar tubercle. It was col- Usia bicolor MACQUART 1855 (Fig. 1d) lected from Palestine chamomile, Anthemis Material: 64 specimens: Aira 21.111.94 (1 9); Al pakstina REUT. (Compositae). Jubayhah 25.1V.96 (I9); Al Mukhybah Al Fawqa 9.III.02 (2 99 +3cc); Al Mukhybah At Tahta 11.111.02 (1O-+1 9); Ayn Aqraba 25.111.02 Subfamily Toxophorinae SCHINER (2 99 +2CO-); Ar Rumman 6.V.02 (lcr); As Salt 13.1V.O2 (3099+14CO-); 16.1V.02 (399 + Tribe Toxophorini SCHINER 2o-CT);Zail.V.93(lo-). Toxophora fasciculata (VILLERS 1789) Distribution: Palaearctic: Bosnia-Herce- (Fig. 1h) govina, Croatia, Egypt, Greece, Italy, Kyrgyz Material: Two specimens: Al Mafraq 29.IV.O1 Republic, Lebanon, Palestine, Slovenia, (1 9 ) (Al Al Bayt University); Al Mukhaybah al Syria, Tajikistan, Turkey, Turkmenistan, Fawqa 31.VII.O2 (19) flying. Uzbekistan, Yugoslavia. Distribution: Afrotropical: Sudan. It is a common species of this genus co- Palaearctic: Algeria, Armenia, Austria, existing with other two species in the high- Azerbaijan, Bulgaria, Croatia, Cyprus, land and Jordan Valley. It is the smallest Czech Republic, Egypt, France, Germany,

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Distribution: Palaearctic: Palestine, Spain, Syria.

The males of this species are characterised by the black hairs on the face, while the females have white hairs and can be confused with Geron krymensis or Geron m- tonsus (GREATHEAD 2001).

Geron olivieri MACQUART 1840 Material: 10 specimens: AI Kafrayn 16.XI.95 (lor); As Salt 2.1X.O2 (6crcr) flying; Yajuz 14X02 (lcr+19) flying; Az Zrqa 13.X.1995 (19)- Distribution: Palaearctic: Cyprus, Greece, Iran, Italy, Lebanon, Palestine, Ro- mania, Saudi Arabia, Syria, Turkey, Ukraine.

Geron erythropus BEZZI 1925 (Fig. 2b) Material: Two specimens: Amman 21.IX.81 (19) (Al Yarmouk University); As Salt l.VI- 11.93 (lor). Distribution: Palaearctic: Egypt.

The above female may be the undescribed female of this species. The size, colour, and pale legs are the same as the male. It has been reared from Palpita union- alis HÜBNER (Pyralidae) (YEATES & GREAT- HEAD 1997).

Geron smirnovi ZAITZEV 1978 (Fig. 2c) Material: One specimen: Zahar 17.VI.O2 (lor). Distribution: Palaearctic: Greece, Tajik- istan, Turkmenistan. Fig. 1: Greece, Gruzia, Hungary, Iran, Italy, Kaza- a: Apolysis sp. 1 This species collected from Blue weed khstan, Kyrgyz Republic, Lebanon, Libya, b: Usia ignorata Echium judaeum LACAITA (Boraginaceae). c: Usia aenea Moldova, Morocco, Palestine, Romania, d: Usia bicolor Russia, Slovakia, Slovenia, Spain, Switzer- Geron krymensis PARAMONOV 1929 e: Parageron gratus land, Tajikistan, Turkey, Turkmenistan, f: Parageron sp. 1 (Fig. 2d,e) Ukraine, Uzbekistan. g: Parageron sp. 2 Material: 17 specimens: Ash Shajarah 22.VII.02 h: Toxophora fasciculata It appears to be a rare species. The two (499 )• As Salt 13.VIII.93 (1 9 ); Dibbin 13.V.02 specimens were collected near water. This (1099); Jarash 15.IV.O2 (19); Hummrit As species is easily identified among the Jor- Sahin3.V1.02(l9). danian Bombyliidae by its well-developed Distribution: Palaearctic: Algeria, Ar- pronotum and long antenna. It is a parasite menia, Austria, Azerbaijan, Bosnia-Herze- of Eumenes pompiformis (FABRICIUS) (EU- govina, Bulgaria, Croatia, Cyprus, Czech menidae) DU MERLE (1975). Republic, France, Greece, Gruzia, Hungary, Iran, Italy, Lebanon, Kyrgyz Republic, Tribe Gerontini HESSE Macedonia, Moldova, Palestine, Portugal, Romania, Russia, Slovakia, Slovenia, Spain, Geron mystacinus BEZZI 1924 (Fig. 2a) Syria, Tajikistan, Turkey, Turkmenistan, Material: Two specimens: Irbid, Busailah 2.1V.94 Ukraine, Uzbekistan, Yugoslavia. (2crcr).

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It was collected from Carthamus persicus WlLLD (Compositae), with Amktus validus LOEW.

Subfamily Phthiriinae BECKER

Tribe Phthiriini BECKER Phthiria sp. 1 Material: One specimen: As Salt 7.X.O2 (1 9 ). This specimen cannot be identified to the species because it was partially damaged.

Subfamily Bombyliinae LATREILLE

Tribe Conophorini BECKER Conophorus glaucescens (LOEW 1863) (Fig. 2f) Material: Five specimens: Wadi Shu'ayb, unknown date (19); Yarqa 9.111.98 (4ccr). Distribution: Palaearctic: Albania, Ar- menia, Azerbaijan, Bulgaria, Croatia, Greece, Gruzia, Italy, Moldova, Palestine, Russia, Turkey, Ukraine, Yugoslavia. This species can be easily recognised among the Jordanian Conophorus by having 3 submarginal cells.

Conophorus nobilis (LOEW 1873) (Fig. 2g) Material: Five specimens: Al Jubayhah 23.IV.78 (ley); Al Mujib, Faqu'a 18.111.02 (lcr) on ground; An Nu'ayyimah 11.V.83 (IC+I9); Dibbln 1.V.93 (la). Distribution: Palaearctic: Armenia, Azerbaijan, Gruzia, Iran, Russia, Turkey, Turkmenistan. Fig. 2: All specimens of this species differ from a: Geron mystacinus ENGEL'S (1937) description in having some Legnotomyia cineracea AUSTEN 1937 b: Geron erythropus fine black hairs among the golden hair at (Fig. 3a) c: Geron smirnovi the apex of the abdomen. d, e: Geron krymensis Material: Three specimens: Abu Nusayr 3.VII.02 f: Conophorus glaucescens (lcr) flying; Ayn Almuallaqah 28.V.01 (la); A: g: Conophorus nobilis Legnotomyia trichorhoea (LOEW 1855) Zarqa5.VII.92(l9). h: Legnotomyia trichorhoea. (Fig. 2h) Distribution: Palaearctic: Palestine, Material: Four specimens: Ajlun 5.V.83 (la); Jarash 25.IV85 (19); Muzayrib 24.1V.95 (lcr); Saudi Arabia. Unknown data (lcr). Its antennae and abdomen are similar to Distribution: Palaearctic: Greece, Iraq, the above species except that the abdomen Italy, Macedonia, Palestine, Syria, Yu- brownish instead of black and the yellow goslavia. bands are narrower. This species has antennal pedicel with a projection in males while the female's pedi- Tribe Bombyliini LATREILLE cel has no projection. Its spherical abdomen Systoechus longirostris BECKER 1916 banded with yellow stripes against the black (Fig. 3b) abdomen makes it difficult to be confused Material: 10 specimens: Hummrit As Sahin with any other species of Bombyliidae in 22.1V.O2 (l9+6cra), 3.VI.02 (19); Dana, Ar Jordan. Rummanah 28.IX.O2 (2 99 ).

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(5 99+2cra) (Al Al Bayt University), 12.IV.80 (2oa); Kurayyimah 8.1V.74 (1 9 +la); Muzayrib 24.IV.87 (19) Anthems pokstma, 24.X.95 (19). unknown data (la) (Al Yarmouk University); Na'orl6.IV.74(la+l9). Distribution: Palaearctic: Albania, Ar- menia, Austria, Azerbaijan, Belarus, Bel- gium, Bosnia-Hercegovina, Bulgaria, Croat- ia, Czech Republic, Egypt, Estonia, France, Gibraltar, Greece, Gruzia, Hungary, Iran, Italy, Kazakhstan, Kuwait, Kyrgyz Republic, Latvia, Lebanon, Lithuania, Macedonia, Malta, Oman, Palestine, Poland, Romania, Russia, Saudi Arabia, Slovakia, Slovenia, Spain, Switzerland, Syria, Tajikistan, Turkey, Turkmenistan, Ukraine, United Arab Emirates, Uzbekistan, Yugoslavia.

It appears to be common in Jordan during spring season. Hairs on the lateral sides of first abdominal tergite could be white or black.

Bombylella simulans (AUSTEN 1937) (Fig. 3d) Material: Two specimens: Ar Rumaymin 17.1V.86 (la); Muzayrib 24.IV.95 (la) on Anthimus pcdestina REUT. (Compositae). Distribution: Palaearctic: Lebanon, Palestine.

This species seems to be less common Fig. 3: than B. ana. This species was collected from a: Legnotomyia cineracea Palestine chamomile Anthemis pakstina b: Systoechus longirostris Distribution: Palaearctic: Armenia, REUT. (Compositae). c: Bombylella atra Azerbaijan, Bosnia-Herzegovina, Bulgaria, d: Bombylella simulans Croatia, France, Germany, Greece, Gruzia, e: Bombomyia discoidea Iran, Italy, Kazakhstan, Macedonia, Moldo- Bombomyia discoidea (FABRICIUS 1794) f: Bombomyia stictica (Fig. 3e) g: Bombylius (Bombylius) fuscus va, Portugal, Russia, Slovenia, Spain, h: Bombylius (Bombylius) medius Ukraine, Yugoslavia. Material: Two specimens: Ghawr As Safi 3.XI.81 (l9);TabaqtFahllO.X.99(la). This species has yellowish hairs and its Distribution: Afrotropical: Botswana, wing pattern is sexually dimorphic, females Burundi, Chad, Congo, Eritrea, Ethiopia, with hyaline wings and males with brown Gambia, Ghana, Kenya, Malawi, Mali, infuscation at base. It is very conspicuous Mozambique, Namibia, Niger, Nigeria, during hovering in the field. Senegal, South Africa, Swaziland, Tanzania, Togo, Uganda, Yemen, Zambia, Zimbabwe. Bombylella atra (SCOPOLI 1763) (Fig. 3c) Palaearctic: Algeria, Armenia, Austria, Material: 40 specimens: Al Aridah 29.111.81 Azerbaijan, China, Cyprus, Egypt, France, (la); AI Karak 1.IV.02 (2crcr); Al Mshar'a Greece, Gruzia, Hungary, Iran, Italy, Kaza- 28.11.93 (1 9 ) (Al Yarmouk University), 2.IV.87 khstan, Kyrgyz Republic, Lebanon, Moldo- (19); Al Mujib Reserve 13.111.02 (2ocr); Al va, Mongolia, Morocco, Oman, Palestine, Mukhaybah al Fawqa 9.III.02 (499+5crcr); Ar Russia, Spain, Syria, Tajikistan, Tunisia, Rumaymin 4-V.93 (19); Ruwayhar al Ghawr Turkey, Turkmenistan, Ukraine, Uzbekistan. 15.111.99 (19); As Sukhnah 28.IV.74 (2crcr); Badhan 1.IV.02 (la); Dayr Alia 31.111.74 (1 9 )- It is characterised by white hairs at dis- 8.IV.74 (la), 7.VI1.89 (la); Jarash 2.1V.01 tal extremity of the abdomen. AUSTEN

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(1937) mentioned that this species was seen (la), 21.IV.90 (la), 13.V.74 (19), 7-IX.81 feeding on or poising above white flowers of (19), Unknown data (1 9); AI Mashr'a 31.I.87 HeUotropium vittosum WlLLA (Boragi- (la); Al Mujib Reserve, Faqu'a 18.111.02 (2 99); naceae). Our female specimens have grey AI Mushaqqar 14.V.90 (la+19); Amman pubescence on the head and thorax, while 22.11.94 (19), 12.111.96 (la), 19.111.93 (la), in the specimens from Oman, the grey 20.111.96 (19), 7.1V.97 (19); Ayn Aqraba 25.111.02 (1 9); As Salt 25.111.98 (la); At Tafilah colour is completely replaced by orange 7.IV.97 (1 9); Dayr Alia 14.11.90 (2aa), 23.11.81 brown pubescence (GREATHEAD 1980a). (2aa), 18.111.74 (2 99), 19.111.74 (299); Dead Sea 29.1V.94 (19); Hummrit As Sahin 12.11.01 Bombomyia stictica (FABRICIUS 1794) (la) (Al Balqa University), 13.11.01 (I9) (Al (Fig. 3f) Balqa University), 19.11.00 (la+19) (Al Balqa Material: Four specimens: As Salt 21.474 (1 9); University), 6.111.00 (la) (Al Balqa University); Dayr Alia 6.V.74 (19); Kafr Asad 9.IV.01 (lcr); 20.111.00 (19) (Al Balqa University), 23.111.00 WadiShu'ayb24.1V.97(lcr). (2 99 ) (Al Balqa University), 27.111.00 (la) (Al Balqa University); Jarash 27.11.01 (1 9 ), 11.1V.88 Distribution: Palaearctic: Afghanistan, (la); Rajib 18.IV.00 (I9); Suwaylih 1.111.85 Albania, Algeria, Armenia, Austria, Azer- (19), 2.111.97 (la), 21.111.93 (1 9); Wadi As Sir baijan, Czech Republic, Egypt, France, Ger- 27.111.97 (19), 6.V.84 (la), Unknown data many, Greece, Gruzia, Hungary, Iran, Iraq, (4aa+2 99). Italy, Kazakhstan, Kuwait, Kyrgyz Republic, Distribution: Palaearctic: Afghanistan, Lebanon, Libya, Moldova, Morocco, Pales- Albania, Algeria, Armenia, Azerbaijan, Be- tine, Poland, Portugal, Romania, Russia, larus, Belgium, Bosnia-Herzegovina, Bulgar- Slovakia, Spain, Syria, Tajikistan, Tunisia, ia, Croatia, Cyprus, Czech Republic, Egypt, Turkey, Turkmenistan, Ukraine, United Estonia, France, Germany, Gibraltar, Greece, Arab Emirates. Gruzia, Hungary, Iran, Iraq, Italy, Kaza- This species characterised by two rows khstan, Kyrgyz Republic, Latvia, Lebanon, of white spots on the underside of the ab- Libya, Lithuania, Macedonia, Malta, Moldo- domen. va, Morocco, Netherlands, Palestine, Poland, Portugal, Romania, Russia, Slovakia, Slove- Bombylius (Bombylius) fuscus FABRICIUS nia, Spain, Sweden, Switzerland, Syria, 1781(Fig. 3g) Tajikistan, Tunisia, Turkey, Turkmenistan, Ukraine, Uzbekistan, Yugoslavia. Material: Two specimens: Ayn Almuallaqah 3.111.97 (10"); unknown location and date (la). B. medius is very common in Jordan. Distribution: Palaearctic: Armenia, Large numbers were collected during March Azerbaijan, Egypt, Greece, Gruzia, Iran, and April. This species exhibits variation in Iraq, Italy, Palestine, Portugal, Spain, Turk- length ranging from 7,2-11,3 mm, and in menistan. the amount and extent of black hairs on the body. However, it can be easily recognised It seems to be a rare species in Jordan. It by the pattern of its spotted wings. There is can be recognised by the completely black one record from the nest of an Andrena sp. body without any white or pale marks, also (Andrenidae) (DU MERLE 1975). by having completely black infuscated wings. Bombylius {Bombylius) trichurus PALLAS 1818 (Fig. 4a) Bombylius (Bombylius) medius Material: Two specimens: As Salt 19.1V.00 (la); LINNAEUS 1758 (Fig. 3h) AlJubayhahlO.IV.79 (la). Material: 85 specimens: AI Badhiyah 13.1II.O2 Distribution: Afrotropical: Yemen. (2 99); AlJubayhah6.II.95 (4aa), 9.II.95 (lc), Palaearctic: Albania, Armenia, Austria, 22.11.93 (lo-), 23.11.81 (2 99 +lcr), 27.11.90 (lcr), 3.111.90 (1O-), 3.I1I.O2 (la), 5.111.95 (1 9), 7.111.8 Azerbaijan, Bulgaria, Croatia, Cyprus, (lo-), 10.111.90 (1O-), i4.iii.90 (lcr), 17.111.89 Czech Republic, France, Greece, Gruzia, (2 99), 19.111.94 (1 9 + 1^) on grasses, 21.111.89 Hungary, Iran, Italy, Lebanon, Macedonia, (la), 22.111.89 (2aa), 22.111.95 (la), 23.111.94 Moldova, Oman, Palestine, Poland, Roma- Ocrcr), 25.111.74 (19). 27.111.89 (la), 27.111.97 nia, Russia, Saudi Arabia, Slovakia, Spain, (299), 4.IV.87 (la), ll.IV? (la), 13.IV.97 Syria, Turkey, Turkmenistan, Ukraine.

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Distribution: Palaearctic: Afghanistan, Armenia, Austria, Azerbaijan, Belarus, Bel- gium, Cyprus, Czech Republic, Egypt, Esto- nia, France, Germany, Greece, Gruzia, Hun- gary, Iran, Italy, Kazakhstan, Kyrgyz Repub- lic, Latvia, Lithuania, Moldova, Morocco, Palestine, Poland, Romania, Russia, Slova- kia, Spain, Syria, Tajikistan, Tunisia, Turkey, Turkmenistan, Ukraine, Uzbekistan, Yugoslavia.

B. fimbriatus is a common species in Jor- dan. Its wing pattern has a sharp fore border and transparent area in the marginal cell. It is very close to B. major but the latter lacks the transparent area in the marginal cell.

Bombylius {Bombylius) major LINNAEUS 1758 (Fig. 4c) Material: Two specimens: Al Jubayhah 22.IV.97 (la); Amman 24.11.95 (1 9 ) collected from al- mond flowers. Distribution: Nearctic: Canada, USA, Mexico. Oriental: Bangladesh, India, Nepal, Pakistan, and Thailand. Palaearctic: Alba- nia, Algeria, Armenia, Austria, Azerbaijan, Belarus, Belgium, Bosnia-Herzegovina, Bul- garia, China, Croatia, Cyprus, Czech Re- Fig. 4: public, Denmark, Egypt, Estonia, Finland, a: Bombylius (Bombylius) trichurus France, Germany, Greece, Gruzia, Hungary, b: Bombylius {Bombylius) fimbriatus This species seems to be rare in Jordan. Ireland, Italy, Japan, Kazakhstan, Korea, c: Bombylius {Bombylius) major The black hairs on the underside of ab- Latvia, Libya, Lithuania, Luxembourg, Mal- d: Bombylius {Bombylius) modestus domen can easily separate this species from e: Bombylius {Bombylius) pumilus ta, Macedonia, Moldova, Mongolia, Moroc- f: Bombylius {Bombylius) fulvescens similar ones in Jordan that have pale hairs. co, Netherlands, Norway, Poland, Portugal, g: Bombylius {Bombylius) canescens It has been reared from Evylaeus nigripes (LE- Romania, Russia, Slovakia, Slovenia, Spain, h: Bombylius {Bombylius) mendax PELETIER) (DU MERLE 1975). Sweden, Switzerland, Tajikistan, Tunisia, Turkey, Turkmenistan, United Kingdom, Bombylius (Bombylius) fimbriatus Uzbekistan, Yugoslavia. MEIGEN 1820 (Fig. 4b) This species seems to be a rare species in Material: 40 specimens: AI Juhayhah VIII.02 Jordan. In the United States, it is very abun- (la), 15.111.99 (19), 19.111.93 (19), 19.111.98 dant on plum trees and on Cardamine bul- (la), 22.111.95 (2aa+l9), 30.111.93 (la), bosa (SCHREB). The immature stages attack 1.IV.82 (la), 2.IV.97 (2aa), 3.IV.93 (la), 4.1V.98 (la), 6.1V.99 (la), 14.IV.74 (la), Andrena spp. (Andrenidae) and Halictus 15.1V.75 (I9), 19.IV.98 (la), 25.IV.93 (la); (Halictidae) (DU MERLE 1975, YEATES & I.V.? (la), 2.V.98 (la+1 9 ), 3.V.98 (1 9); Am- GREATHEAD 1997). man 3.1V.90 (19); Ayn Aqraba 25.111.02 (2 99+2oa); Ayn Almuallaqah 3.1V.97 (la); Bombylius (Bombylius) modestus Dana 4.1V.94 (la); Dayr Alia 23.111.88 (la), LOEW 1873 (Fig. 4d) 20.IV.98 (la), 22.IV.84 (la); Irbid 24.111.84 Material: Seven specimens: AI Mujib 2411I99 (19) (AI Yarmouk University); Marka 8.IV.94 (19). 3.V.99 (19); Ar Rumman 6.V.02 (19); Tabaqt fahl 22.111.00 (la); Wadi Shu'ayb (la+1 9) on the ground; Ayn esh Shallaleh 1.V.97 (la); Zai I.V. 93 (1 9 ); Unknown data 24.VI.02 (la) flying; Na'or 5.VIII.O2 (1 9 ) on (2oa+l 9). the ground; Unknown data (1 9 ).

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Distribution: Afrotropical: Ethiopia, Su- Al Mashari'a 1.111.83 (la); Al Mukhaybah al dan, Yemen. Oriental: Pakistan, Palaearctic: Fawqa 9.111.02 (8oo+4aa); Ayn Aqraba Afghanistan, Armenia, Azerbaijan, Cyprus, 25.III.O2 (2 99+la); As Salt 5.111.74 (19); Egypt, Greece, Gruzia, Iran, Italy, Kaza- Dayr Alia 5.IV.98 (19); Jarash 15.IV.O2 (1 9). khstan, Macedonia, Russia, Saudi Arabia, Distribution: Palaearctic: Albania, Ar- Tajikistan, Turkmenistan, Uzbekistan, Yu- menia, Austria, Azerbaijan, Belarus, Bel- goslavia. gium, Bosnia-Hercegovina, Bulgaria, Croat- ia, Cyprus, Czech Republic, Estonia, France, Only the males of this species have Germany, Greece, Gruzia, Hungary, Ireland, white scales at apex of abdomen, while in B. Italy, Latvia, Libya, Lithuania, Malta, posticus both sexes have such scales. Moldova, Netherlands, Poland, Romania, Russia, Slovakia, Slovenia, Spain, Switzer- Bombylius (Bombylius) pumilus land, Syria, Turkey, Turkmenistan, United MEIGEN 1820 (Fig. 4e) Kingdom, Ukraine, Yugoslavia. Material: Four specimens: Al Mukhaybah al Fawqa 9.1II.02 (la); Jarash 12.IV.80 (la); Muza- Bombylius canescens appears to be com- yrib 24.1V.95 (la); Unknown data (la). mon during spring season where many spec- Distribution: Afrotropical: Yemen. imens were collected in March. This species Palaearctic: Albania, Algeria, Armenia, has been reared from nests of Odynerus reni- Azerbaijan, Egypt, France, Germany, formis (GMELIN) (Eumenidae), also several Greece, Gruzia, Hungary, Iran, Iraq, Italy, species of Halictidae (DU MERLE 1975). Kuwait, Lebanon, Macedonia, Moldova, Oman, Palestine, Poland, Portugal, Roma- Bombylius (Bombylius) mendax nia, Russia, Saudi Arabia, Spain, Syria, AUSTEN 1937 (Fig. 4h) Turkey, Ukraine, United Arab Emirates. Material: 22 specimens: Al Jubayhah 3.111.02 (la), 8.111.89 (la), 22.111.95 (la), 23.111.94 This species has the shortest proboscis Ocra), 8.1V.93 (la), 15.V1I.97 (la); As among all Jordanian Bombylius. The wing Sukhnah l.IV? (la); Ayn Almuallaqah 3.IV.94 patterns are sexually dimorphic; females (la); Dana 4.1V.94 (3crcr); Dayr Alia 14.11.90 with hyaline wings and males with basal in- (la); Irbid 6.111.84 (la) (Al Yarmouk Universi- fuscation. ty); Jarash 7.111.90 (2aa+l9); Sayl Al Karak 1.IV.02 (19); Wadi As Sir 21.111.96 (la), Bombylius (Bombylius) fulvescens 27.111.97 (19); Yajuz 5.IV.88 (la). WIEDEMANN 1820 (Fig. 4f) Distribution: Palaearctic: Palestine. Material: Two specimens: As Salt 31.V.93 (1 9 ); DayrAUa8.IV.74(la). Bombylius mendax is known so far from Jordan and Palestine, which appears native Distribution: Palaearctic: Albania, Ar- to this area. The black hairs along notopleu- menia, Austria, Azerbaijan, Bosnia-Herze- ra distinguish this species from B. canescens, govina, Bulgaria, China, Croatia, Czech Re- and presence of black hairs on the postocu- public, Greece, Gruzia, Hungary, Iran, Iraq, lar margin distinguish it from B. cinerascens. Italy, Kazakhstan, Macedonia, Malta, Moldova, Palestine, Poland, Portugal, Ro- mania, Russia, Saudi Arabia, Slovakia, Bombylius (Bombylius) cinerascens Slovenia, Spain, Switzerland, Tajikistan, MIKAN 1796 (Fig. 5a) Tunisia, Turkey, Turkmenistan, Ukraine, Material: 11 specimens: Al Jubayhah 17.111.95 Uzbekistan, Yugoslavia. (la), 1O.1V.79 (19), 23.IV9O (la); Al Mukhay- bah al Fawqa 9.111.02 (399); Ayn Aqraba It is very similar to B. pumilus but it can 25.111.02 (la); As Salt 24.IV.94 (19); As be distinguished from it by its longer pro- Sukhnah 10.IV.79 (19); Ayn esh Shallaleh boscis. 2O.IV.98 (la); Dayr Alia 8.IV.74 (19). Distribution: Palaearctic: Albania, Ar- Bombylius (Bombylius) canescens menia, Austria, Azerbaijan, Belgium, MIKAN 1796 (Fig. 4g) Bosnia-Herzegovina, Bulgaria, Croatia, Material: 24 specimens: Al Jubayhah 23.11.95 Czech Republic, Egypt, France, Germany, (lo+la); 19.111.94 (Hj+lc); 22.111.95 (la); Greece, Gruzia, Hungary, Italy, Macedonia,

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garia, Croatia, Cyprus, Czech Republic, Denmark, Egypt, France, Germany, Gibral- tar, Greece, Gruzia, Hungary, Iran, Iraq, Italy, Kazakhstan, Macedonia, Moldova, Morocco, Poland, Portugal, Romania, Rus- sia, Slovakia, Slovenia, Spain, Switzerland, Syria, Tajikistan, Tunisia, Turkey, Turk- menistan, Ukraine.

The white scales at apex of abdomen may be difficult to see when the specimens are in good condition (with many hairs) (AUSTEN 1937). The larvae were found par- asitising Panurgus denapes LATREILLE and P. cakaratus (SCOPOLI) (Andrenidae) and also Evylaeus nigripes LEPELETIER (Halictidae) (DU MERLE 1975).

Bombylius (Zephyrectes) quadrifarius LOEW 1855 (Fig. 5c) Material: Four specimens: Al Mafraq 19.111.01 (lor); Al Mujib 13.111.02 (2 99 +lcr). Distribution: Palaearctic: Albania, Ar- menia, Azerbaijan, Bulgaria, China, Czech Republic, Greece, Gruzia, Hungary, Iran, Italy, Macedonia, Moldova, Palestine, Ro- mania, Russia, Slovakia, Tajikistan, Turkey, Ukraine, Yugoslavia.

This species has two longitudinal stripes formed by white hairs on the mesonotum, and transverse bands formed by white hairs at the posterior margin of abdominal ter- Moldova, Palestine, Poland, Romania, Rus- gites. Wings are sexually dimorphic; females sia, Saudi Arabia, Slovakia, Slovenia, have hyaline ones and males with basal in- Spain, Switzerland, Turkey, Ukraine, Yu- fuscation. goslavia. Fig. 5: a: Bombylius (Bombylius) cinerascens This species seems to be less common Dischistus syriacus (VILLENEUVE 1912) b: Bombylius (Bombylius) posticus than B. canescens and B. mendax since few (Fig. 5d) c: Bombylius (Zephyrectes) quadrifarius d: Dischistus syriacus specimens were collected. These three Material: 23 specimens: AI Adasiyah 11.VI1.94 e. Neobombylodes giganteus species are very similar in general appear- (lcr); Al Aridah 29.111.81 (lor); Al Ghawr f: Anastoechus trisignatus ance, but B. cinerascens lacks black hairs on 5.1II.78 (la); As Sukhnah 28.IV.74 (2aa); Ayn g: Anastoechus bahirae the postocular margin. The wings pattern is Aqraba 29.IV.02 (la+19); Ayn Qantarh h: -Anastoechus nitidulus 16.V.87 (la) (Al Yarmouk University); Dayr Al- sexually dimorphic. The wing base is mum- ia 19.111.78 (la), 27.111.74 (1 9), 31.111.74 (1 9); my-brown in males and light sepia-coloured Hummnt As Sahin 3.VI.02 (19); Jarash in females. 22.IV.81 (la), 6.V.02 (2aa+l9); Kafr Asad 9.IV.01 (1 9 ) (Al Al Bayt University); Wadi Al Bombylius (Bombylius) posticus Arab 3.V.82 (2oa) (Al Yarmouk University); FABRICIUS 1805 (Fig. 5b) Wadi Shu'ayb ?.V97 (2 99), Unknown data (2aa). Material: Six specimens: AI Jubayhah 8.V.84 (la); Al Wala 8.IV.02 (1 9 +4oa). Distribution: Palaearctic: Lebanon, Palestine. Distribution: Palaearctic: Afghanistan, Albania, Algeria, Armenia, Austria, Azer- Dischistus syriacus appears to be a native baijan, Belgium, Bosnia-Hercegovina, Bul- species in the Eastern Mediterranean. It is a

364 © Biologiezentrum Linz/Austria; download unter www.biologiezentrum.at common species in Jordan. It is sexually di- 30.111.00 (19), Um Ishrin 31.111.00 (2crar), morphic; males have whitish hairs while fe- 1.1V.00 (2O-O-+1 9 ); Wadi An Nawatif 27.IX.O2 males have yellowish hairs. (lcr+3 99); Wadi shuqayq 24.IX.O2 (3o*o-+l 9); Unknown data (2crcr).

Neobombylodes giganteus (VILLENEUVE Distribution: Palaearctic: Afghanistan, 1920) (Fig. 5e) Austria, Belgium, Bosnia-Herzegovina, Bul- Material: Seven specimens: Al Aridah 18.111.79 garia, China, Croatia, France, Germany, Greece, Hungary, Iran, Italy, Japan, Kaza- (3 99); Al Mujib 18.111.02 (2o-o-); Dayr Alia 31.111.74 (lcr); Hummrit As Sahin 26.111.00 khstan, Kyrgyz Republic, Macedonia, Mon- (19)- golia, Portugal, Romania, Russia, Slovenia, Spain, Slovakia, Turkey. Distribution: Palaearctic: Algeria, Egypt, Libya. It appears to be common in dry regions This robust species is sexually dimor- of Jordan. It was collected from Loctuca ori- phic; males have elongate oval abdomen entalis BoiSS (Compositae) and Polygonum while females have short spherical ab- equisetiforme SlBTH (Polygonaceae). The lar- domen. vae of this species were reared from Locusta migratoria (LINNAEUS) and other species of Acrididae (DU MERLE 1975). Anastoechus trisignatus (PORTSCHINSKY 1881) (Fig. 5f) Anastoechus exalbidus Material: One specimen: Yajuz 14X.02 (1 9) on (WIEDEMANN 1820) (Fig. 6a) the ground. Material: Five specimens: Rum, Urn Ishrin Grid Distribution: Afrotropical: Sudan, 87 31.111.00 (2 99), Wadi Marsad 27.111.00 Yemen. Palaearctic: Algeria, Armenia, (2 99 ), 1.IV.00 (1 9 ) (Al Al Bayt University). Azerbaijan, Bosnia-Herzegovina, Croatia, Distribution: Afrotropical: Sudan, Egypt, Greece, Gruzia, Italy, Jordan, Kuwait, Yemen. Palaearctic: Algeria, Armenia, Lebanon, Libya, Macedonia, Morocco, Azerbaijan, Egypt, France, Gruzia, Italy, Jor- Oman, Palestine, Qatar, Saudi Arabia, dan, Kuwait, Lebanon, Oman, Palestine, Slovenia, Syria, Tunisia, Yugoslavia. Portugal, Saudi Arabia, Spain, Syria, This species is easily recognised by a Tunisia, Yugoslavia. longitudinal median stripe of oppressed It is common in desertic regions of Jor- scales on the body. This species was record- dan. It was previously recorded from Jordan ed previously from Jordan (EVENHUIS &. (EVENHUIS & GREATHEAD 1999). GREATHEAD 1999).

Subfamily Ecliminae HALL Anastoechus bahirae BECKER 1915 (Fig. 5g) Eclimus gracilis LOEW 1844 (Fig. 6b) Material: 12 specimens: AI Muwaqqar 16.IX.O2 Material: One specimen: Kufrinjah, unknown (1099), 7.X.O2 (19); Dana, Ar Rummanah date and month. 99 (lc). 28.1X.O2 (la). Distribution: Palaearctic: France; Distribution: Afrotropical: Kenya, Su- Greece, Italy, Libya, Macedonia, Morocco, dan. Palaearctic: Algeria, Egypt, Libya, Palestine, Turkey, Yugoslavia. Palestine, Saudi Arabia, Spain, Syria, This species seems to be a rare species. It Tunisia. is black with an extremely narrow body. The It seems to be common in the desertic head is semibilobate but there is no concav- regions of Jordan. It was collected from Poly- ity. It is almost mosquito like in appearance gonum equisetiforme SlBTH (Polygonaceae). except for the spotted wings.

Subfamily Cythereinae BECKER Anastoechus nitidulus (FABRICIUS 1794) (Fig. 5h) Callostoma fascipenne Material: 41 specimens: Al Muwaqqar 7.X.O2 MACQUART 1840 (Fig. 6c) (6CTO-+13 99); Ar Rummanah 28.IX.O2 Material: Eight specimens: Al Aridah 4-V.81 (2O-O-+2 99); Dana 23.111.95 (lcr+19); Rum (19+lOf); Al Balqa University 28.111.01

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Distribution: Palaearctic: Albania, Ar- menia, Austria, Azerbaijan, Bosnia-Herie- govina, Bulgaria, Croatia, Czech Republic, France, Greece, Gruzia, Iran, Italy, Lebanon, Macedonia, Moldova, Morocco, Palestine, Portugal, Russia, Slovakia, Slove- nia, Spain, Syria, Tajikistan, Turkey, Turk- menistan, Ukraine, Yugoslavia.

This species exhibits unstable venation in some specimens, some have a supernumerary anterior transverse veins in each wing (AUSTEN 1937). It is recorded as a predator on the eggs of CaMptamus italicus (LINNAEUS) and Dociostaurus maroccanus (THUNBERG) (DU MERLE 1975).

Cytherea aureus FABRICIUS 1794 (Fig. 6e) Material: Three specimens: Dayr Alia 8.IV.74

(ltf+1 o ); AI Mujib 13.1II.02 (lcr). Distribution: Palaearctic: Algeria, Egypt, Italy, Jordan, Libya, Morocco, Pales- T/v tine, Tunisia.

This species is readily recognisable ow- ing to its unusual brownish yellow tawny colour; and the black hairs are confined to ocellar tubercle (AUSTEN 1937). It was recorded from Jordan previously (EvENHUIS & GREATHEAD 1999).

Cytherea fenestrata (LOEW 1873)

Fig. 6: (Fig. 6f) a: Anastoechus exalbidus Material: Nine specimens: Al Jubayhah 13.V74 b: Eclimus gracilis (19), 23.V.92 (19); Amman 15.V.97 (lcr), c: Callostoma fascipenne + 16.V.78 (1 9 ); Dayr Alia 8.IV.74 (1 9 ); Humm- d: Cytherea obscura (Icr+19); Ar Rumman 13.V.91 Ü9 l°"). rit As Sahin 21.111.01 (lcr), 21.IV.00 (I9) (Al e: Cytherea aureus 6.VI.88 (icy); Jarash 1.VI.80 (1 9 ). f: Cytherea fenestrata Balqa University); Wadi Shu'ayb 14.IV? (1 9); g: Cytherea nucleorum Distribution: Palaearctic: Afghanistan, unknown data (1 9 ). h: Cytherea delicata Armenia, Azerbaijan, Greece, Gruzia, Iran, Distribution: Afrotropical: Yemen. Palae- Italy, Macedonia, Palestine, Turkey, Turk- arctic: Egypt, Kuwait, Morocco, Oman, Sau- menistan, Uzbekistan, Yugoslavia. di Arabia, Syria, Tajikistan, Turkey, Turkme- Adults of this species suck the nectar of nistan, United Arab Emirates, Uzbekistan. flowers especially of the Pink Scabious and This species is separated by the heavy Thistle (AUSTEN 1937). The larvae of these black bristles on the abdomen from C. bar- flies are important consumers of egg pods of bora, which has pale bristles. locusts (Acrididae) (AUSTEN 1937, DU MER- LE 1975). Cytherea nucleorum (BECKER 1902) (Fig. 6g) Cytherea obscura FABRICIUS 1794 Material: Five specimens: Al Mujib Natural Re- (Fig. 6d) serve 13.111.02 (4crcr); Dayr Alia 19.111.74 (1 9 ). Material: Three specimens: AI Jubayhah 13.V.74 Distribution: Palaearctic: Egypt, Jordan, (lcr); Ar Rumman 18.V.01 (lor); Nahlah Kuwait, Libya, Palestine, Saudi Arabia, Syr- 9.VII.74 (!

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This species is characterised by the mot- tled wings. It was collected from the ground surface. It was recorded from Jordan previously (EVENHUIS & GREATHEAD 1999).

Cytherea delicata (BECKER 1906) (Fig. 6h) a Material: One specimen: AI Bahhath 20.V.02 (19). Distribution: Afrotropical: Yemen. Palaearctic: Algeria, Egypt, Iran, Jordan, Kuwait, Morocco, Oman, Palestine, Saudi Arabia, Tunisia, Turkey, United Arab Emi- rates. This species characterised by coarse, broad linear and truncate scales, thick sparse bristles, and by the white scales on the pleura. It was recorded from Jordan previously (EVENHUIS & GREATHEAD 1999).

Cytherea dispar (LOEW 1873) (Fig. 7a) Material: Nine specimens: AI Jubayhah 3.V.92 (lor), 12.V.O2 (la); AI Mujib, Faqu'a 18.I1I.02 (19): Al Wala 8.IV.02 (lc); As Sukhnah 28.1V74 (2oa+l 9); Dayr Alia 19.111.74 (lor); Yajuz28.IV.74 (la). Distribution: Palaearctic: Afghanistan, Algeria, Armenia, Azerbaijan, Gruzia, Iran, Jordan, Kyrgyz Republic, Morocco, Tajik- istan, Tunisia, Turkey, Turkmenistan, \ > Uzbekistan. Chalcochiton pallasii This species was recorded previously (LOEW 1856) (Fig. 7c,d) from Jordan (EVENHUIS & GREATHEAD Material: 26 specimens: Amman 4.V.90 (1 9 ); 1999). As Salt 29.1V.94 (la) Ant/i«mis polestina R.EUT; Ayn Aqraba 25.111.02 (699+3oa), 29.IV.02 (la+1 9 ); Ar Rumaymin 17.IV.86 (1 9 ); Dayr Fig. 7: Chakochiton speciosus (LOEW 1844) Alia 25.111.90 (la); Jarash 15.IV.O2 (la), a: Cytherea dispar (Fig. 7b) 25.IV.85 (la), 3.V.90 (la); Kurayyimah 8.IV.74 b: Chalcochiton speciosus Material: Eight specimens: AI Aridah 4.III.O2 cd: Chalcochiton pallasii (4oa); Murayrib 24.IV.95 (2 99) Anthemis (2 99); Al Mujib 18.111.02 (la); Ayn Aqraba t: Chalcochiton syriacus pakitma REUT.; Na'or 16.IV.74 d9); Wadi f: Amictus obliquenotatus 25.111.02 (lcr+2 99 ); Jarash 15.IV.O2 (la); Wadi Shu'ayb 14.IV. ?(la). g: Amictus validus Shu'ayb 1.V.97 (la). h: Amictus sp. near tigrinus Distribution: Palaearctic: Armenia, Distribution: Palaearctic: Afghanistan, Azerbaijan, Bulgaria, Greece, Gruzia, Iran, Iran, Turkey. Italy, Kyrgyz Republic, Macedonia, Moldo- Males of this species have dense white va, Morocco, Poland, Romania, Russia, scales at all abdominal tergites, while fe- Slovenia, Syria, Tajikistan, Turkmenistan, males have white scales confined to 1" ter- Ukraine, Uzbekistan. gum and small spots at sides of 2nd, 6th and 7th terga, the remainder is black with bronzy This species is common in the highlands glint. and Jordan Valley. One specimen of this species was collected from Anthemis pakstina REUT. Females of C. pallasii are easily recognised by the orange hairs at the lateral sides

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Amictus validus LOEW 1869 (Fig. 7g) Material: 29 specimens: AI Airaq Natural Re- serve 17.V.O2 (19); Ar Ramtha 29.VI1.02 (3

This species is a common species in the northern mountains of Jordan. It was collected from Carthamus persicus WlLLD (Compositae). Males have a remarkably long bristle on the under side of the fröre tarsus just in front of the tip, while in the females there is a pair of bristles, which are longer. Males have dense black hairs surrounding the genitalia which can be easily seen without dissection. This species exhibits variation in the length from 5,9-11,5 mm.

Amictus sp. near tigrinus AUSTEN 1937 (Fig. 7h) Material: One specimen: AI Bahhath 2O.V.O2 l! or l abdominal tergite, while males have (19). nJ th white hairs at sides of 2 to 6 abdominal Distribution: Palaearctic: Palestine. tergites and with black hairs on the last two It seems to be rare species in Jordan. tergites. This species may be looked upon as inter- Fig. 8: a: Amictus sp. near zinamominus mediate (according to the first posterior b: Amktus virgatus Chalcochiton syriacus (LOEW 1869) cell) between A. virgatus, which has first c: Amiaus sp. near p/crus (Fig. 7e) posterior cell open and A. validus, which has d: Lomatia abbreviata Material: One specimen: AI Wala 8.IV.02 (1 9 )• first posterior cell closed and stalked. How- e: Lomatia polyzona f: Lomatia tysiphone Distribution: Palaearctic: Libya, Pales- ever, A. tigrinus has the first posterior cell closed near the wing margin (AUSTEN g: Cononedys stenura tine, Syria, Turkey. h: Cononedys inornata 1937). This specimen may be aberrant since it has three submarginal cells in the left Amictus obliquenotatus AUSTEN 1937 wing and four in right wing. (Fig. 7f) Material: 13 specimens: Kufrinjah 27.V.02 (1 9 ); Amictus sp. near zinamominus BECKER AI Wala 8.IV.02 (19); Ar Rumman 27.V.O2 1906 (Fig. 8a) (lcr+7 99 ); Dibbin 13.V.02 (2crer+l 9 ). Material: Five specimens: AI Wala 8.IV.02

Distribution: Palaearctic: Palestine. (2 99+2ocf);Zahar 17.VI.O2 (lor). This species has previously only been Distribution: Palaearctic: Algeria, recorded from Palestine. It is distinguishable Libya, Tunisia. by the ground colour of the body, which is This species was described from Algeria black or blackish brown while other species and not reported so far further east than of Jordanian Amictus have yellowish brown. Libya. Possibly this is a new species.

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Amictus virgatus AUSTEN 1937 (Fig. 8b) Lomatia tysiphone LOEW 1860 (Fig. 8f) Material: Four specimens: Ar Rumman 6.V.O2 Material: Ten specimens: Al Jubayhah 13.V.74 (2 99); Zahar 17.VI.O2 (2crcr). (lcr), 22.V.88 (3crcr), 23.V.92 (lcr); Amman 7.IV.96 (lcr); Birqish 14.V.01 (1 9 ) (Al Al Bayt Distribution: Palaearctic: Afghanistan; University); Jarash 12.V.83 (la); Rihaab 7.V.01 Palestine; Turkey. (1 9 +la) (Al Al Bayt University). Distribution: Palaearctic: Albania, Al- Amictus sp. near pictus LOEW 1869 geria, Armenia, Azerbaijan, Bulgaria, (Fig. 8c) France, Gibraltar, Greece, Gruzia, Hungary, Material: Two specimens: Hummrit As Sahin Italy, Libya, Macedonia, Moldova, Pales- 3.V1.O2 (1 9); Al Mafraq 21.V.01 (1 9). tine, Romania, Russia, Spain, Syria, Turkey, Distribution: Palaearctic: Algeria, Ar- Turkmenistan, Ukraine, Yugoslavia. menia, Azerbaijan, Bosnia-Hercegovina, The yellow bands on abdominal tergites Bulgaria, Croatia, Greece, Gruzia, Italy, in both sexes are interrupted in the middle. Macedonia, Slovenia, Yugoslavia.

It is the only Amictus species in Jordan Subfamily Anthracinae LATREILLE that has black scales. The specimens are not in sufficiently good condition to allow for Tribe Aphoebantini BECKER positive identification to the specific level. Cononedys stenura (LOEW 1871) (Fig. 8g) Subfamily Lomatiinae SCHINER Material: Ten specimens: Al Jubayhah 11.VI.79 (19), 13.V1.93 (lcr); Ar Rumaymin 3.VII.O2 Tribe Lomatiini SCHINER (5O-CT+19); Ayn esh Shallaleh 16.V97 (la);

Lomatia abbreviata VILLENEUVE 1911 Mahis 15.VII.02 (lcr). (Fig. 8d) Distribution: Oriental: Pakistan. Material: Ten specimens: AI Aridah 18.I1I.79 Palaearctic: Palestine, Tajikistan, Turk- (2oa), 13.1V.79 (lcr); Hummrit As Sahin menistan, Uzbekistan. 22.IV.O2 (la); Kafr Asad 9.IV.01 (3 99 ) (AI AI Bayt University); Kurayyimah 8.1V.74 (3O-O-). Cononedys inornata (GREATHEAD 1967) Distribution: Palaearctic: Algeria, (Fig. 8h) Greece, Italy, Morocco, Palestine, Romania, Material: Four specimens: Dana 23.IX.O2 Spain, Syria, Tajikistan, Tunisia, Turkey, (2 99 + lcr). AI Muwaqqar 7.X.O2 (1 9 ) Polygon- um argenstavum SlBTH. Turkmenistan. Distribution: Palaearctic: Eritrea, Saudi This species is distinguished from other Arabia. Lomatia in Jordan by black hairs on male's body and by the orange lateral extremity of Tribe Prorostomatini HULL the female's 2nd abdominal tergite. Stomylomyia europaea (LOEW 1869) (Fig. 9a) Lomatia polyzona LOEW 1869 (Fig. 8e) Material: Three specimens: Ayn Aqraba 29.IV.02 Material: Seven specimens: AI Jubayhah 23.V.92 (2 99 ); Hummrit As Sahin 3.V1.O2 (1 9 )• (19); Al Mushaqqar 28.V.90 (I9); Amman Distribution: Palaearctic: Afghanistan, 12.VI.78 (19); Dayr Alia 25.V.74 (19); Irbid Albania, Armenia, Azerbaijan, Egypt, 6.V.86 (2 99); Marj AI Hamam 15.IV.98 (1 9 ). France, Greece, Gruzia, Hungary, Iran, Italy, Distribution: Palaearctic: Albania, Ar- Macedonia, Moldova, Palestine, Romania, menia, Azerbaijan, Greece, Gruzia, Italy, Russia, Turkey, Ukraine, Yugoslavia. Macedonia, Romania, Syria, Turkey, Yu- goslavia. Jordanian specimens vary in having mostly yellow legs and all bristles on the The yellow bands in this species are uni- thorax in front of wings are pale. This form in width while the other Lomatia species is difficult to see in the field since its species known from Jordan has interrupted colour resembles the ground colour. It has bands. great variation in size (AUSTEN 1937).

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Distribution: Oriental: Pakistan. Palaearctic: Afghanistan, Egypt, Iran, Saudi Arabia.

Xeramoeba sabulonis (BECKER 1906) (Fig. 9e) a Material: Five specimens: Al Bahhath 20.V.02 (2 9 9); Dana 23.IX.O2 (19); Hummrit As Sahin 3.VI.02 (1 9 ); 22.1V.O2 (1 9 ). Distribution: Afrotropical: Sudan. Palaearctic: Algeria, Egypt, Greece, Mace- donia, Palestine, Tunisia, Turkey, Yu- goslavia.

Xeramoeba salwae EL HAWAGRY 2001 Material: Two specimens: Al Bahhath 2O.V.O2 (19+10'). Distribution: Palaearctic: Egypt.

Petrorossia hespera (Rossi 1790) (Fig. 9f) Material: 19 specimens: Al Badhyah 25.IX.O2 (1 9 ); AI Husson 19.VI1I.02 (lor); Al Mukhay- bah al Fawqa 31.V1I.O2 (lcr); Al Mujib 3.V.99 (lor); AI Muwaqqar 16.IX.02 (1 9 ); Ayn Aqraba 29.IV.02 (2 99+2O-O-); Ar Rumaymin 17.IV.86 (10.+1O-); Ash Shajarah 22.VI1.02 (lcr); Ash Shawbak 7.V111.O2 (1 9 ); As Salt 2.IX.02 (lcr); Fig. 9: Hummrit As Sahin 22.IX.02 (1 9 ); Ma'ain 5.VI- a: Stomylomyia europaea Plesiocera algira MACQUART 1840 11.02 (2 99+lor); Western Bayudah 26.VIII.02 b: Plesiocera algira (Fig. 9b) (19). c: Desmatoneura sp. 1 d: Xeramoeba semirufa Material: Four specimens: AI Bahhath 20.V.02 Distribution: Palaearctic: Algeria, Ar- (lcr+19); Ma'ain 5.VIII.02 (1?); Western e: Xeramoeba sabulonis menia, Austria, Azerbaijan, Bosnia-Herce- f: Petrorossia hespera Bayudah26.VIII.02d9). g: Petrorossia letho govina, Bulgaria, Canary Is, Croatia, Egypt, h: Petrorossia albula Distribution: Palaearctic: Algeria, Italy, France, Greece, Gruzia, Italy, Kyrgyz Repub- Lebanon, Palestine, Spain, Syria, Tunisia. lic, Libya, Macedonia, Malta, Moldova, Mo- This species was collected flying near rocco, Palestine, Portugal, Romania, Russia, ground surface between vegetation. Spain, Tajikistan, Tunisia, Turkey, Turk- menistan, Ukraine, Uzbekistan, Yugoslavia. Tribe Xeramoebini HULL It appears to be common in the high- Desmatoneura sp. 1 (Fig. 9c) lands and the Jordan Valley. It has been Material: One specimen: Ad Disah 20.VII.96 reared from the nests of several species of (lor). sphecid wasps (DU MERLE 1975). The frons and face of this species is orange with gold scaling. Dr. Greathead ex- Petrorossia letho (WIEDEMANN 1828) amined one specimen with such characters (Fig. 9g) from Oman among a series originally identi- Material: 15 specimens: AI Jubayhah 3.IX.02 fied by him as D. sica. (19 +lcr); Ash Shumari 23.VII.O2 (699 +4ccr) Retma raeum (FORSSK) (Leguminosae); Al Mujib Xeramoeba semirufa (SACK 1909) 24.IX.02 (lcr); Wadi Shuqayq 24.1X.02 (Fig. 9d) (1 9 + lcr) on Venhemia iphionouks. Material: One specimen: Ash Shajarah 2.VI.O2 Distribution: Afrotropical: Benin, (Kr). Chad, Eritrea, Ethiopia, Kenya, Somalia,

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Sudan. Palaearctic: Afghanistan, Algeria, Austria, Bosnia-Hercegovina, Bulgaria, Croatia, Egypt, France, Iran, Italy, Libya, Macedonia, Russia, Saudi Arabia, Slovenia, Spain, Tajikistan, Turkey, Turkmenistan, Uzbekistan, Yugoslavia.

It was collected from white broom Ret- ma raetam (FORSK.) (Leguminosae).

Petrorossia albula ZAITZEV 1962 (Fig. 9h) Material: Five specimens: Al Mukhaybah al Fawqa 31.1X.02 (2oo+2oa); Sama As Sarhan 29.V1I.02 (19). Distribution: Afrotropical: Benin, Ghana, Kenya, Nigeria. Palaearctic: Alge- ria, Armenia, Azerbaijan, Gruzia, Kaza- khstan, Kyrgyz Republic, Mongolia, Moroc- co, Saudi Arabia, Tajikistan, Turkmenistan, United Arab Emirates, Uzbekistan.

It was collected from white broom Rei- ma roetom (FöRSK.) (Leguminosae).

Pipunculopsis stackelbergi ZAITZEV 2000 (Fig. 10a) Material: Four specimens: As Safawi 14.V11I.02 (40T0P). Fig. 10: Distribution: Palaearctic: Palestine. a: Pipunculopsis stackelbergi Spogostylum ocyale (WIEDEMANN 1828) b: Pipunculopsis sp. 1 It was found only so far from the Eastern (Fig. 10d) c: Spogostylum sordidum Desert of Jordan. d: Spogostylum ocyale Material: 11 specimens: Al Jubayhah 5.X.95 e: Spogostylum griseipenne (19), 23.X.96 (19); Amman 6.VII1.97 (19), f: Spogostylum sp. 1 Pipunculopsis sp. 1 (Fig. 10b) 25.XII.98 (la); As Salt 3.IX.93 (ley); Dana, An g: Spogostylum nitidum Material: One specimen: As Safawi 14.VHI.02 h: Spogostylum perpusillum Nawatif 27.IX.O2 (2aa+l 9 ); Dayr Alia 20.VI- (19). 11.94 (la) Light trap; Petra MV.95 (1 9); Wadi This species is found only so far from the Shua'yb26.VlII.96(la). Eastern Desert of Jordan. Distribution: Afrotropical: Somalia, Su- dan, Yemen. Palaearctic: Egypt, Iran, Jor- Tribe Anthracini LATREILLE dan, Libya, Oman, Palestine, Saudi Arabia.

Spogostylum sordidum SACK 1909 This species was recorded from Jordan (Fig. 10c) previously (EVENHUIS &. GREATHEAD 1999). Material: Four specimens: AI Muwaqqar 7.X.02 (la+19) Mating; Dayr Alia 5.X.95 (lcr); As Spogostylum griseipenne (MACQUART Safawi 13.VIII.02 (19). 1850) (Fig. 10e) Distribution: Palaearctic: Afghanistan, Material: Three specimens: Ma'an 20.VII.96 Egypt, Iran, Oman, Saudi Arabia, Tajik- (la); Irbid 21.IX.95 (la); Al Jubayhah unknown istan, Turkmenistan, Uzbekistan. date and month. 90(1?). This species is a large robust species, Distribution: Afrotropical: Mauritania, completely black, but the females have Niger. Palaearctic: Egypt, Iraq, Libya, Pales- white spots on 6th and 7th abdominal ter- tine, Saudi Arabia, Tunisia, United Arab gites. Emirates.

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Spogostylum candidum (SACK 1909) (Fig. 11a) Material: Two specimens: Jarash 21.V.0O (lor); WadiShuqayq24.IX.02(lc). Distribution: Oriental: Pakistan. Palaearctic: Egypt, Iran, Turkey, United Arab Emirates.

The individuals of this species exhibit considerable variation in size, wing venation and colour vestiture (AUSTEN 1937). This species is similar to S. isis, but the body is not completely black and the tibia is brown (GREATHEAD 1988). It is distinguished from other similar species by dense white scales at last abdominal tergites.

Spogostylum isis (MEIGEN 1820) (Fig. 11b) Material: 17 specimens: Ajlun 27.V.O2 (1 9 ); Al Jubayhah 28.VI.94 (la), 26.VIII.90 (lcr); Al Husson 5.V1.97 (lor+19); 17.V1.02 (lcr+19); Amman 5.VI.97 (2aa+l9); Ayn Aqraba 25.111.02 (1 9 ); Ar Rumaymin 3.VII.O2 (1 9 ); Ar Rumman 13.V.91 (la); Hummrit As Sahin 22.IV.O2 (1 9 ); Jarash 6.V.02 (1 9 ); Ma'ain 5.V1- Fig. 11: This spcac» i_jn be separated from the 11.02 (19); Zai 12.VIII.93 (lcr). a: Spogostylum candidum Spogostylum ocyale by its wing pattern Distribution: Afrotropical: Mali. Pa- b: Spogostylum isis (GREATHEAD 2003). c. Anthrax virgo laearctic: Algeria, Armenia, Austria, Azer- d: Anthrax dentata baijan, Belgium, Croatia, Cyprus, Egypt, e: Anthrax aethiops Spogostylum sp. 1 (Fig. 10f) France, Greece, Gruzia, Italy, Kazakhstan, f. Anthrax greatheadi Kyrgyz Republic, Libya, Macedonia, Malta, g: Anthrax sticticus Material: One specimen: Irbid 28.IX.86 (1 9 ). Morocco, Palestine, Portugal, Saudi Arabia, h: Anthrax niger This species has completely infuscated Spain, Syria, Tajikistan, Tunisia, Turkey, wings. It may be a dark form of S. areni- Turkmenistan, Uzbekistan, Yugoslavia. vagum AUSTEN which in turn may be a syn- onym of S. hippolyta (WlEDEMANN). This species is a common species in Jor- dan. It has considerable variation in wing

Spogostylum nitidum AUSTEN 1937 venation, colour of vestiture, and in size (Fig. 10g) (4,9-9,6 mm). Immature stages of this species were recorded as predators on egg Material: One specimen: Al Mujib, Faqu'a pods of Dociostaurus maroccanus (THUN- 28.IX.02 (19). BERG) (Acrididae) but the correctness of Distribution: Palaearctic: Palestine. this record is in doubt (YEATES & GREAT- This species seems to be rare in Jordan. HEAD 1997). It is easily recognised by the yellowish or whitish spots, fonned by patches of scales on Anthrax virgo EGGER 1859 (Fig. 11c) the dorsum of the abdomen (AUSTEN 1937). Material: 14 specimens: Aira 15.V11.O2 (19); Ajlun 27.V.O2 (lcr+19); Ar Rumman 6.V.02 (19); Ash Shawbak 7.VIII.O2 (lor+19); Ash Spogostylum perpusillum AUSTEN 1937 Shunah 15.1X.87 (1 9 ); Dayr Alia 25.V74 (lcr); (Fig. 10h) Dibbin 13.V.O2 (lcr+2 99 ); Jarash 8.V.02 (lor); Material: Three specimens: Ar Rumaymin Kafr Huda 2.IX.O2 (lcr); Mahis 15.VIII.O2 (la). 3.V1I.02 (lor); Al Bahhath 2O.V.O2 (1 9 +1C-). Distribution: Palaearctic: Albania, Al- Distribution: Palaearctic: Palestine. geria, Armenia, Austria, Azerbaijan, Bulgar-

372 © Biologiezentrum Linz/Austria; download unter www.biologiezentrum.at ia, Croatia, France, Greece, Gruzia, Hun- Anthrax sticticus KLUG 1832 (Fig. 11g) gary, Italy, Libya, Malta, Morocco, Pales- Material: Three specimens: As Salt 1.IX.93 tine, Saudi Arabia, Spain, Tunisia, Turkey. (19); Wadi Shuqayq 24.1X.O2 (2aa). This species is common in Jordan. It is Distribution: Afrotropical: Chad, Er- characterised by the absence of scales on the itrea, Guinea-Bissau, Nigeria, Senegal. body and by the absence of recurrent appen- Palaearctic: Afghanistan, Armenia, Azer- dices to veins in the wings, and by presence baijan, Cyprus, Greece, Gruzia, Iran, Italy, of hairs instead of scales on the body. It has Kyrgyz Republic, Macedonia, Oman, Pales- been reared from Anthocopa spp. (Megachil- tine, Syria, Tajikistan, Turkey, Turk- idae) (DU MERLE 1975). menistan, Uzbekistan, Yugoslavia.

This species is characterised by presence Anthrax dentata (BECKER 1906) of more than three spots on the wings. (Fig. 11d) GREATHEAD (1967) suggested that this Material: One specimen: Dayr Alia 13.XI.O2 species is a parasite of mud dauber wasps (19)- (Sphecidae) (EL-HAWAGRY et al. 2000) and Distribution: Afrotropical: Aldabra, has also been reared from megachilid bees Chad, Eritrea, Kenya, Tanzania, Yemen. (DU MERLE 1975). Palaearctic: Algeria, Egypt, Hungary, Italy, Libya, Malta, Morocco, Palestine, Saudi Anthrax niger (AUSTEN 1937) (Fig. 11 h) Arabia, Spain, Tunisia. Material: Two Specimens: Dead Sea 3O.X.O2 This species seems to be a rare species in (19) during flying; Wadi Shuqayq 24.IX.O2 Jordan. (1 9) on the ground. Distribution: Palaearctic: Palestine. Anthrax aethiops (FABRICIUS 1781) It is almost uniformly black coloration (Fig. 11e) of the hairy covering of the body, combined Material: Three specimens: Ayn esh Shallaleh with the presence of only two dark flecks in 24.VI.O2 (lor); Hummrit As Sahin 22.1V.O2 each wing; prevent this species from being (la); Khdair 30.V11I.81 (la) (Al Yarmouk Uni- confused with other species recorded in Jor- versity). dan. Distribution: Palaearctic: Algeria, Ar- menia, Austria, Azerbaijan, Bosnia-Herce- Anthrax candidapex (AUSTEN 1937) govina, Bulgaria, Croatia, Cyprus, Czech (Fig. 12a) Republic, Egypt, France, Germany, Greece, Material: Four specimens: AI Wala 8.1V.02 (la); Gruzia, Hungary, Iran, Italy, Kyrgyz Repub- AI Mujib 13.1II.02 (la); Hummrit As Sahin lic, Libya, Morocco, Palestine, Poland, Por- 22.IV.O2 (2aa). tugal, Romania, Russia, Slovakia, Slovenia, Spain, Switzerland, Tajikistan, Tunisia, Distribution: Palaearctic: Palestine. Turkmenistan, Ukraine, Uzbekistan. The dense white scales at the end of ab- This species is recognisable by the broad domen distinguish A. candidapex from simi- scales at the tip of its abdomen. AUSTEN lar Anthrax ssp. in Anthrax of Jordan. (1937) mentioned that a male hatched from a nest of Osmia paUicomis FRIESE (Megachil- Tribe Villini HULL idae) in Egypt and it has also been reared Oestranthrax brunnescens from other Megachilidae (DU MERLE 1975). (LOEW 1857) (Fig. 12b) Material: One specimen: Dana, Wadi An Anthrax greatheadi EL HAWAGRY 1998 Nawatif27.IX.02(l9). (Fig. 11f) Distribution: Palaearctic: Egypt, Kyrgyz Material: Two specimens: Ar Rumman 6.V.O2 Republic, Palestine, Syria, Tajikistan, Turk- (lcr);AsSalt28.V93(lor). menistan, Uzbekistan. Distribution: Palaearctic: Egypt. This species seems to be rare in Jordan. This species seems to be a rare species in Jordan.

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Distribution: Palaearctic: Armenia, Austria, Azerbaijan, Bulgaria, Croatia, Cyprus, Czech Republic, Egypt, France, Greece, Gruzia, Hungary, Iran, Italy, Libya, Moldova, Morocco, Poland, Romania, Rus- sia, Slovakia, Slovenia, Spain, Turkey, Switzerland, Ukraine.

The development of white or silver scales on the apical abdominal terga of males of this species is variable, but all have identical genitalia. This species is characterised by having white hairs only surrounding the abdomen.

Villa sp. near laevis BECKER 1915 (Fig. 12f) Material: Three specimens: Aljubayhah 1.VI.88 (l9);KafrHuda2.1X.02(lo+lcr). Distribution: Palaearctic: Egypt, Moroc- co, Tunisia. The scales on abdomen of the available specimens were not well preserved which made it difficult to confirm the identification.

Villa insignis AUSTEN 1937 (Fig. 12g) Material: One specimen. No data, (lcr). Distribution: Palaearctic: Palestine, Syr- ia. Fig. 12: Oestranthrax sp. near alfierii a: Anthrax candidapex This species has distinctive yellow hairs PARAMONOV 1931 (Fig. 12c) b: Oestranthrax brunnescens and scales on the abdomen. c: Oestranthrax sp. near alfierii Material: One specimen: Wadi Al Arab (lcr). d: Oestranthrax sp. near pallifrons •: W//a ixion Distribution: Palaearctic: Egypt. Villa atricauda AUSTEN 1937 (Fig. 12h) f: V///a sp. near laevis It appears to be rare in Jordan. The only Material: Two specimens: Ar Rumaymin 3.VI1.02 g: W//a insignis (l9);Zaharl7.V1.02(lcr). h: W//a atricauda specimen was collected buried in the sand under Rema raetam (FÖRSK). Distribution: Palaearctic: Palestine, Syr- ia. Oestranthrax sp. near pallifrons BEZZI 1926 (Fig. 12d) This species was described from females only. The undescribed male Jordanian spec- Material: One specimen: Al Mafraq 10.V.95 imen agrees with the description of the fe- (lcr). male but has silver patagium. Distribution: Palaearctic: Libya, Spain.

The Jordanian specimen fits the female Villa sp. near stenozona (LOEW 1869) description of this species, but it does not (Fig. 13a) have brown infuscation in the costal cell. Material: Two specimens: As Salt 28.VIII.93 While the male is not described yet, it may (IC), 12.1X.93 (lcr). be the undescribed male of this species. Distribution: Palaearctic: Algeria, Egypt, France, Greece, Italy, Turkey, Yu- Villa ixion (FABRICIUS 1794) (Fig. 12e) goslavia. Material: Three specimens: Al Mushaqqar 28.V.90 (lcr); Al Mafraq, Rihaab 7.V.01 (lor); Jordanian specimens differ in having Wadi Shuqayq 24.IX.02 (lcr). white hairs and scales on all sternites while

374 © Biologiezentrum Linz/Austria; download unter www.biologiezentrum.at these are black on apical segment of V. stenozona.

Villa niphobleta (LOEW 1869) (Fig. 13b) Material: Four specimens: Al Mafraq 27.IX.95 (1 9); Al Mujib 3.V.99 (1 9); Amman 15.V.97 (19 ); Hummrit As Sahin 3.VI.02 (1 9). Distribution: Palaearctic: Afghanistan, Algeria, Armenia, Azerbaijan, Bulgaria, Cyprus, France, Greece, Gruzia, Italy, Kyr- gyz Republic, Morocco, Romania, Russia, Spain, Tajikistan, Tunisia, Turkey, Turkme- nia, Uzbekistan, Yugoslavia.

Villa bivirgata AUSTEN 1937 (Fig. 13c) Material: 37 specimens: Ajlun 22.X.92 (lcr); Al Husson 17.VI.O2 (19); Al Jubayhah 22.VII.92 (lcr), 29.X.89 (19), unknown date and month 1995 (lcr); Al Muwaqqar 7.X.O2 (3 99); Al Mukhaybah al Fawqa 21.X.02 (2 99); Amman 27.IX.78 (19); Ar Ramtha 29.VI1.02 (2 99); Ash Shajarah 22.VII.O2 (19); Ash Shawbak

7.VIII.O2 (3 99+2crcr); Ayn Almuallaqah 28.V.01 (l9+3c?cr) (Al Al Bayt University); Dana, Wadi Ar Rummanah 28.IX.02 (19); Hummrit As Sahin 22.V1.02 (19); Irbid 19.V1I.75 (1 9); Kafr Asad 9.IV.01 (1 9 ) (Al Al Bayt University); Sama As Sarhan 29.V1I.02 (lcr), unknown data (lcr); Wadi Ar Rayyan Fig. 13:

8.X.O2 (lcr+499); Yajm 14X02 (2 99 + !

Thyridanthrax lotus (LOEW 1869) (Fig. 13g) Material: 28 specimens: Abu Nussayr 3.VII.O2 (299+lcr); Aira 15.VII.O2 (lcr); Al Husson 17.VI.O2 (2 99 ); AI Muwaqqar 16.IX.02 (499 ); 7.X.O2 (2 99 ); Ar Ramtha 29.V1I.02 (1 9 ); Ash

Shajarah 22.VI1.O2 (2crcr); Ash Shawbak 7.V1- 11.02 (19+lcr); Kafr Khal 22.V1I.O2 (3 99); Maain 5.VII1.02 (5 99); Muzayrib 31.VH.O2 (19); Sama As Sarhan 29.VII.02 (2 99 ). Distribution: Palaearctic: Afghanistan, Armenia, Azerbaijan, Cyprus, Egypt, Greece, Gruzia, Iran, Italy, Kazakhstan, Kyr- gyz Republic, Spain, Tajikistan, Turkey, Turkmenistan, Uzbekistan.

This species is a common species in Jor- dan. It is unusual for Thyridanthrax to have completely hyaline wings. Sama As Sarhan specimens differ in having the dorsal surface of thorax and abdomen with yellow-brown scales and shiny bristles and are smaller size than most T. lotus individuals.

Thyridanthrax perspicillaris ssp. perspicillaris (LOEW 1869) (Fig. 13h) Material: 24 specimens: AI Jubayhah 11.V.94 (19), 22.V.92 (19), 19.VII.93 (la), 31.VI1.90 (lcr),6.VIII.90(l9). 16.IX.90 (2crcr), 17.IX.90 (lor), 18.IX.90 (lcr), 23.IX.90 (\cr), 24.1X.90 (IQ+Icr), 26.1X.89 (Ig + lcr); Al Mafraq, Urn Bottema 8.X.93 (19); Ash Shawbak 7.VII1.O2 Fig. 14: (3 99 +2crcr); Jarash 8.IV.96 (1 9 ); Ma'ain 5.VI- a: Thyridanthrax perspicillaris ssp. idolus Slovakia, Slovenia, Spain, Switzerland, Syr- 11.02 (lcr); Jabaqt Fahl 10.X.99 (19); Wadi al b: Thyridanthrax polyphemus ia, Tunisia, Turkey, Turkmenistan, Ukraine, Arab Dam 9.IX.99 (lc) (Al Balqa University). c: Thyridanthrax sp. near griseolus Yugoslavia. d: Thyridanthrax incanus Distribution: Afrotropical: Eritrea, e: Thyridanthrax elegans This species is common in Jordan. It was Ethiopia, Gambia, Ghana, Mali, Maurita- f: Pachyanthrax fulvifacies collected from Acacia (Leguminosae). It is a nia, South Africa, Sudan. Oriental: Pak- g: Pachyanthrax telamon h: ? Pachyanthrax nimrodicus hyperparasitoid of the pupae of the dipterous istan. Palaearctic: Afghanistan, Albania, and hymenopterous parasitoids Thaume- Armenia, Austria, Azerbaijan, Bulgaria, topoeapityocampa (DENIS & SCHIFFERMÜLLER) Croatia, Cyprus, Egypt, France, Germany, (Thaumetopoeidae) (DU MERLE 1975). Greece, Gruzia, Iran, Iraq, Italy, Kazakhstan, Kyrgyz Republic, Libya, Morocco, Mongo- lia, Palestine, Romania, Russia, Saudi Ara- Caecanthrax arabicus (MACQUART bia, Spain, Tajikistan, Turkey, Turk- 1840) (Fig. 13f) menistan, United Arab Emirates, Uzbek- Material: Two specimens: AI Bahhath 2O.V.O2 istan, Yugoslavia. (19); unknown data (lcr). This subspecies is a very common sub- Distribution: Oriental: Pakistan. species in Jordan. It has 2 submarginal cells Palaearctic: Afghanistan, Armenia, Azer- while the following subspecies has 3 sub- baijan, France, Greece, Gruzia, Iran, Italy, marginal cells. Jordan, Kyrgyz Republic, Saudi Arabia, Tajikistan, Turkey, Turkmenistan, United Thyridanthrax perspicillaris ssp. idolus Arab Emirates, Uzbekistan, Yugoslavia. HESSE 1956 (Fig. 14a) It was recorded previously from Jordan Material: One specimen: Tabaqt Fahl 10.X.99 (EVENHUIS & GREATHEAD 1999). (1 9 ) (AI Balqa University).

Distribution: Afrotropical: Botswana, 22.V1I.O2 (la); Ma'ain 5.VI1I.O2 (19); Muza- Eritrea, Ethiopia, Kenya, Mali, Namibia, yrib 31.VI1.O2 (299); Om As Summaq 19.X.92 South Africa, Uganda, Yemen, Zimbabwe. (H); Sama As Sarhan 29.VII.02 (5 99 +la); Zai Palaearctic: Saudi Arabia. 8.VI11.90 (19); unknown data (1?) (Al Balqa University); unknown data (1?) (Al Yarmouk This subspecies has 3 submarginal cells, University). which distinguish it from the above sub- Distribution: Oriental: Pakistan. Palae- species. arctic: Afghanistan, Algeria, Armenia, Azerbaijan, Cyprus, Egypt, France, Greece, Thyridanthrax polyphemus Gruzia, Iran, Italy, Lebanon, Libya, Macedo- (WIEDEMANN 1819) (Fig. 14b) nia, Morocco, Palestine, Syria, Tunisia, Material: 31 specimens: AI Husson 17.VI.O2 Turkey, Turkmenistan, Yugoslavia. (2 99), 19.V11I.02 (1 9); Al Muwaqqar 27.IX.95 (2 99 +2CTO-) (Al Al Bayt University), 16.1X.02 It is a common species in Jordan. It was (19), 5.IX.95 (1O-) (Al Al Bayt University), collected from Bishop's Weed, Ammi majus 30.1X.95 (la); Amman 5.V.95 (lcr), unknown L. (Umbelliferae). data (1?); Ar Rumman 6.V.O2 (1 9); Ash Sha- jarah 22.V11.O2 (19); Hummrit As Sahin Thyridanthrax elegans 25.X.O1 (19) (Al Balqa University); Khdier (WIEDEMANN 1818) (Fig. 14e) 25.11.82 (19) (Al Yarmouk University); Kafr Material: 24 specimens: Aira 15.VI1.O2 Khal 22.VI1.O2 (8 99+la); Ma'ain 5.VI1I.O2 (19+la); Al Azraq Natural Reserve 2O.V.O2 (lc); Urn Bottema 8.X.93 (1 9), 10.X.95 (3co-) ; (19); Al Muwaqqar 16.IX.02 (3 99), 27.1X.95 Amman 15.VU.95 (la). (19) (Al Al Bayt Univ.); Al Mushaqqar Distribution: Palaearctic: Armenia, 16.1X.02 (3aa); Ash Shajarah 22.VI1.O2 (19); Azerbaijan, France, Greece, Gruzia, Iran, Ash Shawbak 7.VII1.O2 (la); As Salt 2.IX.O2 Italy, Libya, Malta, Morocco, Palestine, Por- (la); Eastern Bayudah 26.V1II.O2 (19+la); tugal, Spain, Syria, Tunisia, Turkey, Turk- Kafr Huda 2.1X.O2 (2aa); Na'or 5.VI1I.O2 (2 99+la); Ma'ain 5.VI1I.O2 (2 99); Muzayrib menistan, Uzbekistan, Yugoslavia. 31.VILO2 (1 9 ); Zai 3.VII.O2 (lcr). This species is a very common species in Distribution: Palaearctic: Armenia, Jordan. Austria, Azerbaijan, Bosnia-Hercegovina, Bulgaria, Croatia, Cyprus, Egypt, France, Thyridanthrax sp. near griseolus (KLUG Greece, Gruzia, Iran, Italy, Kazakhstan, Kyr- 1832) (Fig. 14c) gyz Republic, Lebanon, Libya, Malta, Mol- Material: Two specimens: Wadi Shu'ayb 15.VI- dova, Mongolia, Morocco, Poland, Portugal, 11.02 (1 9); Al Mukhaybah al Fawqa 31.V1I.O2 Russia, Spain, Syria, Tajikistan, Turkey, (19)- Turkmenistan, Ukraine, Uzbekistan, Yu- Distribution: Palaearctic: Egypt, Iran, goslavia. Lebanon, Libya, Saudi Arabia, Tunisia. T. elegans is a common species collected Male specimens are needed to make a from the yellow Heliotropium digynum reliable identification of the species. (FÖRSKAL) (Boraginaceae).

Thyridanthrax incanus (KLUG 1832) Pachyanthrax fulvifacies (AUSTEN 1937) (Fig. 14d) (Fig. 14f) Material: 49 specimens: Aira 15.V1I.O2 (la), Al Material: One specimen: Ad Disah 20.V1I.96 Ghawr 3.V.96 (la), Al Husson 17.VI.O2 (I9), (la). Al Jubayhah 3.1X.90 (19), 22.V.74 (19); Al Muwaqqar 22.V1I.95 (2 99) (Al Al Bayt Uni- Distribution: Palaearctic: Palestine. versity), 5.IX.95 (2 99 )(A1 Al Bayt University); This species appears to be endemic in Al Mushaqqar-16.lX.02 (2 99); Ar Ramtha Palestine and Jordan so far. 29.V11.O2 (19); Ar Rumaymin 8.VIII.90 (19+la), 3.VI1.O2 (la); Ash Shajarah 22.VII.O2 (19+la); Ash Shawbak 7.VII1.O2 Pachyanthrax telamon (LOEW 1869) (699 +4aa), 4.1V.89 (la); As Safawi 13.V1II.O2 (Fig. 14g) (la); As Salt 26.V1I.94 (1 ?); Hummrit As Sahin Material: 24 specimens: AI Arida 4.V.81 (2aa);

22.IV.O2 (2 99); Irbid 4.IX.88 (1 9 ); Kafr Khal Al Husson 17.VI.O2 (3aa); Amman 15.IV.90

dm to bifurcation of R4. Antenna and wings as figures in ZAITZEV (1998). The Jordanian specimen differs in having 1" posterior cell open while it is closed on the wing margin in the Palestinian specimen. Examination of male genitalia is required for generic alloca- tion but it is definitely not a Pachyanthrax and possibly it is a new genus (GREATHEAD in litt.).

Exhyalanthrax afer (FABRICIUS 1794) (Fig. 15a) Material; 13 specimens: Ajloun 27.V.O2 (19); Al Jubayhah 22.V.79 (19); Al Muwaqqar 16.1X.02 (19); AI Wala 8.IV.02 (19); Ayn Aqraba 29.IV.02 (19); As Salt 2.IX.O2 (I9); Dana 23.IX.02 (19); Dayr Alia 8.1V.74 Ü9); Hummrit As Sahin 3.VI.O2 (2 99), 22.V1.O2 (1 9 ); Kafr Huda 2.IX.O2 (1 9 ); Khdair 6.1X.81 (1 9 ) (Al Yarmouk University). Distribution: Afrotropical: Chad, Er- itrea, Ghana, Kenya, Yemen. Oriental: Pak- istan. Palaearctic: Afghanistan, Armenia, Austria, Azerbaijan, Belgium, Bulgaria, China, Croatia, Cyprus, C:ech Republic, Denmark, Egypt, France, Germany, Gibral- tar, Greece, Gruzia, Hungary, Iran, Italy, Kazakhstan, Kyrgyz Republic, Libya, Mace- f \ (19), 15.V.97 (lg+5crar), 16.V.9C (2o-cr); Ar donia, Malta, Mongolia, Morocco, Nether- lands, Oman, Palestine, Poland, Portugal, Fig. 15: Rumman 6.V.02 (1 9 +lcr); Ayn Aqraba 29.IV.02 a: Exhyalanthrax afer (la); Dibbin 13.V.02 (lor); Hummrit As Sahin Romania, Russia, Saudi Arabia, Slovakia, b: Exhyalanthrax melanchlaenus 3.V1.02 (lcr); Jarash 12.V.83 (19); Kafr Asad Slovenia, Spain, Switzerland, Tajikistan, c: Exhyalanthrax contrarius 9.1V.01 (2crcr) (AI AI Bayt University); Rihaab Turkey, Turkmenistan, Ukraine, United d,«: Exhyalanthrax muscarius 7.V.01 (lc) (AI AI Bayt University); Wadi al Arab Emirates, Uzbekistan, Yugoslavia. f: Veribubo anus Arab 16.IV.01 (lcr) (AI AI Bayt University). g: Veribubo saffra E. afer is a common species in Jordan. It h: Veribubo misellus Distribution: Palaearctic: Armenia, is very close to E. melanchlaenus except for Azerbaijan, Cyprus, France, Greece, Gruzia, wing pattern, which does not exceed distal Iran, Italy, Kyrgy: Republic, Palestine, Syria, extremity of 2nd basal cell while it exceeds Tajikistan, Turkey, Turkmenistan, Uzbek- the distal extremity of this cell in E. istan, Yugoslavia. melanchlaenus. It is a hyperparasitoid of the It is a common species in Jordan. pupae of the dipterous and hymenopterous parasitoids Thaumetopoea pityocampa (DENIS

? Pachyanthrax nimrodicus ZAITZEV & SCHIFFERMÜLLER) (Thaumetopoeidae) 1998 (Fig. 14h) but it is also a parasitoid in the puparia of muscoid flies (DU MERLE 1975). Material: One specimen: Zubia 8.VII.02 (1 9). Distribution: Palaearctic: Palestine. Exhyalanthrax melanchlaenus This species was described from a single (LOEW 1869) (Fig. 15b) female specimen collected in Palestine. It Material: Six specimens: Ajloun 27.V.O2 (10*); has the head as Exhyalanthrax, the body Amman 13.IV.9? (I9); Wadi An Nawatif black with brownish yellow scaling on 27.IX.02 (2 99); Zai 3.V1I.02 (lcr+1 9). nd mesonotum, 1" tergite and base of 2 and Distribution: Palaearctic: Armenia, rf 3 . R2<, with angular dip before apical bend. Azerbaijan, Cyprus, Greece, Gruzia, Italy, Wings infuscated in middle from the base of Palestine, Portugal, Spain, Syria, Turkey.

This species is similar to E. afer but the Distribution: Palaearctic: Saudi Arabia. wing pattern exceeds the 2nd basal cell and This species appears to be a desertic has entirely black hair on the pleura. species. Exhyalanthrax contrarius (BECKER 1916) (Fig. 15c) Veribubo misellus (LOEW 1869) (Fig. 15h) Material: One specimen: Kafr Huda 2.IX.O2 Material: Four specimens: Kafr Huda 2.1X.O2 (3 99+lcr). Distribution: Palaearctic: Croatia, Greece, Italy. Distribution: Palaearctic: Armenia, Azerbaijan, Egypt, Greece, Gruzia, Iran, It is the only species with completely Italy, Kyrgyz Republic, Libya, Tajikistan, hyaline wings among the Jordanian species Turkmenistan, Uzbekistan, Yugoslavia. of this genus. It is similar to E. perpusillus (AUSTEN) but has different pattern of ab- The length range of this species is 5,5- dominal scales. 8,5 mm.

Exhyalanthrax muscarius (PALLAS 1818) Tribe Exoprosopini BECKER (Fig. 15d,e) Material: 16 specimens: AI Bahhath 2O.V.O2 Micomitra iris (LOEW 1869) (Fig. 16a) (lcr+2 00); Al Jubayhah 1.X.91 (lcr); 13.V.79 Material: One specimen: Unknown data (1 9). (la), 9.X.02 (lcr); Ayn Aqraba 25.111.02 (lcr); Distribution: Palaearctic: Armenia, As Salt 2.IX.O2 (lcr); Dana 23.IX.O2 (lor); Dib- Austria, Azerbaijan, Bulgaria, Croatia, bin 13.V.02 (19+la); Kafr Huda 2.IX.O2 France, Greece, Gruzia, Iran, Italy, Kaza- (2cror); Ma'ain 5.VII1.O2 (la); Yajuz 14X02 khstan, Kyrgyz Republic, Moldova, Pales- (1 9); Zai 22.VIII.93 (lor). tine, Romania, Russia, Spain, Tajikistan, Distribution: Oriental: Pakistan. Turkey, Turkmenistan, Ukraine, Uzbekistan. Palaearctic: Afghanistan, Armenia, Austria, Azerbaijan, Croatia, Egypt, France, Ger- Micomitra sp. near chrystallina many, Gibraltar, Greece, Gruzia, Hungary, (BEZZI 1924) (Fig. 16b) Iran, Italy, Kyrgyz Republic, , Moldova, Material: One specimen: Al Mujib 28.IV.99 Poland, Portugal, Romania, Russia, Slove- (19). nia, Spain, Switzerland, Macedonia Tajik- istan, Tunisia, Turkey, Turkmenistan, Distribution: Afrotropical: Somalia, Ukraine, Uzbekistan, Yugoslavia. Yemen. Palaearctic: Saudi Arabia. Micomitra chrystallina has black hairs on This species has a wide range in length the prosternum unlike M. iris and M. stupi- (2,9-7,6 mm). da (Rossi).

Veribubo anus (WIEDEMANN 1828) (Fig. 15f) Heteralonia (Zygodipla) singularis (MACQUART 1840) (Fig. 16c) Material: Four specimens: Hummrit As Sahin 22.IV.O2 (499). Material: One specimen: Jabir As Sarhan 21 .V.01 Distribution: Afrotropical: Sudan. (19). Palaearctic: Egypt, Libya, Saudi Arabia, Distribution: Afrotropical: Yemen. Palae- Tunisia, United Arab Emirates. arctic: Egypt, Libya, Morocco, Oman, Pales- tine, Saudi Arabia, United Arab Emirates. Jordanian specimens have hyaline wings instead of milky wings, which are known to This species appears to be a rare species. occur in specimens collected from other It is the only Jordanian species which has a countries. supernumerary cross vein in r5 cell.

Veribubo saffra GREATHEAD 1981 Heteralonia {Zygodipla) bagdadensis (Fig. 15g) (MACQUART 1840) (Fig. 16d) Material: One specimen: AI Muwaqqar 16.IX.02 Material: Five specimens: Ayn esh Shallaleh 21.V.01 (lcr) (Al Al Bayt University); Sumayya

This species appears to occur in desertic regions of Jordan. One specimen was collected from a vinegar pit fall trap near An- abasis sp.

Heteralonia (Acrodisca) suffusa (KLUG 1832) (Fig. 16g) Material: 50 specimens: Al Jubayhah 5.VIH.9O (1 9 ), 6.VIII.90 (lcr), unknown data (1 9 ); Ar Rumaymin 3.V1.02 (lcr+2 99 ), 8.VI1I.90 (1 9 );

Ash Shajarah 22.V1I.02 (699+3crcr); Ash Shawbak 4.III. 94 (4oa), 7.VIII.O2 d9); As Salt 1.VIII.02 (la), 18.VIII.93 (19); Ayn Al- muallaqah U.VI.Ol (2orcr) (Al Al Bayt Univer- sity); Eastern Bayudah 26.V1I1.02 (2 99); Irbid 27.XII.98 (la); Kafr Khal 31.VII.O2 (3 99); Ma'ain 5.VII1.O2 (I9); Mahis 15.V11.O2 (lcr); Muzayrib 31.V1I.O2 (2 99+5ccr); Na'or 5.V1- 11.02 (2 99); Wadi Shua'yb 27.VIII.9O (I9), 26.IX.93 (19); Wadi esh Shallaleh 21.V.01 (2 99 ) (Al Al Bayt University), 16.V1.97 (1 9 ); Western Bayudah 26.V111.02 (lcr); Zai 22.VI- 11.93 (la); 19.V1I.93 (la).

Distribution: Afrotropical: Kenya, So- malia. Palaearctic: Afghanistan, Armenia, Azerbaijan, Cyprus, Greece, Gruzia, Iran, Italy, Kazakhstan, Moldova, Palestine, Rus- sia, Saudi Arabia, Syria, Tunisia, Turkey, Fig. 16: As Sarhan 28.V.01 (3 99 ) (AI AI Bayt Universi- Ukraine, Yugoslavia. a: Micomitra iris ty); Unknown data (la). b: Micomitra sp. near chrystallina c: Heteralonia (Zygodipla) singularis Distribution: Palaearctic: Armenia, H. suffusa is very common during sum- d: Heteralonia (Zygodipla) bagdadensis Azerbaijan, Iran, Iraq, Mongolia, Oman, mer in Jordan except in desertic regions. It e: Heteralonia (Zygodipla) hermani is easily recognized in the field, flying near f: Heteralonia (Zygodipla) mucorea Palestine, Saudi Arabia, United Arab Emi- ground surface. It has completely infuscated g: Heteralonia (Acrodisca) suffusa rates. h: Heteralonia (Homolonia) megerlei wings ranging from brown to black. This species is characterised by the r, cell, which closes far from the wing margin. Heteralonia (Homolonia) megerlei (MEIGEN 1820) (Fig. 16h) Heteralonia (Zygodipla) hermani Material: Two specimens: Ayn esh Shallaleh (FRANCOIS 1967) (Fig. 16e) 21.V.01 (1 9 ); Wadi Shu'ayb 10.111.84 (19). Material: Five specimens: Ayn esh Shallaleh Distribution: Afrotropical: Chad, Gam- 21.V.01 (3 99+lcr) (Al Al Bayt University); bia, Ghana, Mauritania, Niger, Nigeria, Sumayya As Sarhan 28.V.01 (19)- Senegal, Sudan, Yemen. Oriental: Pakistan. Distribution: Palaearctic: Palestine. Palaearctic: Albania, Armenia, Austria, Azerbaijan, Bulgaria, Croatia, Cyprus, Heteralonia (Zygodipla) mucorea Egypt, France, Greece, Gruzia, Iran, Italy, (KLUG 1832) (Fig. 16f) Kazakhstan, Kyrgyz Republic, Libya, Mace- Material: Four specimens: Wadi Rum 24.IV.OO donia, Malta, Moldova, Oman, Palestine, (19), 28.IV.00d 9+lcr), 11.V.OO(19). Romania, Russia, Saudi Arabia, Slovenia, Switzerland, Tajikistan, Tunisia, Turkey, Distribution: Palaearctic: Afghanistan, Turkmenistan, Ukraine, United Arab Emi- Algeria, Egypt, Iran, Kuwait, Kyrgyz Repub- rates, Uzbekistan, Yugoslavia. lic, Oman, Saudi Arabia, Syria, Tajikistan, Tunisia, Turkmenistan, United Arab Emi- The wing pattern is variable in extent rates, Uzbekistan. and intensity.

Exoprosopa pandora (FABRICIUS 1805) (Fig. 17a) Material: 18 specimens: AI Ghawr 3.V.93 (1 9 ); Ar Rumman 13.V.91 (I9). 18.V.00 (19), 6.V1.88 (lcr); Ash Shawbak 7.VIH.O2 (1 9 + lor); Rihaab 7.V-01 (8 99 +2oro-) (AI AI Bayt Univer- sity); Yajuz 5.V.95 (lcr), unknown data (lo-). Distribution: Palaearctic: Algeria, Croa- tia, Egypt, France, Greece, Iran, Italy, Libya, Macedonia, Morocco, Palestine, Slovenia, Turkey, Yugoslavia. This species varies in the length from 6,4 to 10,4 mm. Exoprosopa sp. near pusi'Ila MACQUART Fig. 17: Exoprosopa minos (MEIGEN 1804) 1840 (Fig. 17d) a: Exoprosopa pandora (Fig. 17b) Material: Two specimens: Amman 9.V.84 (2 99 )• b: Exoprosopa minos Material: One specimen: Al Bahhath 2O.V.O2 Distribution: Afrotropical: Chad, c: Exoprosopa rivularis (19)- Ethiopia, Gambia, Ghana, Guinea-Bissau, d.Exoprosopa sp. near pusilla Distribution: Palaearctic: Algeria, Ar- Ivory Coast, Kenya, Nigeria, Senegal, Ugan- menia, Austria, Azerbaijan, Croatia, Czech da, Yemen. Palaearctic: Saudi Arabia. Republic, Egypt, France, Germany, Greece, Jordanian specimens of this species have Gruzia, Hungary, Iran, Italy, Kazakhstan, a more extensive wing patterns compared to Kyrgyz Republic, Lebanon, Libya, Moldova, E. pusiUa reaching the apex of the wings on Morocco, Palestine, Poland, Romania, Rus- the fore margin, and clear area in middle of sia, Slovakia, Spain, Syria, Tajikistan, discal cell is much narrower. Also it has Tunisia, Turkey, Turkmenistan, Ukraine, white scales band at base of 2nJ tergite. Uzbekistan. This species seems to be rare in Jordan. Discussion Exoprosopa rivularis (MEIGEN 1818) This is the first study of the Bombyliidae (Fig. 17c) of Jordan. A total of 132 species of Bombyli- Material: 33 specimens: Aira 15.V11.O2 (1 9 ); Al idae are recorded, 124 of them for the first Jubayhah 9.IX.02 (lcr); Al Mukhaybah Al Fawqa time. EVENHUIS & GREATHEAD (1999) 31.VII.O2 (lcr), Ar Ramtha 29.V11.02 (2cfor), Ar recorded 14 species, eight of them were also Rumman 6.V.02 (1 9 ); Ar Rumaymin 3.VII.O2 (1 9+lor); Ash Shajarah 22.VI1.O2 (4ccr); Ash recorded in this study, however, six of them Shawbak 7.VIII.O2 (2crcr); Ayn Almuallaqah were not found during our collection trips. 11.VI.01 (1 9 ) (Al Al Bayt University); Ayn esh These are: Anastoechus nivifrons (WALKER), Shallaleh 16.VI.97 (2 99 +2crcr), 24.VI.O2 (lcr); Bombylisorw flavibarbum (LOEW), Bombylius Dayr Alia 18.X1.74 (\Q+lCf); Hummrit As argentifacies AUSTEN, Chakochium argyro- Sahin 3.VI.O2 (lor); Kafr Khal 22.VII.02 cephalus (MACQUART), Cytherea alboUneata (1 9 +lcr); Ma'ain 5.VIII.O2 (3 99 +300-); Na'or (BEZZl), and Cytherea barbara SACK. There- 5.VIII.02 (lor); Wadi Shua'yeb 27.VII1.90 (lcr). for, the species known to occur in Jordan are Distribution: Palaearctic: Armenia, currently 138. They belong to eight subfam- Austria, Azerbaijan, Croatia, France, ilies. The largest subfamily is Anthracinae, Gruzia, Iran, Italy, Kyrgyz Republic, Libya, which includes 70 species in six tribes and 20 Macedonia, Mongolia, Morocco, Palestine, genera. The Bombyliinae includes 28 species Portugal, Saudi Arabia, Slovenia, Spain, in two tribes and nine genera. The Cythere- Tajikistan, Turkey, Turkmenistan, Uzbek- inae include 16 species in four genera. The istan, Yugoslavia. Usiinae includes four identified species and This species is a very common in Jordan three unidentified species in two tribes and during summer except in desertic areas. Fe- three genera. The Toxophorinae includes six males are variable in the intensity of wing species in two tribes and two genera. The patterns. Lomatiinae includes three species in one

tribe and one genus. The smallest subfami- naceae), Anthemis palaestina REUTER, Lactu- lies according to the number of species are ca orientalis BOISS, Verthimia iphionoides Ecliminae with one species and Phthirinae BOISS & BLANCHE, Canhamus persicus with one unidentified species. WlLLD (Compositae), Acacia sp. and Rerma raetam (FöRSK.) (Leguminosae), Linum pu- The large number of genera with a small bescens BANKS & SOL. and Linum peyronii average number of species recorded in this POST (Linaceae), Ammi majus L. (Umbellif- study (3,1 species per genus) probably re- erae), Papaver syriacum BoiSS & BLANCHE flects the great diversity in climatic condi- (Papaveraceae) and Polygonum equisenforme tions and phytogeographical regions in our SlBTH (Polygonaceae). area which include: the North Mediter- ranean zone, the Southern Mediterranean Grasshoppers, wasps and bees appear to zone, the Irano Turanian belt surrounding be the known hosts of the Jordanian imma- the Mediterranean zone, the North Jordan ture Bombyliidae. Valley, the Eastern Desert, and the South- According to the collecting dates (Table ern Desert. The largest genus is Bombylius 2), it appears that most of 'Homeophthal- with 13 species, followed by Villa with nine mae' appear earlier in the season (23 species species, Spogostylum with eight species, An- in March and 25 species in April), while thrax with seven species, Cytherea, Amictus, most of the 'Tomophthalmae' group appear Thyridanthrax and Heteralonia each with six later (48 species in May). The lowest num- species, Geron with five species, Exoprosopa, bers of specimens were collected in January Anastoechus and Exhyalanthrax each with where only one specimen of Usia aenea was four species, Usia, Pamgeron, Chalcochiwn, collected. In December, three specimens Lomatia, Petrorossia, Oestranthrax, Pachyan- from Anthracinae (one of Spogostylum thrax and Veribubo each with three species. ocyale, one of Heteralonia suffusa and one of The other genera each have only one or two Hemipenthis veluana) were collected. More species. specimens were collected from the 'To- From a Zoogeographie point of view, the mophthalmae' (737) than from the 'Home- known Jordanian Bombyliidae are mostly ophthalmae' (674) (Table 3). Palaearctic (86 species, 69,4 %). Twenty-six Some species are common in Jordan such species (21 %) occur in the Palaearctic and as Usia aenea, U. bicolor, Bombylius medius, B. Afrotropical regions. Seven species (5,65%) are distributed in the Palaearctic and the fimbriatus, Heteralonia suffusa, Thyridanthrax Oriental regions. Only four species (3,2 %) incanus, T. polyphemus, and Hemipenthes ve- (Bombylius modestus, Heteralonia megerlei, luana. BODENHEIMER (1935) recorded similar Exhyalanthrax afer and Thyridanthrax perspic- results, he considered that Bombylius, Exo- illaris) are distributed in the Palaearctic, the prosopa (including what is known now as Ex- Afrotropical and the Oriental regions. One oprosopa, Heteralonia, Micomirra, Pachyan- species (0,81%), Bombylius major, is widely thrax), Anthrax and Usia the most common distributed in the Palaearctic, the Nearctic genera in Palestine. Other species so far, are and the Oriental regions. rare such as Desmatoneura sp., Toxophora fasciculata, Caecanthrax arabicus, Micomitra ssp., Most species were collected from the Oestranthrax ssp., and Eclimus gracilis. How- ground surface since the Bombyliidae spend ever, further collecting may prove their oc- most of the time basking on rocks, bare currence in greater numbers. ground and dry vegetation or searching for hosts for oviposition. Few species of Bom- The species list of Bombyliidae of Jordan byliidae were collected from plants during is far from complete. More species are ex- feeding. However, all specimens of Usia and pected to be found after further extensive Parageron were collected from flowers be- collecting for a longer period of time. For cause these flies are weak fliers compared to example, in the neighbouring Palestine, other species and they rest inside the flowers (AUSTEN 1937) recorded 128 species and during feeding. The plants on which the bee ZAITZEV (1995, 1996, 1997, 1998 1999) flies were observed to feed belong to seven added new records raising the number of families; Echium judaeum LACAITA and He- species to 229. Of these, 153 species are not liotropium digynum (FÖRSKAL) (Boragi- yet recorded in Jordan. And there are 41

382 © Biologiezentrum Linz/Austria; download unter www.biologiezentrum.at species recorded from Jordan not yet record- Table 2: Number of species collected from the 'Homeophthalmae' and the 'Tomophthalmae' in each month of the year. ed in Palestine. From Arabia, 137 species were recorded (GREATHEAD 1980b, 1988) Month many of them are not known to occur in Group 1 2 3 4 5 6 7 8 9 10 11 12 Jordan so far. In particular more species of Species No. of the subfamily Phthiriinae are expected to 'Homeophthalmae' 1 7 23 25 15 5 6 3 6 7 2 0 Species No. of occur in Jordan, since only one species is Tomophthalmae' 0 1 16 29 48 23 25 23 33 11 2 3 recorded in this study while 8 species have Total 1 8 39 54 63 28 31 26 39 18 4 3 been recorded from Palestine. Table 3: Number of specimens collected from the 'Homeophthalmae and the In addition to the identified species, 'Tomophthalmae' in each month of the year. there are 8 unidentified species belonging to Month Apolysis (1 species), Parageron (2 species), Group 1 2 3 4 5 6 7 8 9 10 11 12 Phthiria (1 species), Spogostylum (1 species), Specimens No. of Villa (1 species), Desmatoneura (1 species), 'Homeophthalmae' 1 35 336 177 45 5 10 3 33 27 2 0 and Pipunndopsis (1 species). There were Specimens No. of some difficulties in the identification of 'Tomophthalmae' 0 1 40 88 152 52 142 105 100 45 9 3 species belonging to Parageron. This genus Total 1 36 376 265 197 57 152 108 133 72 11 3 needs revision because in the past new melschwebern durchgeführt und dabei mehr species have been described that were found als 870 Belege an 72 Fundorten gesammelt. later to be variants of the same species. In In der Studie wurden auch früher gesam- addition, it was difficult to identify species melte Belege von Hummelschwebern aus of Apolysis, Geron and Petrorossia. Species of der Insektensammlung der Universität von these genera have few external characters Jordanien und aus anderen jordanischen and need the examination of male and fe- Sammlungen berücksichtigt. Die Belege male genitalia for correct identification. repräsentieren 132 Arten in 41 Gattungen This study can be considered as a base und 8 Unterfamilien. Davon wurden 124 line for further studies on the Bombyliidae Arten erstmals in Jordanien nachgewiesen. of Jordan. More species are expected to be Für jede Art werden Anzahl der untersucht- collected. Studies on the distribution, adult en Belege, Aufsammlungsorte und -zeiten in and immature hosts, biology and ecology of Jordanien sowie Anmerkungen angeführt. most species need to be done. References

Acknowledgments AL-HOUTY W.(1989): Insect Fauna of Kuwait. 1" edi- We thank Dr. Greathead, Imperial Col- tion. — University of Kuwait: 1-187. lege London, for his help in the identifica- ANONYMOUS (1990): Gazetteer of Jordan. 2nd edi- tion of specimens; Dr. Neal Evenhuis, Bishop tion. — Defense Mapping Agency, Washing- ton. D. C: 1-337. Museum, Hawaii, USA and Dr. David Yeates, AUSTEN E.E.(1937): Bombyliidae of Palestine. — CSIRO Entomology, Black Mountain Labo- British Museum of Natural History, London: 1- ratories, Australia; and Dr. M. El-Hawagry, 188. Cairo University, for providing literature. Al- BODENHEIMER F.S.(1935): Animal Life in Palestine. — so we thank Mr. Suhail Ismael, Natural His- Jerusalem: 1-279. tory Museum, Al Yarmouk University; Mrs. BODENHEIMER F.S.(1937): Prodromus Faunae Pales- Fatima AI Umoush, AI AI Bayt University, tine. — Memoires de L'lnstitut d'Egypte Cairo: and Curator of the Al Balqa'a Applied Uni- 1-286. versity Collection for providing specimens. ou MERLE P. (1975): Les notes et les stades pre- imaginaux des Dipteres Bombyliidae. — Re- vue bibliographique annotee. Bulletin SROP Zusammenfassung 1975 (4): 1-289.

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