PSAMMONALIA

The Newsletter of the International Association of Meiobenthologists

Number 168, December 2017

Composed and Printed at Lab. of Coastal Benthic Ecology P.P. Shirshov Institute of Oceanology RAS

DONT FORGET TO RENEW YOUR MEMBERSHIP IN IAM! THE APPLICATION CAN BE FOUND AT: http://www.meiofauna.org/appform.html This newsletter is mailed electronically. Paper copies will be sent only upon request

This Newsletter is not part of the scientific literature for taxonomic purposes 1

Executive Committee Vadim Mokievsky, Chairperson P.P. Shirshov Institute of Oceanology, Russian Hanan Mitwally (term expires 2019) Academy of Sciences, 36 Nakhimovskiy Faculty of Science, Oceanography, University of Prospect, 117218 Moscow, Russia. Alexandria, Moharram Bay, 21151, Egypt . [email protected] [email protected] Wonchoel Lee, Past Chairperson Gustavo Fonseca (term expires 2019) Lab Of Biodiversity, (#505), Department of Life Universidade Federal de São Paulo, Instituto do Science, College of Natural Sciences, Hanyang Mar, Av. AlmZ Saldanha da Gama 89, 11030- University, Seoul, South Korea. 400 Santos, Brazil. [email protected] [email protected] Daniel Leduc (term expires 2022) Ann Vanreusel, Treasurer National Institute of Water and Atmospheric Lab Morphologie, Universiteit Gent, Research, Private Bag 14-901, Wellington, New Ladengancjstraat 35, B-9000 Gent, Belgium. Zealand. [email protected] [email protected] Nabil Majdi (term expires 2022) Jyotsna Sharma, Assistant Treasurer Bielefeld University, Ecology, Department of Biology, University of Texas at Konsequenz 45, 33615, Bielefeld, Germany. San Antonio, San Antonio, TX 78249-0661, [email protected] USA. [email protected]

Ex-Officio Executive Committee (Past Chairpersons)

• 1966-67 Robert Higgins - Founding Editor • 1987-89 John Fleeger • 1968-69 W. Duane Hope • 1990-92 Richard Warwick • 1970-71 John Gray • 1993-95 Paul Montagna • 1972-73 Wilfried Westheide • 1996-98 Magda Vincx • 1974-75 Bruce Coull • 1999-2001 Yoshihisa Shirayama • 1976-77 Jeanne Renaud-Mornant • 2002-2004 John Lambshead • 1978-79 William Hummon • 2005-2007 Keith Walters • 1980-81 Robert Higgins • 2008-2010 Paulo Santos • 1982-83 Carlo Heip • 2011-2013 Nikolaos Lampadariou • 1984-86 Olav Giere • 2014-2016 Wochoel Lee

Board of Correspondents

Er Hua College of Marine Life Science, Ocean University of China, 5 Yushan Road, Qingdao 266003, P.R.C. [email protected] Jeroen Ingels Plymouth Marine Laboratory, Prospect Place, Marten Sørrensen The Hoe, Plymouth PL1 3DH, UK. Natural History Museum, Denmark. [email protected] [email protected] Cheng Chen Samuel Gomez University of Malaysia Sabah, Malaysia. Universidad Nacional Autónoma de México. Joel [email protected] Montes Camarena s/n, 82040 Mazatlán, Sinaloa, México. [email protected]

This Newsletter is not part of the scientific literature for taxonomic purposes 2 Editor in Chief: Editorial Vadim Mokievsky Cover page: Dear Colleagues, members of International Elena Chertoprood, Lesia Garlitska Association of Meiobenthologists and all who are interested in microscopic creatures, Contents I’m glad to send you my warmest seasonal Editorial ………………………………….3 greetings and wish you every success in life Upcoming conferences ..……………...….4 and science in New Year! The last year was rich by events and activities 17th International Conference on Meiofauna in meiobenthic science. Several conferences 14th International Symposium on devoted to various groups of meiofauna held Tardigrada in different places gave us a good opportunity International Crustacean Congress to see each other again and share results of European Society of Nematologists new researches. Many expeditions, coastal, Conference 2018 marine and freshwater gathered new data and Ocean Sciences Meeting in Portland, replenished our collections. Many new Oregon papers were printed out since June, electronic resource grew up and new conferences are After the conferences ….………………..5 proposed for the coming months. Some Fourth International Congress on information on each of these topics you can Invertebrate Morphology found on following pages. Twelfth International Symposium of the I’m glad to welcome new members of Russian Society of Nematologists Association and wish you to enjoy your Conference "Crustacea: diversity, ecology studies on these nice microscopic and evolution" and also wish you fruitful contacts with your colleagues. I would be glad if IAM and our Eхpeditions ……………..……………….8 communication tools will help you on this way. Studying the meiofauna of the world’s Concerning IAM routine, we are working on deepest environments the webpage and mailing list. The number of mistakes was fixed, and I have to express my New members...... 8 gratitude to Jeff Baguley and Board Members Remembering Bill Hummon ….………10 for significant improvement of list of members. However, some gaps and mistakes Electronic resources…………………….12 are exist there and I still would like to have information about any incorrectness. It Meiobenthic groups within the World appears also, that in some cases messages Register of Marine Species from “Psammonalia” mailbox may lost because of spam filters of users. Please, Recent Literature……………………….13 check it. Membership Renewal Form……………18 All previous issues of PLAMMONALIA are available at: http://www.meiofauna.org/psammona.html I have to thank all of members of

Association who help me with preparation of this issue. Happy New Year! Yours Vadim Mokievsky

This Newsletter is not part of the scientific literature for taxonomic purposes 3 Upcoming conferences

17th MEIOFAUNA CONFERENCE 14th International Symposium on SeventhIMCO Tardigrada 30 July - 03 August, 2018 7-12th July 2019 (Évora, Portugal) We are very pleased to welcome you to the 14th International Symposium on Tardigrada, which will be held in Copenhagen, Denmark, 30 July - 03 August 2018.

From the 7th till the 12th July 2019, the Information about the Symposium is University of Évora, the Marine and available at the website: Environmental Sciences Centre (MARE) and https://www.tardigrada2018.org the International Association of

Meiobenthologists (IAM) will organize the Please note that the deadline for Early 17th edition of the Meiofauna Conference Registration and Abstract submission is 30 (SeventhIMCO) to be held at Évora, March 2018. Late Registration is open until 1 Portugal. June 2018. Information about registration The conference aims to discuss and and abstract preparation etc. can be found at contribute to the advance of the the Symposium website. understanding on the current state of meiofauna diversity and its interactions in We are looking forward to seeing you at the marine and freshwater systems. It will be a Copenhagen Symposium in 2018! dynamic conference focused on innovative Please forward this message to anyone who research topics and providing a set of might be interested. interesting keynote sessions, contributed On behalf of the Organising Committee, talks and posters and thematic workshops Chair of Tardigrada 2018, addressing meiofauna across a wide range of relevant research challenges, namely Nadja Møbjerg analytical methods, conservation, ecosystem interaction and services, marine and freshwater ecology, frontiers areas and International Crustacean Congress policies. May 22-25, 2018 At present, we are developing a program of inspirational speakers and will announce International Crustacean Congress (ICC9) more details in the webpage very soon. that will take place May 22-25, 2018, in Please share this event with your global Washington DC, USA. The Congress hosted network using your newsletters, websites and by the Smithsonian Institution and the social media. Renaissance Hotel. This will be the first time

that an ICC has been held within the United We hope to see you in Évora! States and the first time to be hosted by the

Smithsonian Institution. The ICC 9 meeting On behalf of the SeventhIMCO Organizing is also co-hosted by The Crustacean Society Committee and is dedicated to dissemination of all Helena Adão aspects of crustacean biology to local and e-mail: [email protected] international audiences. University of Évora and MARE- Marine and For registration, early bird deadline is Environmental Sciences Centre January 15. Abstract deadline is http://www.mare-centre.pt/en March 1. http://www.evora-portugal.com/index.html e-mail:[email protected]

This Newsletter is not part of the scientific literature for taxonomic purposes 4 For detailed information, please visit the ICC9 website: Other information and list of papers of the http://www.birenheide.com/ICC2018/index. session are here: php https://agu.confex.com/agu/os18/meetingapp .cgi/Session/28611 European Society of Nematologists Conference 2018 General information about the Ocean 9-13th September, Ghent, Belgium Sciences Meeting is here: https://osm.agu.org/2018/ Main topics of the Conference are: - Systematics (, Morphology, Registration Deadline: 10 February Phylogeny, Barcoding) Housing Deadline: 17 January 2018 - Ecology - Management After the conference: participants’ brief - Biocontrol (including entomoparasitic notes ) - Plant- interactions Fourth International Congress on - Animal parasites Invertebrate Morphology - Model organisms The large-scale Congress took place at M.V. Deadline abstract submission applicants Lomonosov Moscow State University, student bursaries 15/03/2018. Moscow from August 18 to 23, 2017. Among Early registration 15/05/2018. many other thematic sessions in the symposia Deadline abstract submission 15/05/2018 structure, the special session “Minute Information avalable at: metazoans: Little pigeons can carry great https://www.esn-online.org/conference messages” was devoted to micrometazoans and partly to tiny meiobenthic organisms Ocean Sciences Meeting in Portland, fitting in the scope of Psammonalia. Katrine Oregon. Worsaae and oth. (Denmark) gave a lecture Session: Benthic Meiofauna Structure, “A meiofauna perspective on nervous system Pattern, and Function in Ocean Basins evolution” emphasizing that meiofauna February 13, 2018 lineages may constitute sister clades to the Special session proposed for meiofauna major groups of Bilateria: Xenacoelomorpha studies. Session will be in Oregon as sister to other bilaterians, Scalidophora – Convention Center - Poster Hall at Tuesday, to remaining ecdysozoans, Gnathifera – to February 13, 2018 04:00 PM - 06:00 PM remaining spiralians and thus necessity for a In this topical session, submissions are comparative approach and reconstruction of solicited in the area of benthic meiofauna ancestry using phylogenetic tools. Three community structure, their biogeography communications concerned free-living across ocean basins as well studies nematodes. Vladimir Yushin (Russia) and investigating specific role of benthic oth. (“Male gametes and evolution in meiofauna in marine ecosystems including nematodes …”) particularly accentuated biogeochemical cycling. In particular, problems of incongruence of sperm submissions are welcome in the emerging morphology to certain nematode lineages and areas of benthic meiofauna research that enigmatic phenomenon of spern dimorphism incorporates new tools and techniques in some taxa. Myriam Clayes (Belgium) and including next generation sequencing. oth. showed an easy-to-use and low-cost Session is organized by Holly M. Bik method of self-pressurised rapid freezing; (University of California, Riverside), this method tested on free-living nematodes Punyasloke Bhadury (Indian Institute of preserves finely all the cell structures. Izabela Science Education and Research, Kolkata) B. Poprawa and oth. (Poland) reported on and Jeroen Ingels (Florida State University). oogenesis in a parthenogenetic eutardigrade.

This Newsletter is not part of the scientific literature for taxonomic purposes 5 Magdalena Rost-Roszkowska and oth. Russian or in Russian with explicit comments (Poland) described a variety of functions of and remarks in English. Most midgut epithelium in eutardigrades – not only communications concerned zoo-, phyto- and secretion, absorbtion, accumulation of entomoparasitic as well as terrestrial reserve material but also defense against nematodes but a few lectures and poster external factors. Nataliya Budaeva and presentations dealt with marine free-living Conrad Helm (Norway) reported on nematodes. Thus, Nguyen Dinh Tu and oth. enigmatic Charlie Chaplin annelids of the (Hanoi) presented a study on impact of a Histriobdellidae living as commensals on typhoon on nematode communities in coastal lobsters – using a variety of approaches, the mangrove zone in Vietnam. Maria Fedyaeva authors found histriobdellids close to some and Darja Portnova (Moscow) had an oral taxa within Eunicida. Several presentations presentation on distribution of nematode were devoted to structural peculiarities of diversity across the shelf of the Arctic Laptev microarthropods: wing apparatus and fight as Sea. For me, the most interesting detail in this well as head morphology of tiniest beetles study turned out to be finding of numerous (Alexey Polilov and oth., Margarita Cryonema sp. juveniles in bottom sediments: Yavorskaya and oth., Russia), brain structure Cryonema species until recently were known of tiny dipterans (Anastasia Makarova and to be only associated with long-term and Alexey Polilov, Russia). Alexandra seasonal Arctic sea ice. Alexei Tchesunov Petrunina and Gregory Kolbasov (Russia) and Vadim Mokievsky (Moscow) gave a described anatomy of Tantulocarida, smallest lecture on nematode assemblages and crustaceans ectoparasitizing other tardigrades within bioclastic carbonate sand microscopic marine host crustaceans; the on summit plateau of the giant isolated authors suggest that so-called honeycomb- seamount Great Meteor in the north-east like structure filling in the anterior part of the Atlantic Ocean. Nematode diversity and male cephalothorax represent actually a male taxonomic composition generally resembles gonad. Yoko Matsumura and oth. (Germany) those of coastal coarse sands but portions of presented a very fine study on biomechanics some taxa are increased while other taxa are of a hyper-elongated structure of a beetle lowered or absent. Thus, an unusually intromittent organ. The presentations diversity of the predatory genus Latronema demonstrated a high diversity of methods (Selachinematidae) was revealed. Eight allowing to solve problems concerning morphologically distinct new Latronema functional morphology, physiology, life species share some structural features in cycles and phylogenetic relationships of a common (stoma armature reduction) that broad range of micrometazoans. enables to hypothesize their endemic Alexei Tchesunov speciation. Maria Fedyaeva presented a wide study of the intestine ultrastructure in a number of marine free-living nematode Twelfth International Symposium of the species in order to test an expectation that the Russian Society of Nematologists midgut characters might be linked with the “Nematodes and other Ecdysozoa under feeding type of species. The most variable the growing ecological footprint on structure among species proves to be ecosystem” extracellular coating over microvillar brush, The Symposium has been held on July 30 – a gycocalyx – its thickness and structural August 6, 2017 at N.I. Lobachevsky complexity may be connected with diet: University in Nizhni Novgorod, a pictorial species ingesting coarse sediment particles large city on Volga River, formerly (e.g. Daptonema spp.) tend to have more considered as a “third capital of Russia”. The thick multilayered glycocalyx than those total number of participants was 90 persons, swallowing fine particles or piercing and at that twelve participants were from abroad. sucking protists or multicellular organisms. All the presentations were held either in Among poster presentations, the poster of English with subsequent comments in Nic Smol (Ghent) and oth. “An introductory

This Newsletter is not part of the scientific literature for taxonomic purposes 6 guide to NeMys …” drew the most vivid Crustacean Society was proposed, with the attention of participants. aim to create a Russian Crustacean Society Alexei Tchesunov associated with the Russian Academy of Sciences. The coordinators of such efforts are Conference "Crustacea: diversity, ecology A.A. Kotov, I.N. Marin (A.N. Severtsov and evolution" in Moscow and creation of Institute of Ecology and Evolution, the Russian Crustacean Society Moscow), N.V. Aladin (Zoological Institure, St.-Petersburg), G.A. Kolbasov, V.N. A conference "Crustacea: diversity, ecology Ivanenko (M.V. Lomonosov Moscow State and evolution" was held at A.N. Severtsov University, Moscow), V.A. Spiridonov (P.P. Institute of Ecology and Evolution, Moscow, Shirshov Institute of Oceanology, Moscow), Russia on October 30th to November 2nd of V.V. Takhteev (Irkutsk State University, 2017 as the A.N. Severtsov Institute of Irkutsk) and I.S. Turbanov (Inland Water Ecology and Evolution of Russian Academy Biology Institute, Borok). Our final aim is to of Sciences. More than 100 investigators met create the Russian Crustacean Society with a in Moscow, and 72 researchers from 16 legal judicial status in Russia, being a non- different towns of Russia and also Norway commercial "Interregional public association and Belarus presented 15 plenary lectures, 40 without creation of a legal entity" with the oral presentations and 42 posters grouped following objectives: into five sections: (1) Taxonomy, (1) Overcoming a disunity of the zoogeography, faunistics: recent approaches carcinologists dealing with different groups to taxonomic revisions and inventories of the by different methods in different Regions of biodiversity; (2) Morphology, anatomy, Russia by exchanging of information (i.e. by embryology and physiology; (3) Ecology and specially created Web portal behaviour; (4) Palaeolimnology, www.crustacea.ru, organization of palaeontology and evolutionary conferences and workshops; reconstructions; (5) Recent methods of the (2) Help in the planning and conducting of crustacean investigations: achievements, carcinological investigations, publication of problems and perspectives. This conference results and introduction of fundamental was mainly aimed at an involving junior results of the Society members for practical biologists in crustacean studies. Special use; prizes were awarded to junior scientists for 3) Popularization of the crustaceans as the best oral communications and the best objects of any scientific works, attraction of posters. The Organization Committee also junior scientists to the crustacean problems; assigned a special prize to two students of a 4) Informing about possible fund sources for primary and secondary school in the town of scientific activity in carcinology; Blagoveshchensk (Amur Area) for their 5) Promotion of a cooperation with poster "Triopsids, living fossils from the crustacean investigators from other countries. water bodies of the Amur Area". Resolving any questions concerning the We were happy to explore that investigators formal association with the Russian from several regional Russian institutes and Academy of Sciences (and administrative universities are conducting projects of a high formalities concerned such association) is international rank in several directions of addressed to Dr. A.A. Kotov, a carcinological research. But we need to note Corresponding Member of the Russian that many regional Russian scientific Academy of Sciences. organizations, with a good potential for In May of 2018 a next National Conference crustacean studies, have problems with on the Crustacea will be organized at the quality of their publication, access to recent Inland Water Biology Institute, Borok, literature and recent methods of scientific Yaroslavl Area. This conference will be at the work. And scientists from large cities of same time a 1st Congress of the Russian Russia have a chance to help them. At the Crustacean Society where it will be created as conference, the voluntary Russian a legal non-commercial public organization,

This Newsletter is not part of the scientific literature for taxonomic purposes 7 also its chapter will be adopted and its suggest a considerable degree of authority for the time before the next heterogeneity in benthic metabolism along Congress will be elected. the deepest part of the trench axis (>9000 m), as well as clear evidence of turbidites, which Alexey Kotov likely have an influence on meiofaunal communities. Analyses of metazoan Eхpedition meiofauna from the trench, which will be led by Daniel Leduc at NIWA, will focus on Studying the meiofauna of the world’s community ecology, molecular sequencing deepest environments and environmental DNA approaches, as well as food web biomarkers. Analyses of foraminiferans will be conducted by Hiroshi Kitazato (JAMSTEC). The next voyage, due to depart from Chile in March 2018 and led by Frank Wenzhöfer from Max Planck Institute, will explore the Atacama Trench with Daniela Zeppilli from Ifremer leading the sampling and analyses of metazoan meiofauna. The research is primarily funded by a 5-year European Research Council Advanced Grant University of Southern Denmark’s profiler (ERC adG 2014 669947), with additional lander being retrieved from the depths of the support from various national research Kermadec Trench by NIWA’s research programmes (including from NIWA). vessel Tangaroa. Daniel Leduc

A voyage to the Kermadec Trench, the first New members of a series of expeditions exploring selected Prof. Juan I. Cañete trenches of the Pacific Ocean, has recently Research interests: benthic meiofaunal been completed. This voyage is part of the polychaetes, meroneustonic meiofauna HADES-ERC project, which will provide the (Polygordius, Microsetella) first detailed, combined analysis of benthic diagenesis and microbial/meiofaunal ecology I'm Chilean marine biologist. My of some of the deepest oceanic trenches on professional title was obtained in the Catholic Earth. Voyage leaders were Ronnie Glud University of North of Chile (www.ucn.cl)and my Ms Sc. postgraduate from the University of Southern Denmark, studies was obtained in the University of and Ashley Rowden from NIWA. Concepcion (www.udec.cl). In the first During this voyage, a range of sophisticated university my thesis was referred to the autonomous landers as well as traditional taxonomy, ecology and biogeography of sampling methods were used to sample benthic polychaetes from Easter island and in meiofauna down to Scholl Deep, the deepest the second university my thesis was on the larval and early postlarval ecology of the point of the Kermadec Trench. The team was edible squat-lobster Pleuroncodes monodon able to obtain the deepest sediment samples on the continental shelf of Concepcion Bay, ever retrieved using wire cable-deployed central Chile. My interest on marine instruments: a boxcorer at 9994 m, and a meiofauna began in 1997 when I participate multicorer at 9690 m. Preliminary results the International Atacama Trench Expedition

This Newsletter is not part of the scientific literature for taxonomic purposes 8 (ATIE) at the team of Norberto Della Crocce, meiofauna and ecosystem services provided Chilean benthologist and oceanographer. In by this interesting group. this cruise an important hotspot of deep-sea nematodes biodiversity was discovery (see Prof. Juan I. Cañete some papers of Danovaro et al). Since 2010, University of Magallanes I began to study the neustonic pelagic Punta Arenas – Chile communities of the estuarine zone dominated by channels and fjords of the southern Chile. Tania Figueroa Delgado Some important discovery allow me to map Research interests: benthic macrofauna, the relationship between salinity and spatial meiofauna, neuston and marine pollution. distribution of some meiofaunistic Polygordius/Microsetella larvae along I´am a Chilean marine biologist. I obtained Puerto Montt (40°S) to Navarino Island my professional title in the University of (56°)coast, in this important biogeographic Magallanes in Punta Arenas, Chile; my pre- subantarctic province (sensu Spalding et al., graduate thesis was about marine pollution, 2007). I worked and studied in four Chilean specifically in how the heavy metals affect universities: the expression of the gene of protein in the 1989-1991: Catholic University of North Antarctic sea urchin Sterechinus neumayeri, 1992-1994: University of Concepcion I obtained a scholarship to work in the 1994-1998: University of Valparaiso Antarctic with the INACH (Chilean Antarctic 1999-to present: University of Magallanes Institute). After that, I worked in other (UMAG). research institute implementing a molecular In the UMAG, now I teach on Invertebrate biology lab and finally I returned to zoology, Vertebrate zoology Universidad de Magallanes, were now I (Protochordates), Environment Management, work as a research assistant with the Integrated Management of Coastal Zone and professor Ivan Cañete. This year I started to Biological Oceanography. In the class I study again to obtain my master degree in the establish the role of marine benthic program of this university (Magister en communities in studies of biodiversity and as Ciencias con mención enmanejo y environmental tool for monitoring pollution conservación de recursos naturales and marine benthic resources. subantárticos),and now I am developing a Through of the financial support of Chilean thesis project about the composition and Agency of Sciences and Technology, this abundance of marine meiofauna in the year I can to fund the visit of the member of Magallanes region; meiofauna has not been International Association of studied much in this region. Meiobenthologists, German meiobenthologist Dr. Andreas Schmidt- Tania Figueroa Delgado Rhaesa to work on meiobenthic communities Lic.en Ciencias del Mar located along Chilean sandy beach. In this Bióloga Marina initiative also participate meiobenthologists Matthew Lee and Javier Sellanes. The first is Dr. Nigel J. Marley (UK, Plymouth) also member of IAM. We expect to build a Interests: Returning to the Association after South American scientific net to study the many years mostly due to health issues. Still special features of the Chilean marine mainly working on Tardigrada from all meiofauna. The first step of this will be the environments but also interested in all visit of my student Mrs. Tania Figueroa to the meiofauna. Currently focusing on developing Laboratory of Andreas Schmidt-Rhaesa in integrated diagnostic keys and nomenclature December 2017. I hope follow searching lists of names. financial support to increase the number of students and research on subantarctic and Dr. John Moverley Australia, Camberwell Chilean marine and estuarine biodiversity of Research interests: Papua New Guinea deep sea meiofauna communities.

This Newsletter is not part of the scientific literature for taxonomic purposes 9 Hummon who died at home in Athens, Ohio José Cerca de Oliveira (Norway, Oslo) (USA), on January 1, 2017. He was born on July 27, 1932, in Akron, I am interested in the evolutionary history of Ohio, the son of Dale Levi Hummon and meiofauna and invertebrates in general. This Lorena May Payne Hummon. On December includes studying intraspecific (demographic 24, 1958 he married Julia Margaret (Meg) history, selection - broadly population Raper at her family home in Oakton, genomics) and interspecific (clade Virginia. They have two daughters, Julia diversification and speciation) levels. Margaret Hummon of Honolulu, Hawaii, and Additionally, I am interested at patterns of Cheryl Hummon of Salem, Oregon. molecular and genome evolution (e.g. After earning a BA in Philosophy at the genomic architecture). To study these, I University of Montana, Missoula (1955), Bill employ population genomic tools such as was a conscientious objector to military RAD-seq and genome assembly. Following service, and served as an attendant at Apple these interests I am currently a PhD at the Creek State Hospital in Ohio in lieu of University of Oslo, under Torsten Struck. I military service (1955-58). He then returned am quantifying genomic and morphological to the University of Montana for a BS in divergence in a complex of cryptic wildlife biology (1960), and an MA in meiofaunal annelids (Stygocapitella Teaching of Biology (1961). Subsequently, subterranea) and delving into the Bill received a PhD in Zoology at the evolutionary history of this complex. University of Massachusetts, Amherst José Cerca de Oliveira (1969), where he conducted seminal research on marine gastrotrichs of New England, described in his thesis “Distributional ecology of marine interstitial Gastrotricha Remembering Bill Hummon from Woods Hole, Massachusetts, with (1932-2017) taxonomic comments on previously described species”. His Ph.D. research marked the beginning of a life-long dedication to the Gastrotricha, that enigmatic group of beautiful meiofaunal invertebrates that have captured the attention of zoologists worldwide, and continue to inspire scientists in their quests to understand some of the greatest mysteries of animal evolution. Bill went on to become an inspiring professor of marine biology and zoology at Ohio University, Athens in 1969 (until his retirement in 2002). During his tenure, he was a recipient of several fellowships and Bill Hummon at St. John Island in 2011, scholarships which made it possible to him to continuing research on the Caribbean travel to and work in England, Scotland and gastrotrichs (see Hummon W. D. 2010. Egypt. Bill’s impact on the discipline grew Marine Gastrotricha of the Caribbean Sea: a with each passing year; he supervised eleven review and new descriptions. Bulletin of MS students, two Ph.D. students, three Marine Science, 86: 661-708. postdoctoral researchers, and produced 110 publications on invertebrate morphology, It is with a deep sorrow and a feeling of big taxonomy, ecology, ecotoxicology, and the loss that we are writing this note to philosophy of science. Bill was active in commemorate our friend, mentor, teacher several scientific societies, acted as President and collaborator, William (Bill) Dale for the International Association of Meiobenthologists (IAM, 1978-79) and was

This Newsletter is not part of the scientific literature for taxonomic purposes 10 elected a Fellow of the American Association charge) via the Internet. He provided the first for the Advancement of Science. video recording (SVHS format) of a new Over 43 illustrious years, he worked with taxon, Prostobuccantia broca Evans & gastrotrichs, copepods, crayfish, insects, Hummon, 1991, which was deposited in the rotifers and tardigrades, and described 2 National Museum of Natural History, families, 6 genera, and 105 species of Washington, DC (USNM 235577). Since gastrotrichs along the way. Bill’s focus on then, Bill has made innumerable video marine Gastrotricha allowed him the rare recordings (in digital format) of new and (and envious) opportunity to travel across known species from across the globe, and some of the most historic and beautiful provided his followers hours (and gigabytes) coastal landscapes of the Azores, Belgium, of video data, photographic images, GPS France, Germany, Greece, Ireland, Israel, coordinates, maps, and taxon lists that remain Italy, Poland, Portugal, the United Kingdom, as influential as any DNA barcode. the Caribbean, and Central and South Originally posted on a dedicated server, America. And while his quest to understand Bill’s works are now part of the Gastrotricha gastrotrichs almost always had a global World Portal component, this did not mean he ignored his (http://www.gastrotricha.unimore.it/moviega own backyard. Bill’s research on fauna of the llery.htm). east and west coasts of the United States As well as being a dedicated and remains some of his most influential involved scientist whom we admired and taxonomic work on the phylum, and even learned from, Bill was – to us also a great, included studies of the effects of pollutants, close and supportive friend, appreciative of such as DDT and mine acids, on freshwater good conversation over a dram of single malt, species. But whether Bill was traversing the fond of music (and cats), and ready to bring USA in his mobile lab (his vehicle full of people together. portable microscopes and equipment) or Bill’s influence on our scientific traveling across vast oceans and seas, he understanding and appreciation of always kept one eye on the sand and one eye Gastrotricha and the meiofauna in general on the microscope. cannot be overstated. He inspired scientists Bill’s importance to across the globe, many of whom have never research went well beyond his influential had the chance to meet this extraordinary publications on systematics and ecology. He man, yet, will continue to learn from him well also highlighted the very real and very into the 21st century and beyond. Bill’s legacy practical problem of conserving type will live in the annals of scientific discovery, specimens for natural history museums, and in the fond memories of those who knew which is nothing short of impossible when him and who will never forget him. dealing with soft-bodied meiofauna. Bill’s hand-drawn illustrations of species always M. Antonio Todaro made identification relatively easy for the Maria Balsamo uninitiated, but without good physical John Fleeger specimens in museum collections, one could Teresa Radziejewska never be sure if the animal found on one Rick Hochberg beach was the same as that on another. Nowadays, molecular sequencing is the preferred choice for species synonymy (and A list of Bill’s publication can be found at differentiation), but this requires a dedicated http://www.gastrotricha.unimore.it/news.ht lab, skills, and most importantly, funding, m which can be in short supply for those who work on meiofauna. Bill’s solution to this dilemma predated the molecular revolution and in fact was much simpler, and importantly, globally accessible (free of

This Newsletter is not part of the scientific literature for taxonomic purposes 11 The World Register of Marine Species Electronic resources (WoRMS) is available at www.marinespecies.org. The text on the Meiobenthic groups within the World WoRMS pages is open-access under the Register of Marine Species terms of the Creative Commons Attribution License (CC-BY), which permits unrestricted The aim of a World Register of Marine use as long as the provided citation is used. Species (WoRMS) is to provide an Upon request by the editors-in-charge, a authoritative and comprehensive list of dedicated portal can be created for a names of marine organisms, including taxonomic group – referred to as a Global information on synonymy. While the highest Species Database – giving more visibility to priority goes to valid names, other names in their group and recognition to the involved use are included so that this register can serve editors. It also allows editors to elaborate as a guide to interpret taxonomic literature. more on their group and - in some cases – a Next to taxonomy, the system can also store dedicated news feed and twitter messages. distribution information, literature, attributes An overview of the available registers for or traits, specimen information including phyla containing meiobenthic representatives type localities and images. Total number of species names (accepted & unaccepted) available in each group (including all environments, excluding fossil species) Taxa # species Portal Cnidaria 22 560 through WoRMS

Platyhelminthes 24 602 http://www.marinespecies.org/turbellarians/

Nemertea 2 725 http://www.marinespecies.org/nemertea/ Gnathostomulida 113 through WoRMS Kinorhyncha 363 through WoRMS Loricifera 28 through WoRMS

Nematoda 10 855 http://nemys.ugent.be/ Rotifera 317 through WoRMS Gastrotricha 1 069 through WoRMS Priapulida 21 through WoRMS Sipuncula 1 294 through WoRMS through WoRMS (in part: Annelida 25 439 http://www.marinespecies.org/polychaeta)

Arthropoda - Copepoda 23 071 http://www.marinespecies.org/copepoda/ Arthropoda - 1462 through WoRMS Halacaroidea

Arthropoda - Ostracoda 14 521 http://www.marinespecies.org/ostracoda/ Arthropoda - 14 through WoRMS Mystacocarida Arthropoda - 42 through WoRMS Tantulocarida

Tardigrada 1588 http://www.marinespecies.org/tardigrada/

Mollusca 113 831 http://www.molluscabase.org/ Bryozoa 8 169 through WoRMS Brachiopoda 679 through WoRMS through WoRMS (in part: Echinodermata 21 176 http://www.marinespecies.org/echinoidea & http://www.marinespecies.org/asteroidea) Chordata 59 753 through WoRMS Sarcomastigota 435 through WoRMS Ciliophora 4 188 through WoRMS

This Newsletter is not part of the scientific literature for taxonomic purposes 12 is given in the table. Next to the marine Bastami, Kazem Darvish, et al. "Nematode representatives, editors can also document community structure in relation to metals in the non-marine taxa in their group, thereby the southern of Caspian Sea." Acta creating a single entry-point for their cross- Oceanologica Sinica 36.10 (2017): 79-86. environment group. Within the attributes section of WoRMS, it is Caridi, Francesca, et al. "A new Arctic the goal to document the size class to which seepage site? Preliminary evidence from each species belongs, so a general filter can benthic community." EGU General be applied to group e.g. all meiofaunal Assembly Conference Abstracts. Vol. 19. species across taxa. This is still a work in 2017. progress. In addition, it will be documented whether species belong to the benthos or the Cesaroni, Lucia, et al. "Scanning electron pelagos, taking into account the life stage. microscopy in the taxonomical study of free- Keeping WoRMS and its Global Species living marine nematodes." Microscopie 28.2 Databases up-to-date is a continuous process. (2017): 31-38. New information is entered daily by the taxonomic editors and by the members of our Cvitković, Ivan, et al. "Structure of data management team. No database of this epibiontic and sediment meiofauna in the size is without errors and omissions. area invaded by invasive alga Caulerpa Although we cannot promise to make no taxifolia." Marine Biology 164.1 (2017): 4. errors, but we do promise to follow up and give feedback on any communications de Jesús-Navarrete, Alberto. "Littoral free pointing out errors. All feedback is very living nematode fauna of Socorro Island, welcome through [email protected]. Colima, Mexico." Hidrobiológica 17.1 (2017): 61-66. Leen Vandepitte Fu, Sujing, et al. "Nematode responses to the invasion of exotic Spartina in mangrove wetlands in southern China." Estuaries and Coasts 40.5 (2017): 1437–1449. Recent Literature Fu, Sujing, Guy Boucher, and Lizhe Cai. Marine and general "Two new ovoviviparous species of the Annapurna, C., et al. "Lorcifera, an under family Selachinematidae and known phyla: first record of higgins larva of Sphaerolaimidae (Nematoda, Chromadorida Armorloricus (Loricifera: Nanaloricidae) & Monhysterida) from the northern South from Indian waters." 46.02 (2017): 317-321. China Sea." Zootaxa 4317.1 (2017): 95-110. Ansari, Kapuli Gani Mohamed Thameemul, Gao, Qun, and Yong Huang. "Oncholaimus and Punyasloke Bhadury. "An updated zhangi sp. nov.(Oncholaimidae, Nematoda) species checklist for free-living marine from the intertidal zone of the East China nematodes from the world’s largest Sea." Chinese Journal of Oceanology and mangrove ecosystem, Limnology 35.5 (2017): 1212-1217. Sundarbans." Zootaxa 4290.1 (2017): 177- Garraffoni, André Rinaldo Senna. 191. "Checklist of Gastrotricha from Mato Bartels, Paul J., Lilvia J. Bradbury, and Grosso do Sul state, Brazil." Iheringia. Série Diane R. Nelson. "Marine tardigrades from Zoologia 107 (2017): 1-4. South Carolina, USA." Journal of the South Garraffoni, André RS, Maikon Di Carolina Academy of Science 15.1 (2017): Domenico, and Rick Hochberg. "New 7. records of marine Gastrotricha from São Sebastião Island (Brazil) and the description

This Newsletter is not part of the scientific literature for taxonomic purposes 13 of a new species." Marine Biodiversity47.2 Argentine coast." Zootaxa 4250.4 (2017): (2017): 451-459. 347-357. Gąsiorowski, Ludwik, et al. "New insights Montagna, Paul A., et al. "Persistent impacts on the musculature of filospermoid to the deep soft‐bottom benthos one year Gnathostomulida." Zoomorphology 136.4 after the Deepwater Horizon (2017): 413-424. event." Integrated environmental assessment and management 13.2 (2017): 342-351. Gomes-Júnior, Edivaldo, et al. "A new species of Ligiarctus (Tardigrada, Morad, Tzachy Y., Zvy Dubinsky, and Arthrotardigrada) from the Brazilian David Iluz. "Meiobenthos Assemblages as continental shelf, Southwestern Atlantic Bioindicators for Coastal Pollution Ocean." Marine Biodiversity (2017): 1-8. Assessment." Science 7 (2017): 409-423. Hochberg*, Rick, et al. "A Tribute to Netto, Sergio A., and Gustavo Fonseca. William Hummon–Gastrotrich Biologist "Regime shifts in coastal lagoons: Evidence Extraordinaire." Proceedings of the from free-living marine nematodes." PloS Biological Society of Washington 130.1 one12.2 (2017): e0172366. (2017): 113-119. Nguyen, Dinh Tu, et al. "Meiobenthos Janakiraman, A., et al. "Ecological communities for different mangrove types in restoration assessment of Adyar creek and Can Gio Biosphere Reserve, estuary using meiofaunal communities as Vietnam." Proceedings of DalRybVtuz ecological indicators for aquatic (Vladivostok). 41 (2017): 74-84 pollution." Regional Studies in Marine Oh, Je Hyeok, et al. "Effect of increased p Science 9 (2017): 135-144. CO2 in seawater on survival rate of different Jouili, Soufiane, et al. "Environmental developmental stages of the harpacticoid quality assessment of El Bibane lagoon copepod Tigriopus japonicus." Animal Cells (Tunisia) using taxonomic and functional and Systems 21.3 (2017): 217-222. diversity of meiofauna and Pape, Ellen, et al. "Limited Spatial and nematodes." Journal of the Marine Temporal Variability in Meiofauna and Biological Association of the United Nematode Communities at Distant but Kingdom 97.8 (2017): 1593-1603. Environmentally Similar Sites in an Area of Kånneby, Tobias, and M. Antonio Todaro. Interest for Deep-Sea Mining." Frontiers in "A new species of Aspidiophorus Marine Science 4 (2017): 205. (Gastrotricha: ) from the Pereira, Tiago José, et al. "Patterns of Swedish west coast." Zootaxa 4290.2 Spatial Variation of Meiofauna in Sandy (2017): 390-394. Beaches of Northwestern Mexico with Kiko, Rainer, et al. "Colonization of newly Contrasting Levels of forming Arctic sea ice by meiofauna: a case Disturbance." Thalassas: An International study for the future Arctic?." Polar Journal of Marine Sciences (2017): 1-11. Biology 40.6 (2017): 1277-1288. Prasath, D., et al. "New record of two free- Martelli, Antonela, et al. "Two new species living marine nematode species, of free-living marine nematodes of the Sphaerolaimus balticus and Sphaerolaimus family Oxystominidae Chitwood, 1935 islandicus (Nematoda: Sphaerolaimaidae) () with a review of the genus from Sipphighat mangrove region, South Thalassoalaimus de Man, 1893 from the Andaman." 46 (2017): 1105-1109.

This Newsletter is not part of the scientific literature for taxonomic purposes 14 Radziejewska, Teresa, Jonne Kotta, and Shi, Benze, and Kuidong Xu. Lech Kotwicki. "Sandy coasts." Biological "Spirobolbolaimus undulatus sp. nov. in Oceanography of the Baltic Sea. Springer intertidal sediment from the East China Sea, Netherlands, 2017. 457-482. with transfer of two Microlaimus species to Molgolaimus (Nematoda, Revkova, Tatiana N. "First records of Desmodorida)." Journal of the Marine Microlaimus tenuispiculum de Man, 1922 Biological Association of the United (Nematoda: Microlaimidae) from the Black Kingdom97.6 (2017): 1335-1342. Sea." Ecologica Montenegrina 14 (2017): 74-79. Stark, Jonathan S., et al. "The effects of hydrocarbons on meiofauna in marine Rosli, Norliana, et al. "Review of recent sediments in Antarctica." Journal of trends in ecological studies of deep-sea Experimental Marine Biology and meiofauna, with focus on patterns and Ecology 496 (2017): 56-73. processes at small to regional spatial scales." Marine Biodiversity (2017): 1-22. Struck, Torsten H., et al. "Two new species in the annelid genus Stygocapitella Rubal, Marcos, et al. "A new Batillipes (Orbiniida, Parergodrilidae) with comments (Tardigrada, Heterotardigrada, on their biogeography." Zootaxa 4286.3 Batillipedidae) from North Portugal (2017): 301-332. (Atlantic Ocean)." Marine Biodiversity 47.3 (2017): 921-928. Tarhan, Lidya G. "Meiofauna mute the Cambrian Explosion." Nature Ecology & Ruiz-Abierno, Alexei, and Maickel Evolution 1.10 (2017): 1423. Armenteros. "Coral reef habitats strongly influence the diversity of macro-and Tchesunov, Alexei V. "Free-living meiobenthos in the Caribbean." Marine nematodes of the genus Syringolaimus de Biodiversity 47.1 (2017): 101-111. Man, 1888 (Nematoda, Enoplida, Ironidae) from the Angola Basin, South-East Atlantic Sahoo, Gobardhan, et al. "Defaunation of Abyss." Zootaxa 4306.4 (2017): 478-500. meiofauna in Mumbai bay (India)-A severely polluted area." Regional Studies in Thai, Tran Thanh, et al. "MEIOFAUNA IN Marine Science 16 (2017): 98-108. THE MANGROVE–SHRIMP FARMS PONDS, CA MAU PROVINCE." Journal Sahoo, Gobardhan, et al. "Epibiotic of Science and Technology 55.3 (2017): 271. communities (microalgae and meiofauna) on the pneumatophores of Avicennia officinalis Yanko-Hombach, Valentina V., et al. (L.)." Estuarine, Coastal and Shelf "Meiobenthos as an indicator of gaseous Science (2017). hydrocarbon reservoirs under floor of the Black sea." (2017). Joint Plenary Santos, T. M. T., and V. Venekey. Conference and Field Trip of IGCP 610 and "Meiofauna and free-living nematodes in INQUA IFG POCAS Palermo, Italy, 1-9 volcanic sands of a remote South Atlantic, October 2017: 224-229 oceanic island (Trindade, Brazil)." Journal of the Marine Biological Association of the Yin, Shengle, et al. "Seasonal dynamics of United Kingdom (2017): 1-16. meiofaunal distribution in the Dagu River Estuary, Jiaozhou Bay, China." Acta Sergeeva, Nelli G., and Derya Ürkmez. Oceanologica Sinica 36.12 (2017): 79-86. "Current views on the diversity and distribution of deep-water meiobenthos at Zainuri, Muhammad, Sutrisno Anggoro, and the Turkish shelf (Black Sea)." Ecologica Hermin Pancasakti Kusumaningrum. Montenegrina 14 (2017): 60-73. "Proximate Content of

This Newsletter is not part of the scientific literature for taxonomic purposes 15 “Klekap”(Microphytobenthos and Their on their occurrence, distribution and Associated Meiofauna) from Milk-Fish ecological relevance. Nematology 19 Pond." IOP Conference Series: Earth and (6):627-644 Environmental Science. Vol. 55. No. 1. IOP Garraffoni ARS (2017) Checklist of Publishing, 2017. Gastrotricha from Mato Grosso do Sul state, Freshwater Brazil. Iheringia Série Zoologia 107:doi: 10.1590/1678-4766e2017104 Arican C, Traunspurger W, Spann N (2017) The influence of thiacloprid on the feeding Garraffoni ARS, Araujo TQ, Lourenço AP, behaviour of the copepod, Diacyclops Guidi L, Balsamo M (2017) A new genus bicuspidatus, preying on nematodes. and new species of freshwater Nematology 19:1201-1215 Chaetonotidae (Gastrotricha: ) from Brazil with phylogenetic position Baranov V, Milosevic D, Kurz MJ, inferred from nuclear and mitochondrial Zarnetske JP, Sabater F, Marti E, Robertson DNA sequences. Systematics and AL, Brandt T, Sorolla A, Lewandowski J Biodiversity 15 (1):49-62 (2017) Helophyte impacts on the response of hyporheic invertebrate communities to Gusakov, Vladimir A., and Vladimir G. inundation events in intermittent streams. Gagarin. "An annotated checklist of the Ecohydrology doi: 10.1002/eco.1857 main representatives of meiobenthos from inland water bodies of Central and Southern Bozkurt A (2017) First Record of Vietnam. I. Roundworms Epactophanes richardi Mrazek, 1893 (Nematoda)." Zootaxa 4300.1 (2017): 1-43. (Copepoda, Harpacticoida, Camptocamptidae) for Turkish Inland Haegerbaeumer A, Höss S, Heininger P, Waters. Turkish Journal of Fisheries and Traunspurger W (2017) Is Caenorhabditis Aquatic Sciences 17 (1):25-29 elegans representative of freshwater nematode species in toxicity testing? Chambord S, Tackx M, Chauvet E, Escolar Environmental Science and Pollution G, Colas F (2017) Two microcrustaceans Research doi: 10.1007/s11356-017-0714- affect microbial and macroinvertebrate- 7:1-10 driven litter breakdown. Freshwater Biology 62 (3):530-543 Haegerbaeumer A, Höss S, Heininger P, Traunspurger W (2018) Response of da Silva GL, Metzelthin MH, da Costa T, nematode communities to metals and PAHs Rocha MS, Silva DE, Ferla NJ, da Silva OS in freshwater microcosms. Ecotoxicology (2017) Responses of water mite assemblages and environmental safety 148:244-253 (Acari) to environmental parameters at irrigated rice cultivation fields and native Haegerbaeumer A, Höss S, Ristau K, Claus lakes. Zoologia (Curitiba) 34:doi: E, Heininger P, Traunspurger W (2017) The 10.3897/zoologia.3834.e19988 use of meiofauna in freshwater sediment assessments: Structural and functional Drazina T, Spoljar M, Primc B, Habdija I responses of meiobenthic communities to (2017) Distribution of rotifers and other metal and organics contamination. meiofauna in the bryophytes and hyporheic Ecological Indicators 78:512-525 zone of a karst hydrosystem-an example of a nested community. Marine and Freshwater Kolicka M (2017) New Arctic Gastrotrich Research 68 (1):43-52 — Three Chaetonotus (Chaetonotidae) from Longyearbyen (Spitsbergen, Svalbard Du Preez G, Majdi N, Swart A, Archipelago). Annales Zoologici 67:131- Traunspurger W, Fourie H (2017) 180 Nematodes in caves: a historical perspective

This Newsletter is not part of the scientific literature for taxonomic purposes 16 Kolicka M, Gadawski P, Dabert M (2017) A with descriptions of three new species. new species of freshwater Chaetonotidae Zootaxa 4318 (1):1-46 (Gastrotricha, Chaetonotida) from Obodska Spoljar M, Zhang C, Drazina T, Zhao G, Cave (Montenegro) based on morphological Lajtner J, Radonic G (2017) Development of and molecular characters. European Journal submerged macrophyte and epiphyton in a of Taxonomy 354:1-30 flow-through system: Assessment and Kreuzinger-Janik B, Brinke M, modelling predictions in interconnected Traunspurger W, Majdi N (2017) Life reservoirs. Ecological Indicators 75:145-154 history traits of the free-living nematode, Tsiafouli MA, Bhusal DR, Sgardelis SP Plectus acuminatus Bastian, 1865, and (2017) Nematode community indices for responses to cadmium exposure. microhabitat type and large scale landscape Nematology 19 (6):645-654 properties. Ecological Indicators 73:472-479 Mendoza G, Traunspurger W, Palomo A, Veras, Tatiane Barbosa, et al. "Interação rio- Catalan J (2017) Nematode distributions as aquífero e a meiofauna do ambiente spatial null models for macroinvertebrate hiporreico." Águas Subterrâneas 31.1 species richness across environmental (2017): 20-35. gradients: A case from mountain lakes. Ecology and evolution 7 (9):3016-3028 Wiackowski K, Kocerba-Soroka W (2017) Selective predation by a harpacticoid Olson DS, Janssen J (2017) Early feeding of copepod on ciliates in phytotelmata: a round goby (Neogobius melanostomus) fry. laboratory experiment. Hydrobiologia 790 Journal of Great Lakes Research 43 (4):728- (1):13-22 736

Parry LA, Boggiani PC, Condon DJ, Garwood RJ, Leme JM, McIlroy D, Brasier Forthcoming. MD, Trindade R, Campanha GAC, Pacheco As a rule, the list of publications contains LAF (2017) Ichnological evidence for printed papers only, not “on-line first”. As meiofaunal bilaterians from the terminal an exception, we put here the reference to Ediacaran and earliest Cambrian of Brazil. important review, which will open the paper Nature Ecology & Evolution 1 (10):1455- collection on a topic of great importance: 1464 Zeppilli, D., Leduc, D., Fontanier, C. et al. Ptatscheck C, Putzki H, Traunspurger W “Characteristics of meiofauna in extreme (2017) Impact of deposit-feeding marine ecosystems: a review” Marine chironomid larvae (Chironomus riparius) on Biodiversity (2017). meiofauna and protozoans. Freshwater https://doi.org/10.1007/s12526-017-0815-z Science 36 (4):796-804 Rosati M, Rossetti G, Cantonati M, Pieri V, Roca JR, Mesquita-Joanes F (2017) Are aquatic assemblages from small water bodies more stochastic in dryer climates? An analysis of ostracod spring metacommunities. Hydrobiologia 793 (1):199-212 Shearn R, Schön I, Martens K, Halse S, Krawiec J, Koenders A (2017) Patterns of genetic divergence in the Ilyodromus amplicolis lineage (Crustacea, Ostracoda),

This Newsletter is not part of the scientific literature for taxonomic purposes 17 INTERNATIONAL ASSOCIATION OF MEIOBENTHOLOGISTS

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This Newsletter is not part of the scientific literature for taxonomic purposes 18