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Quaternary International 603 (2021) 40–63
Quaternary International 603 (2021) 40–63 Contents lists available at ScienceDirect Quaternary International journal homepage: www.elsevier.com/locate/quaint Taxonomy, taphonomy and chronology of the Pleistocene faunal assemblage at Ngalau Gupin cave, Sumatra Holly E. Smith a,*, Gilbert J. Price b, Mathieu Duval c,a, Kira Westaway d, Jahdi Zaim e, Yan Rizal e, Aswan e, Mika Rizki Puspaningrum e, Agus Trihascaryo e, Mathew Stewart f, Julien Louys a a Australian Research Centre for Human Evolution, Environmental Futures Research Institute, Griffith University, Nathan, Queensland, 4111, Australia b School of Earth and Environmental Sciences, The University of Queensland, St Lucia, Queensland, 4072, Australia c Centro Nacional de Investigacion´ Sobre la Evolucion´ Humana (CENIEH), Burgos, 09002, Spain d Department of Earth and Environmental Sciences, Macquarie University, Sydney, New South Wales, Australia e Geology Study Program, Institut Teknologi Bandung, Jawa Barat, 40132, Indonesia f Extreme Events Research Group, Max Planck Institutes for Chemical Ecology, the Science of Human History, and Biogeochemistry, Jena, Germany ARTICLE INFO ABSTRACT Keywords: Ngalau Gupin is a broad karstic cave system in the Padang Highlands of western Sumatra, Indonesia. Abundant Taxonomy fossils, consisting of mostly isolated teeth from small-to large-sized animals, were recovered from breccias Taphonomy cemented on the cave walls and unconsolidated sediments on the cave floor.Two loci on the walls and floorsof Cave Ngalau Gupin, named NG-A and NG-B respectively, are studied. We determine that NG-B most likely formed as a Pleistocene result of the erosion and redeposition of material from NG-A. The collection reveals a rich, diverse Pleistocene Southeast Asia Hexaprotodon faunal assemblage (Proboscidea, Primates, Rodentia, Artiodactyla, Perissodactyla, Carnivora) largely analogous ESR and U-series dating to extant fauna in the modern rainforests of Sumatra. -
Endangered Species (Protection, Conser Va Tion and Regulation of Trade)
ENDANGERED SPECIES (PROTECTION, CONSER VA TION AND REGULATION OF TRADE) THE ENDANGERED SPECIES (PROTECTION, CONSERVATION AND REGULATION OF TRADE) ACT ARRANGEMENT OF SECTIONS Preliminary Short title. Interpretation. Objects of Act. Saving of other laws. Exemptions, etc., relating to trade. Amendment of Schedules. Approved management programmes. Approval of scientific institution. Inter-scientific institution transfer. Breeding in captivity. Artificial propagation. Export of personal or household effects. PART I. Administration Designahem of Mana~mentand establishment of Scientific Authority. Policy directions. Functions of Management Authority. Functions of Scientific Authority. Scientific reports. PART II. Restriction on wade in endangered species 18. Restriction on trade in endangered species. 2 ENDANGERED SPECIES (PROTECTION, CONSERVATION AND REGULA TION OF TRADE) Regulation of trade in species spec fled in the First, Second, Third and Fourth Schedules Application to trade in endangered specimen of species specified in First, Second, Third and Fourth Schedule. Export of specimens of species specified in First Schedule. Importation of specimens of species specified in First Schedule. Re-export of specimens of species specified in First Schedule. Introduction from the sea certificate for specimens of species specified in First Schedule. Export of specimens of species specified in Second Schedule. Import of specimens of species specified in Second Schedule. Re-export of specimens of species specified in Second Schedule. Introduction from the sea of specimens of species specified in Second Schedule. Export of specimens of species specified in Third Schedule. Import of specimens of species specified in Third Schedule. Re-export of specimens of species specified in Third Schedule. Export of specimens specified in Fourth Schedule. PART 111. -
Investigating Sexual Dimorphism in Ceratopsid Horncores
University of Calgary PRISM: University of Calgary's Digital Repository Graduate Studies The Vault: Electronic Theses and Dissertations 2013-01-25 Investigating Sexual Dimorphism in Ceratopsid Horncores Borkovic, Benjamin Borkovic, B. (2013). Investigating Sexual Dimorphism in Ceratopsid Horncores (Unpublished master's thesis). University of Calgary, Calgary, AB. doi:10.11575/PRISM/26635 http://hdl.handle.net/11023/498 master thesis University of Calgary graduate students retain copyright ownership and moral rights for their thesis. You may use this material in any way that is permitted by the Copyright Act or through licensing that has been assigned to the document. For uses that are not allowable under copyright legislation or licensing, you are required to seek permission. Downloaded from PRISM: https://prism.ucalgary.ca UNIVERSITY OF CALGARY Investigating Sexual Dimorphism in Ceratopsid Horncores by Benjamin Borkovic A THESIS SUBMITTED TO THE FACULTY OF GRADUATE STUDIES IN PARTIAL FULFILMENT OF THE REQUIREMENTS FOR THE DEGREE OF MASTER OF SCIENCE DEPARTMENT OF BIOLOGICAL SCIENCES CALGARY, ALBERTA JANUARY, 2013 © Benjamin Borkovic 2013 Abstract Evidence for sexual dimorphism was investigated in the horncores of two ceratopsid dinosaurs, Triceratops and Centrosaurus apertus. A review of studies of sexual dimorphism in the vertebrate fossil record revealed methods that were selected for use in ceratopsids. Mountain goats, bison, and pronghorn were selected as exemplar taxa for a proof of principle study that tested the selected methods, and informed and guided the investigation of sexual dimorphism in dinosaurs. Skulls of these exemplar taxa were measured in museum collections, and methods of analysing morphological variation were tested for their ability to demonstrate sexual dimorphism in their horns and horncores. -
Lieberman 2001E.Pdf
news and views Another face in our family tree Daniel E. Lieberman The evolutionary history of humans is complex and unresolved. It now looks set to be thrown into further confusion by the discovery of another species and genus, dated to 3.5 million years ago. ntil a few years ago, the evolutionary history of our species was thought to be Ureasonably straightforward. Only three diverse groups of hominins — species more closely related to humans than to chim- panzees — were known, namely Australo- pithecus, Paranthropus and Homo, the genus to which humans belong. Of these, Paran- MUSEUMS OF KENYA NATIONAL thropus and Homo were presumed to have evolved between two and three million years ago1,2 from an early species in the genus Australopithecus, most likely A. afarensis, made famous by the fossil Lucy. But lately, confusion has been sown in the human evolutionary tree. The discovery of three new australopithecine species — A. anamensis3, A. garhi 4 and A. bahrelghazali5, in Kenya, Ethiopia and Chad, respectively — showed that genus to be more diverse and Figure 1 Two fossil skulls from early hominin species. Left, KNM-WT 40000. This newly discovered widespread than had been thought. Then fossil is described by Leakey et al.8. It is judged to represent a new species, Kenyanthropus platyops. there was the finding of another, as yet poorly Right, KNM-ER 1470. This skull was formerly attributed to Homo rudolfensis1, but might best be understood, genus of early hominin, Ardi- reassigned to the genus Kenyanthropus — the two skulls share many similarities, such as the flatness pithecus, which is dated to 4.4 million years of the face and the shape of the brow. -
Chapter6(PDF)
The University Museum The University of Tokyo Bulletin No. 47 KONSO-GARDULA RESEARCH PROJECT Volume 1 Paleontological Collections: Background and Fossil Aves, Cercopithecidae, and Suidae Edited by Gen Suwa, Yonas Beyene, and Berhane Asfaw ITY RS MU E S IV E U N M U U N I V O E Y R K SI O TY OF T 2014 TOKYO Editorial Board Yoshihiro Nishiaki (Editor-in-chief; Archaeology) Hiroshi Ikeda (Botany) Mari Kuroki (Marine Biology) Takenori Sasaki (Paleontology) Gen Suwa (Physical Anthropology) Eisei Tsurumi (Cultural Anthropology) Masaya Yago (Entomology) All communications pertaining to this Bulletin should be addressed to the Editorial Board, the University Museum, the University of Tokyo, 7-3-1 Hongo, Bunkyo-ku, Tokyo 113-0033, Japan. Issued March 20, 2014 ISSN 0910-481 X © The University Museum, The University of Tokyo Printed by Akita Kappan Printing Co., Ltd. CONTENTS Acknowledgements Chapter 1. Introduction (Gen Suwa, Yonas Beyene, and Berhane Asfaw) 1 Chapter 2. The Konso Formation Paleontological Assemblages: Collecting and Documentation Methodologies (Gen Suwa, Hideo Nakaya, and Berhane Asfaw) 5 Chapter 3. Stratigraphic and Chronologic Context of the Konso Formation Paleontology (Shigehiro Katoh, Gen Suwa, Hideo Nakaya, and Yonas Beyene) 11 Chapter 4. Fossil Birds of the Konso Formation (Antoine Louchart) 25 Chapter 5. Fossil Cercopithecidae of the Konso Formation (Stephen R. Frost) 41 Chapter 6. Fossil Suidae of the Konso Formation (Gen Suwa, Antoine Souron, and Berhane Asfaw) 73 Appendix 1. Aves Referred Materials 89 Appendix 2. Cercopithecidae Referred Materials 91 Appendix 3. Suidae Referred Materials and Dental Metrics 97 Related Archival Materials: The KGA paleontological collection record plots (on file, available for viewing upon request) CHAPTER 6 Fossil Suidae of the Konso Formation Gen Suwa1, Antoine Souron2, and Berhane Asfaw3 1The University Museum, The University of Tokyo, Hongo, Bunkyo-ku, Tokyo, 113-0033 Japan. -
New Skulls of Kolpochoerus Phacochoeroides (Suidae: Mammalia) from the Late Pliocene of Ahl Al Oughlam, Morocco
New skulls of Kolpochoerus phacochoeroides (Suidae: Mammalia) from the late Pliocene of Ahl al Oughlam, Morocco Denis Geraads UPR 2147 du CNRS, 44 rue de l’Amiral Mouchez, 75014 PARIS, France E-mail: [email protected] Received 10 August 2004. Accepted 20 December 2004. The discovery of two male skulls of Kolpochoerus phacochoeroides from the late Pliocene of Ahl al Oughlam in Morocco, and the revision of the whole collection from this locality, allows us to extend the description of this North African form, to estimate its sexual dimor- phism and the extent of individual variation in a large isochronous sample, to reveal some ontogenic changes, and to confirm its distinc- tion as a species on its own, as its cranial proportions (large occipital, short snout) and tooth characters (lack of enamel on upper canines, reduced incisors and premolars, complicated third molars) set it clearly apart from the East and South African forms. A cladistic analysis shows that K. phacochoeroides and Hylochoerus are the terminal branches of the Kolpochoerus clade, which is the sister-group of Potamochoerus. Keywords: Africa, Pliocene, Pleistocene, Suidae, Mammalia, Kolpochoerus, cladistics. INTRODUCTION SYSTEMATIC PALAEONTOLOGY The late Pliocene site of Ahl al Oughlam in Morocco has been excavated under the author’s leadership, as part of Genus Kolpochoerus van Hoepen & van Hoepen, 1932 the ‘Programme Casablanca’ of the Institut National des Mesochoerus Shaw & Cooke, 1941 Sciences de l’Archéologie et du Patrimoine of Rabat. It is Omochoerus Arambourg, 1943 the richest fossil locality of the North African Neogene, Promesochoerus Leakey, 1967 with about 55 species of mammals (Raynal et al. -
The First Occurrence of Eurygnathohippus Van Hoepen, 1930 (Mammalia, Perissodactyla, Equidae) Outside Africa and Its Biogeograph
TO L O N O G E I L C A A P I ' T A A T L E I I A Bollettino della Società Paleontologica Italiana, 58 (2), 2019, 171-179. Modena C N O A S S. P. I. The frst occurrence of Eurygnathohippus Van Hoepen, 1930 (Mammalia, Perissodactyla, Equidae) outside Africa and its biogeographic signifcance Advait Mahesh Jukar, Boyang Sun, Avinash C. Nanda & Raymond L. Bernor A.M. Jukar, Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington DC 20013, USA; [email protected] B. Sun, Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China; University of Chinese Academy of Sciences, Beijing 100039, China; College of Medicine, Department of Anatomy, Laboratory of Evolutionary Biology, Howard University, Washington D.C. 20059, USA; [email protected] A.C. Nanda, Wadia Institute of Himalayan Geology, Dehra Dun 248001, India; [email protected] R.L. Bernor, College of Medicine, Department of Anatomy, Laboratory of Evolutionary Biology, Howard University, Washington D.C. 20059, USA; [email protected] KEY WORDS - South Asia, Pliocene, Biogeography, Dispersal, Siwalik, Hipparionine horses. ABSTRACT - The Pliocene fossil record of hipparionine horses in the Indian Subcontinent is poorly known. Historically, only one species, “Hippotherium” antelopinum Falconer & Cautley, 1849, was described from the Upper Siwaliks. Here, we present the frst evidence of Eurygnathohippus Van Hoepen, 1930, a lineage hitherto only known from Africa, in the Upper Siwaliks during the late Pliocene. Morphologically, the South Asian Eurygnathohippus is most similar to Eurygnathohippus hasumense (Eisenmann, 1983) from Afar, Ethiopia, a species with a similar temporal range. -
To Changing Environments in the Plio-Pleistocene: Tooth Wear, Diets and Habitats
2019 Pigs, hogs or boars (suids) were not the subjects I thought to research during my PhD before AND OFGEOSCIENCES GEOGRAPHY DEPARTMENT I started, but I definitely fell in love with them during the journey. The same applies for Africa, though I have had the dream of working with something related to the wildlife of Africa since my childhood. In the end I was able to combine them with the second secret dream, being a palaeontologist in Finland. In this thesis I have investigated the peculiar case of the Plio-Pleistocene African suids, which show shifting from omnivorous diet to grazing in three different lineages. I have conducted experimental work on dental wear by different food items with a mechanical masticator. My work also provides insights for abundances of the Turkana Basin suids in relation to climate changes in the Plio-Pleistocene and identifying a relationship between dental topography and diet preferences in present-day suids and applying the results for the extinct suids. A75 DEPARTMENT OF GEOSCIENCES AND GEOGRAPHY A75 JANINA RANNIKKO JANINA Department of Geosciences and Geography A ISSN-L 1798-7911 ISSN 1798-7911 (print) ISBN 978-951-51-4917-6 (paperback) Adaptations of the Turkana Basin pigs ISBN 978-951-51-4918-3 (PDF) http://ethesis.helsinki.fi/I (Suidae) to changing environments in Painosalama the Plio-Pleistocene: tooth wear, diets Turku 2019 and habitats JANINA RANNIKKO Adaptations of the Turkana Basin pigs (Suidae) to changing environments in the Plio-Pleistocene: tooth wear, diets and habitats JANINA RANNIKKO ACADEMIC DISSERTATION To be presented with the permission of the Faculty of Science of the University of Helsinki, for public examination in auditorium E204 Physicum, Kumpula, on 20th May 2019, at 12 noon. -
To My Mind, Not Due So Much to the Lack of Change in the Teeth As to The
19291 Matthew, Critical Observations upon Siwalik Mammals 553 to my mind, not due so much to the lack of change in the teeth as to the rapidity of change in the skull, and the whole family derives from late Miocene paleomerycines, an antiquity decidedly less than most mam- malian families. According to Pilgrim' the Giraffidae afford " one of our most complete proofs for the Pontian age of the Dhok Pathan." But his argument will not stand analysis. "There we find side by side with G. attica is in fact distinctly smaller and Giraffa punjabiensis, paralleled by more brachydont than punjabiensis. Giraffa attica at Pikermi, the large genus What Pilgrim called 'Helladotherium' HeUadotherium allied to the Pikermi grande is in fact Lydekker's Hydaspi- species H. duvernoyi and the almost therium 7negacephalum. HeUadotherium equally large Vishnutherium. duvernoyi is much more primitive; com- pares with "Vishnutherium," but then " Vishnutherium" occurs in both the middle and lower Siwalik. "The palaeotragine group, including the Giraffokeryx is a member of it, and is genera Palzeotragus, Sanwtherium and considered by Bohlin to be a species of Alcicephalus, has not so far been Paleotragus. One species of Giraffokeryx, recognized in India. however, if not both, carries four horns, and is generically distinct from Paleo- tragus, although clearly related. "On the other hand, allied forms posses- As we should expect if the Dhok Pathan sing a complex horn development occur and Perim Island beds are later than in the genus Hyda pitherium in the Dhok Pikermi, Samos or Northern China. Pathan zone, and as Bramatherium in the slightly older Perim Island beds. -
The Phylogeny and Taxonomy of Hippopotamidae (Mammalia: Artiodactyla): a Review Based on Morphology and Cladistic Analysis
Blackwell Science, LtdOxford, UKZOJZoological Journal of the Linnean Society0024-4082The Lin- nean Society of London, 2005? 2005 143? 126 Original Article J.-R. BOISSERIEHIPPOPOTAMIDAE PHYLOGENY AND TAXONOMY Zoological Journal of the Linnean Society, 2005, 143, 1–26. With 11 figures The phylogeny and taxonomy of Hippopotamidae (Mammalia: Artiodactyla): a review based on morphology and cladistic analysis JEAN-RENAUD BOISSERIE1,2* 1Laboratoire de Géobiologie, Biochronologie et Paléontologie Humaine, UMR CNRS 6046, Université de Poitiers, 40 avenue du Recteur, Pineau 86022 Cedex, France 2Laboratory for Human Evolutionary Studies, Department of Integrative Biology, Museum of Vertebrate Zoology, University of California at Berkeley, 3101 Valley Life Science Building, Berkeley, CA 94720-3160, USA Received August 2003; accepted for publication June 2004 The phylogeny and taxonomy of the whole family Hippopotamidae is in need of reconsideration, the present confu- sion obstructing palaeoecology and palaeobiogeography studies of these Neogene mammals. The revision of the Hip- popotamidae initiated here deals with the last 8 Myr of African and Asian species. The first thorough cladistic analysis of the family is presented here. The outcome of this analysis, including 37 morphological characters coded for 15 extant and fossil taxa, as well as non-coded features of mandibular morphology, was used to reconstruct broad outlines of hippo phylogeny. Distinct lineages within the paraphyletic genus Hexaprotodon are recognized and char- acterized. In order to harmonize taxonomy and phylogeny, two new genera are created. The genus name Choeropsis is re-validated for the extant Liberian hippo. The nomen Hexaprotodon is restricted to the fossil lineage mostly known in Asia, but also including at least one African species. -
Faunal and Environmental Change in the Late Miocene Siwaliks of Northern Pakistan
Copyright ( 2002, The Paleontological Society Faunal and environmental change in the late Miocene Siwaliks of northern Pakistan John C. Barry, MicheÁle E. Morgan, Lawrence J. Flynn, David Pilbeam, Anna K. Behrensmeyer, S. Mahmood Raza, Imran A. Khan, Catherine Badgley, Jason Hicks, and Jay Kelley Abstract.ÐThe Siwalik formations of northern Pakistan consist of deposits of ancient rivers that existed throughout the early Miocene through the late Pliocene. The formations are highly fossil- iferous with a diverse array of terrestrial and freshwater vertebrates, which in combination with exceptional lateral exposure and good chronostratigraphic control allows a more detailed and tem- porally resolved study of the sediments and faunas than is typical in terrestrial deposits. Conse- quently the Siwaliks provide an opportunity to document temporal differences in species richness, turnover, and ecological structure in a terrestrial setting, and to investigate how such differences are related to changes in the ¯uvial system, vegetation, and climate. Here we focus on the interval between 10.7 and 5.7 Ma, a time of signi®cant local tectonic and global climatic change. It is also the interval with the best temporal calibration of Siwalik faunas and most comprehensive data on species occurrences. A methodological focus of this paper is on controlling sampling biases that confound biological and ecological signals. Such biases include uneven sampling through time, differential preservation of larger animals and more durable skeletal elements, errors in age-dating imposed by uncertainties in correlation and paleomagnetic timescale calibrations, and uneven tax- onomic treatment across groups. We attempt to control for them primarily by using a relative-abun- dance model to estimate limits for the ®rst and last appearances from the occurrence data. -
Microstonyx Antiquus Aus Dem Alt-Pliozän Mittel-Europas
ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Annalen des Naturhistorischen Museums in Wien Jahr/Year: 1972 Band/Volume: 76 Autor(en)/Author(s): Thenius Erich Artikel/Article: Microstonyx antiquus aus dem Alt-Pliozän Mittel-Europas. Zur Taxonomie und Evolution der Suidae (Mammalia). 539-586 ©Naturhistorisches Museum Wien, download unter www.biologiezentrum.at Ann. Naturhistor. Mus. Wien 76 539-586 Wien, April 1972 Microstonyx antiquus aus dem Alt-Pliozän Mittel-Europas. Zur Taxonomie und Evolution der Suidae (Mammalia) Von Erich THENIUS X) (Mit 10 Textabbildungen und 3 Tafeln) Manuskript eingelangt am 15. Juni 1970 Inhaltsübersicht Seite Zusammenfassung 539 Summary 540 Einleitung 540 Fundort und Altersstellung der Fundschichten 542 Beschreibung von Schädel und Gebiß 543 Taxonomische und phylogenetische Stellung von Microstonyx 555 Zur funktioneilen Analyse des Microstonyx-Sch&dels 566 Lebensraum und Lebensweise von Microstonyx 574 Die zeitliche und räumliche Verbreitung von Microstonyx antiquus 575 Zur Evolution der Suidae 576 Ergebnisse 581 Literaturverzeichnis 581 Zusammenfassung Aus altpliozänen (pannonischen) Schottern von Stratzing N Krems in der Molassezone Niederösterreichs wird ein Schädel von Microstonyx antiquus be- schrieben. Es ist der erste Schädelrest dieser Art, die damit erstmalig sicher aus dem österreichischen Tertiär nachgewiesen ist. Ein Vergleich mit fossilen und rezenten Suiden zeigt die generische Sonderstellung von Microstonyx. Sie wird als Endform der Hyotheriinen und nicht als Angehörige der Suinae ge- deutet und vom miozänen Hyotherium abgeleitet. Die Gattungs- und Art- Diagnose wird neu definiert. Microstonyx antiquus ist als Mitglied der Hipparionenfaunen vom Eppels- heim-Typ eine Waldform, M. major dagegen ein Savannenbewohner und Ange- höriger von Pikermi-Faunen.