Cotinga 13 An inventory of the of Segua Marsh, Manabí, Ecuador Bernabé López-Lanús and Paola Gastezzi 13 (2000): 59–64

Dentro del marco de las políticas de acción del Programa de Manejo de Recursos Costeros se encuentra declarar al humedal La Ciénaga de La Segua (1742 ha), Manabí, Ecuador, como sitio oficial Ramsar. Los autores realizaron un inventario, listado comentado y cuantificación de la población de aves del lugar, incluyendo en esta nota el listado de aves ( 156 especies ), comentarios de 13 especies raras o con problemas de conservación, y presentando una nueva especie para Ecuador: la Polluela Pechiamarilla Porzana flaviventer. El sitio funciona como refugio de aves acuáticas (el 44% de las especies inventariadas) siendo de importancia vital en la estación seca.

Introduction December–April. The area is surrounded by low hills A proposal prepared by the Programa de Manejo that are farmed and grazed. The marsh originally de Recursos Costeros (hereafter PMRC) for the formed part of the Chone river estuary and was environmental management of Segua Marsh states covered by mangroves but, as the watershed was its decision to undertake studies necessary for the deforested, the mouth of the Chone river silted, Ecuadorian government to declare the marsh a blocking the entry of tidal waters but permitting Ramsar site1. Given that the Ramsar Convention the river to enter the sea. The salinity of the water is the principal inter-governmental forum for the in Segua decreased and local mangroves died and promotion of international cooperation for the were logged. At present, there are large areas of conservation of wetlands, this declaration should standing fresh water with abundant floating increase the possibility of obtaining funds and vegetation (Eichornia sp. Pontedereacea) international technical assistance for the surrounded by ricefields, pastures, and brushland sustainable development of this area. For this (Figs. 1 & 2). Our report to PMRC8 includes a more reason, a series of visits were made to the area in detailed description of the area. October 1996–April 1997 to produce an annotated list of the birds that use the marsh and their Results numbers. The full results of this work were included This annotated list of the birds of Segua marsh in a report to PMRC8 that recommended declaration includes a discussion of rare and threatened species, as a Ramsar site. together with details of the first Ecuadorian record of Yellow-breasted Crake Porzana flaviventer. The Methods inventory is principally based on our records from Ornithologists first visited this marsh in 1993 and visits on 4–5 October 1996, 22 October–3 November 199513,15. Subsequently, the area has been included 1996, 21–30 November 1996, 26 December 1996–2 in two site guides promoting ecotourism9,14, and has January 1997, 27 February–7 March 1997, 12–23 been visited by a number of birdwatchers who have April 1997 and 10–18 May 1997, together with other prepared unpublished lists of species observed at supplementary data (see Introduction). In this area, the site. In the present study, six visits of 10 days the dry season is in October-December and the each were planned to cover both the dry and wet rains in February–April. A total of 156 species, of seasons, to monitor the arrival and departure of 46 families (30 non- and 17 strictly migrants, the presence and absence of resident aquatic), were recorded. A total of 44% of the species species at different times of year, and reproductive encountered were aquatic or associated with flooded behaviour, as well as to quantify species abundances . The following notes refer to the most via sampling quadrats. The species included were important records. identified via direct observation, photographs, sound recordings, published lists or unpublished Southern Pochard Netta erythrophthalma sightings by competent observers. A group of at least 14 (approximately equal numbers of males and females) was observed on 30 October Study site 1996, dispersed across a patch of open water with Segua Marsh covers 1742 ha13, at the confluence of numerous floating plants (Eichornia sp.). The next the Carrizal and Chone rivers, in Manabi province, day, 20 were seen in nearby flooded ricefields, Ecuador. The Carrizal river and the roads from frequently flushing due to human disturbance. The Bahía de Caráquez and Portoviejo meet at the only other sighting during the fieldwork was a male village of San Antonio and form the limits of this on 21 November 1996. The species has previously permanent marsh and peripheral areas that are been recorded here, on 28 August 19953, and in the temporarily flooded during the rainy season, in general area13 (M. Jácome pers. comm.). There are

59 Cotinga 13 Inventory o f the birds o f Segua Marsh, Ecuador few records in Ecuador and it is considered Very Porzana carolina rare’11 and ‘critically endangered’7 in the country. One on 27 February 1997. Considered rare11 in Ecuador. Muscovy Cairina moschata One was seen in November 1981 by F. Man Ging Yellow-breasted Crake Porzana flaviventer (pers. comm.) and another on 28 August 19953. This Seen twice, on 23 November 1996 and 27 February species is considered rare11 in Ecuador. 1997, in clumps of floating vegetation (Eichornia sp.). In both cases, the was walking on floating Rufescent Tiger- Tigrisoma lineatum vegetation, permitting brief views before it flew. One adult on 29 December 1996. There are previous Identification was based on its distinct plumage: records at Segua2,6. This species is considered rare11 crown and eye-lines black, whitish superciliary line, in western Ecuador. yellowish underparts (and feet), brown back mottled with whitish and cinnamon, white flanks with Least Ixobrychus exilis blackish bars. This is the first record in Ecuador Observed 2–5 times during each visit. Always and anywhere west of the Andes12. Included in associated with dense clumps of floating vegetation Ridgely et al.11 and Ridgely & Greenfield10 on the (Eichornia sp.). This species is considered rare11 in basis of these sightings. Ecuador. Heliornis fulica Botaurus pinnatus Heard by BLL on 27 October 1996 and 21 April 1997. Common in October 1996 (c.15), but only 2–5 during Previously known from the area2,6. The species is other visits. Always associated with dense clumps considered rare11 in western Ecuador. of floating vegetation (Eichornia sp.). Previously recorded at Segua3,6. This species is considered Calidris canutus rare11 in Ecuador. One record, a group of 32 birds on 6 March 1997 flying over cultivated rice fields. A migrant in this Glossy Plegadis falcinellus area with no further records during the study Solitary or in groups of up to four, with a flock of 35 period. on 31 December 1996. Present on all visits, probably the same birds. Relatively common in April 1997, Grey-headed Larus cirrocephalus with a flock of seven seen. Identification was based One on 28 February 1997. Among the many on soft-part coloration: bill greyish horn, legs greyish Laughing Larus atricilla present, it is horn to slate-grey, bare skin at base of bill slate- probable that others could have gone unnoticed. The grey, iris reddish. This species is ‘very rare’11 in species has not previously been recorded this far Ecuador. north on the Ecuadorian coast (M. Robbins pers. comm.). Falco columbarius Observed twice during each visit, except in Red-masked Parakeet Aratinga erythrogenys December. This species is ‘rare’11 in Ecuador. Commonly observed in November–December 1996 and March–April 1997, probably involving the same Clapper Rallus longirostris flock of c. 20 birds, in cornfields at the edge of the The only record is by Granizo6, who found three in marsh or flying over. The species is considered February 1993. The species is associated with globally Near-threatened4,5 and vulnerable in flooded, low-lying tidal areas, usually mangroves. Ecuador7. Its presence in freshwater is remarkable, although M. Cheek and BLL found six at Fincacua Fork-tailed Flycatcher Tyrannus savana shrimp ponds, c. 20 km west of Guayaquil, on 20 Two to five seen in March-April 1997. The species December 1997. Although these were indirectly is rare11 in western Ecuador. associated with mangroves, via supply canals to the shrimp ponds, it appears that the species does Carduelis siemiradzkii utilise open areas such as those at Segua. It is worth A pair on 30 December 1996. The male’s mantle remarking that, until a few years ago, Segua was olivaceous-yellow, including the upper part of consisted of mangrove (see Study site) and the the nape, and lacked dark olive spotting. The birds highest tides still reach within c. 2 km of the marsh. perched in an isolated, leafless bush c. 1 m high, There are very few records of this species, which is surrounded by floating vegetation (Eichornia sp.). considered ‘vulnerable to extinction’7 and ‘very A few minutes later, they flew to an extensive, 3-m- rare’11 in Ecuador. high scrub nearby. As this was the only record, it appears that a permanent population does not occur

60 Cotinga 13 Inventory o f the birds o f Segua Marsh, Ecuador

Figures 1–2. Segua Marsh, Manabí, from the air (left) and ground level (B. López-Lanús) in the area. The species is considered Vulnerable at References national and global levels4,5,7. This record is the first 1. Altamirano, M. & Gastezzi, P. (1996) Propuestas in Manabi province. de políticas para el manejo ambiental de La Segua, un pantano de Ecuador. Programa de Conclusions Manejo de Recursos Costeros (PMRC). Segua Marsh is an important refuge for waterbirds, Guayaquil, Ecuador. Unpublished report. especially during the dry season8, and is therefore 2. Briones, E., Flachier, A., Gómez, J., Tirira, D., of vital importance for migrant and resident species. Medina, H., Jaramillo, I. & Chiriboga, C. (1997) The presence of a number of species that are rare Inventario de Humedales del Ecuador. Primera or threatened in Ecuador increases its importance. Parte: humedales lénticos de las provincias de Yellow-breasted Crake is unknown elsewhere in Esmeraldas y Manabí. EcoCiencia/INEFAN/ Ecuador or west of the Andes. The presence of Convención de Ramsar. migrant Nearctic shorebirds further enhances the 3. Cisneros, D. & Caceref, F. (1995) Lista de aves conservation value of the area and justifies its observadas en los humedales de La Segua. 28 protection. This inventory and the report submitted August 1995. Unpublished report. to PMRC8 represent key information for the 4. Collar, N. J., Gonzaga, L. P., Krabbe, N., preparation of a management plan for the Segua Madroño-Nieto, A., Naranjo, L. G., Parker, T. Marsh by PMRC, the first step toward formal A. & Wege D. C. (1992) Threatened birds of the declaration as a Ramsar site. The report submitted Americas: the ICBP/IUCN Red Data Book. to the PMRC8 is unique within the region and should Cambridge, UK: International Council for Bird be of particular interest to Ecuadorian and South Preservation. American ornithology, as it furnishes details of the 5. Collar, N. J., Crosby, M. J. & Stattersfield, A. J. avifauna of a region of the country that is still little (1994) Birds to Watch 2: the world list of known. threatened birds. Cambridge, UK: BirdLife International (Conservation Series 4). Acknowledgements 6. Granizo, T. ( 1993) Lista y censo de aves acuáticas We are grateful for the continuous support of the observadas en La Sabana, Manabí. Febrero y main office of the PMRC (Guayaquil), especially agosto de 1993. Unpublished list. Segundo Coello, Héctor Ayón, Manfred Altamirano, 7. Granizo, T, Guerrero, M., Pacheco, C., Phillips, Ricardo Noboa, Juan Carlos Avilés and Guillermo R., Ribadeneira, M. B. & Suárez, L. (1997) Lista Baños. We also thank the team of the Bahía de de aves amenazadas de extinción en el Ecuador. Caráquez Special Management Zone (PMRC, Quito: IUCN-Sur, CECIA, INEFAN, Bahía): Ramón Zambrano, Javier Alvarado, John EcoCiencia y BirdLife International. Herbert (Peace Corps Volunteer from the USA) and 8. López-Lanús, B. & Gastezzi, P. (in prep.) Howar Vera. The communities of La Segua and La Inventario, listado comentado y cuantificación Sabana were very helpful. Niels Krabbe and Paul de la población de aves de la ciénaga de La Greenfield made a number of helpful suggestions. Segua, Manabí, Ecuador. Guayaquil: Marco Jácome (MECN), Félix Man Ging Programa de Manejo de Recursos Costeros (MFCNUG), and Francisco Sánchez (Jardín (PMRC). Botánico de Bogotá) provided useful information. 9. Green, C. (1996) Birding Ecuador. Second Chris Canaday and Paul Salaman reviewed the text edition. Privately published. and translated it into English. 10. Ridgely, R. S. & Greenfield, P. J. (in press) A guide to the birds of Ecuador. Ithaca, NY: Cornell University Press.

61 Cotinga 13 Inventory o f the birds o f Segua Marsh, Ecuador

11. Ridgely, R. S., Greenfield, P. J. & Guerrero G. Netto erythrophthalma Porrón Sureño, Southern Pochard (1998) An annotated list of the birds of Catrina moschata Pato Real, Muscovy Duck mainland Ecuador. Quito: Fundación Oxyura dominica Pato Enmascarado, Masked Duck

Ornitológica del Ecuador, CECIA. ARDEIDAE 12. Taylor, B. & van Perlo, B. (1998) Rails. A guide Ardea cocoi Garzón Cocoi, to the rails, crakes, gallinules and coots of the Ardea alba Garceta Grande, Great world. Robertsbridge: Pica Press. Egretta thula Garceta Nívea, 13. Velásquez, M., Burgos, M., Zambrano, N. & Egretta caerulea Garceta Azul, Gastezzi, P. (1993) Visión global de La Segua, Egretta tricolor Garceta Tricolor, Tricolored Heron Butorides striatus Garcilla Estriada, Striated Heron un humedal de Ecuador. Guayaquil: Programa Bubulcus ibis Garceta Bueyera, de Manejo de Recursos Costeros (PMRC). Nycticorax nycticorax Garza-Nocturna Coroninegra, Black- Unpublished report. crowned Night-Heron 14. Williams, R. S. R., Best, B. & Heijnen, T. (1996) Nyctanassa violacea Garza-Nocturna Coroniamarilla, Yellow- A guide to birdwatching in Ecuador and the crowned Night-Heron Tigrisoma lineatum Garza-Tigre Castaña, Rufescent Tiger-Heron Galápagos Islands. Otley: Biosphere Ixobrychus exilis Mirasol Menor, Publications. Botaurus pinnatus Mirasol Pinado, Pinnated Bittern 15. Zambrano, N. & Flores, P. (1994) Hábitos alimenticios y contenido estomacal de aves acuáticas en una chaînera de La Segua, Eudocimus albus Ibis Blanco, W hite Ibis Manabí. Universidad de Guayaquil, Facultad Plegadis falcinellus Ibis B ronceado, de Ciencias Naturales, Escuela de Biología. CICONIIDAE Mycteria americana Cigüeña Americana, W ood Bernabé López-Lanús Proyecto Ognorhynchus, Carrera 4. Nro. 5-48, CATHARTIDAE Salento, Quindio, Colombia. Coragyps atratus Gallinazo Negro, E-mail: [email protected] / Cathartes aura Gallinazo Cabecirrojo, [email protected]. Pandion haliaetus Águila Pescadora, Paola Gastezzi Gampsonyx swainsonii Elanio Perla, Pearl (two) Programa de Manejo de Recursos Costeros (PMRC), Rostrhamus sociabilis Elanio Caracolero, Av. Quito y Padre Solano, Ed. Buteogallus meridionalis Gavilán Sabanero, Savanna Buteogallus subtilis Gavilán Manglero, Mangrove Black Hawk MAC piso 20, P.O.Box 09-01-5850, Guayaquil Parabuteo unicinctus Gavilán Alibayo, Harris’ Hawk Ecuador. E-mail: [email protected]. Buteo magnirostris Gavilán Caminero,

FALCONIDAE Appendix: Inventory of birds of Segua Marsh, Caracara plancus Caracara Crestado, Manabí, Ecuador Herpetotheres cachinnans Halcón Reidor, Falco columbarius Esmerejón, Merlin Falco peregrinus Halcón Peregrino, follows Ridgely et a l.11. M aximum totals are given for selected species, not included in the RALLIDAE species accounts, in brackets. Laterallus albigularis Polluela Goliblanca, W hite-throated Crake Rallus longirostris Rascón M anglero, C lapper Rail Aramides sp. rascón-montés, wood-rail sp. FREGATIDAE Porzana carolina Sora, Sora Fregata magnifícens Fragata Magnífica, Magnificent Porzana flaviventer Polluela Pechiamarilla, Yellow-breasted Crake Porphyrula martinico Gallareta Púrpura, Purple Gallinule PHALACROCORACIDAE Gallínula chloropus Gallareta Común, Phalacrocorax brasilianus Cormorán Neotropical, Neotropic C o rm o ra n t Heliornis fulica Ave-Sol Americano, Sungrebe ANHINGIDAE anhinga Aninga, Anhinga ARAMIDAE Aramus guarauna Carrao, PELECANIDAE Pelecanus occidentalis Pelícano Pardo, Brown Jacana jacana Jacana Carunculada, W attled Jacana (60+) Dendrocygna bicolor Pato-Silbón Canelo, Fulvous Whistling-Duck SCOLOPACIDAE Dendrocygna autumnalis Pato-Silbón Ventrinegro, Black-bellied melanoleuca Patiamarillo Mayor, (17) Whistling-Duck Tringa flavipes Patiamarillo Menor, (26) Anas bahamensis Ánade Cariblanco, White-cheeked Pintail Tringa solitaria Playero Solitario, Solitary (two) Anas discors Cerceta Aliazul, Blue-winged Teal Actitis macularia Playero Coleador, (seven)

62 C o t i n g a 1 3 Inventory o f the birds o f Segua Marsh, Ecuador

Numenius phaeopus Zarapito Trinador, Whimbrel (13) ALCEDINIDAE Arenaria interpres Vuelvepiedras Rojizo, (two) Megaceryle torquata Martín Pescador Grande, Ringed Calidris canutus Playero Rojo, Red Knot (23) Chloroceryle americana Martín Pescador Verde, Calidas pusilla Playero Semipalmado, (65) Calidris mauri Playero Occidental, (58) PICIDAE Calidris minutilla Playero Menudo, (56) Piculus rubiginosus Carpintero Olividorado, Golden-olive Calidris melanotos Playero Pectoral, (nine) Calidris himantopus Playero Tarsilargo, (19) Dryocopus lineatus Carpintero Lineado, Lineated Woodpecker Limnodromus griseus Agujeta Piquicorto, Short-billed Veniliornis callonotus Carpinterito Dorsiescarlata, Scarlet-backed (seven) Woodpecker Steganopus tricolor Falaropo Tricolor, Wilson’s (20) FURNARIIDAE Furnarius cinnamomeus Hornero del Pacífico, Pacific Hornero Himantopus mexicanus Cigüeñuela Cuellinegra, Black-necked Stilt (17) DENDROCOLAPTIDAE Lepidocolaptes souleyetii Trepatroncos Cabecirrayado, Streak­ headed Woodcreeper Charadrius semipalmatus Chorlo Semipalmeado, Semipalmated (two) THAMNOPHILIDAE Charadrius collaris Chorlo Collarejo, (14) Sakesphorus bernardi Batará Collarejo, Collared Antshrike Charadrius vociferus Chorlo Tildio, (five) Pyriglena leuconota Ojo-de-Fuego Dorsiblanco, White-backed Fire- eye LARI DAE Larus atricilla Gaviota Reidora, RHINOCRYPTIDAE Larus cirrocephalus Gaviota Cabecigris, Grey-headed Gull Melanopareia elegans Pecholuna Elegante, Elegant Crescentchest Larus pipixcan Gaviota de Franklin, Franklin’s Gull (two) Sterna nilotica Gaviotin Piquigrueso, Gull-billed TYRANNIDAE Sterna hirundo Gaviotin Común, Camptostoma obsoletum Tiranolete-Silbador Sureño, Southern Beardless-Tyrannulet Elaenia flavogaster Elaenia Penachuda, Yellow-bellied Elaenia Columba cayennensis Paloma Ventripálida, Pale-vented Pigeon Euscarthmus meloryphus Tirano-Enano Frentileonado, Tawny- Zenaida auriculata Tórtola Orejuda, crowned Pygmy-Tyrant Zenaida meloda Tórtola Melódica, West Peruvian Dove Lophotriccus pileatus Cimerillo Crestiescamado, Scale-crested buckleyi Tortolita Ecuatoriana, Ecuadorian Ground Dove Pygmy-Tyrant Columbina cruziana Tortolita Croante, Croaking Ground Dove Todirostrum cinereum Espatulilla Común, Common Tody-Flycatcher Leptotila verreauxi Paloma Apical, White-tipped Dove Tolmomyias sulphurescens Picoancho Azufrado, Yellow-olive Flycatcher Myiophobus fasciatus Mosquerito Pechirrayado, Bran-colored Aratinga erythrogenys Perico Caretirrojo, Red-masked Parakeet Flycatcher Forpus coelestis Periquito del Pacífico, Pyrocephalus rubinus Mosquero Bermellón, Vermilion Flycatcher Muscigralla brevicauda Tiranito Colicorto, Short-tailed Field-Tyrant CUCULIDAE Fluvicola nengeta Tirano-de-Ciénega, Masked Water-tyrant Coccyzus melacoryphus Cuclillo Piquioscuro, Dark-billed Myiarchus phaeocephalus Copetón Coronitiznado, Sooty-crowned Piaya cayana Cuco Ardilla, Squirrel Cuckoo Flycatcher Crotophaga Garrapatero Piquiliso, Smooth-billed Ani Megarynchus pitangua Mosquero Picudo, Boat-billed Flycatcher Crotophaga sulcirostris Garrapatero Piquiestriado, Groove-billed Myiozetetes cayanensis Mosquero Alicastaño, Rusty-margined Ani Flycatcher Tapera naevia Cuclillo Listado, Myiozetetes similis Mosquero Social, Myiodynastes maculatus Mosquero Rayado, Streaked Flycatcher TYTONIDAE Myiodynastes bairdii Mosquero de Baird, Baird’s Flycatcher Tyto alba Lechuza Campanaria, Barn Tyrannus savana Tijereta Sabanera, Fork-tailed Flycatcher Tyrannus melancholicus Tirano Tropical, STRIGIDAE Tyrannus niveigularis Tirano Goliníveo, Snowy-throated Kingbird Glaucidium peruanum Mochuelo del Pacífico, Pacific Pygmy-Owl Pachyramphus homochrous Cabezón Unicolor, One-coloured Becard NYCTIBIIDAE Nyctibius griseus Nictibio Común, Common VIREONIDAE Cyclarhis gujanensis Vireón Cejirrufo, Rufous-browed Peppershrike CAPRIMULGIDAE olivaceus Vireo Ojirrojo, Red-eyed Vireo Chordeiles acutipennis Añapero Menor, Lesser Nighthawk Nyctidromus albicollis Pauraque, Pauraque MIMIDAE Mimus longicaudatus Sinsonte Colilargo, Long-tailed APODIDAE Streptoprocne zonaris Vencejo Cuelliblanco, White-collared HIRUNDINIDAE Panyptila cayennensis Vencejo Tijereta Menor, Lesser -tailed Progne chalybea Martin Pechigris, Gray-breasted Martin Swift Notiochelidon cyanoleuca Golondrina Azul y Blanca, Blue-and-white Swallow TROCHILIDAE Stelgidopteryx ruficollis Golondrina Alirrasposa Sureña, Southern Amazilia amazilia Amazilia Ventrirrufa Amazilia, Rough-winged Swallow

63 Cotinga 13 Inventory o f the birds o f Segua Marsh, Ecuador

Riparia riparia Martín Ribereño, Bank Swallow CARDINALIDAE Hirundo rustica Golondrina Tijereta, striatipectus Saltador Listado, Streaked Saltator Hirundo rufocollaris Golondrina Ruficollareja, Chestnut-collared Pheucticus chrysogaster Picogrueso Amarillo, Sureño Southern Swallow Yellow-Grosbeak Cyanocompsa cyanoides Picogrueso Negriazulado, Blue-black TROGLODYTIDAE Grosbeak fasciatus Soterrey Ondeado, Fasciated Thryothorus superciliaris Soterrey Cejón, Superciliated Wren EMBERIZIDAE Troglodytes musculus Soterrey-Criollo Sureño, Southern House Rhodospingus cruentus Pinzón Pechicarmesi, Crimson - Wren Volatinia jacarina Semillerito Negriazulado, Blue-black Grassquit Sporophila aurita Espiguero Variable, Variable Seedeater POLIOPTILIDAE Sporophila peruviana Espiguero Pico de Loro, -billed Polioptila plumbea Perlita Tropical, Tropical Seedeater Sporophila telasco Espiguero Gorjicastaño, Chestnut-throated PARULIDAE Seedeater Parula pitiayumi Parula Tropical, Tropical Parula Dendroica aestiva (D. petechia group) Reinita Amarilla, Yellow ICTERIDAE Warbler Molothrus bonariensis Vaquero Brilloso, Geothlypis auricularis Antifacito Lorinegro, Black-lored Yellowthroat Cacicus cela Cacique Lomiamarillo, Yellow-rumped Cacique Dives warszewiczi Negro Matorralero, Scrub Blackbird THRAUPIDAE Quiscalus mexicanus Clarinero Coligrande, Great-tailed Grackle Euphonia saturata Eufonía Coroninaranja, Orange-crowned Icterus mesomelas Oriol Coliamarillo, Yellow-tailed Oriole Euphonia Sturnella bellicosa Pastorero Peruano, Peruvian Meadowlark Euphonia laniirostris Eufonía Piquigruesa, Thick-billed Euphonia Thraupis episcopus Tangara Azuleja, Blue-grey Tanager FRINGILLIDAE Carduelis siemiradzkii Jilguero Azafranado, Saffron Siskin

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