ANNALS OF AGRARIAN SCIENCE, vol. 13, no. 1, 2015 ИЗВЕСТИЯ АГРАРНОЙ НАУКИ, Том 13, Ном. 1, 2015

AGRONOMY AND AGROECOLOGY АГРОНОМИЯ И АГРОЭКОЛОГИЯ

RENEWED CHECKLIST OF BEES (HYMENOPTERA: APOIDEA) FROM

G. J.Kirkitadze*, G. O. Japoshvili ** *Agricultural University of Georgia, Institute of Entomology 240, David Agmashenebeli Alley, , 0159, Georgia;[email protected] **Invertebrate Research Center 26, Agladze Str., Tbilisi, 0119, Georgia; [email protected], [email protected] Received: 19.12.14; accepted: 02.02.15

Bees are one of the most diverse insect group in the world. Bees have a special importance for ecosystems. Almost 75% of agricultural production depends on pollination intensivity. Many of bees are used in green houses for increasing yield. Bees are not well protected against infl uence of different factors, which can affect on their number decline. For conservation of bees the knowledge of their importance, their distributional data completeness are very important. During last 30 years taxonomic status of bees distributed in Georgia had not been revised. Complete list of Georgian bees was incorporated in the list of Caucasus bees in 1981. Since that time we made a fi rst complete list of all bees distributed in Georgia, with taxonomic changes had been done during last 33 years. Totally it made 356 species belonging to 46 different genera and 6 families. We made more then 150 changes in the names of families, genera and species recorded and published from Georgia. The most abundant family was family Apidae with 107 species. Must be noted that 41 species of Bombus have been registered in Georgia, which are composing Bombus fauna of Caucasus with additional 3 species. We were not able to fi nd validity of Bombus georgicus Vogt. Therefore we did not include this species in the list.

INTRODUCTION synonymized and changed their status. These changes was Bees are one of the most diverse insect group in the world. refl ected in Fauna Europea and Discover life webs [8,9]. More than 20 000 species are described until now [1]. Complete list of Georgian bees was incorporated in the list Bees have a special importance for ecosystems. Almost of Caucasus bees in 1981[10]. Since that time complete 75% of agricultural production depends on pollination renewed checklist of Georgian bees had not been done, intensivity[2] and it’s cost is amost 153 bilion [3,4].The only some family list was published[11-23]. most important pollinators are bees and Genus Bombus is most importantemong them , Which are known as a best OBJECTIVES AND METHODS pollinators in the world. Many of them are used in green We analized all existed literature about bee distribution in houses for increasing yield [5,6]. Must be noted also genus Georgia [10-37], all received results we compared with Andrena, which is also extremely important for fruit orchards Michener [1] and related web resources [8,9]. pollination [7]. Besides this lot of plant’s reproduction and distribution are depended on bees in the wild nature, which RESULTS AND ANALYSIS makes clear that bees are important not only for agricultural Family - Colletidae pruductivity, but for stability of biodiversity as well. Bees Genus - Colletes Latreille, 1802 are not well protected against infl uence of different factors, C. albomaculatus (Lucas, 1849) (=Colletes spectabilis) which can affect on their number decline. Increasing of Distribution: Tbilisi [19,33]. agricultural fi eld areas, pesticides and habitat degradation C. carinatus Radoszkowski, 1891 greatly affect on ecosystems and fi naly they damage bees. Distribution: [10,19,27]. For conservation of bees the knowledge of their importance, C. cunicularius (Linnaeus, 1761) their distributional data completeness are very important. Distribution: Tbilisi, Sairme, Paliastomi [19]. During last 30 years taxonomic status of bees distributed C. daviesanus Smith, 1846 in Georgia had not been revised. Many species had been Distribution: Tsodoreti, , Shroma,

20 Matsimi, Rachisubani, Khizabavra, Glola, Distribution: , [10, 23, 26,27]. [10,19,27,28,32,33,35]. H.communis Nylander, 1852 (=Prosopis communis) C. eous Morice, 1904 Distribution: Borjomi reserve, Tsalka, Tsaishi, Akhalsopeli, Distribution: Tbilisi [19]. Lagodekhi reserve, Utsera, Seva, Vashlovani National Park C. Floralis Eversmann, 1852 (=Colletes montanus, Colletes reserve [10,11,16,19,26,27,28,30- 35]. mountainous) H. confusus Nylander, 1852 (=Prosopis confusa) Distribution: Pasanauri, [19]. Distribution: Borjomi, , Liakhvi national park C. fodiens (Fourcroy, 1785) [10,19,27]. Distribution: Tbilisi, Mt Mamadaviti, Rv Katsalkhevi H. cornutus Curtis, 1831 (=Prosopis cornuta) () [19]. Distribution: Tbilisi, Lagodekhi park, Sartichala, Diklo, C. hylaeiformis Eversmann, 1852 [10, 19, 27]. Distribution: Tbilisi [10,19,27]. H. difformis (Eversmann, 1852) (=Prosopis difformis, P. C. Lebedewi Noskiewicz, 1936 subfasciata) Distribution: Tbilisi [19]. Distribution: Pasanauri, [19]. C. maidli Noskiewicz, 1936 H. dilatatus (Kirby, 1802) (=Prosopis dilatata) Distribution: Bagdati, Ajameti restriction [19]. Distribution: Georgia [27]. C. marginatus Smith, 1846 H. euryscapus Förster, 1871 (=Prosopis euryscapa, P. Distribution: Tsodoreti, Shroma, Glola, Kaspi rimosa) [10,11,19,27,31,32,35]. Distribution: Tbilisi [19]. C. mlokossewiczi Radoszkowski, 1891 H. gibbus Saunders, 1850 (=Prosopis gibba) Distribution: Tbilisi, Lagodekhi [10,19,31]. Distribution: Tbilisi, Lagodekhi park [10,19]. C. Nasutus Smith, 1853 H. hungaricus (Alfken, 1905) (=Prosopis hungarica, P. Distribution: Atskuri, Tbilisi [19]. affi nis) C. Pallescens Noskiewicz, 1936 Distribution: Borjomi [19]. Distribution: Lagodekhi reserve, Shroma, Rachisubani, H. hyalinatus Smith, 1842 (=Prosopis hyalinata, Parakheti, Kumuros khevi [10,19,27,28,31-35]. P.hyalinatus,P. armillatus) C. similis Schenck, 1853 Distribution: Borjomi, Kazbegi [10,19,27]. Distribution: Borjomi [10,19,27]. H. ibex Morawitz, 1878 (=Prosopis ibex) C. Tuberculatus Morawitz, 1894 Distribution: Gudauri, Kazbegi [19]. Distribution: Tbilisi [19]. H. incongruus Förster, 1871 (=Prosopis genalis) Genus - Hylaeus Fabricius, 1793 Distribution: Geoirgia (Western Palaearctic) [27] H. angustatus (Schenck, 1861) (=Prosopis angustata H. lineolatus (Schenck, 1861) (=Prosopis lineolata) Schenck) Distribution: Tbilisi, Borjomi, Lagodekhi, Kojori, Distribution: Tbilisi, Lagodekhi park, Kojori [19]. [19,27]. H. Annularis (Kirby, 1802) (=Prosopis annularis) H. nigritus (Fabricius, 1798) (=Prosopis nigrita, Hylaeus Distribution: Borjomi, Lagodekhi, Chkaduashi, propinquus) Letsurtsume, Datvis khevi (Dusheti) [10,19,27,30]. Distribution: Kazbegi, Roshka pass (Dusheti) [19]. H. Annulatus (Linnaeus 1758) (=Prosopis annulata, P. H. pictipes Nylander, 1852 (=Prosopis pictipes) borealis) Distribution: Borjomi, Lagodekhi, Tbilisi [19]. Distribution: Lagodekhi, Rachisubani, matsimi, H. punctulatissimus Smith, 1842 (=Prosopis Gurgeniani, , Lidzava, Kasristskali, kumuros khevi punctulatissima, P. obscurata, P. obscuratus) [10,11,19,27,28,31-34,36]. Distribution: Borjomi, Tbilisi [10,19,27]. H. bifasciatus (Jurine, 1807) (=Prosopis bifasciata) H. signatus (Panzer, 1798) (=Prosopis pratensis, P.signata, Distribution: Tbilisi [19]. P. bipunctata) H. bisinuatus Foerster, 1871 (=Prosopis bisinuata) Distribution: Borjomi, Tbilisi [10,19,27]. Distribution: Lagodekhi park, Besleti (Sokhumi), Gumista, H. sinuatus (Schenck, 1853) (=Prosopis minuta, P. sinuata) , Ritsa, [19,27,31,33,34]. Distribution: Tbilisi, Glola (Oni) [19,27]. H. brevicornis Nylander, 1852 (=Prosopis brevicornis, H. variegatus (Fabricius, 1798) (=Prosopis variegate, P. Prosopis breviceps) variegata) Distribution: Akhalsopeli, Garakha, Khizabavra, Distribution: Vashlovani National Park Park, Omalo Shroma, Lagodekhi reserve, Kutaisi, [10,19,27,32,36]. [10,19,27,30,31,33,34]. H. clypearis (Schenck, 1853) (=Halictus clypearis) Family - Andrenidae

21 Genus –Andrena Fabicius, 1775 A. dorsata (Kirby, 1802) (=Andrena propinqua,A. dorata) A. aeneiventris Morawitz, 1872 (=Andrena aeniventris) Distribution: Chkhorotsku, Digomi, Tsodoreti, Lagodekhi, Distribution: Patara , Eldari valley, Vashlovani Gurgeniani, Khimshi, , Vashlovani National Park, National Park, Kumuros khevi, Kaspi, Gardabani Seriboseli (), Kintrishi, , Tskhumaldi, [10,18,22,26,27,32, 34-36]. Sasashi [10,14,18,22,27,28,30-36]. A. albopunctata (Rossi, 1792) A. erberi Morawitz, 1871 Distribution: Kojori, Vashlovani National Park, Gardabani Distribution: Tbilisi, Vashlovani National Park [18,22,36]. (Udabno) [10,18,22,27,32]. A. falsifi ca Perkins, 1915 A. assimilis Radoszkowski, 1876 Distribution: (Bogdanovka), Chkhorotsku Distribution: Tbilisi [18,22]. [18,22]. A. bicolor Fabricius, 1775 A. fi gurata Morawitz, 1866 Distribution: Mtiskalta (Oni), Kazbegi, Tbilisi, Distribution: Vashlovani National Park, Kaspi , -Azvareti, Rtskhmeluri, Lentekhi [10,18,22,27,32,36]. [10,14,18,22,27,35]. A. fl avipes Panzer, 1799 A. bimaculata (Kirby, 1802) Distribution: Lisi, Dmanisi, Digomi, Tsodoreti, Mtskheta, Distribution: , Lagodekhi [22]. Vashlovani National Park, Kasristskali, Motsmao, Teleti, A. bisulcata Morawitz, 1877 Kutaisi, Parakheti, Shaori, Tlugi, Cheliagele, Mtiskalta, Distribution: Sartichala, Datvis khevi (Vashlovani National Skhvava, Udabno, Davit Gareji, Korugi, Pantisharas Park), Digomi [18,22]. khevi, Kaspi, Dusheti, Borjomi, Lentekhi, Khaishi A. braunsiana Friese, 1887 [10,14,15,18,22,26-29,32,35,36]. Distribution: Vashlovani National Park [18,22,36]. A. fl oricola Eversmann, 1852 A. carantonica Perez, 1902 Distribution: Magaroskari, Tetritskaro-Tbisi, Tbilisi Distribution: [18,22]. [10,18,22,27,29]. A. carinata Morawitz, 1877 A. fulvago (Christ, 1791) Distribution: Atskuri, Akhaltsikhe [24,25,27,36]. Distribution: Kizilkilisa, Gudrukhis khevi, Akhalkalaki, A. chrysopyga Schenck,1853 Mts Tetrobi, Lagodekhi, Borjomi [16,18,22,26-28]. Distribution: Chkaduashi (), Matsimi A. fuscosa Erichson, 1835 [10,18,22,27,30-32]. Distribution: Borjomi, Kojori, Vashlovani National Park A. chrysosceles (Kirby, 1802) [22,36]. Distribution: Vashlovani National Park, Matsimi A. hattorfi ana (Fabricius, 1775) [10,11,18,22,27,31,32,36]. Distribution: Borjomi, Lagodekhi [10,18,22,16]. A. cineraria (Linnaeus, 1758) A. haemorrhoa (Fabricius, 1871) (=Andrena haemorhoa, Distribution: Pantiani, Kazbegi [18,22]. A. albicans) A. cingulata (Fabricius, 1775) Distribution: Digomi, Lagodekhi, Gardabani, Korugi, Distribution: Shuapkho, Lentekhi, Tekali [10,27]. Mt Akhuni (Dusheti), Tvalivi (Dusheti), Kazbegi, Gutsa A. coitana (Kirby, 1802) (=Andrena shawella) (Kazbegi) [10,18,22,27,29,32-34]. Distribution: Gardabani, Tbilisi, Gudauri, Kazbegi A. hesperia Smith, 1853 [10,18,22,27,32]. Distribution: Vashlovani National Park National Park, A. colletiformis Morawitz, 1873 Akhaltsikhe [18,22,36]. Distribution: Vashlovani National Park, Kasristskali, A. humilis Imhoff, 1832 Kutaisi [10,18,22,27,32,36]. Distribution: Mtiskalta, Borjomi, Kizilkilisa, Akneli, Kmo A. combaella Warncke, 1966 sti[10,18,20,22,26,27,29,33-36]. Distribution: Akhalkalaki, Lagodekhi park, Khizabavra A. hystrix Schmiedeknecht, 1883 [18,22]. Distribution: Akhalkalaki, Vashlovani National Park A. combinata (Christ, 1791) [18,22,36]. Distribution: Tbilisi [18,22]. A. incisa Eversmann, 1852 A. cordialis Morawitz, 1877 Distribution: Tbilisi, Marneuli [10,18,27]. Distribution: Marneuli, Digomi, Vashlovani National Park A. labialis (Kirby, 1802) (=Andrena labiatus) [18,22]. Distribution: Kustba, Vashlovani National Park, Borjomi, A. curvungula Thomson, 1870 Tbilisi, Gardabani, Keshishi, Kojori, Akhalkalaki, Mt Distribution: Borjomi, Marneuli [18,22]. Tetrobi [10,18,22,32]. A. decipiens Schenk, 1861 A. labiata Fabricius, 1781 Distribution: Borjomi, Tbilisi [10,22,27]. Distribution: Shuapkho, Akhalkalaki, Tbilisi [18,22,29].

22 A. lateralis Morawitz, 1876 A. proxima (Kirby, 1802) Distribution: Tbilisi, Marneuli, Vashlovani National Park, Distribution: Shuapkho, Tbilisi [10,18,22,27,29]. Mamachais khevi, Aspindza [10,18,22,27,32,36]. A. pyropygia Kriechbaumer, 1873 A. leucorhina Morawitz, 1876 (=Andrena leycorhina) Distribution: Vashlovani National Park National Park Distribution: Chkaduashi [10,27]. [18,22,36]. A. lepida Schenk, 1861 A. rufi zona Imhoff, 1834 Distribution: Aspindza, Akhaltsikhe [18,22]. Distribution: Kazbegi [18,22]. A. limbata Eversmann, 1852 A.schencki Morawitz, 1866 Distribution: Borjomi, Marneuli [18,22]. Distribution: Borjomi [18,22]. A. limata Smith, 1853 A. scita Eversmann, 1852 Distribution: Zekari pass, Gardabani, Korugi [18,22]. Distribution: Vashlovani National Park, Marneuli, Tbilisi, A. magunta Warcke, 1965 Khertvisi [10,18,22,27,32-34,36]. Distribution: Akhalkalaki, Kaspi, Shiomgvime [18,22]. A. semirubra Morawitz, 1876 A. marginata Fabricius, 1777 Distribution: Akhalkalaki [10,18,22,27]. Distribution: Vashlovani National Park, Matsimchai A. stoeckhertella Pittioni, 1948 (=Andrena stöckhertella) [11,27,31,31,36]. Distribution: Vashlovani National Park National Park A. minutula (Kirby, 1802) (=Andrena parvula) [18,36]. Distribution: Tbilisi, , Lisi, Kiketi, Katsalkhevi, A. tecta Radoszkowski, 1876 Shuapkho, Kmosti, Likokis khevi, Digomi, Lagodekhi, Distribution: Atskuri [18,22]. Matsimi, Glola, Teleti [10,18,22,27,29,33-35]. A. thoracica (Fabricius, 1775) A. minutuloides Perkins, 1914 (=Andrena minutoloides) Distribution: Digomi, Korugi, Akhaltsikhe, Dusheti, Distribution: Shuapkho, Mt Shilda (Dmanisi), Goderdzi Gardabani, Mtis kalta (Oni) [10,18,22,27,32-35]. pass [10,18 22,27]. A. tibialis (Kirby, 1802) A. morio Brulle, 1832 Distribution: Digomi [18,22]. Distribution: Tbilisi, Borjomi, Kojori [18,22]. A. taraxaci Giraud, 1861 A. nigroaenea (Kirby, 1802) (=Andrena nigoaenea) Distribution: Borjomi [10,18]. Distribution: Lagodekhi, Vashlovani National Park National A. transitoria Morawitz, 1871 Park, Kumuros khevi, Mamachais khevi [18,22,36]. Distribution: Datvis khevi, Mamachais khevi, Eldari valley, A. nitida (Muller, 1776) (=Andrena pubescens) Vashlovani National Park, Akhaltsikhe, Tbilisi, Kasris Distribution: Borjomi, Digomi, Betania, Mamachais khevi, tskali [10,18,22,27,32-34,36]. Keshishi, Kaspi [16,18,22,27,32-34,36]. A. truncatilabris Morawitz, 1877 (=Andrena truncatilebris) A. nitidiuscula Schenk, 1853 Distribution: Vashlovani National Park, Telavi, Datvis Distribution: Tbilisi [18,22]. khvreli, Kumuros khevi, Marneuli [10,18,22,27,29,32- A. nobilis Morawitz, 1874 34,36]. Distribution: Vashlovani National Park [10,18,22,27,32,36]. A. tscheki Morawitz, 1872 A. orientana Warncke, 1965 (Andrena taraxaci) Distribution: Borjomi [22]. Distribution: Borjomi [22,27]. A. vaga Panzer, 1799 A. ovatula (Kirby, 1802) (=Andrena albofasciata) Distribution: [18,22]. Distribution: Chkaduashi, Taia, Akhaltsikhe, Magaroskari, A. ventralis Imhoff, 1832 Gardabani, Tbilisi, Kazbegi, Tsalka, Shiomgvime, Kaspi, Distribution: Akhaltsikhe [18,22]. Vashlovani National Park National Park, Kintrishi A. vetula Lepeletier, 1841 [10,18,22,24,29,30,32-34,36]. Distribution: Tbilisi, Vashlovani National Park, Digomi A. panurgimorpha Mavromoustakis, 1957 [10,27,28,32-34,36]. Distribution: Vashlovani National Park [18,22,36]. A. wilkella (Kirby, 1802) A. pilipes Fabricius,1781 (=Andrena carbonaria) Distribution: Shuapkho, Barisakho [10,18,22,27,29,33,34]. Distribution: Digomi, Lisis tba, David Gareji, Udabno, Genus –Panurginus Nylander, 1848 Vashlovani National Park, Kumuros khevi, Kaspi, P. alticolus Morawitz, 1876 (=Panurginus alticola) Abastumani, Marneuli, Tbilisi, Dusheti [10,18,22,27,28,32- Distribution: Kazbegi [10,27]. 34,36]. P. halictoides Giraud, 1861 A. polita Smith, 1847 Distribution: Pantisharas khevi, Vashlovani National Park, Distribution: Borjomi [18,22,27]. Kus tba, Mamachais khevi [10,27,32,33,36]. A. praecox (Scopoli, 1763) P. labiatus Eversmann, 1852 Distribution: Manglisi [18,22]. Distribution: Kus tba, Kumuros khevi, Vashlovani National

23 Park National Park [10,18,22,27,32,33,36]. H. maculatus Smith,1848 (=Halictus maculates) P. montanus Giraus, 1861 Distribution: Tsalka, Akhalsopeli, Taia, Tbilisi, Zugdidi, Distribution: Akneli (Barisakho) [10,27,29]. Tsodoreti, Lagodekhi National Par, Borjomi, Sharakhevi P. punctiventris Morawitz, 1876 (Dusheti), Putieti, Shaori, Tlugi, Khimshi (), Distribution: Tbilisi [10,27]. Bari [10,16,23,26-30,34,35]. Genus – Panurgus Panzer, 1806 H. mucoreus Eversmann,1852 P. banksianus (Kirby, 1802) Distribution: Lagodekhi [20,23,27]. Distribution:Gudauri, Kazbegi [22]. H. patellatus Morawitz,1873 P. calcaratus (Scopoli, 1763) (=Andrena calcaratus) Distribution: Tbilisi, Tetritsklebi, Gardabani, Tsodoreti, Distribution: Nikortsminda, Mtis kalta (Oni), Putieti (Oni), Matsevani, Skhvava, Gomi (Oni), Nikortsminda, Shaori, Bari (Oni) [10,18,22,27,33,35]. Putieti, Mtiskalta, Keshishi, Vashlovani National Park, Genus – Melitturga Latreille, 1809 Kumuros khevi, Rtskhmeluri forestry production M. clavicornis (Latreille, 1808) (=Andrena chavicornis) (Lentekhi) [10,14,20,23,27,28,32-36]. Distribution: Akhalkalaki, Akhaltsikhe, Khertvisi, Atskuri, H. quadricinctus (Fabricius, 1776) [10,18,22,24,25,27]. Distribution: Tbilisi, Tsodoreti, Dedoplistskaro, Kumuroskhevi, Kaspi [10,20,23,32,36]. Family - Halictidae H. resurgens Nurse, 1903 (=Halictus holtzi) Genus –Rophites Spinola, 1808 Distribution: Tbilisi [10,27]. R. caucasicus Morawitz, 1875 (=Rhophites caucasicus) H. rubicundus (Christ, 1791) Distribution: Kobi, Lagodekhi [10,18,23,27]. Distribution: Roshka, Kmosti [10,20,23,27,29,33,34]. R. quinquespinosus Spinola, 1808 (=Rhophites H. seladonius (Fabricius, 1794) (=Halictus geminatus, H. quinquespinosus) geminates) Distribution: Tsodoreti, Glola [10,18,23,27,33,34]. Distribution: Nakiani (Chkhorotsku), Zugdidi Genus -SystrophaI lliger, 1806 [10,23,27,30]. S. planidens Giraud,1861 H. senilis (Eversmann, 1852) Distribution: Borjomi, Akhaltsikhe [10,20,23]. Distribution: Digomi, Pantisharas khevi, Vashlovani Genus –Pseudapis Kirby, 1900 National Park, Kasris tskali, Kus tba, Tsodoreti, Keshishi, P. diversipes (Latreille,1806) (=Nomia diversipes) Gardabani [10,20,23,27,28,32-34,36]. Distribution:Akhali Atoni, Pitsunda, Kumuros khevi, H. simplex Blüthgen, 1923 Vashlovani National Park [10,20,23,32,33,34,36]. Distribution: Mtiskalta [35]. P. bispinosa (Brulle, 1832) (=Nomia rufi cornis) H. sexcinctus (Fabricius,1775) Distribution: Gardabani [20]. Distribution: Borjomi preserve, Uraveli, Akhaltsikhe, Genus –Nomioides Schenck, 1866 Tsodoreti, Oni, Mtiskalta, Pitsunda, Vashlovani National N. minutissimus (Rossi, 1790) Park [10,16,20,23-28,32-34]. Distribution: Tbilisi [10,20,23]. H. subauratus (Rossi,1792) Genus -Ceylalictus Strand, 1913 Distribution: Gomareti, Tsodoreti, Kasris tskali, C. variegatus (Olivier, 1789) (=Nomoiodes variegata) Rachisubani, Kutaisi, Putieti, Kelasuri, Besleti, Gudauta, Distribution: Tbilisi [10,23,27]. Lidzava (Gagra), Pitsunda [10,20,23,26-35]. Genus - Halictus Latreille, 1804 H. tetrazonianellus Strand, 1909 (=Halictus H. carinthiacus Bluthgen, 1936 (=Halictus brachyceros) tetrazonienellus) Distribution: Chkhorotsku [23,30]. Distribution: Lagodekhi [10,20,23,27]. H. compressus (Walckenaer,1802) (=Halictus eurygnathus) H. tetrazonius (Klug,1817) Distribution: Bediani, Sharakhevi, Gudrikhis khevi, Distribution: Tbilisi [10,20,23,27]. Katsalkhevi, Datvis khevi (Magaroskari), Akneli, Tsalka, H. tomentosus (Schenck, 1853) Dusheti, Mt Kudo [10,23,27,29,33,34]. Distribution: Mtskheta [10,20,23,27]. H. confusus Smith, 1853 (=Halictus perkinsi) H. tumulorum (Linnaeus,1758) Distribution: Lugela (Chkhorotsku), Dusheti, Magaroskari Distribution: Kramis khevi, Akhalsopeli, Saguramo, [10,20,23,30]. Kiketi, Katsalkhevi, Shuapkho, Roshka, Glola, Utsera, H. lucidipennis Smith,1853 (=Halictus varipes) Besleti, Gudauta, Pitsunda, Nikortsminda, Tsalka, Tbilisi, Distribution: Lisi, Vashlovani National Park, Kelasuri, Lentekhi, Tskhumali [10,14,20,23,26-30,33-35]. Gudauta, Datvis khevi, Kaspi [10,20,23,27,32-34,36]. Genus -Lasioglossum Curtis, 1833 H. lucidulus Schenck, 1859 L. albipes (Fabricius,1781) (=Halictus albipes) Distribution: Kobuleti, Kintrishi [10,20,23,27,37]. Distribution: Shipiaki, Magaroskari, Khomisdziri,

24 Katsalkhevi, Shuapkho, Akneli, Roshka, Kmosti, Matsmao, L. malachurum (Kirby,1802) (=Halictus malachurus) Tsalka, Dusheti [20,23,26-29,33-35]. Distribution: Ukangori, Tsaishi, Chkaduashi, L. calceatum (Scopoli,1763) (=Halictus calceatus, H. Akhalsopeli, Chkhorotsku, Taia, Magaroskari, Dmanisi calcatus) [10,20,23,26,27,29,30,33,34]. Distribution: Chkhorotsku, GUdrukhis khevi, Vashlovani L. marginatum (Brulle,1832) (=Halictus marginatus) National Park, Kasris tskali, Magaroskari, Mt Akhuni, Distribution: Tbilisi, Lisi, Tbisi, Matsevani, Vashlovani Katsalkhevi, Shuapkho, Akneli, Likokis khevi, Matsmao, National Park, Digomi, Kasris tskali, Kus tba, Shaori, Datvis khevi, David Gareji,Mawimi, Saguramo, Seva, Cheliagele, Nikortsminda, Tlugi, Bari, Putieti, Pitsunda, Cheliagele, , Nikortsminda, GudauTa, Pitsunda, David Gareji, Datvis khevi, Lagodekhi, Gardabani, Tsiteltskaro, Vashlovani National Park, Gardabani, Mtiskalta, Kutaisi, Ambrolauri, Glola, Lentekhi, Laskadura Ambrolauri, Kasris tskali, Akhilei (Dusheti), Bekhvintis revine, Kheledula revine [10,11,20,23,27,28,32-36]. park (Apkhazeti), Mtiskalta, Khopuri forestry production, L. morio (Fabricius,1793) (=Halictus morio) Muashi [10,11,14,20,23,27-29,30-36]. Distribution: Lugela, Shuapkho, Nakiani, Letsurtsume, L. corvinum (Morawitz, 1876) (=Halictus corvinus) Roshka, Datvis khevi, Sharakhevi, Chkhorotsku Distribution: Dusheti, Sharakhevi [23]. [10,20,23,27-30,33,35]. L.costulatus (Kriechbaumer,1873) (=Halictus costulatus) L. pauxillum (Schenck,1853) (=Halictus pauxillus) Distribution: Kizilkilisa (Dmanisi), Shuapkho, Dusheti Distribution: Tsaishi, Chkaduashi, Akhalsopeli, [10,23,27,29,30]. Chkhorotsku, Garakha, Nakiani, Letsurtsume L. discum (Smith,1853) (=Halictus morbillosus) [10,20,23,27,28,30,33,34]. Distribution: Borjomi reserve, Patara Dmanisi, Dmanisi, L. politum (Schenck,1853) (=Halictus politus) Vashlovani National Park, Skhvava, Putieti, Lekis tskali Distribution: Nakiani, Kutaisi, Gulripshi [10,11,20,23,26-28,32-36]. [10,20,23,27,30,33,34]. L. elegans (Lepeletier, 1841) (=Halictus elegans) L. pincticolle (Morawitz,1871) (=Halictus puncticollis) Distribution: Borjomi, Atskuri [20]. Distribution: Vashlovani National Park [10,20,23,27,32,36]. L. fulvicorne (Kirby,1802) (=Halictus fulvicornis) L. sexnotatum (Kyrby,1802) (=Halictus sexnotatus) Distribution: Digomi, Tsodoreti, Mt Akhuni, Katsalkhevi, Distribution: Lagodekhi Reserve [10,20,27,31]. Shuapkho, Akneli, Roshka, Kmosti, Likokiskhevi, L. spinolae (Reed, 1892) (=Halictus minutes) Matsmao, Datvis khevi [10,23,27-29,33,34]. Distribution: Borjomi [20]. L. glabriusculum (Morawitz, 1872) (=Halictus L. subfasciatum (Imhoff, 1832) (=Halictus vulpinus) glabriusculus,H. glabrinsculus) Distribution: Borjomi, Atskuri [20]. Distribution: Lagodekhi, Letsurtsume, Akhalsopeli, L. truncaticolle (Morawitz,1877) (=Halictus truncaticollis) Chkhorotsku, Nakiani, Gulripshi, Kelasuri [10,27,30,33,34]. Distribution: Akhaltsikhe, Mtskheta, Tbilisi [20,23,27]. L. griseolum (Morawitz, 1872) (=Halictus griseolus) L. villosulum (Kirby, 1802) (=Halictus villosulus) Distribution: Kutaisi [10,23,27]. Distribution: Akhalsopeli, Nakiani, Letsurtsume, L. interruptum (Panzer, 1798) (=Halictus interruptus) Magaroskari, Chkhorotsku [10,20,23,27,29,30,33,34]. Distribution: Mtiskalta [34]. L. xanthopus (Kirby, 1802) (=Halictus xanthopus) L. laevigatum (Kirby, 1802) (=Halictus laevigatus) Distribution: Balkho, Ukangori, Kasris tskali, Kus tba, Distribution: Likokis khevi (Barisakho), Dusheti, Kazbegi Vashlovani National Park [10,20,24-27,32-34,36]. [10,23,27,30]. L. zonulum (Smith,1848) (=Halictus zonulus) L. laeve (Kirby, 1802) (=Halictus laevis) Distribution: Barisakho (Datvis khevi) [10,20,23,29]. Distribution: Kazbegi [10,27]. Genus -Sphecodes Latreille, 1805 L. laticeps (Schenck ,1870) (=Halictus laticeps) S. albilabris (Fabricius,1793) (=Sphecodes fuscipennis) Distribution: Chkhorotsku, Sharakhevi, Matsmao Distribution: Vashlovani National Park [10,20,23,32,36]. [10,23,27,29,30,33,34]. S. ephippius (Linne,1767) L. leucozonius (Schrank, 1781) (=Halictus leucozonius) Distribution: Borjomi, Tbilisi [10,20,23,27]. Distribution: Tsaishi, Chkaduashi, Tsodoreti, Tbilisi, Kus S. gibbus (Linnaeus,1758) tba, , Nikortsminda, Gulripshi, Pitsunda, Vashlovani Distribution: Borjomi [10,20,23]. National Park, Datvis khevi, Kumuros khevi, Gardabani, S. reticulatus Thomson, 1870 (=Sphecodes reticulates) Tetriskaro, Tbisi [10,15,20,23,27,28,30,32-36]. Distribution: Borjomi, Sokhumi [10,20,27]. L. longirostre (Morawitz,1876) (=Halictus longirostris) Distribution: Vashlovani National Park [10,20,23,32,36]. Family - Melittidae L. majus (Nylander,1852) (=Halictus major) Genus – Dasypoda Latreille, 1802 Distribution: Borjimi, Adigeni [24,25,27]. D. argentata Panzer,1809

25 Distribution: Shatili [10,19,27]. Distribution: Borjomi reserve, Lisi, Algeti, Matsevani, D. hirtipes (Fabricius,1793) (=Dasypoda plumipes) Vashlovani National Park, Kumuros khevi, Lentekhi, Distribution: Tbilisi, Kintrishi National Park [19,37]. Laskadula revine [10,14,16,27,28,32,36]. Genus - Macropis Panzer, 1809 O. cephalotes Morawitz,1870 (=Osmia longiceps) M. fulvipes (Fabricius, 1804) Distribution: Borjomi [10,27]. Distribution: Orule (Zugdidi) [10,19,27]. O.cerinthidis Morawitz,1876 (=Osmia cerinthides) M. labiata (Fabricius, 1804) Distribution: Tskhumaldi, Mulakhi, Ushguli [14]. Distribution: Lagodekhi [10,19,30]. O. cornuta (Latreille,1805) Genus -Melitta Kirby, 1802 Distribution: Tbilisi [10]. M. haemorrhoidalis (Fabricius,1775) O. leaiana (Kirby,1802) Distribution: Borjomi, Abastumani [10,19,27]. Distribution: Borjomi, Lagodekhi [10]. M. leporina (Panzer,1799) O. rufa (Linnaeus,1758) (=Ceratosmia rufa) Distribution: Borjomi [10,19,27]. Distribution: Dmanisi, Tbilisi, Lisi, Digomi [10,26,34,28]. O. rufohirta (Latreille, 1811) (=Acanthosmia rufohirta) Family - Megachilidae Distribution: Lisi, Pantishara revine, Korugi [10,27,36]. Genus – Lithurgus Latreille, 1825 Genus -Protosmia Ducke, 1900 L. chrysurus Fonscolombe, 1834 P. tifl ensis (Morawitz,1876) Distribution: Tsodoreti, Skhvava, Kaspi [10,27,33,35]. Distribution: Tbilisi [10]. L. cornutus (Fabricius,1787) (=Lithurgus fuscipennis) Genus - Anthidiellum Cockerell, 1904 Distribution: Pasanauri, Kaspi [10,27,32]. A. strigatum (Panzer,1805) (=Paranthidiellum strigatum) Genus – Chelostoma Latreille, 1809 Distribution: Tsodoreti, Gardabani [10,27,32]. C. emarginatum (Nylander,1856) Genus -Anthidium Fabricius, 1804 Distribution: Lagodekhi [10]. A. cingulatum Latreille,1809 (=Anthidium cungulatum) C. fl orisomne (Linnaeus,1758) (=Chelostoma maxillosum) Distribution: Tsodoreti, Vashlovani National Park Distribution: Borjomi reserve, Ukangori, Matsevani, Tbisi [10,27,32,33,36]. [10,16,26,33,34]. A. diadema Latreille,1809 C. foveolatum (Morawitz,1868) Distribution: Kumuros khevi [10,27,32,36]. Distribution: Utsera, Shaori, Kasris tskali [10]. A. fl orentinum (Fabricius, 1775) C. rapunculi (Lepeletier, 1841) (=Chelostoma fuliginosum) Distribution: , Teleti, Kintrishi [10,27,37]. Distribution: Kasris tskali, Utsera, Shaori A. manicatum (Linnaeus,1758) [10,27,32,33,34,36]. Distribution: Borjomi reserve, Tmogvi, Tsodoreti Genus -Heriades Spinola, 1808 [10,16,24,25,27,33]. H. crenulatus Nylander,1856 A. oblongatum (Illiger, 1806) (=Paroanthidium oblongatum) Distribution: Tsaishi [10,27,28,30,33,34]. Distribution: Borjomi [27]. H. truncorum (Linnaeus,1758) Genus -Pseudoanthidium Friese, 1898 Distribution: Borjomi reserve, Bediani, Tsaishi, Chkaduashi, P. melanurum (Klug, 1832) (=Anthidium nigricolle) Akhalsopeli, Lagodekhi preserve, Utsera, Shaori, Besleti, Distribution: Kaspi [10,27,32]. Gudauta, Vashlovani National Park [10,16,26-28,30-36]. Genus –Rhodanthidium Isensee, 1927 Genus -Hoplitis Klug, 1807 R. christophi (Morawitz, 1884) (=Meganthidium H. bicallosa (Morawitz, 1876) (=Heriades bicallosa, christophi) Osmia (acanthosmia) bicallosa) Distribution: Gardabani [10]. Distribution: Borjomi, Georgia [10,27]. R. ducale (Morawitz,1876) (=Anthidium ducale) H. leucomelana (Kirby 1802) (=Acanthosmia Distribution: Borjomi [10,27]. leucomelaena) Genus –Stelis Panzer, 1806 Distribution: Borjomi, Mtskheta, Tbilisi [10,27]. S.breviuscula Nylander,1848 Genus -Osmia Panzer, 1806 Distribution: Lagodekhi [10,27]. O. andrenoides (Spinola, 1808) (=Osmia (Pseudosmia) S. phaeoptera (Kirby,1802) anthrenoides) Distribution: Borjomi, Mtskheta [10,27]. Distribution: Borjomi reserve, Kintrishi, Kumuros khevi S. punctulatissima (Kirby,1802) (=Stelis aterrima) [16,27,36,37]. Distribution: Mtskheta[10,27]. O. bicolor (Schrank,1871) (=Amblys bicolor) Genus -Trachusa Panzer, 1804 Distribution: Borjomi [10]. T. pubescens (Morawitz, 1872) (=Archianthidium O. caerulescens (Linnaeus, 1758) (=Osmia coerulescens) pubenscens, Anthidium pubescens)

26 Distribution: Vashlovani National Park [10,27,32,36]. M. willughbiella (Kirby, 1802) T. byssina (Panzer, 1798) (=Trachusa serratulae) Distribution: Borjomi [10,27]. Distribution: Shovi [27,35]. M. parietina (Geoffroy, 1785) (=Chalicodoma parietinum) Genus - Coelioxys Latreille, 1809 Distribution: Uraveli, Abastumani, Ardoti, Mutso [10]. C. acanthura (Illiger, 1806) Distribution: [10,27]. Family – Apidae C. afra Lepeletier, 1841 Genus -Xylocopa Latreille, 1802 Distribution: Tbilisi [10,27]. X. iris (Christ, 1791) C. aurolimbata Forster, 1853 Distribution: Lagodekhi [10]. Distribution: Borjomi [10,27]. X. valga Gerstacker, 1872 C. conoidea (Illiger, 1806) Distribution: Borjomi reserve, Minadze, Igoeti, Aspindza, Distribution: Borjomi [10,27]. Akhaltsikhe, Abastumani, Uraveli, Patara Dmanisi, C. inermis (Kirby, 1802) Ukangori, Orozmani, Bediani, Tbilisi, Sharakhevi, Distribution: Borjomi [10,27]. Magaroskari, Shuapkho, Chrdilis khevi, Tsodoreti, Kasris C. mandibularis Nyander, 1848 tskali, Saguramo, Vashlovani National Park reserve, Distribution: Vashlovani National Park [10,27,32,36]. Mtiskalta, David GAreji, , Ninuas kutkhe, Paki, C. rufescens Lepeletier and Adinet-Serville, 1825 [10,11,14-16,24-29,31-34,36]. Distribution: Borjomi [10,27]. X. violacea (Linnaeus, 1758) Genus - Megachile Latreille, 1802 Distribution: Igoeti, Ukangori, Kutaisi, Lagodekhi [10,24- M. apicalis Spinola, 1808 27,32-34]. Distribution: Georgia [10]. X. olivieri (Lepeletier, 1841) (=Proxylocopa olivieri) M. bombycina Radoszkowski, 1874 (=Megachile Distribution: Tbilisi [10,27]. bombycinae) Genus -Ceratina Latreille, 1802 Distribution: Nakiani, Mukhuri [10,27,30]. C.acuta Friese, 1896 M. centuncularis (Linnaeus, 1758) Distribution: Tbilisi [10,27]. Distribution: Borjomi, Tbilisi, Lagodekhi, Tsodoreti C. callosa (Fabricius, 1794) [10,27]. Distribution: Atskuri, Akhaltsikhe [10,24-27,33,34]. M. circumcincta (Kirby, 1802) (=Megaxhile circumcinta) C. chalcites Germar, 1839 Distribution: Borjomi reserve, Uraveli, Adigeni, Goderdzi Distribution: Tbilisi [10,27]. pass, Vashlovani National Park, DAtvis khevi, Mamachais 255.C. cyanea (Kirby, 1802) khevi, Kintrishi [10,16,24,25,27,32-34,36,37]. Distribution: Mukhuri [10,27,30,33,34]. M. communis Morawitz, 1875 C. dallatorreana Friese, 1896 Distribution: Tbilisi, Lagodekhi [10]. Distribution: Sokhumi [10]. M. dorsalis Perez, 1879 C. dentiventris Gerstacker, 1869 Distribution: Vashlovani National Park [10,27,32,36]. Distribution: Tbilisi [10,27]. M. fl avipes Spinola, 1838 (=Archimegachile fl avipes) C. gravidula Gerstacker, 1869 Distribution: Tbilisi [27,33,34]. Distribution: Tbilisi [10,27]. M. leacella Curtis, 1828 (=Megachile argentata) C. loewi Gerstacker, 1869 (=Ceratina löwi) Distribution: Georgia [27,34]. Distribution: Lagodekhi [10,27]. M.lefebvrei (Lepeletier, 1841) Genus - Nomada Scopoli, 1770 Distribution: Georgia [27]. N. alboguttata Herrich-Schaffer, 1839 M. leucomalla Gerstacker, 1869 (=Megachile excellens) Distribution: Gardabani [10,27,32]. Distribution: Bakuriani [10]. N. armata Herrich-Schaffer, 1839 M. octosignata Nylander, 1852 Distribution: Borjomi [10,27]. Distribution: Borjomi [10,27]. N. distinguenda Morawitz, 1874 M. pilicrus Morawitz, 1877 Distribution: Borjomi [10,27]. Distribution: Georgia [27]. N. ferruginata (Linne, 1767) M. pilidens Alfken, 1924 Distribution: Tbilisi, Korugi [10,27,32]. Distribution: Tsodoreti, Gardabani [20,27,32]. N. fl avoguttata (Kirby, 1802) (=Nomada minuta) M. rotundata (Fabricius, 1787) Distribution: Borjomi reserve, Gardabani [10,16,27,32]. Distribution: Tsodoreti, Lagodekhi [10,11,27]. N. fl avopicta (Kirby, 1802) M. saussurei Radoszkowski, 1874 Distribution: Borjomi [10,27]. Distribution: Tbilisi [10,27]. N. fucata Panzer, 1798

27 Distribution: Digomi, Korugi [10,27,32,33]. Distribution: Tbilisi [10,27]. N. fulvicornis Fabricius, 1793 (=Nomada lineola) Genus -Amegilla Friese, 1897 Distribution: Borjomi, Tbilisi, Vashlovani National Park A. albigena (Lepeletier, 1841) [10,27,28,32,36]. Distribution: Tbilisi surroundings [10]. N. lathburiana (Kirbby, 1802) A. garrula (Rossi, 1790) Distribution: Gardabani [10,27,32]. Distribution: Gagra, Tbilisi, Lagodekhi [10,27]. N. mutabilis Morawitz, 1870 A. magnilabris (Fedtschenko, 1875) Distribution: Akhalkalaki [10,27]. Distribution: Tbilisi [10,27]. Genus - Epeolus Latreille, 1802 A. quadrifasciata (de Villers, 1789) E. transitorius Eversmann, 1852 Distribution: Vashlovani National Park [10,27,32,36]. Distribution: Tbilisi, Lagodekhi [10,27]. Genus -Anthophora Latreille, 1803 E. variegatus (Linnaeus, 1758) A. aestivalis (Panzer, 1801) Distribution: Lagodekhi [10]. Distribution: Uraveli [24,25]. Genus - Cubitalia Friese, 1911 A. affi nis Brulle, 1832 (=Anthophora liturata) C. tristis (Morawitz, 1876) (=Eucera tristis) Distribution: Mamachais khevi, Vashlovani National Park, Distribution: Borjomi [10,27]. Mt Zuldashi, Keshishi [10,27,32-34,36]. Genus -Eucera Scopoli, 1770 A. astragali (Morrawitz, 1877) E.cineraria Eversmann, 1852(=Eucera cinerea) Distribution: Pantishara khevi, Vashlovani National Park Distribution: Vashlovani National Park, Kumuros khevi, [10,27,28,32,36]. Gardabani, Datvis khevi [10,27,32-34,36]. A. biciliata Lepeletier, 1841 (=Anthophora mucida, E. clypeata Erichson, 1835 (=Eucera similis,E. clypeara) Anthophora caucasica) Distribution: Borjomi, Tbilisi, Vashlovani National Park Distribution: Vashlovani National Park, David Gareji, [10,27,32,36]. Borjomi [10,27,32-34,36]. E. interrupta Bar, 1850 A. crinipes Smith, 1854 Distribution: Vashlovani National Park, David Gareji Distribution: Vashlovani National Park, Keshishi, Khertvisi [10,27,32,36]. [10,27,32-34,36]. E. longicornis (Linnaeus, 1758) A. dufourii Lepeleteir, 1841 Distribution: Betania, Tskhumaldi [10,14,27,33,34]. Distribution: Vashlovani National Park, Kumuros khevi E. nigrifacies Lepeletier, 1841 [10,27,32,36]. Distribution: Kasris tskali, Kumuros khevi [10,27,32,36]. A. erschowi Fedtschenko, 1875 (=Anthophora ershawi) E. sogdiana Morawitz, 1875 (=Eucera melanostoma) Distribution: Vashlovani National Park [10,27,32,36]. Distribution: Udabno, Mtskheta, Tskhumaldi [10,14,27,32]. A. fulvitarsis Brulle, 1832 E. spectabilis (Morawitz, 1875) (=Tetralonia spectabilis) Distribution: Vashlovani National Park, David GAreji, Distribution: Borjomi [10,16,27]. Datvis khevi, Udabno [10,27,32-34,36]. E. vicina (Morawitz, 1876) (=Tetraloniavicina) A. furcata (Panzer, 1798) (= Clisodon caucasicus, C. Distribution: Borjomi [10]. furcatus) Genus -Tetralonia Spinola, 1838 Distribution: Bediani [10,26,27,33]. T. malvae (Rossi, 1790) A. nigriceps Morawitz, 1886 Distribution: Bakuriani [10,27]. Distribution: Vashlovani National Park, Udabno T. rufi collis (Friese, 1911) [10,27,28,32,36]. Distribution: Kasris tskali, Datvis khevi, Vashlovani A. plagiata (llliger,1806) (=Anthophora parietina) National Park, Lekis tskali, Mamachais khevi, Kumuros Distribution: Aspindza, Lagodekhi [10,27]. khevi [10,27,32-34,36]. A. plumipes (Pallas, 1772) (=Anthophora acervorum) T. salicariae (Lepeltier, 1841) Distribution: , Akhaltsikhe, Tbilisi, Saguramo, Kasris Distribution: Shaori, Gulripshi, Akhalsopeli, Akhali aponi tskali, Betania, Kutaisi, Telavi, Vashlovani National Park, [10,33-35]. Gardabani, Mtskheta, Datvis khevi, Shio mgvime, Keshishi T. tricincta (Erichson, 1835) [10,24,25,27,28,32,34,36]. Distribution: Tbilisi [10,27]. A. radoszkowskyi Fedtschenko, 1875 (=Anthophora Genus -Tetraloniella Ashmead, 1899 radoszkouski, Anthophora radoskowskii) T. dentata (Germar, 1839) (=Tetralinia dentata) Distribution: Vashlovani National Park, Udabno, Distribution: Lekis tskali, Vashlovani National Park Pantisharas khevi [27,32-34,36]. [10,27,32,36]. A. retusa (Linnaeus, 1758) (=Anthophora monacha) T. rufi cornis (Fabricius, 1804) (Tetralonia rufi cornis) Distribution: Sokhumi, Vashlovani National Park

28 [10,27,28,32-34,36]. Shevardeni, Ketrisi, Abano, Zemo Okrokana, Ressi, A. robusta (Klug, 1845) [10,13,27,28,31,33,34]. Distribution: Borjomi [10,27]. B. humilis Illiger, 1806 (=Bombus tristis, B. helferanus) A. quadrimaculata (Panzer, 1798) (=Anthophora vulpina) Distribution: Borjomi reserve, Uraveli, Aspindza, Adigeni, Distribution: Kazbegi [10]. MInadze, Balkho, Tba Pantiani, Beshtasheni, Dashbashi, A. gracilipes (Morawitz, 1873) (=Paramegilla gracilipes) Bediani, Shipiaki, Tsalka, Kaburi, Chivtkilisa, Abranlo, Distribution: Keshishi (Gardabani) [10,27,32]. Khando, Tikilisa, , Tsodoreti, Shuapkho, Kobulo, A. bimaculata (Panzer, 1798) (=Heliophila bimaculata) Parakheti, Shovi, Shaori, Mtiskalta, Nikortsminda, Mt Distribution: Tbilisi [10,18,20,27,28]. Ushba foot, Shirakha Tourist base surroundings, Ushguli, Genus -Habropoda Smith, 1854 Jibiani, Majdieri [10,13-17,24-29,32-34]. H. tarsata (Spinola, 1838) B. hortorum (Linnaeus, 1761) Distribution: Lagodekhi [10,27]. Distribution: Zekari pass, Goderdzi pass, Merenia, Genus - Melecta Latreille, 1802 Olodauri, Tskavashi, Skuri, Khomisdziri, Jojo, Tsablana, M. albiforns (Forster, 1771) (=Melecta armata) Danisparauli, Gogadzeebi, Zeribiseli, Tsodoreti, Kumurdo, Distribution: Mandlisi, Borjomi [10,27]. Kobulo, Matsmao, Beshumi, Parakheti, Bari, Seva, M. luctuosa (Scopoli, 1770) Shkmeri, Mtiskalta, Lagodekhi reserve, Kintrishi reserve, Distribution: Tbilisi [10,27]. Tskhumaldi, Shirakha tourist surroundings, Adishi, Genus -Thyreus Panzer, 1806 Mulakhi, Mujali, Majdieri [10,11,13,14,16,17,24,25,27- T. histrionicus (Illiger, 1806) (=Thyreus histrionucus) 31,33-36]. Distribution: Tbilisi, Eldari valley [10,27]. B. hypnorum (Linnaeus, 1758) T. ramosus (Lepeletier, 1841) Distribution: Lagodekhi reserve, Akhaltsikhe, Ninotsminda Distribution: Lagodekhi [10,27]. (Bogdanovka) [10,13,17,27]. T. truncatus (Perez,1883) B. incertus Morawitz, 1881 Distribution: Tbilisi [10,27]. Distribution: Ninotsminda (Bogdanovka), , Apindza, Genus - Bombus Latreille, 1802 Akhaltsikhe [10,24,25,27]. B. argillaceus (Scopoli, 1763) B. jonellus (Kirby, 1802) Distribution: Uraveli, Akhaltsikhe, Igoeti, Atskuri, Distribution: Oni [10,13,17,27,35]. Abastumani, Akhalkalaki, Ninotsminda (Bogdanovka), B. keriensis Morawitz, 1886 (=Bombus alagesianus) Vashlovani National Park, Tsodoreti, Betania, Aspindza, Distribution: Java, Kepi, Ressi, Abano, Shevardeni, Gimara, Mtskheta, Borjomi [10,13,17,24,25,27,28,32-34,36]. Ketrisi, Zemo Okrokana, Mtiskalta, Bari, Bakuriani, Oni, B. armeniacus Radoszkowski, 1877 Mulakhi, Majdieri, Mujali [10,13,14,27,28,33-35]. Distribution: Zekari pass [10,13,17,24,25,27]. B. lapidarius (Linnaeus, 1758) (=Bombus eriophorus) B. barbutellus (Kirby, 1802) (=Psithyrus barbutellus) Distribution: Zekari pass, Goderdzi pass, Atskuri, Tsablana, Distribution: Borjomi reserve, Shipiaki, Katsalkhevi Gogadzeebi, Nagvarevi, Chimaga, Java, Skhvava, [10,16,26 27,29]. Cheliagele, Parakheti, Bari, Tsodoreti, Kaspi, Garikula, B. brodmannicus Vogt, 1909 (=Bombus brodmanicus) Oshora, Kintrishi, Oni, Ambrolauri, Tbilisi surroundings, Distribution: Tikilisa, Zemo Okrokana, Ressi, Shevardeni, Borjomi, Akhaltsikhe, , Khulo, Mtiskalta, Gimara, Shovi, Oni, Lagodekhi, reserve, Tsalka Shkmeri, Ushguli, Jibiani [10,13-15,17,21,24,25,27-29,33- [10,13,17,26,27,33-35]. 35,37]. B. cullumanus (Kirby, 1802) (=Bombus apollineus) B. laesus Morawitz, 1875 Distribution: Uraveli, Akhalkalaki, Arali, Balkho, Igoeti, Distribution: Tba Pantiani, Beshtasheni, Shipiaki, Tba Sulda, Merenia, Ninotsminda (Bogdanovka), Avranlo, Santa, Tikilisa, Lagodekhi reserve, Matsimi, Dmanisi, Tikilisa [10,13,17,24,25,27,28,30,33,34]. Tsalka [10,11,13,17,26-28,31,33,34] B. fragrans Pallas, 1771 B. lucorum (Linnaeus, 1761) Distribution: Tbilisi [10,13,17,27]. Distribution: Borjomi reserve, Zekari pass, Balkho, B. haematurus Kriechbaumer, 1870 Merenia, Uraveli, Oladauri, Kariaki, Chivtkilisa, Tba Distribution: Uraveli, Tsablana, Digomi, Skuri, Santa, Khulo, Gorderdzi pass, Tsablana, Magaroskari, Tsvermagala, Gudrukhis khevi, Chargali, Magaroskari, Mt Khomisdziri, Mt Kudo, Gudani, Zeriboseli, Akneli, Roshka, Kudo, Nagvarevi, Betania, Lagodekhi reserve, Skhvava, Kobuli, Beshumi, Matsmao, Chrdilis khevi, Shatili, Mutso, Shovi, Kintrishi, Oni, Ambrolauri, Akhaltsikhe, Shuakhevi Arkhoti, Giorgitsminda, Ardoti, Akhieli, Chimga, Amga, [10,11,13,15,17,24,25,27-31,33-35]. Lagodekhi reserve, Mtiskalta, Shkmeri, Parakheti, Abano, B. handlirschianus Vogt, 1909 Bari, Seva, Shovi, Ressi, Ketrisi, Kintrishi, Ateni, Ailama, Distribution: Kepi, Lagodekhi reserve, Gimara, Lentekhi second forestry productuction, Mujali, Mulakhi,

29 Majdieri [10,11,13-15,17,24-29,31,33,34,36,37]. Distribution: Skuri, Tikilisa [26,30,11]. B. mesomelas Gerstacker,1869 (=Bombus alboluteus) B. sichelii Radszkowski, 1859 (=Bombus cazurroi) Distribution: Zekari pass, Goderdzi pass, Akhalkalaki, Distribution: Zemo Okrokana, Kvemo Okrokana, Bari, Balkho, Tba Pantiani, Tsalka, Shipiaki, Kaburi, Khando, Mtiskalta, Gimara, Kepi, Abano, Ressi, Shevardeni Avarlo, Tikilisa, Likokis khevi Oshora, Danisparauli, [27,28,33-35]. Kintrishi, Dmanisi, Ushguli, Jibiani, Mt Ushba foot, B. soroeensis (Fabricius, 1776) (=Bombus soröensis) Shirakha tourist Base, Adishi, [10,13-15,24-29,33,34,37]. Distribution: BOrjomi reserve, Uraveli, Tba Lelovani, B. mlokosievitzii Radoszkowski, 1877 (Bombus Igoeti, Abastumani, Zekari pass, Goderdzi pass, Adigeni, mlokosiewitzi) Balkho, Avranlo, Shatili, Tsablana, Khulo, Mutso, Distribution: Borjomi reserve, Zekari pass, Dashbashi, Ardoti, Arkhoti, Akhieli, Saguramo, Seva, Tsodoreti, Khando, Khando, Tsalka, KAriaki, Kaburi, Avarlo, Lagodekhi reserve, Betania, Bari, Mtiskalta, Parakheti, Tsablana, Gogadzeebi, Chargali, Khomisdziri, Gudani, Shovi, Skhvava, Kintrishi, Mtskheta, Garikula, Borjomi, Kobulo, Roshka, Datvis khevi, Matsmao, Chrilis khevi, Ertatsminda, Kaspi, Lentekhi second forestry production, Mtiskalta, Bari, Lagodekhi reserve, Kazbegi, Seva, Glola, Ailama, Mt Ushba foot, Shirakha Tourist base, Mt Zuruldi Kintrishi [10,11,13,15-17,24,26-29,31,33-35,37]. surroundings, Ushguli, Adishi, Mujali, Majdieri, Mulakhi B. muscorum (Linnaeus, 1758) [10,11-16,24-29,31-35,37]. Distribution: Gardabani [10,13,17,21,27]. B. subterraneus (Linnaeus, 1758) B. niveatus Kriechbaumer, 1870 (=Bombus vorticosus) Distribution: Tba Pantiani, Tsalka, Trialeti, Shipiaki, Distribution: Bakuriani [10,13,17,27]. Khando, Gogadzeebi, Mt Kudo, Kmosti, Chimga, B. pascuorum (Scopoli, 1763) (=Bombus rehbinderi, B. Mtiskalta, Shkmeri, Abano, Shovi, Lagodekhi reserve, agrorum) Kvemo Okrokana, Barisakho, Magaroskari, Shuakhevi, Distribution: Borjomi reserve, Atskuri, Abastumani, Zekari Gori, Borjomi, Akhaltsikhe, Oni, Kazbegi [10,11,15,17,24- pass, Balkho, Uraveli, Goerdzi pass, Olodauri, Skuri, 29,31,33-35,37]. , Nakiani, Danisparauli, Tsablana, Gogadzeebi, B. sylvarum (Linnaeus, 1761) (=Bombus daghestanicus, B. Jojo, Tsoniarisi, Kintrishi, Zeriboseli, Tsvermagala, Digomi, silvarum) Tsodoreti, Betania, Kobulo, Chrdiliskhevi, Matsimchai, Distribution: Borjomi reserve, Manglisi, Uraveli, Lagodekhi, Parakheti, Bari, Skhvava, Mtiskalta, Shovi, Akhaltsikhe, Arali, Igoeti, Aspindza, Atskuri, Zekari pass, Khaishi, Java, Oshora, Bavari, Lentekhi second forestry Adigeni, Akhalkalaki, Balkho, Khando, Merenia, Minadze, production, Mt Ushba foot, Shirakha tourist base, Mt Abastumani, Goderdzi pass, Kizilkilisa, Orozmani, Zurulda, Mulakhi [10,11,13-17,24-31,33-35,37]. Boslebi, Tba Pantiani, Patara Dmanisi, Tsalka, Beshtasheni, B. persicus Radoszkowski, 1881 Chivtkilisa, Kariaki, Kaburi, Avranlo, Khrami, Tikilisa, Distribution: Kaburi, Khando, Mutso, Shaori, Kala, Mazeri, Gomareti, Tskavashi, Danisparuli, Tsodoreti, Amga, Ushba surroundings [10,13,14,26,27,29,33-35]. Akhieli, Mt Kudo, Tsablana, Abano, Parakheti, Skhvava, B. pomorum (Panzer, 1805) (= Bombus albopauperatus) Gimara, Shevardeni, Nikortsminda, Mtiskalta, Java, Distribution: Mt Kudo, Giorgitsminda, Roshka, Gimara, Ressi, Shaori, Seva, Betania, Matsevani, Ketrissi, Kaspi, Ressi, Kepi, Abano, Mtiskalta, Seva, Shkmeri, Ketrisi, Ertatsminda, Kintrishi, Oni, Ambrolauri, Mtskheta, Tbilisi, Mukhrani [10]. Adishi, Mujali, Mulakhi, Majdieri, Choloshi, Ushguli B. portchinsky Radoszkowski, 1883 (=Bombus [10,13-17,24-30,33-35,37]. portschinskii) B. sylvestris (Lepeletier, 1832) (=Psithyrus silvestris) Distribution: Akhalkalaki, Shipiaki, Tba Santa, Khando, Distribution: Tskavashi [10,17,27,30,33,34]. Mt Kudo, Roshka, Kmosti, Mt Sachali, Akhieli, Mtiskalta, B. terrestris (Linnaeus, 1758) Kepi, Abano, Lagodekhi reserve, Gimara, Shevardeni Distribution: Dmanisi, Ukangori, Bediani, Tsodoreti, [10,13,24-29,33-35]. Skhvava, Bari, Mtiskalta, Parakheti, Seva, Shkmeri, B. pratorum (Linnaeus, 1761) Shovi, Kazbegi, Kintrishi, Ambrolauri, Oni, Tsalka, Tbilisi Distribution: Borjomi reserve, Nagvarevi, Beshumi, surroundings [10,13,17,26,27,28,33-35,37]. LAgodekhi reserve, Kvemo Okrokana, Kintrishi, Skuri, B. velox (Shkorikov, 1914) Khulo, Tsalka, Bakuriani [10,11,13,16,28,31,33,34,37]. Distribution: Ninotsminda (Bogdanovka), Kaburi, Ketrisi, B. quadricolor (Lepeletier, 1832) (=Psithyrus quadricolor) Ressi, Shevardeni, Gimara, Kazbegi, Tsalka, Tbilisi Distribution: Georgia [10,27]. surroundings, lake surroundings [10,13,17,24- B. ruderarius (Muller, 1776) (=Bombus simulatilis) 27,33,34]. Distribution: Zekari pass, Java, Borjomi B. wurfl enii Radoszkowski, 1859 (=Bombus alpigenus) [10,13,17,24,25,27,33,34]. Distribution: Balkho, Merenia, Akhalkalaki, Mt Kudo, B. skorikovi (Popov, 1927) Akneli, Roshka, Kobuli, Barisakho, Giorgitsminda, Mutso,

30 Amga, Lagodekhi reserve, Zemi Okrokana, Shkmeri, Landscapes for World Crops // Proceedings of the Royal Parakheti, Bari, Abano, Gimara, Kepi, Mtiskalta, Paravani Society B , 274, 2007, pp. 303-313. lake surroundings, Majdieri, Mt Ushba foot, Turbaza 3. Gallai N. Jean Michel Salles, J.Settle, Shirakha surroundings [10,13,14,21,24,27-29,31,35]. B.E.Vaissiere. Economic Valuation of the Vulnerability B. bohemicus (Seidl, 1838) (=Psithyrus bohemicus) of World Agriculture Confronted with Pollinator Decline // Distribution: Khando, Katsalkhevi, Kmosti, Chrdilis khevi Ecological Economics. 68(3), 2009, pp. 810-821. [10,26,27,29,33,34]. 4. Lautenbach S, Seppelt R, Leibscher J, Dormann. B. campestris (Panzer,1801) (=Psithyrus campestris) Spatial and Temporal Trends of Global Pollination Benefi t Distribution: Borjomi reserve, Atskuri, Adigeni, Kintrishi // PLoS ONE, 7(4), 2012, pp. 1-16. [10,16,17,24,25,27,37]. 5. BurdM. Bateman’s Principle and Reproduction: the B. maxillosus Klug,1817 (=Psithyrus maxillosus) Role of Pollinator Limitation in Fruit and Seed Set // Bot. Distribution: Minadze, Arali, Igoeti, Atskuri, Akhaltsikhe, Rev. 60, 1994, pp. 83-139. Kaspi [10,17,24,25,27,28,32,33,34]. 6. Garibaldi A. Global Growth and Stability of B. rupestris (Fabricius,1793) (=Psithyrus rupestris) Agricultural Yeld Decrease with Pollinator Dependence // Distribution: Kintrishi, Akhalkalai, Atskuri, Barisakho PNAS, 18, 2011, pp. 5909-5914. [10,24,25,27,29,33,34,37]. 7. Klug M. and G. Büneman. Pollination: wild Bees Genus –Apis Linnaeus, 1758 as an Alternative to the Honey Bee // Acta Horticulturae, A. mellifera Linnaeus, 1758 139, 1983, pp. 59–64. Distribution: Throughout all Georgia. 8. Mitrou, M. Fauna Europaea: Hymenoptera. Fauna List of 356 species belonging to 46 genera and 6 families has Europaea version 2.6 // 2013, http://www.faunaeur.org. been completed after analizing all existed literature about 9. Pickering J. Discover life // 2013, http://www. bees distribution in Georgia. Validity of all species has been discoverlife.org/, checked and compared with Fauna Europea [8]and Discover 10. Skhirtladze I. Bees of Caucasus (Hymenoptera, life [9]web sources. Fortyone species were reperesenting Apoidea) // GNAS, National Museum of Georgia, Tbilisi, family Colletidae, Family Andrenidae was represented 1981, 147 pp. (in Russian) with 82 species, Family Halictidae with 60 species, Family 11. Skhirtladze I. For Bee Vertical Zonal Distribution Melitidae with 6 species, Family Megachilidae with 58 Studies in Batsara Reserve // Nature Protection of Georgia, species, and fi nally the most abundant family was family 10, 1982, pp 98-105 (in Georgian). Apidae with 107 species. Must be noted that 41 species of 12. Skhirtladze I. Bees (Hymenoptera, Apoidea) Bombus have been registered in Georgia, while in Caucasus of Georgia and Their Protection // Nature Protection of there are only three more species recorded: B. melanurus Georgia, 11, 1983, pp. 164–176 (in Georgian). Lepelietier, 1836, B. zonatus Smith, 1854 and B. vestalis 13. Skhirtladze I. Bombus Distribution in Georgia (Geoffroy, 1785) [10]. We were not able to fi nd validity of and Some Aspect of Their Protection // Nature Protection Bombus georgicus Vogt. Therefore we did not include this of Georgia, 13, 1985, pp 59–68 (in Georgian). species in the list. 14. Skhirtladze I. Apidofauna of Svaneti // Nature Halictus cervinus F.Mor.[29] most must be H. cephalicus Protection of Georgia, Bulletin of Georgian Academy of Morawitz, 1873, however as we were not able to examine Sciences, 1986, pp. 112-121(in Georgian). voucher specimens we excluded this specie sfrom the list. 15. Skhirtladze I. For Some Aspects of Landscape We were not able to fi nedany information about species Distribution of Bees from Algeti Reserve // Reserves of Halictus pditus Schck.[30], also we could net check voucher Georgia, 4, 1987, pp. 263-267 (in Georgian). specimens, therefore we excluded this species from the list. 16. Skhnirtladze I. For Study of Vertical-Zonal Distribution of Bees from Borjomi State Reserve // ACKNOWLEDGMENT Reserves of Georgia, 6, 1987, pp. 268-273 (in Georgian). We would like to thank to Mr Øistein Berg, Norway and Dr 17. Skhirtladze I. The Key for Bees from Caucasus Ryuki Murao, Kyushu University, Japan for their kind help Isthmus // Academy of Sciences of Georgian SSR, Tbilisi, for valid name for some problematic species. 1988, pp. 1-33 (in Russian). 18. Skhirtladze I. List of Short Tramp Bees REFERENCES (Hymenoptera, Andrenidae) of Georgia // Proceedings of 1. Michener C.D. The Bees of the World // Baltimore, the Institute of Zoology, 21, 2002, pp. 214-221. 2007, 953 pp. 19. Skhirtladze I. The List of Bees of Caucasus 2. Klein A., B. E. Vaissiere, J.H.Cane, I.Steffan- (Hymenoptera: Colletidae, Melittidae) // Caucasian Dewenter, S.A. Cunninham, C.Kremen and T. messenger. 8, 2003, http://www.nplg.gov.ge/caucasia/ Tscharntke. Importance of Pollinators in Changing messenger/Eng/Page1.htm

31 20. Skhirtladze I. Faunistic List of Bees of Caucasus 30. Skhirtladze I. To Study Egrisi Gorges Southern (Hymenoptera:Apoidea,Halictidae). Caucasology, 5, 2004, Slope Bees (Zugdidi, Chkhorotsku, Tsalenjikha) // Bulletin pp 71-80. of Acad. S. Janashia State Museum of Georgia, 29 (A), 21. Skhirtladze I. Some Kinds of Wild Bees Subject 1976, pp. 134-153 (in Georgian). to Protection (Hymenoptera, Apidae). Russian Academy of 31. Skhirtladze I. The Materials to Study Bees Sciences, Institute of Linguistics Caucasology, 8, 2005,pp. of Lagodekhi Reserve // Proceedings of Reserves and 42-46 (in Russian). Hunting Company of Forestry Production Committee of 22. Skhirtladze I. Faunistik List of Bees (Hymenoptera, Ministry Board of Georgian SSR, 4, 1976, pp. 337-347 (in Andrenidae) of Transcaucasus // Caucasology, 11, 2006, Georgian). pp. 46-60. (In Russian) 32. Skhirtladze I. The Materials of Bee (Hymenoptera: 23. Skhirtlade I. List of Bees (Hymenoptera, Apoidea) Fauna of Transcaucasus Arid Districts // Halictidae) of Georgia // Proceedings of the Institute of Some Groups of Anikmals from Transcaucasus Arid Zoology. 23, 2008, pp.153-158. Districts.1979, pp.115-145 (in Russian). 24. Skhirtladze I. The Materials to Study Bees of 33. Skhirtladze I. Ecological discussion of Bees Meskhet-Javakheti // Bulletin of Geogian Academy of (Hymenoptera: Apoidea) of Georgia // Bulletin of Acad. S. Sciences, 46, 1967, pp. 215-222 (in Georgian). Janashia State Museum of Georgia, 30 (A), 1980, pp.176- 25. Skhirtladze I. To study Hymenopterans of Minor 189 (in Russian). Caucasus // Bulletin of Acad. S. Janashia State Museum of 34. Skhirtladze I. The New Data to Study Bee Georgia, 26-27 (A), 1970, pp.191-205 (in Georgian). (Hymenoptera: Apoidea) Settlements in Georgia // Bulletin 26. Skhirtladze I. To Study Bees of Minor Caucasus of Georgian Academy of Sciences. 98(3), 1980, pp. 701- (Tsalka and Dmanisi Districts) // Bulletin of Acad. S. 704. (in Russian) Janashia State Museum of Georgia, 28 (A), 1975, pp. 270- 35. Skhirtladze I. To Study Bees (Hymenotpera, 290 (in Georgian). Apoidea) of Racha (Oni and Ambrolauri Districts) // 27. Skhirtladze I. Faunistic Complexes of Bees Bulletin of Acad. S. Janashia State Museum of Georgia, 31 and Their Distribution in Georgia // Bulletin of Acad. S. (A), 1981, pp.181-201 (in Russian). Janashia State Museum of Georgia, 28 (A), 1975, pp. 413- 36. Skhirtladze I. To Study Fauna of Vashlovani 429 (in Russian). National Park Reserve Bees (Hymenoptera: Apoidea) // 28. Skhirtladze I. Bees (Hymenoptera: Apoidea) of Proceedings of Reserves of Georgia // 5, 1981, pp.145-160 Georgia (Fauna, Distribution, Ecological and Ladscape (in Georgian). Characteristics) // Thesis of PhD, 1975, 26 pp. (in Russian). 37. Skhirtladze I. To Study Vertical-Zonal Distribution 29. Skhirtladze I. To Study Bees (Hymenoptera: of Apidofauna (Hymenoptera: Apoidea) in Kintrishi Apoidea) of Pshav-Khevsureti // Bulletin of Acad. S. Reserve // Proceedings of Reserves of Georgia, 5, 1981, Janashia State Museum of Georgia, 29 (A), 1976, pp. 110- pp.161-165 (in Georgian). 133 (in Georgian).

ОБНОВЛЕННЫЙ СПИСОК ПЧЕЛИНЫХ (HYMENOPTERA: APOIDEA) ГРУЗИИ

Г. Дж. Киркитадзе, Г.О. Джапошвили

Пчелы являются одной из наиболее разнообразных групп насекомых в мире. Пчелы имеют особое значение для экосистем. Почти 75% сельскохозяйственного производства зависит от интенсивности опыления. Многие из пчелиных используются в теплицах для повышения урожайности. Пчелы не очень хорошо защищены от влияния различных факторов, которые могут повлиять на их количество. Для сохранения пчел знание важности и данных их распространения очень важны. За последние 30 лет не был пересмотрен таксономический статус пчел, распространённых в Грузии. Полный список пчел Грузии был интегрирован в списк пчел Кавказа в 1981 году. С тех пор составлен полный список всех пчел Грузии, с таксономическими изменениями, сделанными за последние 33 года. Всего было внесено 356 вида, принадлежащих к 46 различным родам и 6 семействам. Было внесено более 150 изменений в названия семейств, родов и видов, зарегистрированных и опубликованных в Грузии. Наиболее распространенным семейством является Apidae с 107 видами. Следует отметить, что 41 вид Bombus был зарегистрирован в Грузии, что дополняет фауну Bombus Кавказа еще тремя видами. Нам не удалось подтвердить валидность Bombus georgicus Vogt. Поэтому этот вид не включен в список.

32