DOCSLIB.ORG

Mammalia, Bovidae) from the Turolian of Hadjidimovo, Bulgaria, and a Revision of the Late Miocene Mediterranean Boselaphini

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Mammalia, Bovidae) from the Turolian of Hadjidimovo, Bulgaria, and a Revision of the Late Miocene Mediterranean Boselaphini Tragoportax Pilgrim, 1937 and Miotragocerus Stromer, 1928 (Mammalia, Bovidae) from the Turolian of Hadjidimovo, Bulgaria, and a revision of the late Miocene Mediterranean Boselaphini Nikolai SPASSOV National Museum of Natural History, 1 blvd Tsar Osvoboditel, 1000 Sofia (Bulgaria) [email protected] Denis GERAADS UPR 2147 CNRS, 44 rue de l’Amiral Mouchez, F-75014 Paris (France) [email protected] Spassov N. & Geraads D. 2004. — Tragoportax Pilgrim, 1937 and Miotragocerus Stromer, 1928 (Mammalia, Bovidae) from the Turolian of Hadjidimovo, Bulgaria, and a revision of the late Miocene Mediterranean Boselaphini. Geodiversitas 26 (2) : 339-370. ABSTRACT KEY WORDS A taxonomic revision of the late Miocene Boselaphini is proposed on the basis Mammalia, Bovidae, of the description of abundant Turolian material from the locality of Boselaphini, Hadjidimovo, Bulgaria. The genus Tragoportax Pilgrim, 1937 as usually Tragoportax, Miotragocerus, understood is divided into two genera – Tragoportax and Miotragocerus Pikermicerus, Stromer, 1928 – the latter itself divided into two subgenera – M.(Mio- late Miocene, tragocerus) Stromer, 1928 and M. (Pikermicerus) Kretzoi, 1941. The sexual Bulgaria, taphonomy, dimorphism and the paleoecology of the taxa are discussed as well as the ecology. taphonomy of Tragoportax from Hadjidimovo. RÉSUMÉ Tragoportax Pilgrim, 1937 et Miotragocerus Stromer, 1928 (Mammalia, Bovidae) du Turolien de Hadjidimovo, Bulgarie, et révision des Boselaphini du Miocène supérieur de Méditerranée. MOTS CLÉS À partir de la description de l’abondant matériel turolien de la localité de Mammalia, Bovidae, Hadjidimovo en Bulgarie, nous proposons une révision des Boselaphini du Boselaphini, Miocène supérieur. Le genre Tragoportax Pilgrim, 1937 tel qu’il est habituel- Tragoportax, Miotragocerus, lement compris est divisé en deux genres – Tragoportax et Miotragocerus Pikermicerus, Stromer, 1928 – ce dernier lui-même divisé en deux sous-genres M.(Mio- Miocène supérieur, tragocerus) Stromer, 1928 and M. (Pikermicerus) Kretzoi, 1941. Le dimor- Bulgarie, taphonomie, phisme sexuel et la paléoécologie des taxons, ainsi que la taphonomie de écologie. Tragoportax de Hadjidimovo sont aussi discutés. GEODIVERSITAS • 2004 • 26 (2) © Publications Scientifiques du Muséum national d’Histoire naturelle, Paris. www.geodiversitas.com 339 Spassov N. & Geraads D. INTRODUCTION which comes mainly from the locality Hadjidimovo-1 (Hadjidimovo-Girizite) and is The Bovids included in recent times in the genus still mostly unpublished, makes it the richest Tragoportax Pilgrim, 1937 were among the most upper Miocene site of Bulgaria and among the widespread in oriental and southeastern Europe, richest of the Eastern Mediterranean. Intensive as well as in Western Asia (the “Balkano-Iranian” investigations of the fauna and especially of the province: Spassov 2002; Geraads et al. 2003), in ungulates of the locality have started recently the upper Miocene. Numerous European, (Spassov & Ginsburg 1999; Geraads et al. 2001; Asiatic, and even African sites, have yielded Kostopoulos et al. 2001; Hristova et al. 2002; remains of various taxa linked to Tragoportax, Geraads et al. 2003). The faunal list of the loca- that were described during the last 150 years. lity Hadjidimovo-1 includes about 30 mammal These remains, unfortunately often incomplete species. The faunal complex of Hadjidimovo-1 or poorly preserved, are now stored in many shows similarities in taxonomic composition and scientific collections. The opinions expressed by faunal associations with several Hipparion locali- authors are often incomplete or contradictory, ties of the Balkano-Iranian region, and of the and numerous taxonomic and nomenclatorial Northern Paratethys, indicating a middle/late problems remain unsolved. In spite of the wealth Maeotian (middle Turolian) age. Hadjidimovo is of scientific papers on the subject, recent revi- somewhat earlier than Pikermi and it could be sions do not provide a satisfactory frame of the placed at the MN11/12 boundary, or perhaps at taxonomy of this group of fossil boselaphines. the beginning of MN12 (Spassov 2002). We describe here the rich and well preserved sample of boselaphines from the Turolian site of ABBREVIATIONS Hadjidimovo-1 (southwestern Bulgaria). This HD Hadjidimovo; MNHN Muséum national d’Histoire naturelle, also gives us the opportunity to review critically Paris; the taxonomic features mentioned in the litera- NHMW Naturhistorisches Museum Wien; ture, and to propose our own conceptions on NMNH National Museum of Natural History, the taxonomic frame of the “Tragoportax- Sofia; PIK Pikermi; Miotragocerus complex” as well as a new revision PIM Paläontologisches Institut Münster; of the late Miocene Mediterranean Boselaphini. SAM Samos. Upper teeth are in upper case, lower teeth in lower THE LOCALITY HADJIDIMOVO-1: case. LOCATION, FAUNA AND BIOCHRONOLOGY The Hadjidimovo fossiliferous area is situated in the Mesta river valley near the Hadjidimovo SYSTEMATICS town (Gotse Delchev district) and the Bulgarian- Greek border; it is a late Miocene fossil site with Order ARTIODACTYLA Owen, 1848 four localities of vertebrate fauna. The sediments Family BOVIDAE Gray, 1821 (light clayey sands) belong to the Nevrokop Subfamily BOVINAE Gray, 1821 Formation (Vatsev 1980). The site was first men- tioned by Nikolov (1973, 1985) but its richness Tribe BOSELAPHINI Knottnerus-Meyer, 1907 was only fully acknowledged after the excavations of Dimitar Kovachev between 1985 and 1998. Sokolov (1953) created a new subtribe Trago- More than 19,000 bone remains from cerina for the fossil antelopes Tragocerus Gaudry, Hadjidimovo (D. K. collection) are presently 1861, Miotragocerus Stromer, 1928, Para- stored in the Assenovgrad Museum – a paleonto- tragocerus Sokolov, 1949 and Sivaceros Pilgrim, logical division of the National Museum of 1937, but the monophyly of this group is Natural History – Sofia. This huge collection, questionable. Moyá-Solá (1983) defines the 340 GEODIVERSITAS • 2004 • 26 (2) Boselaphini (Mammalia, Bovidae) of Bulgaria Boselaphini by several features as follows: subtri- ?Tragoportax curvicornis Andree, 1926. Samos angular basal section on the horn-cores related to (= T. browni Pilgrim, 1937, Siwaliks, Pakistan, Turolian). the presence of distinct anterior and postero- Tragoportax salmontanus Pilgrim, 1937. Siwaliks external keels; presence of rugosities and crests on (Pakistan). (About 8.1 to 7.9 Ma, but this absolute the anterior part of the parietals. In fact, only the date is doubtful: Barry et al. 2002). presence of the anterior keel is a constant feature Tragoportax maius Meladze, 1967. Bazaleti, Georgia (late Turolian). Perhaps a synonym of the poorly for all taxa included in the Boselaphini. The known Tragoportax eldaricus (Gabashvili, 1956), type other ones are only strong tendencies that may no species of the genus Mirabilocerus Hadjiev, 1961, from be expressed in some species, or variably the late Vallesian/early Turolian of Eldar in expressed within the same species. Azerbaijan. ?Tragoportax cyrenaicus Thomas, 1979, Sahabi (Libya), probably late Miocene. Tragoportax acrae (Gentry, 1974). Langebaanweg Genus Tragoportax Pilgrim, 1937 (South Africa), Mio-Pliocene. Tragoportax macedoniensis Bouvrain, 1988. Dytiko Tragocerus Gaudry, 1861: 298 (type species: Capra (Greece; MN13). amalthea Roth & Wagner, 1854) (non Tragocerus de A number of other specific names, several of them Jean, 1821). from Asia, often based upon fragmentary material, are Tragoportax Pilgrim, 1937: 774. of doubtful validity; we will mention them in the comparison and discussion but we will not try to revise Pontoportax Kretzoi, 1941: 341 (type species: Trago- them here. cerus parvidens Schlosser, 1904). STRATIGRAPHIC AND GEOGRAPHIC DISTRIBUTION. — ?Gazelloportax Kretzoi, 1941: 341 (type species: G. gal- From the Vallesian/Turolian boundary (or late licus Kretzoi, 1941). Vallesian?) to the end of the Turolian, perhaps earli- est Pliocene in Africa. From southeastern Europe and ?Mirabilocerus Hadjiev, 1961: 3 (type species: the northern Paratethys region through Asia minor Tragocerus eldaricus Gabashvili, 1956). and the Middle East to Africa and the northern part Tragoceridus Kretzoi, 1968: 165 (type species: Capra of the Indian subcontinent (and possibly central amalthea Roth & Wagner, 1854). Asia). Mesembriportax Gentry, 1974: 146 (type species: NEW DIAGNOSIS. — Size generally large, approxi- M. acrae Gentry, 1974). mately that of European Cervus elaphus. The post- cornual fronto-parietal surface is a flat or slightly ?Mesotragocerus Korotkevich, 1982: 10 (type species: concave well defined depressed area, usually bor- M. citus Korotkevich, 1982). dered laterally by well marked temporal ridges and caudally by a step leading to a slightly raised plateau TYPE SPECIES OF TRAGOPORTAX. — Tragoportax salmontanus Pilgrim, 1937 (Pilgrim 1937: 774) by (Fig. 1). The basi-occipital has a longitudinal groove original designation. between the anterior and the posterior tuberosities, in the bottom of which often runs a weak sagittal INCLUDED SPECIES. — Tragoportax amalthea (Roth & crest (Fig. 2). Adult male horn-cores are long and Wagner, 1854). Pikermi, most probably also Samos slender, usually curved backwards, with a triangular and Halmyropotamos (Greece) (= ?T. frolovi to subtriangular cross-section, well marked postero- (M.Pavlow, 1913), Chobruchi [Moldova]); from the lateral keel and flattened lateral sides, but are less end(?) of the early Turolian to the beginning of the
Load more

Recommended publications
  • JVP 26(3) September 2006—ABSTRACTS
    Neoceti Symposium, Saturday 8:45 acid-prepared osteolepiforms Medoevia and Gogonasus has offered strong support for BODY SIZE AND CRYPTIC TROPHIC SEPARATION OF GENERALIZED Jarvik’s interpretation, but Eusthenopteron itself has not been reexamined in detail. PIERCE-FEEDING CETACEANS: THE ROLE OF FEEDING DIVERSITY DUR- Uncertainty has persisted about the relationship between the large endoskeletal “fenestra ING THE RISE OF THE NEOCETI endochoanalis” and the apparently much smaller choana, and about the occlusion of upper ADAM, Peter, Univ. of California, Los Angeles, Los Angeles, CA; JETT, Kristin, Univ. of and lower jaw fangs relative to the choana. California, Davis, Davis, CA; OLSON, Joshua, Univ. of California, Los Angeles, Los A CT scan investigation of a large skull of Eusthenopteron, carried out in collaboration Angeles, CA with University of Texas and Parc de Miguasha, offers an opportunity to image and digital- Marine mammals with homodont dentition and relatively little specialization of the feeding ly “dissect” a complete three-dimensional snout region. We find that a choana is indeed apparatus are often categorized as generalist eaters of squid and fish. However, analyses of present, somewhat narrower but otherwise similar to that described by Jarvik. It does not many modern ecosystems reveal the importance of body size in determining trophic parti- receive the anterior coronoid fang, which bites mesial to the edge of the dermopalatine and tioning and diversity among predators. We established relationships between body sizes of is received by a pit in that bone. The fenestra endochoanalis is partly floored by the vomer extant cetaceans and their prey in order to infer prey size and potential trophic separation of and the dermopalatine, restricting the choana to the lateral part of the fenestra.
    [Show full text]
  • Boselaphus Tragocamelus</I>
    University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln USGS Staff -- Published Research US Geological Survey 2008 Boselaphus tragocamelus (Artiodactyla: Bovidae) David M. Leslie Jr. U.S. Geological Survey, [email protected] Follow this and additional works at: https://digitalcommons.unl.edu/usgsstaffpub Leslie, David M. Jr., "Boselaphus tragocamelus (Artiodactyla: Bovidae)" (2008). USGS Staff -- Published Research. 723. https://digitalcommons.unl.edu/usgsstaffpub/723 This Article is brought to you for free and open access by the US Geological Survey at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in USGS Staff -- Published Research by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. MAMMALIAN SPECIES 813:1–16 Boselaphus tragocamelus (Artiodactyla: Bovidae) DAVID M. LESLIE,JR. United States Geological Survey, Oklahoma Cooperative Fish and Wildlife Research Unit and Department of Natural Resource Ecology and Management, Oklahoma State University, Stillwater, OK 74078-3051, USA; [email protected] Abstract: Boselaphus tragocamelus (Pallas, 1766) is a bovid commonly called the nilgai or blue bull and is Asia’s largest antelope. A sexually dimorphic ungulate of large stature and unique coloration, it is the only species in the genus Boselaphus. It is endemic to peninsular India and small parts of Pakistan and Nepal, has been extirpated from Bangladesh, and has been introduced in the United States (Texas), Mexico, South Africa, and Italy. It prefers open grassland and savannas and locally is a significant agricultural pest in India. It is not of special conservation concern and is well represented in zoos and private collections throughout the world. DOI: 10.1644/813.1.
    [Show full text]
  • Revision of the Boselaphin Bovid Miotragocerus Monacensis STROMER, 1928 (Mammalia, Bovidae) at the Middle to Late Miocene Transition in Central Europe
    N. Jb. Geol. Paläont. Abh. 276/3 (2015), 229–265 Article Stuttgart, June 2015 Revision of the boselaphin bovid Miotragocerus monacensis STROMER, 1928 (Mammalia, Bovidae) at the Middle to Late Miocene transition in Central Europe Jochen Fuss, Jérôme Prieto, and Madelaine Böhme With 19 figures and 8 tables Abstract: During excavations in 2011-2014, new fossil material of the boselaphin Miotragocerus monacensis STROMER, 1928 (Mammalia, Bovidae) was found at the locality Hammerschmiede (Bavaria, Germany), which is dated to ~11.6 Ma (Middle to Late Miocene transition). For the first time, both dentition and postcranial material can be studied on this species. These new findings complete a collection of casts stored in the Bavarian State Collection for Palaeontology and Geology. In addition, the holotype of M. monacensis from Oberföhring (Bavaria, Germany) and further unpublished material from Southern Germany and Lower Austria are newly described in this study. Important new taxonomic characters are emphasized improving our knowledge on the species which was originally described based on one single horn core. M. monacensis can be assigned to the basal Boselaphini based on the plesiomorphic features in the dentition and characters of the postcranial material. Intraspecific variabilities, ontogenetic changes and allometries are identified improving the differenciation to other basal boselaphins like Miotragocerus pannoniae, Austroportax latifrons and Protagocerus chantrei. An improved statement regarding the biostratigraphic range of basal Boselaphines from Central Europe is provided. Key words: Taxonomy, Biostratigraphy, Boselaphini, Miotragocerus monacensis, Middle to Late Miocene transition, Central Paratethys, Southern Germany, Lower Austria. 1. Introduction The genus Miotragocerus STROMER, 1928 was one of the dominant taxa among the Boselaphini during The locality Hammerschmiede (Bavaria, Germany) pro- the Late Miocene in terms of diversity and geographic vides a rare insight into the European palaeoecosystem distribution.
    [Show full text]
  • Unravelling the Positional Behaviour of Fossil Hominoids: Morphofunctional and Structural Analysis of the Primate Hindlimb
    ADVERTIMENT. Lʼaccés als continguts dʼaquesta tesi queda condicionat a lʼacceptació de les condicions dʼús establertes per la següent llicència Creative Commons: http://cat.creativecommons.org/?page_id=184 ADVERTENCIA. El acceso a los contenidos de esta tesis queda condicionado a la aceptación de las condiciones de uso establecidas por la siguiente licencia Creative Commons: http://es.creativecommons.org/blog/licencias/ WARNING. The access to the contents of this doctoral thesis it is limited to the acceptance of the use conditions set by the following Creative Commons license: https://creativecommons.org/licenses/?lang=en Doctorado en Biodiversitat Facultad de Ciènces Tesis doctoral Unravelling the positional behaviour of fossil hominoids: Morphofunctional and structural analysis of the primate hindlimb Marta Pina Miguel 2016 Memoria presentada por Marta Pina Miguel para optar al grado de Doctor por la Universitat Autònoma de Barcelona, programa de doctorado en Biodiversitat del Departamento de Biologia Animal, de Biologia Vegetal i d’Ecologia (Facultad de Ciències). Este trabajo ha sido dirigido por el Dr. Salvador Moyà Solà (Institut Català de Paleontologia Miquel Crusafont) y el Dr. Sergio Almécija Martínez (The George Washington Univertisy). Director Co-director Dr. Salvador Moyà Solà Dr. Sergio Almécija Martínez A mis padres y hermana. Y a todas aquelas personas que un día decidieron perseguir un sueño Contents Acknowledgments [in Spanish] 13 Abstract 19 Resumen 21 Section I. Introduction 23 Hominoid positional behaviour The great apes of the Vallès-Penedès Basin: State-of-the-art Section II. Objectives 55 Section III. Material and Methods 59 Hindlimb fossil remains of the Vallès-Penedès hominoids Comparative sample Area of study: The Vallès-Penedès Basin Methodology: Generalities and principles Section IV.
    [Show full text]
  • Evaluation of 3 Milk Replacers for Hand-Rearing Indian Nilgai (Boselaphus Tragocamelus)
    Evaluation of 3 Milk Replacers for Hand-Rearing Indian Nilgai (Boselaphus tragocamelus) Catherine D. Burch, M. Diehl, and M.S Edwards Zoological Society of San Diego, San Diego Wild Animal Park, San Diego, California Hoofstock neonates are often hand-raised for multiple reasons. Feeding maternal milk is optimal, but is rarely available for hand-rearing. Therefore, suitable substitutes must be identified. Published nutrient composition of maternal milk is frequently used as a guide. In many instances, however, no information is available regarding the milk composition of a particular species. Nilgai have been successfully raised at the Wild Animal Park using several formulas. It has yet to be determined which formula(s) produce the best response. Eight nilgai (3 females, 5 males) were received for hand-rearing and were randomly assigned 1 of the following 3 formulas for a blind study: evaporated cows (EC) :nonfat cows (NFC) 2: 1, EC: Milk Matrix 30/55®:water 10: 3: 12, or EC:SPF-Iac® 2: 1. EC: NFC 2: 1, which has been used for several seasons, was designated as the control. The composition of EC:30/55:H2O 10:3: 12 is comparable to the published composition of eland milk, a species taxonomically related to nilgai. EC:SPF-Iac® 2: 1 has lipid and carbohydrate composition intermediate to the other formulas. Response parameters monitored include weight gain, stool consistency, feeding response, and overall body condition. Based on the results, Indian nilgai can be hand-reared successfully on any of the 3 milk replacers. Of the 3, however, the group fed EC:NFC 2: 1 were the heaviest at weaning and the most vigorous throughout the hand-rearing process.
    [Show full text]
  • Chapter 1 - Introduction
    EURASIAN MIDDLE AND LATE MIOCENE HOMINOID PALEOBIOGEOGRAPHY AND THE GEOGRAPHIC ORIGINS OF THE HOMININAE by Mariam C. Nargolwalla A thesis submitted in conformity with the requirements for the degree of Doctor of Philosophy Graduate Department of Anthropology University of Toronto © Copyright by M. Nargolwalla (2009) Eurasian Middle and Late Miocene Hominoid Paleobiogeography and the Geographic Origins of the Homininae Mariam C. Nargolwalla Doctor of Philosophy Department of Anthropology University of Toronto 2009 Abstract The origin and diversification of great apes and humans is among the most researched and debated series of events in the evolutionary history of the Primates. A fundamental part of understanding these events involves reconstructing paleoenvironmental and paleogeographic patterns in the Eurasian Miocene; a time period and geographic expanse rich in evidence of lineage origins and dispersals of numerous mammalian lineages, including apes. Traditionally, the geographic origin of the African ape and human lineage is considered to have occurred in Africa, however, an alternative hypothesis favouring a Eurasian origin has been proposed. This hypothesis suggests that that after an initial dispersal from Africa to Eurasia at ~17Ma and subsequent radiation from Spain to China, fossil apes disperse back to Africa at least once and found the African ape and human lineage in the late Miocene. The purpose of this study is to test the Eurasian origin hypothesis through the analysis of spatial and temporal patterns of distribution, in situ evolution, interprovincial and intercontinental dispersals of Eurasian terrestrial mammals in response to environmental factors. Using the NOW and Paleobiology databases, together with data collected through survey and excavation of middle and late Miocene vertebrate localities in Hungary and Romania, taphonomic bias and sampling completeness of Eurasian faunas are assessed.
    [Show full text]
  • 71St Annual Meeting Society of Vertebrate Paleontology Paris Las Vegas Las Vegas, Nevada, USA November 2 – 5, 2011 SESSION CONCURRENT SESSION CONCURRENT
    ISSN 1937-2809 online Journal of Supplement to the November 2011 Vertebrate Paleontology Vertebrate Society of Vertebrate Paleontology Society of Vertebrate 71st Annual Meeting Paleontology Society of Vertebrate Las Vegas Paris Nevada, USA Las Vegas, November 2 – 5, 2011 Program and Abstracts Society of Vertebrate Paleontology 71st Annual Meeting Program and Abstracts COMMITTEE MEETING ROOM POSTER SESSION/ CONCURRENT CONCURRENT SESSION EXHIBITS SESSION COMMITTEE MEETING ROOMS AUCTION EVENT REGISTRATION, CONCURRENT MERCHANDISE SESSION LOUNGE, EDUCATION & OUTREACH SPEAKER READY COMMITTEE MEETING POSTER SESSION ROOM ROOM SOCIETY OF VERTEBRATE PALEONTOLOGY ABSTRACTS OF PAPERS SEVENTY-FIRST ANNUAL MEETING PARIS LAS VEGAS HOTEL LAS VEGAS, NV, USA NOVEMBER 2–5, 2011 HOST COMMITTEE Stephen Rowland, Co-Chair; Aubrey Bonde, Co-Chair; Joshua Bonde; David Elliott; Lee Hall; Jerry Harris; Andrew Milner; Eric Roberts EXECUTIVE COMMITTEE Philip Currie, President; Blaire Van Valkenburgh, Past President; Catherine Forster, Vice President; Christopher Bell, Secretary; Ted Vlamis, Treasurer; Julia Clarke, Member at Large; Kristina Curry Rogers, Member at Large; Lars Werdelin, Member at Large SYMPOSIUM CONVENORS Roger B.J. Benson, Richard J. Butler, Nadia B. Fröbisch, Hans C.E. Larsson, Mark A. Loewen, Philip D. Mannion, Jim I. Mead, Eric M. Roberts, Scott D. Sampson, Eric D. Scott, Kathleen Springer PROGRAM COMMITTEE Jonathan Bloch, Co-Chair; Anjali Goswami, Co-Chair; Jason Anderson; Paul Barrett; Brian Beatty; Kerin Claeson; Kristina Curry Rogers; Ted Daeschler; David Evans; David Fox; Nadia B. Fröbisch; Christian Kammerer; Johannes Müller; Emily Rayfield; William Sanders; Bruce Shockey; Mary Silcox; Michelle Stocker; Rebecca Terry November 2011—PROGRAM AND ABSTRACTS 1 Members and Friends of the Society of Vertebrate Paleontology, The Host Committee cordially welcomes you to the 71st Annual Meeting of the Society of Vertebrate Paleontology in Las Vegas.
    [Show full text]
  • Evolution and Biogeography of Haemonchus Contortus: Linking Faunal Dynamics in Space and Time
    View metadata, citation and similar papers at core.ac.uk brought to you by CORE provided by DigitalCommons@University of Nebraska University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln U.S. Department of Agriculture: Agricultural Publications from USDA-ARS / UNL Faculty Research Service, Lincoln, Nebraska 2016 Evolution and Biogeography of Haemonchus contortus: Linking Faunal Dynamics in Space and Time Eric P. Hoberg Agricultural Research Service, USDA, [email protected] D.S. Zarlenga Agricultural Research Service, USDA Follow this and additional works at: https://digitalcommons.unl.edu/usdaarsfacpub Hoberg, Eric P. and Zarlenga, D.S., "Evolution and Biogeography of Haemonchus contortus: Linking Faunal Dynamics in Space and Time" (2016). Publications from USDA-ARS / UNL Faculty. 2243. https://digitalcommons.unl.edu/usdaarsfacpub/2243 This Article is brought to you for free and open access by the U.S. Department of Agriculture: Agricultural Research Service, Lincoln, Nebraska at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Publications from USDA-ARS / UNL Faculty by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. CHAPTER ONE Evolution and Biogeography of Haemonchus contortus: Linking Faunal Dynamics in Space and Time E.P. Hoberg*,1, D.S. Zarlengax *US National Parasite Collection and Animal Parasitic Disease Laboratory, Agricultural Research Service, USDA, Beltsville, MD, United States x Animal Parasitic Disease Laboratory, Agricultural Research Service, USDA, Beltsville, MD, United States 1Corresponding author: E-mail: [email protected] Contents 1. Introduction 2 2. Haemonchus: History and Biodiversity 3 3. Phylogeny and Biogeography: Out of Africa 4 4. Domestication, Geographical Expansion and Invasion 7 5.
    [Show full text]
  • For Review Only 440 IUCN SSC Antelope Specialist Group
    Manuscripts submitted to Journal of Mammalogy A Pliocene hybridisation event reconciles incongruent mitochondrial and nuclear gene trees among spiral-horned antelopes Journal:For Journal Review of Mammalogy Only Manuscript ID Draft Manuscript Type: Feature Article Date Submitted by the Author: n/a Complete List of Authors: Rakotoarivelo, Andrinajoro; University of Venda, Department of Zoology O'Donoghue, Paul; Specialist Wildlife Services, Specialist Wildlife Services Bruford, Michael; Cardiff University, Cardiff School of Biosciences Moodley, Yoshan; University of Venda, Department of Zoology adaptive radiation, interspecific hybridisation, paleoclimate, Sub-Saharan Keywords: Africa, Tragelaphus https://mc.manuscriptcentral.com/jmamm Page 1 of 34 Manuscripts submitted to Journal of Mammalogy 1 Yoshan Moodley, Department of Zoology, University of Venda, 2 [email protected] 3 Interspecific hybridization in Tragelaphus 4 A Pliocene hybridisation event reconciles incongruent mitochondrial and nuclear gene 5 trees among spiral-horned antelopes 6 ANDRINAJORO R. RAKATOARIVELO, PAUL O’DONOGHUE, MICHAEL W. BRUFORD, AND * 7 YOSHAN MOODLEY 8 Department of Zoology, University of Venda, University Road, Thohoyandou 0950, Republic 9 of South Africa (ARR, ForYM) Review Only 10 Specialist Wildlife Services, 102 Bowen Court, St Asaph, LL17 0JE, United Kingdom (PO) 11 Cardiff School of Biosciences, Sir Martin Evans Building, Cardiff University, Museum 12 Avenue, Cardiff, CF10 3AX, United Kingdom (MWB) 13 Natiora Ahy Madagasikara, Lot IIU57K Bis, Ampahibe, Antananarivo 101, Madagascar 14 (ARR) 15 16 1 https://mc.manuscriptcentral.com/jmamm Manuscripts submitted to Journal of Mammalogy Page 2 of 34 17 ABSTRACT 18 The spiral-horned antelopes (Genus Tragelaphus) are among the most phenotypically diverse 19 of all large mammals, and evolved in Africa during an adaptive radiation that began in the late 20 Miocene, around 6 million years ago.
    [Show full text]
  • Copyright by Robert Smith Scott 2004
    Copyright by Robert Smith Scott 2004 The Dissertation Committee for Robert Smith Scott certifies that this is the approved version of the following dissertation: The Comparative Paleoecology of Late Miocene Eurasian Hominoids Committee: John Kappelman, Supervisor Claud Bramblett Deborah Overdorff Raymond L. Bernor Liza Shapiro The Comparative Paleoecology of Late Miocene Eurasian Hominoids by Robert Smith Scott, B.A., M.A. Dissertation Presented to the Faculty of the Graduate School of The University of Texas at Austin in Partial Fulfillment of the Requirements for the Degree of Doctor of Philosophy The University of Texas at Austin August, 2004 Dedication This dissertation is dedicated to my grandmother Alma Dinehart Smith who dedicated her working years to science as a scientific illustrator of considerable skill and who has always been an inspiration to her whole family. Acknowledgements This work was supported by NSF grant BCS-0112659 to J. Kappelman (PI) and R.S. Scott (Co-PI) and a Homer Lindsey Bruce Fellowship from the University of Texas at Austin to R.S. Scott. Thanks to Ray Bernor for the contribution of measurements from his extensive hipparion database funded by NSF grant EAR-0125009. Thanks to Adam Gordon and Dave Raichlen for their many commentaries on and off topic. Additional thanks to my committee John Kappelman, Ray Bernor, Claud Bramblett, Deborah Overdorff, and Liza Shapiro. Much gratitude is due my many hospitable European colleagues who have helped make this work possible. And finally, thanks to my wife Shannon for her patience and support throughout this process. v The Comparative Paleoecology of Late Miocene Eurasian Hominoids Publication No._____________ Robert Smith Scott, Ph.D The University of Texas at Austin, 2004 Supervisor: John Kappelman Abstract: Remains of late Miocene hominoids increasingly indicate both taxonomic and adaptive diversity.
    [Show full text]
  • Migration of Organisms Climate • Geography • Ecology Ashraf M
    Ashraf M.T. Elewa Migration of Organisms Climate • Geography • Ecology Ashraf M. T. Elewa (Editor) Migration of Organisms Climate • Geography • Ecology With 67 Figures 123 Dr. Ashraf M. T. Elewa Professor Minia University Faculty of Science Geology Department Egypt E-mail: [email protected] Library of Congress Control Number: 2005927792 ISBN-10 3-540-26603-8 Springer Berlin Heidelberg New York ISBN-13 978-3-540-26603-7 Springer Berlin Heidelberg New York This work is subject to copyright. All rights are reserved, whether the whole or part of the material is concerned, specifically the rights of translation, reprinting, reuse of illustrations, recitations, broadcasting, reproduction on microfilm or in any other way, and storage in data banks. Duplication of this publication or parts thereof is permitted only under the provisions of the German Copyright Law of September 9, 1965, in its current version, and permission for use must always be obtained from Springer. Violations are liable to prosecution under the German Copyright Law. Springer is a part of Springer Science+Business Media springeronline.com © Springer-Verlag Berlin Heidelberg 2005 Printed in The Netherlands The use of general descriptive names, registered names, trademarks, etc. in this publication does not imply, even in the absence of a specific statement, that such names are exempt from the relevant protective laws and regulations and therefore free for general use. Cover design: Erich Kirchner Production: Luisa Tonarelli Typesetting: Camera-ready by the editor Printed on acid-free paper 30/2132/LT – 5 4 3 2 1 0 Dedication This book is dedicated to all people who Believe in One God Believe in Peace Believe in Migration in the Way of God To my father who died on Sunday, the 10th of April, 2005 Foreword P.
    [Show full text]
  • Late Miocene Large Mammals from Mahmutgazi, Denizli Province, Western Turkey
    Late Miocene large mammals from Mahmutgazi, Denizli province, Western Turkey. Denis Geraads To cite this version: Denis Geraads. Late Miocene large mammals from Mahmutgazi, Denizli province, Western Turkey. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen, E Schweizerbart Science Publishers, 2017, 284 (3), pp.241-257. 10.1127/njgpa/2017/0661. halshs-01794188 HAL Id: halshs-01794188 https://halshs.archives-ouvertes.fr/halshs-01794188 Submitted on 17 May 2018 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. GERAADS, D. 2017. Late Miocene large mammals from Mahmutgazi, Denizli province, Western Turkey. Neues Jahrbuch für Geologie und Paläontologie, 284(3): 241-257. Denis GERAADS, CR2P, Sorbonne Universités, MNHN, CNRS, UPMC, CP 38, 8 rue Buffon, 75231 PARIS Cedex 05, France [email protected] Abstract: The upper Miocene locality of Mahmutgazi in Western Turkey was excavated in the 70s by a German team, but most of its large mammals had never been studied. The collection housed in the Staatliches Museum für Naturkunde, Karlsruhe, contains, besides previously published groups, large samples of Giraffidae (Samotherium), Rhinocerotidae (including a nice complete skull of Ceratotherium neumayri), and Equidae, as well as some Chalicotheriidae (Ancylotherium) and Bovidae (Boselaphini), which are studied here.
    [Show full text]