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NuytsirVol.8, No. 3 (19921

in Humea rccogtlisedby Mucller (1893) and thosc dcscribedlater basedon African material. All recentauthors (e.g. Lcwis & Summerhayes1951, Grieve 1975, Anderbcrg 1989, 1991) have fouowed Bentham(1867) in regardingPithocarpd as a dist.inctgenus. Willis (1967),in discussinga number of taxa related to Cassrria, recognised, and Hqeckeria .ts sepamtegenera distinct from Acomis and,Rutidosrs; his generic hasbeen accepted in Australia. Thc African that had been placed in Calomeria rre now consideredto be epapposespecics of llelichrlsum (Hilliard, and Burfr 1973,Hillird 1983).

In 1896 Mueller and Tate formally publishedHumea gradllima which Muellcr (1893) had previouslymentioned by nameonly; theysugg0sted that it was'closelyallied toA comismaua'. The formcr Llxon was subsequentlydescribed irdependently by Hemsley (1905) as a rew and spccies,Thiseltonio dleri. It differs from the speciescurrently included in Acomis in beirg a delictte minutelyglandular-puberulous annual, in havinglhe outcrflorets female with very slender,l-lobed corollas, in having (in the bisexual florels) truncatestyle appendagcswith a central subulateapex, and in havirg a dilphano uspapillose pericarp that is unitedto the pale brown paperytesta. In Acomis the speciesare more or lcss woolly, the florets arc all bisexual,and the p3ricarp is thick, crustaceous and free from the membrunoustesl,.

The speciescurrcntly placed in Aconiis, Rutidosis, and Thiseltonia haye involucral bracts with similar clcarly demarcatedstereomcs which completely enclosethe vascularstrands, they also hitve similff corollx shapes,and similar small anthersin which the tails are fine and difficult to discem.

AlthoughBentham ( 1867,1873) recognised both Acomis:urdRrridosrs DC. (1838)he considered their sepantion to be a ificial since it was basedsolely on the presence(in Ratidosrs)or absence(in Acrmrs)of apappus.The species ofthe two generacan also be dividedinto two groupsbasedon thcir style morphology. In one group the style appendageis truncrte while in thc other it is narrowly trii]ngular. In the former group arefound the type speciesofboth RulidosrsandAcomrs , vrz. Ruttdosis helichrysoidesDC. rtld Acomis macro F. Muell.; in the latter group are fould Rutidosi.sleucantha F.Mucll. andAcorzrsaconra (F.Muell.) Druce. Further wolk may indicate thatA comis at\d Rittidosis arebcst treatedxs beingcongcncric, or it may suggcstthat the genedccircumscription within the complexshould bebased more on themorphology ofrhe styleapex rnd lesson thepresenceorabsence ofa pappus; if the former ttxoromy werc dccided or then the correct name for thc complex would beRr/irlorrr,ifthe lrtter thena newgcneric neme would bc rcquiredfor thosespccies with acuminate stylcapices including the sirrgleAcozrs speciesdcscribed in this papcr

Anderbcrg(1991) commerts on theclose relationship between Acoais andRatiTosis and on the mffked similarity of lhese genera to Leptofilnchos utd Chrysocephalurn. I agrce with the first obseryalionbut not the secondsincc I considerthat the morphologyof the achenesand of the involucral bmcts in the fint pair ofgenera is soobviously distinct from the morphology ofthe achenes and bracts in the secondp:lir as to providc a clear generic scpffrtion.

Acomis

Acomiskakadu Paul G. Wilson,sp. nov. (Figure1)

Herbaerecta ad 40 cm alta,gossypinr. Folia lineffia, 2-4(8)cm longa.Involucrum hemisphcricum; bracteaemultiserirtae, homomorphicae:lamina ovata,albl, 4-8 mm longa. Receptaculumconicum, 'oureH 'llenhl - 'llonhl (el?I ? C) otu11|:totqDtrauopJ uawor! (E6BI)/VI:6 tuN ucrrolcr^ C '11anv1 :@6gDL9t:91 ulJr"Ilsnvqtnos ros fou rord f suorJ'alcJ ? gow4rto.t8 oawnp

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^-uytsia Vol. 8, No. 3 (1992)

E

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