DOCSLIB.ORG

Bathyraja Panthera, a New Species of Skate (Rajidae: Arhynchobatinae) from the Western Aleutian Islands, and Resurrection

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Bathyraja Panthera, a New Species of Skate (Rajidae: Arhynchobatinae) from the Western Aleutian Islands, and Resurrection NOAA Professional Paper NMFS 11 U.S. Department of Commerce March 2011 Bathyraja panthera, a new species of skate (Rajidae: Arhynchobatinae) from the western Aleutian Islands, and resurrection of the subgenus James W. Orr Duane E. Stevenson Arctoraja Ishiyama Gerald R. Hoff Ingrid Spies John D. McEachran U.S. Department of Commerce NOAA Professional Gary Locke Secretary of Commerce National Oceanic Papers NMFS and Atmospheric Administration Jane Lubchenco, Ph.D. Scientific Editor Administrator of NOAA Richard D. Brodeur, Ph.D. Associate Editor National Marine Julie Scheurer Fisheries Service Eric C. Schwaab National Marine Fisheries Service Assistant Administrator Northwest Fisheries Science Center for Fisheries 2030 S. Marine Science Dr. Newport, Oregon 97365-5296 Managing Editor Shelley Arenas National Marine Fisheries Service Scientific Publications Office 7600 Sand Point Way NE Seattle, Washington 98115 Editorial Committee Ann C. Matarese, Ph.D. National Marine Fisheries Service James W. Orr, Ph.D. National Marine Fisheries Service Bruce L. Wing, Ph.D. National Marine Fisheries Service The NOAA Professional Paper NMFS (ISSN 1931-4590) series is published by the Scientific Publications Office, National Marine Fisheries Service, The NOAA Professional Paper NMFS series carries peer-reviewed, lengthy original NOAA, 7600 Sand Point Way NE, research reports, taxonomic keys, species synopses, flora and fauna studies, and data-in- Seattle, WA 98115. tensive reports on investigations in fishery science, engineering, and economics. Copies The Secretary of Commerce has determined that the publication of of the NOAA Professional Paper NMFS series are available free in limited numbers to this series is necessary in the transac- government agencies, both federal and state. They are also available in exchange for tion of the public business required by other scientific and technical publications in the marine sciences. Professional Papers law of this Department. Use of funds are published online in PDF format at http://spo.nmfs.noaa.gov for printing of this series has been ap- proved by the Director of the Office NOTICE: This series was established in 2003 to replace the NOAA Technical Report of Management and Budget. NMFS series. NOAA Professional Paper NMFS 11 Bathyraja panthera, a new species of skate (Rajidae: Arhynchobatinae) from the western Aleutian Islands, and resurrection of the subgenus Arctoraja Ishiyama James W. Orr Duane E. Stevenson Gerald R. Hoff Ingrid Spies John D. McEachran March 2011 U.S. Department of Commerce Seattle, Washington Suggested reference Orr, James W., Duane E. Stevenson, Gerald R. Hoff, Ingrid Spies, and John D. McEachran. 2011. Bathyraja panthera, a new species of skate (Rajidae: Arhyncho- batinae) from the western Aleutian Islands, and resurrection of the subgenus Arctoraja Ishiyama. NOAA Professional Paper NMFS 11, 50 p. Online dissemination This report is posted online in PDF format at http://spo.nmfs.noaa.gov (click on Professional Papers link). Purchasing additional copies Additional copies of this report may be available for purchase in paper copy or microfiche from the National Technical Information Service, 5285 Port Royal Road, Springfield, VA 22161; 1-800-553-NTIS; http://www.ntis.gov. Copyright law Although the contents of the Professional Papers have not been copyrighted and may be reprinted entirely, reference to source is appreciated. Proprietary products The National Marine Fisheries Service (NMFS) does not approve, recommend, or endorse any proprietary product or proprietary material mentioned in this publication. No reference shall be made to NMFS, or to this publication fur- nished by NMFS, in any advertising or sales promotion which would indicate or imply that NMFS approves, recommends, or endorses any proprietary product or proprietary material mentioned herein, or which has as its purpose an intent to cause directly or indirectly the advertised product to be used or purchased because of this NMFS publication. 1 Abstract—We provide morphological Bathyraja panthera, a new species and molecular evidence to recognize a new species of skate from the North Pa- of skate (Rajidae: Arhynchobatinae) cific, Bathyraja panthera. We also resurrect the skate subgenus Arctoraja Ishiyama, from the western Aleutian Islands, confirming its monophyly and the valid- and resurrection of the subgenus Arctoraja Ishiyama ity of the subgenus. Arctoraja was previ- ously recognized as a distinct subgenus of Breviraja and later synonymized with James W. Orr (contact author)1 Bathyraja (family Rajidae). Although the Duane E. Stevenson1 nominal species of Arctoraja have all been 1 considered synonyms of Bathyraja par- Gerald R. Hoff mifera by various authors, on the basis of Ingrid Spies1 morphometric, meristic, chondrological, John D. McEachran2 and molecular data we recognize four 1 species, including the new species. Spe- Resource Assessment and Conservation Engineering Division cies of Arctoraja are distributed across the Alaska Fisheries Science Center North Pacific Ocean and adjacent seas National Marine Fisheries Service, National Oceanic and Atmospheric Administration from southern Japan to British Colum- 7600 Sand Point Way NE bia. Bathyraja parmifera is abundant in Seattle, Washington 98115 the eastern Bering Sea, Aleutian Islands, Email address for contact author: [email protected] and northern Gulf of Alaska; B. smirnovi 2 Department of Wildlife and Fisheries Sciences is a western Pacific species found in the Texas A&M University Sea of Okhotsk and Sea of Japan; B. si- College Station, Texas 77843 moterus is restricted to waters around the northern and eastern coasts of Hokkaido, Japan; and the new species B. panthera is restricted to the western Aleutian Islands. Bathyraja panthera is diagnosed by its color pattern of light yellow blotches with black Introduction at least 25 species are distributed from spotting on a greenish brown background, Japan to California (Ishihara and high thorn and vertebral counts, chondro- Recent bottom-trawl surveys conduct- Ishiyama, 1986; Ishihara, 1990; Ste- logical characters of the neurocranium ed by the Alaska Fisheries Science venson et al., 2004; Ebert and Com- and clasper, and a unique nucleotide sequence within the mitochondrial cyto- Center (AFSC) of the National Ma- pagno, 2007). Twelve species are now chrome oxidase gene. Furthermore, the rine Fisheries Service (NMFS) have considered valid in Alaskan waters species presently recognized as Bathyraja resulted in the collection and subse- (Stevenson et al., 2004; Stevenson and parmifera exhibits two haplotypes among quent discovery of several new species Orr, 2005; Stevenson et al., 2007) and specimens from Alaska, suggesting the and records of fishes from the Aleu- further work is being conducted to possibility of a second, cryptic species. tian Islands, including liparids (Orr clarify the status of suspected species and Busby, 2001, 2006; Orr, 2004; complexes within the region. One of Orr and Maslenikov, 2007), zoarcids these complexes includes the most (Stevenson and Orr, 2006), and skates common species in Alaska, Bathyraja (Stevenson et al., 2004; Stevenson and parmifera, the Alaska skate (Stevenson Orr, 2005). Among those species that et al., 2008) of the subgenus Arctoraja. have been recognized as new but un- Ishiyama (1958a) erected three described is a skate from the western subgenera, Arctoraja, Bathyraja, and Aleutian Islands previously identified Notoraja, for the Japanese skates of as Bathyraja parmifera (Bean, 1881). the widely distributed genus Brevi- Skates of the family Rajidae, domi- raja Bigelow and Schroeder, 1948. nated by the genus Bathyraja, rep- The subgenus Arctoraja included two resent a substantial portion of the species of the North Pacific region, biomass of incidental species encoun- B. parmifera and B. smirnovi (Solda- tered by fisheries in the eastern Ber- tov and Pavlenko, 1915), as well as ing Sea (Spies et al., 2006; Stevenson Ishiyama’s two new subspecies within et al., 2008). Bathyraja Ishiyama, 1958, B. smirnovi: B. s. smirnovi and B. s. comprising over 50 species (Stehm- ankasube. Later, Ishiyama (1967) de- ann, 2005; Ebert and Compagno, scribed Breviraja (Arctoraja) simoterus 2007), is the most diverse genus of based on specimens he had previ- the family, with its major center of ously identified asB. parmifera, yet he abundance in the North Pacific where did not list B. parmifera as a member 2 Professional Paper NMFS 11 of the subgenus. When Ishiyama and Hubbs (1968) elevated the subgenus Bathyraja to generic status, they considered Arctoraja and Notoraja to be synonyms of Bathyraja. Although no comprehensive revision of the sub- genus Arctoraja has been produced since Ishiyama’s (1958a, 1967) seminal works, taxonomic reviews of North Pacific species ofBathyraja have come to contra- dictory conclusions on the status of Bathyraja parmifera and its close relatives. Ishihara and Ishiyama (1985, 1986) treated B. parmifera, B. smirnovi, and B. simoterus as distinct species, as do all recent Japanese authors (Nakaya, 1983, 1984; Nakaya and Shirai, 1992; Nakabo, 2000, 2002), distinguishing them within the context of dichotomous keys and checklists. In their phylogenetic analysis of the Rajidae, McEachran and Dunn (1998) listed all species in a generic classification of species, as did Stehmann (1986), McEachran and Miyake (1990), Figure 1 Compagno (1999), and Ebert and Compagno (2007) Selected terminology used to describe the external in their distributional analyses
Load more

Recommended publications
  • Notoraja Martinezi Sp. Nov., a New Species of Deepwater Skate and The
    Zootaxa 4098 (1): 179–190 ISSN 1175-5326 (print edition) http://www.mapress.com/j/zt/ Article ZOOTAXA Copyright © 2016 Magnolia Press ISSN 1175-5334 (online edition) http://doi.org/10.11646/zootaxa.4098.1.9 http://zoobank.org/urn:lsid:zoobank.org:pub:C7826AC7-6493-42D3-A0A4-89FEBB5AB3E3 Notoraja martinezi sp. nov., a new species of deepwater skate and the first record of the genus Notoraja Ishiyama, 1958 (Rajiformes: Arhynchobatidae) from the eastern Pacific Ocean FRANCISCO J. CONCHA1,2,7, DAVID A. EBERT3,4,5 & DOUGLAS J. LONG4,6 1Department of Ecology & Evolutionary Biology, University of Connecticut. 75 North Eagleville Road – Unit 3043 Storrs, CT, 06269, USA. E-mail: [email protected] 2Facultad de Ciencias del Mar y de Recursos Naturales, Universidad de Valparaíso. Av. Borgoño 16344, Viña del Mar, Chile. 3Pacific Shark Research Center, Moss Landing Marine Laboratories, Moss Landing, CA 95039, USA. 4Department of Ichthyology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA. 94118, USA. 5South African Institute for Aquatic Biodiversity, Private Bag 1015, Grahamstown, 6140, South Africa. E-mail: [email protected] state.edu. 6Department of Biology, St. Mary’s College, 1928 St. Mary’s Road, Moraga, California 94575. E-mail: [email protected] 7Corresponding author Abstract A new arhynchobatid skate, Notoraja martinezi, sp. nov., is described from four specimens collected from the eastern Central Pacific from Costa Rica to Ecuador and between depths of 1256–1472 m. The new species is placed in the genus Notoraja based on the long and flexible rostrum and its proportionally long tail with respect to total length.
    [Show full text]
  • The Evolutionary Origins of Vertebrate Haematopoiesis: Insights from Non-Vertebrate Chordates
    THE EVOLUTIONARY ORIGINS OF VERTEBRATE HAEMATOPOIESIS: INSIGHTS FROM NON-VERTEBRATE CHORDATES A thesis submitted to the University of Manchester for the degree of MPhil Evolutionary Biology in the Faculty of Life Sciences. 2015 PETER E D MILLS 2 LIST OF CONTENTS List of Contents 2 List of Figures 4 List of Abbreviations 5 Abstract 6 Declaration 7 Copyright Statement 7 Introduction 8 Haematopoiesis 11 In gnathostomes 11 In cyclostomes 18 In ascidians 20 In amphioxi 22 In sea urchins 24 In fruit flies 24 The evolution of haematopoiesis 26 Haematopoiesis and innate immune cells 26 Lymphocytes 26 Erythrocytes 27 Aims 28 The basic aim 28 Choice of genes 30 Homologue identification 31 Analysis of EST collections 31 In situ hybridisations 32 gata1/2/3 rescue experiment 34 gata1/2/3 reporter experiment 36 3 Methods 39 Homologue identification 39 Analysis of EST data 39 Collection of amphioxus embryos 40 Collection of ascidian embryos 40 Collection of zebrafish embryos 40 Cloning genes 41 Whole mount in situ hybridisations 41 Knockdown and rescue of gata1 42 gata1/2/3:mCherry reporter 42 Results Homologues in non-gnathostomes 44 Gene expression in ascidian and amphioxus embryos 48 Knockdown of gata1 and rescue with gata1/2/3 51 gata1/2/3:mCherry reporter expression 52 Discussion Non-chordate gene expression 54 Amphioxus gene expression 54 Ascidian gene expression 56 Cyclostome gene expression 59 gata1/2/3 rescue experiment 60 gata1/2/3 reporter experiment 62 Evolution of erythrocytes 63 Evolution of lymphocytes 67 Conclusion 72 References 75 (Word count: 26,246) 4 LIST OF FIGURES Figure 1 – Phylogenetic relationships between the organisms studied in this thesis.
    [Show full text]
  • Bibliography Database of Living/Fossil Sharks, Rays and Chimaeras (Chondrichthyes: Elasmobranchii, Holocephali) Papers of the Year 2016
    www.shark-references.com Version 13.01.2017 Bibliography database of living/fossil sharks, rays and chimaeras (Chondrichthyes: Elasmobranchii, Holocephali) Papers of the year 2016 published by Jürgen Pollerspöck, Benediktinerring 34, 94569 Stephansposching, Germany and Nicolas Straube, Munich, Germany ISSN: 2195-6499 copyright by the authors 1 please inform us about missing papers: [email protected] www.shark-references.com Version 13.01.2017 Abstract: This paper contains a collection of 803 citations (no conference abstracts) on topics related to extant and extinct Chondrichthyes (sharks, rays, and chimaeras) as well as a list of Chondrichthyan species and hosted parasites newly described in 2016. The list is the result of regular queries in numerous journals, books and online publications. It provides a complete list of publication citations as well as a database report containing rearranged subsets of the list sorted by the keyword statistics, extant and extinct genera and species descriptions from the years 2000 to 2016, list of descriptions of extinct and extant species from 2016, parasitology, reproduction, distribution, diet, conservation, and taxonomy. The paper is intended to be consulted for information. In addition, we provide information on the geographic and depth distribution of newly described species, i.e. the type specimens from the year 1990- 2016 in a hot spot analysis. Please note that the content of this paper has been compiled to the best of our abilities based on current knowledge and practice, however,
    [Show full text]
  • Age, Growth, and Sexual Maturity of the Deepsea Skate, Bathyraja
    AGE, GROWTH, AND SEXUAL MATURITY OF THE DEEPSEA SKATE, BATHYRAJA ABYSSICOLA A Thesis Presented to the Faculty of Alaska Pacific University In Partial Fulfillment of the Requirements For the Degree of Master of Science in Environmental Science by Cameron Murray Provost April 2016 Pro Q u est Nu m b er: 10104548 All rig hts reserv e d INF O RM ATI O N T O ALL USERS Th e q u a lity of this re pro d u ctio n is d e p e n d e nt u p o n th e q u a lity of th e c o p y su b mitt e d. In th e unlik e ly e v e nt th a t th e a uth or did n ot se n d a c o m ple t e m a nuscript a n d th ere are missin g p a g es, th ese will b e n ot e d. Also, if m a t eria l h a d to b e re m o v e d, a n ot e will in dic a t e th e d e le tio n. Pro Q u est 10104548 Pu blish e d b y Pro Q u est LL C (2016). C o p yrig ht of th e Dissert a tio n is h e ld b y th e A uth or. All rig hts reserv e d. This w ork is prot e ct e d a g a inst un a uth orize d c o p yin g un d er Title 17, Unit e d St a t es C o d e Microform Editio n © Pro Q u est LL C .
    [Show full text]
  • Extinction Risk and Conservation of the World's Sharks and Rays
    RESEARCH ARTICLE elife.elifesciences.org Extinction risk and conservation of the world’s sharks and rays Nicholas K Dulvy1,2*, Sarah L Fowler3, John A Musick4, Rachel D Cavanagh5, Peter M Kyne6, Lucy R Harrison1,2, John K Carlson7, Lindsay NK Davidson1,2, Sonja V Fordham8, Malcolm P Francis9, Caroline M Pollock10, Colin A Simpfendorfer11,12, George H Burgess13, Kent E Carpenter14,15, Leonard JV Compagno16, David A Ebert17, Claudine Gibson3, Michelle R Heupel18, Suzanne R Livingstone19, Jonnell C Sanciangco14,15, John D Stevens20, Sarah Valenti3, William T White20 1IUCN Species Survival Commission Shark Specialist Group, Department of Biological Sciences, Simon Fraser University, Burnaby, Canada; 2Earth to Ocean Research Group, Department of Biological Sciences, Simon Fraser University, Burnaby, Canada; 3IUCN Species Survival Commission Shark Specialist Group, NatureBureau International, Newbury, United Kingdom; 4Virginia Institute of Marine Science, College of William and Mary, Gloucester Point, United States; 5British Antarctic Survey, Natural Environment Research Council, Cambridge, United Kingdom; 6Research Institute for the Environment and Livelihoods, Charles Darwin University, Darwin, Australia; 7Southeast Fisheries Science Center, NOAA/National Marine Fisheries Service, Panama City, United States; 8Shark Advocates International, The Ocean Foundation, Washington, DC, United States; 9National Institute of Water and Atmospheric Research, Wellington, New Zealand; 10Global Species Programme, International Union for the Conservation
    [Show full text]
  • Skates and Rays Diversity, Exploration and Conservation – Case-Study of the Thornback Ray, Raja Clavata
    UNIVERSIDADE DE LISBOA FACULDADE DE CIÊNCIAS DEPARTAMENTO DE BIOLOGIA ANIMAL SKATES AND RAYS DIVERSITY, EXPLORATION AND CONSERVATION – CASE-STUDY OF THE THORNBACK RAY, RAJA CLAVATA Bárbara Marques Serra Pereira Doutoramento em Ciências do Mar 2010 UNIVERSIDADE DE LISBOA FACULDADE DE CIÊNCIAS DEPARTAMENTO DE BIOLOGIA ANIMAL SKATES AND RAYS DIVERSITY, EXPLORATION AND CONSERVATION – CASE-STUDY OF THE THORNBACK RAY, RAJA CLAVATA Bárbara Marques Serra Pereira Tese orientada por Professor Auxiliar com Agregação Leonel Serrano Gordo e Investigadora Auxiliar Ivone Figueiredo Doutoramento em Ciências do Mar 2010 The research reported in this thesis was carried out at the Instituto de Investigação das Pescas e do Mar (IPIMAR - INRB), Unidade de Recursos Marinhos e Sustentabilidade. This research was funded by Fundação para a Ciência e a Tecnologia (FCT) through a PhD grant (SFRH/BD/23777/2005) and the research project EU Data Collection/DCR (PNAB). Skates and rays diversity, exploration and conservation | Table of Contents Table of Contents List of Figures ............................................................................................................................. i List of Tables ............................................................................................................................. v List of Abbreviations ............................................................................................................. viii Agradecimentos ........................................................................................................................
    [Show full text]
  • An Introduction to the Classification of Elasmobranchs
    An introduction to the classification of elasmobranchs 17 Rekha J. Nair and P.U Zacharia Central Marine Fisheries Research Institute, Kochi-682 018 Introduction eyed, stomachless, deep-sea creatures that possess an upper jaw which is fused to its cranium (unlike in sharks). The term Elasmobranchs or chondrichthyans refers to the The great majority of the commercially important species of group of marine organisms with a skeleton made of cartilage. chondrichthyans are elasmobranchs. The latter are named They include sharks, skates, rays and chimaeras. These for their plated gills which communicate to the exterior by organisms are characterised by and differ from their sister 5–7 openings. In total, there are about 869+ extant species group of bony fishes in the characteristics like cartilaginous of elasmobranchs, with about 400+ of those being sharks skeleton, absence of swim bladders and presence of five and the rest skates and rays. Taxonomy is also perhaps to seven pairs of naked gill slits that are not covered by an infamously known for its constant, yet essential, revisions operculum. The chondrichthyans which are placed in Class of the relationships and identity of different organisms. Elasmobranchii are grouped into two main subdivisions Classification of elasmobranchs certainly does not evade this Holocephalii (Chimaeras or ratfishes and elephant fishes) process, and species are sometimes lumped in with other with three families and approximately 37 species inhabiting species, or renamed, or assigned to different families and deep cool waters; and the Elasmobranchii, which is a large, other taxonomic groupings. It is certain, however, that such diverse group (sharks, skates and rays) with representatives revisions will clarify our view of the taxonomy and phylogeny in all types of environments, from fresh waters to the bottom (evolutionary relationships) of elasmobranchs, leading to a of marine trenches and from polar regions to warm tropical better understanding of how these creatures evolved.
    [Show full text]
  • Classification and Systematic Arrangement
    Introduction 13 CLASSIFICATION AND SYSTEMATIC ARRANGEMENT Considering that the purpose of this document is to Class Chondrichthyes (cartilaginous fishes) provide a simple user-friendly guide for species identification, no reference will be found here to Subclass Holocephali (chimaeras) dichotomy keys for single species. It is important that the classification used in this guide be defined, Order Chimaeriformes (chimaera and silver sharks) as available literature is not always in agreement Subclass Elasmobranchii (sharks) with this presentation. The classification of this group is still under review as no consensus has Superorder Squalomorphi (squalomorph sharks) been found to reconcile different authors’ positions. Order Hexanchiformes (cow and frilled sharks) For more information and further specific details on the taxonomy and biology of cartilaginous fish Order Squaliformes (dogfish sharks) species, refer to Tortonese, 1956; Hureau and Monod 1979; Whitehead et al., 1984; Fischer et al., Order Squatiniformes (angel sharks) 1987; Fredj and Maurin, 1987; Compagno, 1988, 2005; Nelson, 1994; Shirai, 1996; Mould, 1998. The Order Pristiophoriformes (sawsharks) * consultation of FishBase http://www.fishbase.org Order Rajiformes (batoids) (Froese and Pauly, 2000) proved very useful. The most fundamental references are Compagno’s Superorder Galeomorphi (galeomorph sharks) catalogues issued in 1984 and his recent revision partially issued in 2001. Order Heterodontiformes (bullhead sharks) * This guide follows the systematic organization Order
    [Show full text]
  • Identification Guide to the Deep-Sea Cartilaginous Fishes Of
    Identification guide to the deep–sea cartilaginous fishes of the Southeastern Atlantic Ocean FAO. 2015. Identification guide to the deep–sea cartilaginous fishes of the Southeastern Atlantic Ocean. FishFinder Programme, by Ebert, D.A. and Mostarda, E., Rome, Italy. Supervision: Merete Tandstad, Jessica Sanders (FAO, Rome) Technical editor: Edoardo Mostarda (FAO, Rome) Colour illustrations, cover and graphic design: Emanuela D’Antoni (FAO, Rome) This guide was prepared under the “FAO Deep–sea Fisheries Programme” thanks to a generous funding from the Government of Norway (Support to the implementation of the International Guidelines on the Management of Deep-Sea Fisheries in the High Seas project) for the purpose of assisting states, institutions, the fishing industry and RFMO/As in the implementation of FAO International Guidelines for the Management of Deep-sea Fisheries in the High Seas. It was developed in close collaboration with the FishFinder Programme of the Marine and Inland Fisheries Branch, Fisheries Department, Food and Agriculture Organization of the United Nations (FAO). The present guide covers the deep–sea Southeastern Atlantic Ocean and that portion of Southwestern Indian Ocean from 18°42’E to 30°00’E (FAO Fishing Area 47). It includes a selection of cartilaginous fish species of major, moderate and minor importance to fisheries as well as those of doubtful or potential use to fisheries. It also covers those little known species that may be of research, educational, and ecological importance. In this region, the deep–sea chondrichthyan fauna is currently represented by 50 shark, 20 batoid and 8 chimaera species. This guide includes full species accounts for 37 shark, 9 batoid and 4 chimaera species selected as being the more difficult to identify and/or commonly caught.
    [Show full text]
  • XIV. Appendices
    Appendix 1, Page 1 XIV. Appendices Appendix 1. Vertebrate Species of Alaska1 * Threatened/Endangered Fishes Scientific Name Common Name Eptatretus deani black hagfish Lampetra tridentata Pacific lamprey Lampetra camtschatica Arctic lamprey Lampetra alaskense Alaskan brook lamprey Lampetra ayresii river lamprey Lampetra richardsoni western brook lamprey Hydrolagus colliei spotted ratfish Prionace glauca blue shark Apristurus brunneus brown cat shark Lamna ditropis salmon shark Carcharodon carcharias white shark Cetorhinus maximus basking shark Hexanchus griseus bluntnose sixgill shark Somniosus pacificus Pacific sleeper shark Squalus acanthias spiny dogfish Raja binoculata big skate Raja rhina longnose skate Bathyraja parmifera Alaska skate Bathyraja aleutica Aleutian skate Bathyraja interrupta sandpaper skate Bathyraja lindbergi Commander skate Bathyraja abyssicola deepsea skate Bathyraja maculata whiteblotched skate Bathyraja minispinosa whitebrow skate Bathyraja trachura roughtail skate Bathyraja taranetzi mud skate Bathyraja violacea Okhotsk skate Acipenser medirostris green sturgeon Acipenser transmontanus white sturgeon Polyacanthonotus challengeri longnose tapirfish Synaphobranchus affinis slope cutthroat eel Histiobranchus bathybius deepwater cutthroat eel Avocettina infans blackline snipe eel Nemichthys scolopaceus slender snipe eel Alosa sapidissima American shad Clupea pallasii Pacific herring 1 This appendix lists the vertebrate species of Alaska, but it does not include subspecies, even though some of those are featured in the CWCS.
    [Show full text]
  • Redescription and Taxonomic Status of Dipturus Chilensis (Guichenot, 1848), and Description of Dipturus Lamillai Sp
    Zootaxa 4590 (5): 501–524 ISSN 1175-5326 (print edition) https://www.mapress.com/j/zt/ Article ZOOTAXA Copyright © 2019 Magnolia Press ISSN 1175-5334 (online edition) https://doi.org/10.11646/zootaxa.4590.5.1 http://zoobank.org/urn:lsid:zoobank.org:pub:F484C560-CAE9-4A9E-B408-AEC2C8893DAD Redescription and taxonomic status of Dipturus chilensis (Guichenot, 1848), and description of Dipturus lamillai sp. nov. (Rajiformes: Rajidae), a new species of long-snout skate from the Falkland Islands FRANCISCO J. CONCHA1,2,7, JANINE N. CAIRA1, DAVID A. EBERT3,4,5 & JOOST H. W. POMPERT6 1Department of Ecology & Evolutionary Biology, University of Connecticut, 75 North Eagleville Road, Unit 3043 Storrs, CT 06269 – 3043, USA 2Facultad de Ciencias del Mar y de Recursos Naturales, Universidad de Valparaíso, Av. Borgoño 16344, Viña del Mar, Chile 3Pacific Shark Research Center, Moss Landing Marine Laboratories, Moss Landing, CA 95039, USA 4Department of Ichthyology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA 5South African Institute for Aquatic Biodiversity, Private Bag 1015, Grahamstown 6140, South Africa 6Georgia Seafoods Ltd, Waverley House, Stanley, Falkland Islands, United Kingdom 7Corresponding author. E-mail: [email protected] Abstract Recent molecular evidence has called into question the identity of skates collected in the waters off the Falkland Islands previously identified as Zearaja chilensis. NADH2 sequence data indicate that these specimens are not conspecific with those currently referred to as Z. chilensis from Chile and, in fact, represent a novel cryptic species. This study aimed to investigate this hypothesis based on morphological comparisons of specimens from the coasts of both western and eastern South America.
    [Show full text]
  • Elasmobranch Biodiversity, Conservation and Management Proceedings of the International Seminar and Workshop, Sabah, Malaysia, July 1997
    The IUCN Species Survival Commission Elasmobranch Biodiversity, Conservation and Management Proceedings of the International Seminar and Workshop, Sabah, Malaysia, July 1997 Edited by Sarah L. Fowler, Tim M. Reed and Frances A. Dipper Occasional Paper of the IUCN Species Survival Commission No. 25 IUCN The World Conservation Union Donors to the SSC Conservation Communications Programme and Elasmobranch Biodiversity, Conservation and Management: Proceedings of the International Seminar and Workshop, Sabah, Malaysia, July 1997 The IUCN/Species Survival Commission is committed to communicate important species conservation information to natural resource managers, decision-makers and others whose actions affect the conservation of biodiversity. The SSC's Action Plans, Occasional Papers, newsletter Species and other publications are supported by a wide variety of generous donors including: The Sultanate of Oman established the Peter Scott IUCN/SSC Action Plan Fund in 1990. The Fund supports Action Plan development and implementation. To date, more than 80 grants have been made from the Fund to SSC Specialist Groups. The SSC is grateful to the Sultanate of Oman for its confidence in and support for species conservation worldwide. The Council of Agriculture (COA), Taiwan has awarded major grants to the SSC's Wildlife Trade Programme and Conservation Communications Programme. This support has enabled SSC to continue its valuable technical advisory service to the Parties to CITES as well as to the larger global conservation community. Among other responsibilities, the COA is in charge of matters concerning the designation and management of nature reserves, conservation of wildlife and their habitats, conservation of natural landscapes, coordination of law enforcement efforts as well as promotion of conservation education, research and international cooperation.
    [Show full text]