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022 Genus Eagris Guenee AFROTROPICAL BUTTERFLIES 17th edition (2018). MARK C. WILLIAMS. http://www.lepsocafrica.org/?p=publications&s=atb Genus Eagris Guenée, 1862 In: Maillard, L., Notes sur l’Ile de la Reunion 2: 6, 19 (1-72) Paris. [2nd edition]. Type-species: Thymele sabadius Gray, 1832, by monotypy. = Trichosemeia Holland, 1892. Annals and Magazine of Natural History (6) 10: 294 et nota (284-294). Type-species: Trichosemeia subolivescens Holland, by original designation. Eagris lucetia. Image courtesy Jeremy Dobson. The genus Eagris belongs to the Family Hesperiidae Latreille, 1809; Subfamily Pyrginae Burmeister, 1878; Tribe Tagiadini Mabille, 1878. Other genera of the Tribe Tagiadini in the Afrotropical Region are Procampta, Calleagris, Kobelana, Tagiades, Netrobalane, Caprona, Leucochitonea and Abantis. Eagris (Flats) is an Afrotropical genus of ten species. Apparently the genus is structurally most closely related to Tagiades (Evans, 1937). *Eagris decastigma Mabille, 1891 Purple Flat Eagris decastigma Mabille, 1891. Bulletin de la Société Entomologique de Belgique 35: 62 (59-88, 106-121, 168-187). Type locality: Sierra Leone. Distribution: Guinea, Sierra Leone, Ivory Coast, Ghana, Nigeria, Cameroon, Gabon, Congo, Angola, Democratic Republic of Congo, Sudan, Uganda, Kenya, Tanzania, Zambia. Habitat: Paths and clearings in dense forest (Larsen, 2005a). At altitudes from 900 to 1 000 m in Tanzania (Kielland, 1990d). Habits: In West Africa this is a rare species (Larsen, 2005a). Males perch on leaves on the verges of forest roads, making frequent flights along the roads (Larsen, 1991c). Both sexes visit flowers (Larsen, 1991c). Males are found settling on wet rocks in stream-beds in the darkest part of the forest (Kielland, 1990d) and also visit urine patches (Larsen, 2005a). Early stages: Cock & Congdon, 2011 [pupa of subspecies purpura] Larval food: Nothing published. 1 Eagris decastigma decastigma Mabille, 1891 Eagris decastigma Mabille, 1891. Bulletin de la Société Entomologique de Belgique 35: 62 (59-88, 106-121, 168-187). Type locality: Sierra Leone. Distribution: Guinea, Sierra Leone, Ivory Coast, Ghana, Nigeria, Cameroon. Specific localities: Guinea – Labe (Larsen, 2005a); Nimba Mountains (Larsen, 2005a). Ivory Coast – Tai (Larsen, 2005a); Sassandra (Larsen, 2005a); Gagnoa (Larsen, 2005a). Ghana – Atewa (ABRI, vide Larsen, 2005a); Bobiri Butterfly Sanctuary (Larsen et al., 2007). Nigeria – Ibadan (Larsen, 2005a); Mamu Forest (Larsen, 2005a); Okwangwo (Larsen, 2005a); Oban Hills (Larsen, 2005a). Eagris decastigma fuscosa Holland, 1893 Aegris [sic] fuscosa Holland, 1893. Entomological News 5: 27 (26-31). Type locality: Gabon: “Valley of the Ogové”. Distribution: Nigeria, Cameroon, Gabon, Congo, Angola. Specific localities: Gabon – Ogove Valley (TL); Mondah (Vande weghe, 2010); Bakouaka (Vande weghe, 2010); Bitam (Vande weghe, 2010). Angola – Kwanza Norte Province (Mendes et al., 2013). Note: Larsen (2005a: 477) believes that the taxa fuscosa (Holland, 1893) and purpurea Evans, 1937 belong to a species distinct from decastigma Mabille, 1891. Eagris decastigma purpura Evans, 1937 Eagris decastigma purpura Evans, 1937. A catalogue of the African Hesperiidae indicating the classification and nomenclature adopted in the British Museum: 32 (212 pp.). Eagris decastigma purpura. Male. Left – upperside; right – underside. Wingspan: 33mm. Isombo, Ikelenge, Zambia. 8.III.79. A. Heath. (African Butterfly Research Institute, Nairobi). 2 Eagris decastigma purpura. Female. Left – upperside; right – underside. Ikelenge, Zambia. 18 June 2007. Images M.C. Williams ex Gardiner Collection. Type locality: Uganda: “Durro Forest, Toro, 4,000 ft”. Distribution: Democratic Republic of Congo (north-east – Ituri), Sudan (south), Uganda, Kenya (west), Tanzania (west), Zambia. Specific localities: Uganda – Durro Forest (TL); Semuliki N.P. (Davenport & Howard, 1996). Kenya – Kitale (Larsen, 1991c); Mount Elgon (Larsen, 1991c); Kakamega Forest (Larsen, 1991c; female illustrated above). Tanzania – Kigoma District: Kemfu Forest; Kasye Forest (Kielland, 1990d). Zambia – A single male from the Lisombu River, Ikelenge (figured, above) (Heath et al., 2002). Note: Larsen (2005a: 477) believes that the taxa fuscosa (Holland, 1893) and purpurea Evans, 1937 belong to a species distinct from decastigma Mabille, 1891. *Eagris denuba (Plötz, 1879) Cream Flat Antigonus denuba Plötz, 1879. Stettiner Entomologische Zeitung 40: 361 (353-364). Eagris denuba denuba. Male. Left – upperside; right – underside. Yekepah, Nimba Mountains, Liberia. 10 January 2014. J. Dobson. Images M.C. Williams ex Dobson Collection. Type locality: Ghana: “Aburi”. Distribution: Senegal, Guinea, Sierra Leone, Liberia, Ivory Coast, Ghana, Togo, Nigeria, Cameroon, Gabon, Sudan, Ethiopia, Uganda (Davenport, 1996). Habitat: All types of forest, as well as disturbed forest (Larsen, 2005a). Habits: A common species, which may even be numerous occasionally. In West Africa it is commoner west of the Dahomey Gap than it is in Nigeria (Larsen, 2005a). Males patrol territories in the early morning, periodically perching on leaves with the wings held flat (Larsen, 2005a). Contests between males are conducted at a furious pace. Resting takes place on the underside of leaves (Larsen, 2005a). Both sexes come to flowers and males, additionally, feed from bird droppings and occasionally mud- 3 puddle (Larsen, 2005a). Early stages: Nothing published. Larval food: Nothing published. Eagris denuba denuba (Plötz, 1879) Antigonus denuba Plötz, 1879. Stettiner Entomologische Zeitung 40: 361 (353-364). Eagris denuba denuba. Male. Left – upperside; right – underside. Yekepah, Nimba Mountains, Liberia. 10 January 2014. J. Dobson. Images M.C. Williams ex Dobson Collection. Type locality: Ghana: “Aburi”. Distribution: Senegal, Guinea, Sierra Leone, Liberia, Ivory Coast, Ghana, Togo, Nigeria, Cameroon (west), Gabon. Specific localities: Senegal – Basse Casamance (Larsen, 2005a). Guinea – Fouta Djalon (Larsen, 2005a). Sierra Leone – Freetown (Mabille, 1890). Ghana – Aburi (TL); Draw River (Larsen, 2005a); Bobiri Butterfly Sanctuary (Larsen et al., 2007); Boabeng-Fiema Monkey Sanctuary (Larsen et al., 2009). Gabon – Kangwe (Vande weghe, 2010). decolor Mabille, 1890 (as sp. of Eagris). Bulletin de la Société Entomologique de France (6) 9: 155 (149-150, 155-156, 167-169, 183-184). Sierra Leone: “Free-Town (Afrique)”. Eagris denuba obliterata Carpenter, 1928 Eagris lucetia obliterata Carpenter, 1928. Transactions of the Entomological Society of London 76: 48 (25-54). Eagris denuba obliterata. Male. Left – upperside; right – underside. Boter Becho, Ethiopia. 18 September 1997. Images M.C. Williams ex Gardiner Collection. 4 Eagris denuba obliterata. Female. Left – upperside; right – underside. Kibale Forest, Uganda. 22 October 2014. J. Dobson. Images M.C. Williams ex Dobson Collection. Type locality: Sudan: “Didinga”. Distribution: Sudan (south), Ethiopia, Uganda (Davenport, 1996). Specific localities: Sudan – Didinga (TL). Note: Larsen (2005a) is certain that obliterata is a species distinct from denuba. *Eagris hereus (Druce, 1875) Beautiful Orange Flat Tagiades hereus Druce, 1875. Proceedings of the Zoological Society of London 1875: 417 (406-417). Type locality: Angola. Distribution: Guinea, Sierra Leone, Ivory Coast, Ghana, Togo, Nigeria, Cameroon, Gabon, Congo, Angola, Central African Republic. Habits: Generally scarce and localized (Larsen, 2005a). Males aggressively defend territories from perches on leaves two to four metres above the ground. Individuals often come lower down to feed from flowers (Larsen, 2005a). Early stages: Nothing published. Larval food: Nothing published. Eagris hereus hereus (Druce, 1875) Tagiades hereus Druce, 1875. Proceedings of the Zoological Society of London 1875: 417 (406-417). Type locality: Angola. Type material in NHM, London (Evans, 1937). Distribution: Cameroon (except west), Gabon, Congo, Angola, Central African Republic. Specific localities: Cameroon – Victoria (Strand, 1913). Angola – Kwanza Norte Province (Mendes et al., 2013). birgitta Strand, 1913 (as sp. of Trichosemeia). Archiv für Naturgeschichte 78 (A.12.): 128 (121-131). Cameroon: “Victoria”. Eagris hereus quaterna (Mabille, 1889) Ceratrichia quaterna Mabille, 1889. Bulletin de la Société Entomologique de France (6) 9: 156 (149-150, 155-156, 167-169, 5 183-184). Type locality: Sierra Leone. Distribution: Guinea, Sierra Leone, Ivory Coast, Ghana, Togo, Nigeria, Cameroon (west). Specific localities: Guinea – Conakry (Larsen, 2005a). Ivory Coast – Gagnoa (Larsen, 2005a); Tiassale (Larsen, 2005a); Adiopodoume (Larsen, 2005a); Banco (Larsen, 2005a); Tai (Larsen, 2005a); Nimba area (Larsen, 2005a). Ghana – Bia (Larsen, 2005a); Kakum (Larsen, 2005a); Ankasa (Larsen, 2005a). Nigeria – Agege (Larsen, 2005a); Ilaro (Larsen, 2005a); Benin (Larsen, 2005a); Okwangwo (Larsen, 2005a); Port Harcourt (Larsen, 2005a); Oban Hills (Larsen, 2005a). *Eagris lucetia (Hewitson, [1875]) Resting males of Eagris lucetia. Mabira Forest (left) and Kibale Forest (right), Uganda. Images courtesy Jeremy Dobson (left) and Raimund Schutte (right). Leucochitonea lucetia Hewitson, [1875] in Hewitson, [1872-7]. Illustrations of new species of exotic butterflies 5: 112 (127 pp.) London. Eagris lucetia. Male (Wingspan 33 mm). Left – upperside; right – underside. Mabira Forest, Uganda. 14 June 2009. J. Dobson Images M.C. Williams ex Dobson Collection. Eagris lucetia. Female. Left – upperside; right – underside. 6 Kakamega Forest, Kenya. 16 June 1996. Images M.C. Williams ex
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