Phasmid Study Group Newsletter, 25
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Stick Insects (PDF 124KB)
Handle with care Stick insects Species: Indian stick insect (there are lots of others which could be listed but this is the most common) Scientific names: Carausius morosus Description The Indian stick insect is the most commonly available of all stick insect species. Indian stick insects are normally olive green, but brown and yellow-green colours occur too. Their twig and leaf-like appearance gives them good camouflage. An adult stick insect measures about 7.5cm after about five or six skin moults and will live for about a year. Life in the wild Stick insects live in tropical and semi tropical areas of the world. They are vegetarians and eat the leaves of plants, shrubs and trees. Privet and bramble leaves are favourites. In the wild they are usually eaten by birds, so stick insects tend to feed at night when birds are not around. Source of animals Stick insects can be bought from pet shops or specialist breeders. RSPCA animal centres also occasionally have stick insects needing new homes. Prior knowledge and preparation Before keeping stick insects, it is crucial that any potential keeper finds out about them and how to provide for them in captivity. Only then can an informed decision be made about whether s/he can provide the facilities, time, financial means and long-term commitment to look after them properly. Suitable accommodation, food and the necessary accessories should be bought and prepared before taking the animal home. Novice keepers should also take the time to talk to experienced stick insect keepers and professionals – such as a vet – for further advice. -
ARTHROPODA Subphylum Hexapoda Protura, Springtails, Diplura, and Insects
NINE Phylum ARTHROPODA SUBPHYLUM HEXAPODA Protura, springtails, Diplura, and insects ROD P. MACFARLANE, PETER A. MADDISON, IAN G. ANDREW, JOCELYN A. BERRY, PETER M. JOHNS, ROBERT J. B. HOARE, MARIE-CLAUDE LARIVIÈRE, PENELOPE GREENSLADE, ROSA C. HENDERSON, COURTenaY N. SMITHERS, RicarDO L. PALMA, JOHN B. WARD, ROBERT L. C. PILGRIM, DaVID R. TOWNS, IAN McLELLAN, DAVID A. J. TEULON, TERRY R. HITCHINGS, VICTOR F. EASTOP, NICHOLAS A. MARTIN, MURRAY J. FLETCHER, MARLON A. W. STUFKENS, PAMELA J. DALE, Daniel BURCKHARDT, THOMAS R. BUCKLEY, STEVEN A. TREWICK defining feature of the Hexapoda, as the name suggests, is six legs. Also, the body comprises a head, thorax, and abdomen. The number A of abdominal segments varies, however; there are only six in the Collembola (springtails), 9–12 in the Protura, and 10 in the Diplura, whereas in all other hexapods there are strictly 11. Insects are now regarded as comprising only those hexapods with 11 abdominal segments. Whereas crustaceans are the dominant group of arthropods in the sea, hexapods prevail on land, in numbers and biomass. Altogether, the Hexapoda constitutes the most diverse group of animals – the estimated number of described species worldwide is just over 900,000, with the beetles (order Coleoptera) comprising more than a third of these. Today, the Hexapoda is considered to contain four classes – the Insecta, and the Protura, Collembola, and Diplura. The latter three classes were formerly allied with the insect orders Archaeognatha (jumping bristletails) and Thysanura (silverfish) as the insect subclass Apterygota (‘wingless’). The Apterygota is now regarded as an artificial assemblage (Bitsch & Bitsch 2000). -
Looking Beyond Glacial Refugia ⇑ Katharine A
Molecular Phylogenetics and Evolution 59 (2011) 89–102 Contents lists available at ScienceDirect Molecular Phylogenetics and Evolution journal homepage: www.elsevier.com/locate/ympev Reconciling phylogeography and ecological niche models for New Zealand beetles: Looking beyond glacial refugia ⇑ Katharine A. Marske a,b, , Richard A.B. Leschen a, Thomas R. Buckley a a Landcare Research, Private Bag 92170, Auckland 1142, New Zealand b Center for Macroecology, Evolution and Climate, University of Copenhagen, Universitetsparken 15, DK-2100 Copenhagen Ø, Denmark article info abstract Article history: Mitochondrial DNA (cox1) sequence data and recently developed coalescent phylogeography models Received 19 July 2010 were used to construct geo-spatial histories for the New Zealand fungus beetles Epistranus lawsoni and Revised 12 November 2010 Pristoderus bakewelli (Zopheridae). These methods utilize continuous-time Markov chains and Bayesian Accepted 13 January 2011 stochastic search variable selection incorporated in BEAST to identify historical dispersal patterns via Available online 22 January 2011 ancestral state reconstruction. Ecological niche models (ENMs) were incorporated to reconstruct the potential geographic distribution of each species during the Last Glacial Maximum (LGM). Coalescent Keywords: analyses suggest a North Island origin for E. lawsoni, with gene flow predominately north–south between Colydiinae adjacent regions. ENMs for E. lawsoni indicated glacial refugia in coastal regions of both main islands, con- Epistranus lawsoni Pristoderus bakewelli sistent with phylogenetic patterns but at odds with the coalescent dates, which implicate much older Maxent topographic events. Dispersal matrices revealed patterns of gene flow consistent with projected refugia, Coalescent phylogeography suggesting long-term South Island survival with population vicariance around the Southern Alps. -
Analysis of the Stick Insect (Clitarchus Hookeri) Genome Reveals a High Repeat Content and Sex- Biased Genes Associated with Reproduction Chen Wu1,2,3* , Victoria G
Wu et al. BMC Genomics (2017) 18:884 DOI 10.1186/s12864-017-4245-x RESEARCH ARTICLE Open Access Assembling large genomes: analysis of the stick insect (Clitarchus hookeri) genome reveals a high repeat content and sex- biased genes associated with reproduction Chen Wu1,2,3* , Victoria G. Twort1,2,4, Ross N. Crowhurst3, Richard D. Newcomb1,3 and Thomas R. Buckley1,2 Abstract Background: Stick insects (Phasmatodea) have a high incidence of parthenogenesis and other alternative reproductive strategies, yet the genetic basis of reproduction is poorly understood. Phasmatodea includes nearly 3000 species, yet only thegenomeofTimema cristinae has been published to date. Clitarchus hookeri is a geographical parthenogenetic stick insect distributed across New Zealand. Sexual reproduction dominates in northern habitats but is replaced by parthenogenesis in the south. Here, we present a de novo genome assembly of a female C. hookeri and use it to detect candidate genes associated with gamete production and development in females and males. We also explore the factors underlying large genome size in stick insects. Results: The C. hookeri genome assembly was 4.2 Gb, similar to the flow cytometry estimate, making it the second largest insect genome sequenced and assembled to date. Like the large genome of Locusta migratoria,the genome of C. hookeri is also highly repetitive and the predicted gene models are much longer than those from most other sequenced insect genomes, largely due to longer introns. Miniature inverted repeat transposable elements (MITEs), absent in the much smaller T. cristinae genome, is the most abundant repeat type in the C. hookeri genome assembly. -
Nakanomscthesis.Pdf
Copyright is owned by the Author of the thesis. Permission is given for a copy to be downloaded by an individual for the purpose of research and private study only. The thesis may not be reproduced elsewhere without the permission of the Author. Male mate choice in the stick insect Clitarchus hookeri: sexual vs. parthenogenetic females A thesis presented in partial fulfilment of the requirements for the degree of Master of Science in Zoology at Massey University, Manawatū, New Zealand Photo credit – Mary Morgan-Richards Mari Nakano 2018 i Abstract Learning about reproductive strategies in animals is an important step for understanding the evolution of species. New Zealand stick insect, Clitarchus hookeri include both sexual and parthenogenetic females, and parthenogenetic females occur in the distributional range where males are absent and have a limited capacity of sexual reproduction. Since C. hookeri exhibit a scramble competition mating system with distinctive sex roles where females and males co-occur, it is likely that parthenogenetic females do not exhibit traits that are related to the sex roles. Furthermore, due to limited capacity for parthenogenetic females to reproduce sexually, it is likely that C. hookeri males would benefit from discriminating between sexual and parthenogenetic females. The main purpose of this thesis was to explore the unique reproductive features of Clitarchus hookeri. Specifically, I identified morphological and chemical traits that are likely to be under distinctive sex roles in scramble competition; revealed whether morphological and chemical traits seen in sexual females are also seen in parthenogenetic females; and observed whether males can discriminate between sexual and parthenogenetic females for their pre- and post-copulatory choices. -
Extreme Positive Allometry of Animal Adhesive Pads and the Size Limits of Adhesion-Based Climbing
Extreme positive allometry of animal adhesive pads and the size limits of adhesion-based climbing David Labontea,1, Christofer J. Clementeb, Alex Dittrichc, Chi-Yun Kuod, Alfred J. Crosbye, Duncan J. Irschickd, and Walter Federlea aDepartment of Zoology, University of Cambridge, Cambridge CB2 3EJ, United Kingdom; bSchool of Science and Engineering, University of the Sunshine Coast, Sippy Downs, QLD 4556, Australia; cDepartment of Life Sciences, Anglia Ruskin University, Cambridge CB1 1PT, United Kingdom; dBiology Department, University of Massachusetts Amherst, Amherst, MA 01003; and eDepartment of Polymer Science and Engineering, University of Massachusetts Amherst, Amherst, MA 01003 Edited by David B. Wake, University of California, Berkeley, CA, and approved December 11, 2015 (received for review October 7, 2015) Organismal functions are size-dependent whenever body surfaces animals to climb smooth vertical or inverted surfaces, thereby supply body volumes. Larger organisms can develop strongly opening up new habitats. Adhesive pads have evolved multiple folded internal surfaces for enhanced diffusion, but in many cases times independently within arthropods, reptiles, amphibians, and areas cannot be folded so that their enlargement is constrained by mammals and show impressive performance: They are rapidly anatomy, presenting a problem for larger animals. Here, we study controllable, can be used repeatedly without any loss of perfor- the allometry of adhesive pad area in 225 climbing animal species, mance, and function on rough, dirty, and flooded surfaces (12). This covering more than seven orders of magnitude in weight. Across performance has inspired a considerable amount of work on tech- all taxa, adhesive pad area showed extreme positive allometry and nical adhesives that mimic these properties (13). -
The Antennal Motor System of the Stick Insect Carausius Morosus : Anatomy and Antennal Movement Pattern During Walking
J Comp Physiol A ;2001) 187: 131±144 DOI 10.1007/s003590100183 ORIGINAL PAPER Volker DuÈ rr á Yvonne KoÈ nig á Rolf Kittmann The antennal motor system of the stick insect Carausius morosus : anatomy and antennal movement pattern during walking Accepted: 8 January 2001 / Published online: 22 February 2001 Ó Springer-Verlag 2001 Abstract The stick insect Carausius morosus continu- Key words Antennal movement á Tactile sense á ously moves its antennae during locomotion. Active Antennal muscle á Motoneuron á Limb coordination antennal movements may re¯ect employment of anten- nae as tactile probes. Therefore, this study treats two Abbreviations Ab1±4 abductor motoneuron 1±4 á Ad1±3 basic aspects of the antennal motor system: First, the adductor motoneuron 1±3 á AEP: anterior extreme po- anatomy of antennal joints, muscles, nerves and moto- sition á AL antennal lobe á AMMC antennal mechano- neurons is described and discussed in comparison with sensory and motor complex á aTB anterior tentorial other species. Second, the typical movement pattern of branch á cMD central part of MD á CI common inhibi- the antennae is analysed, and its spatio-temporal coor- tor á CT corpus tentorium á Dp1±5 depressor motoneu- dination with leg movements described. Each antenna is ron 1±5 á dTB dorsal tentorial branch á lMD lateral part moved by two single-axis hinge joints. The proximal ofMDá lML lateral levator muscle á Lv1±4 levator head-scape joint is controlled by two levator muscles motoneuron 1±4 á MD depressor muscle á mMD medial and a three-partite depressor muscle. -
Phasma Gigas from New Ireland Mark Bushell
ISSN 0966-0011 PHASMID STUDIES. volume 8, numbers 1 & 2. December 1999. Editor: P.E. Bragg. Published by the Phasmid Study Group. Phasmid Studies ISSN 0966-0011 volume 8, numbers 1 & 2. Contents Studies of the genus Phalces Stal Paul D. Brock . 1 Redescription of Mantis filiformes Fabricius (Phasmatidae: Bacteriinae) Paul D. Brock . 9 Phasmida in Oceania Allan Harman . 13 A Report on a Culture of Phasma gigas from New Ireland Mark Bushell . 20 Reviews and Abstracts Phasmid Abstracts 25 Cover illustration: Female Spinodares jenningsi Bragg, drawing by P.E. Bragg. Studies of the genus Phalces Stal Paul D. Brock, "Papillon", 40 Thorndike Road, Slough SU ISR, UK. Abstract Phalces tuberculatus sp.n. is described from Eland's Bay, Cape Province, South Africa. A key is given to distinguish the Phalces species. Brief notes are given on behaviour, foodplants, and culture notes in the case of P. longiscaphus (de Haan). Key words: Phasmida, Phalces, Phalcestuberculatus sp.n, Introduction As part of my studies on South African stick-insects, I visited Cape Town in September 1998. My research included an examination of the entomology collection at the South African Museum in Cape Town, in addition to material of Phalces species in various museums, observing P. longiscaphus in the wild and rearing this species in captivity. The observations include the description of Phalces tuberculatus sp.n. and a key to distinguish the three Phalces species (of which a Madagascan insect is unlikely to belong to this genus). Museum codens are given below: BMNH Natural History Museum, London, U.K. NHMW Naturhistorisches Museum, Wien, Austria. -
VKM Rapportmal
VKM Report 2016: 36 Assessment of the risks to Norwegian biodiversity from the import and keeping of terrestrial arachnids and insects Opinion of the Panel on Alien Organisms and Trade in Endangered species of the Norwegian Scientific Committee for Food Safety Report from the Norwegian Scientific Committee for Food Safety (VKM) 2016: Assessment of risks to Norwegian biodiversity from the import and keeping of terrestrial arachnids and insects Opinion of the Panel on Alien Organisms and Trade in Endangered species of the Norwegian Scientific Committee for Food Safety 29.06.2016 ISBN: 978-82-8259-226-0 Norwegian Scientific Committee for Food Safety (VKM) Po 4404 Nydalen N – 0403 Oslo Norway Phone: +47 21 62 28 00 Email: [email protected] www.vkm.no www.english.vkm.no Suggested citation: VKM (2016). Assessment of risks to Norwegian biodiversity from the import and keeping of terrestrial arachnids and insects. Scientific Opinion on the Panel on Alien Organisms and Trade in Endangered species of the Norwegian Scientific Committee for Food Safety, ISBN: 978-82-8259-226-0, Oslo, Norway VKM Report 2016: 36 Assessment of risks to Norwegian biodiversity from the import and keeping of terrestrial arachnids and insects Authors preparing the draft opinion Anders Nielsen (chair), Merethe Aasmo Finne (VKM staff), Maria Asmyhr (VKM staff), Jan Ove Gjershaug, Lawrence R. Kirkendall, Vigdis Vandvik, Gaute Velle (Authors in alphabetical order after chair of the working group) Assessed and approved The opinion has been assessed and approved by Panel on Alien Organisms and Trade in Endangered Species (CITES). Members of the panel are: Vigdis Vandvik (chair), Hugo de Boer, Jan Ove Gjershaug, Kjetil Hindar, Lawrence R. -
Ecological Assessment
ENNOR FARM ISLES OF SCILLY ECOLOGICAL ASSESSMENT January 2021 8128.002 Version 5.0 Document Title Ennor Farm Ecological Assessment Prepared for CampbellReith Prepared by TEP Ltd Document Ref 8128.002 Author Gemma Hassall Date October 2020 Checked Lee Greenhough Approved Lee Greenhough Amendment History Check / Modified Version Date Approved Reason(s) issue Status by by Minor update to reflect design freeze and Final for client 2.0 01/12/2020 RAR LG additional appendix approval Inclusion of CampbellReith Drainage 3.0 16/12/2020 LG RAR Strategy plan (ref 13394-CRH-XX-XX-DR-C- Final 5050-P2 Drainage Strategy) January CampbellReith update of proposed layout 4.0 - - For submission 2021 plan Amendment to reflect additional tree removal Final for 5.0 11/01/2021 RAR LG and replacement required to accommodate Planning Issue visibility splay; phase 1 map correction Ennor Farm St. Mary’s, Isles of Scilly Ecological Assessment Contents Page EXECUTIVE SUMMARY ....................................................................................................... 1 1.0 INTRODUCTION ..................................................................................................... 2 Site Description ....................................................................................................... 2 2.0 METHODS............................................................................................................... 4 Desktop Study ......................................................................................................... 4 Habitat -
The Pregenital Abdominal Musculature in Phasmids and Its Implications for the Basal Phylogeny of Phasmatodea (Insecta: Polyneoptera) Rebecca Klugã, Sven Bradler
ARTICLE IN PRESS Organisms, Diversity & Evolution 6 (2006) 171–184 www.elsevier.de/ode The pregenital abdominal musculature in phasmids and its implications for the basal phylogeny of Phasmatodea (Insecta: Polyneoptera) Rebecca KlugÃ, Sven Bradler Zoologisches Institut und Museum, Georg-August-Universita¨tGo¨ttingen, Berliner Str. 28, 37073 Go¨ttingen, Germany Received 7 June 2005; accepted 25 August 2005 Abstract Recently several conflicting hypotheses concerning the basal phylogenetic relationships within the Phasmatodea (stick and leaf insects) have emerged. In previous studies, musculature of the abdomen proved to be quite informative for identifying basal taxa among Phasmatodea and led to conclusions regarding the basal splitting events within the group. However, this character complex was not studied thoroughly for a representative number of species, and usually muscle innervation was omitted. In the present study the musculature and nerve topography of mid-abdominal segments in both sexes of seven phasmid species are described and compared in detail for the first time including all putative basal taxa, e.g. members of Timema, Agathemera, Phylliinae, Aschiphasmatinae and Heteropteryginae. The ground pattern of the muscle and nerve arrangement of mid-abdominal segments, i.e. of those not modified due to association with the thorax or genitalia, is reconstructed. In Timema, the inner ventral longitudinal muscles are present, whereas they are lost in all remaining Phasmatodea (Euphasmatodea). The ventral longitudinal muscles in the abdomen of Agathemera, which span the whole length of each segment, do not represent the plesiomorphic condition as previously assumed, but might be a result of secondary elongation of the external ventral longitudinal muscles. -
The Stick-Insects of Great Britain, Ireland and the Channel Isles by Malcolm Lee, Gullrock, Port Gaverne, Port Isaac, Cornwall
The Phasmid Study Group SEPTEMBER 2006 NEWSLETTER No 107 This Newsletter looks fantastic ISSN 0268-3806 in full colour. Go to the members' area of the PSG INDEX Website to view &/or download. National Insect Week Page 5 Page Content The Colour Page Editorial, Diary Dates, PSG Committee Stick Talk National Insect Week Eurycantha calcarata Observations The Kettering Show, Phasmids in the News Respiration in Phasmids Collecting in Thailand, Additions to Culture list PSG Summer Meeting Plea for Comments on Future PSG Meetings Database of Natural History Museum in Vienna Word Scramble, New Editor Needed for Newsletter Defensive Tactics in Phasmids Report on Pestival Word Scramble Answers, Survey Results, PSG Merchandise, Wants & Exchanges, Members' Website Password, Helpful Taxonomists Wanted Stick Insects of Great Britain Phaenopharos khaoyaiensis Article , Obituary. PSG Summer Meeting, Page 11. Which is the ant, which the nymph? Page15. NOTICE It is to be directly understood that all views, opinions or theories, expressed in the pages of "The Newsletter", are those of the author(s) concerned. All announcements of meetings, requests for help or information, are accepted as bona fide. Neither the Editor, nor Officers and Committee of "The Phasmid Study Group", can be held responsible for any loss, embarrassment or injury that might be sustained by reliance thereon. September 2006 Website: www.stickinsect.org.uk Newsletter 107.1 THE COLOUR PAGE! sp. by Chris Pull September 2006 Website: www.stickinsect.org.uk Newsletter 107.2 Editorial Welcome to the September PSG Newsletter; please read and enjoy. I am amazed at the wonderful " 'V^-r«, response I had when seeking contributions for this Newsletter.